treatments-xml/data/E9/72/FC/E972FC6A323A5C2AB3E96FF4072666AF.xml
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1147.93824" ID-Pensoft-Pub="1313-2970-1147-1" ID-Pensoft-UUID="FF2B6B317B505F9EA0E1000BDCD16CE7" ID-ZooBank="9BB9881B0076473D8E53155D37CA1F50" ModsDocID="1313-2970-1147-1" checkinTime="1676378625502" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Hunyadi, Andras, Vermeulen, Jaap J., Grego, Jozef, Sutcharit, Chirasak, Reischuetz, Alexander, Dumrongrojwattana, Pongrat, Botta-Dukat, Zoltan, Oerstan, Aydin, Fekete, Judit &amp; Jochum, Adrienne" docDate="2023" docId="E972FC6A323A5C2AB3E96FF4072666AF" docLanguage="en" docName="ZooKeys 1147: 1-177" docOrigin="ZooKeys 1147" docPubDate="2023-02-13" docSource="http://dx.doi.org/10.3897/zookeys.1147.93824" docTitle="Angustopila oostoma Pall-Gergely &amp; Vermeulen 2023, sp. nov." docType="treatment" docUuid="9C120D4D-9473-4CB6-8C19-E02953CD7038" docUuidSource="ZooBank" docVersion="1" id="FF2B6B317B505F9EA0E1000BDCD16CE7" lastPageNumber="1" masterDocId="FF2B6B317B505F9EA0E1000BDCD16CE7" masterDocTitle="Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)" masterLastPageNumber="177" masterPageNumber="1" pageNumber="1" updateTime="1676378625502" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pall-Gergely, Barna</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
<mods:affiliation>Centre for Agricultural Research, Plant Protection Institute, Eoetvoes Lorand Research Network, Herman Otto ut 15, H- 1022 Budapest, Hungary</mods:affiliation>
<mods:nameIdentifier type="email">pallgergely2@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hunyadi, Andras</mods:namePart>
<mods:affiliation>Adria setany 10 G 2 / 5., H- 1148 Budapest, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vermeulen, Jaap J.</mods:namePart>
<mods:affiliation>JK Art and Science, Lauwerbes 8, 2318 AT Leiden, Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Grego, Jozef</mods:namePart>
<mods:affiliation>Horna Micina 219, SK- 97401 Banska Bystrica, Slovakia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sutcharit, Chirasak</mods:namePart>
<mods:affiliation>Animal Systematic Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Reischuetz, Alexander</mods:namePart>
<mods:affiliation>Puechhaimgasse 52, A- 3580 Horn, Austria</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dumrongrojwattana, Pongrat</mods:namePart>
<mods:affiliation>Department of Biology, Faculty of Science, Burapha University, 169 Longhardbangsaen Road, Muang District, Chonburi, 20131, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Botta-Dukat, Zoltan</mods:namePart>
<mods:affiliation>Centre for Ecological Research, Institute of Ecology and Botany, Alkotmany 2 - 4, H- 2600, Vacratot, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Oerstan, Aydin</mods:namePart>
<mods:affiliation>12501 Milestone Manor Lane, Germantown, Maryland, 20876, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fekete, Judit</mods:namePart>
<mods:affiliation>University of Pannonia, Centre of Natural Science, Research Group of Limnology, Egyetem u. 10, H- 8200 Veszprem, Hungary &amp; Centre for Ecological Research, Institute of Aquatic Ecology, Department of Tisza Research, 18 / c Bem square, H- 4026 Debrecen, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jochum, Adrienne</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6624-6412</mods:nameIdentifier>
<mods:affiliation>Naturhistorisches Museum der Burgergemeinde Bern, CH- 3005 Bern, Switzerland &amp; Institute of Ecology and Evolution, University of Bern, CH- 3012 Bern, Switzerland &amp; Senckenberg Forschungsinstitut und Naturmuseum, 60325 Frankfurt am Main, Germany</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-02-13</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1147</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>177</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1147.93824</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1147.93824</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1147-1</mods:identifier>
<mods:identifier type="ZooBank">9BB9881B0076473D8E53155D37CA1F50</mods:identifier>
<mods:identifier type="Pensoft-UUID">FF2B6B317B505F9EA0E1000BDCD16CE7</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:zoobank.org:act:9C120D4D-9473-4CB6-8C19-E02953CD7038" httpUri="http://treatment.plazi.org/id/E972FC6A323A5C2AB3E96FF4072666AF" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="https://zoobank.org/9C120D4D-9473-4CB6-8C19-E02953CD7038" authority="Páll-Gergely &amp; Vermeulen" authorityName="Pall-Gergely &amp; Vermeulen" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Angustopila oostoma" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="oostoma" status="sp. nov.">
Angustopila oostoma
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Vermeulen
</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 51" captionStartId="F51" captionText="Figure 51. Angustopila oostoma Pall-Gergely &amp; Vermeulen, sp. nov. (holotype, HNHM 105289). Apertural (A), apical (B), lateral (C) and ventral (D) sides of the shell; eroded surface showing some microstructure on the protoconch (E), aperture (F), ventral (G) and frontal (H) surface of the body whorl." figureDoi="10.3897/zookeys.1147.93824.figure51" httpUri="https://binary.pensoft.net/fig/808860" pageId="0" pageNumber="1">Figs 51</figureCitation>
<figureCitation captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Angustopila oostoma Pall-Gergely &amp; Vermeulen, sp. nov. (2020 / 28). Apertural (A), ventral (B), apical (C) and lateral (D), sides of the shell; aperture (E), microstructure of the protoconch showing protoconch-teleoconch boundary (F), ventral (G) and frontal (H) surface of the body whorl." figureDoi="10.3897/zookeys.1147.93824.figure52" httpUri="https://binary.pensoft.net/fig/808861" pageId="0" pageNumber="1">, 52</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
: Vietnam • 1 empty shell (H: 0.89 mm, D: 0.81 mm); Ninh Binh Province, Cuc Phuong National Park, path to Fairy Cave, primary forest, limestone outcrops in shale bedrock;
<geoCoordinate degrees="20" direction="north" minutes="13.06" orientation="latitude" precision="9" value="20.217667">20°13.06'N</geoCoordinate>
,
<geoCoordinate degrees="105" direction="east" minutes="53.43" orientation="longitude" precision="9" value="105.8905">105°53.43'E</geoCoordinate>
; 10 October 1998; J.J. Vermeulen &amp; L. Deharveng leg.; HNHM 105289 (original inventory number: JJV 6224).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
: Vietnam • 1 adult shell with imperfectly grown last whorl; Ninh Binh Province, Cuc Phuong National Park, 1 km WNW of Xom Bong, primary forest, small limestone outcrops in clay soil; approximate GPS coordinates:
<geoCoordinate degrees="20" direction="north" minutes="55.57" orientation="latitude" precision="9" value="20.926167">20°55.57'N</geoCoordinate>
,
<geoCoordinate degrees="105" direction="east" minutes="9.24" orientation="longitude" precision="9" value="105.154">105°9.24'E</geoCoordinate>
; 10 Oct. 1998; J.J. Vermeulen &amp; L. Deharveng leg.; JJV 6251 • 2 shells; Ninh Binh Province, Cuc Phuong National Park, 2 km road Xom Bong to Headquarters, primary forest, limestone outcrops in thick clayey soil; 10 Oct. 1998; J.J. Vermeulen &amp; L. Deharveng leg.; JJV 6222.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Additional material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation accessionNumber="JJV 17629" collectingDate="1998-10-10" collectingDateMax="2020-02-11" collectingDateMin="1998-10-10" collectorName="Tan L, Quy H, A. Hunyadi, Xom Bong, Headquarters, J. J. Vermeulen, L. Deharveng" country="Vietnam" county="District" elevation="1300" latitude="20.2935" location="Cuc Phuong National Park" longLatPrecision="12" longitude="105.6685" municipality="Cuc Phuong National Park" specimenCount="3" specimenCount-adult="3" stateProvince="Hoa Binh Province">
<collectingCountry name="Vietnam">Vietnam</collectingCountry>
<specimenCount type="adult">3 adult</specimenCount>
shells;
<collectingRegion country="Vietnam" name="Hoa Binh">
<normalizedToken originalValue="Hòa">Hoa</normalizedToken>
<normalizedToken originalValue="Bình">Binh</normalizedToken>
Province
</collectingRegion>
,
<collectorName>
<normalizedToken originalValue="Tân">Tan</normalizedToken>
L
</collectorName>
ạc
<collectingCounty>District</collectingCounty>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.3" unit="m" value="1300.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.3" unit="m" value="1300.0">1300 m</elevation>
</quantity>
west from
<collectorName>Quy H</collectorName>
ậu on road no. 6, rock wall (locality code: 2020/28);
<geoCoordinate degrees="20" direction="north" minutes="37.93" orientation="latitude" precision="9" value="20.632166">20°37.93'N</geoCoordinate>
,
<geoCoordinate degrees="105" direction="east" minutes="16.17" orientation="longitude" precision="9" value="105.2695">105°16.17'E</geoCoordinate>
;
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.5" unit="m" value="150.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.5" unit="m" value="150.0">150 m</quantity>
a.s.l.
</elevation>
;
<collectingDate value="2020-02-11">11 Feb. 2020</collectingDate>
;
<collectorName>A. Hunyadi</collectorName>
leg.; coll. HA • 3 j/b shells;
<collectingRegion country="Vietnam" name="Ninh Binh">Ninh Binh Province</collectingRegion>
,
<collectingMunicipality>Cuc Phuong National Park</collectingMunicipality>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="km" value="2.0">2 km</quantity>
road
<collectorName>Xom Bong</collectorName>
to
<collectorName>Headquarters</collectorName>
, primary forest, limestone outcrops in thick clayey soil;
<collectingDate value="1998-10-10">10 Oct. 1998</collectingDate>
;
<collectorName>J.J. Vermeulen</collectorName>
&amp;
<collectorName>L. Deharveng</collectorName>
leg.;
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/JJV17629">JJV 17629</accessionNumber>
(ex 6222) • 1 shell;
<collectingRegion country="Vietnam" name="Ninh Binh">Ninh Binh Province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:E972FC6A323A5C2AB3E96FF4072666AF:91C337837B6D48218C3015B47F5BB170" country="Vietnam" county="District" latitude="20.2935" longLatPrecision="12" longitude="105.6685" municipality="Cuc Phuong National Park" name="Cuc Phuong National Park" stateProvince="Hoa Binh Province">Cuc Phuong National Park</location>
,
<location LSID="urn:lsid:plazi:treatment:E972FC6A323A5C2AB3E96FF4072666AF:1A6EEC88A2409EBD257FC7CD2B48718B" country="Vietnam" county="District" latitude="20.2935" longLatPrecision="12" longitude="105.6685" municipality="Cuc Phuong National Park" name="Prehistoric Man Cave" stateProvince="Hoa Binh Province">Prehistoric Man Cave</location>
, steep limestone slope with disturbed forest;
<geoCoordinate degrees="20" direction="north" minutes="17.61" orientation="latitude" precision="9" value="20.2935">20°17.61'N</geoCoordinate>
,
<geoCoordinate degrees="105" direction="east" minutes="40.11" orientation="longitude" precision="9" value="105.6685">105°40.11'E</geoCoordinate>
;
<collectingDate value="1998-10-10">10 Oct. 1998</collectingDate>
;
<collectorName>J.J. Vermeulen</collectorName>
&amp;
<collectorName>L. Deharveng</collectorName>
leg.; JJV 6252
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
A medium-sized
<taxonomicName authorityName="Jochum, Slapnik &amp; Pall-Gergely" authorityYear="2014" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila</emphasis>
</taxonomicName>
species with a conical shell, an ovate aperture and a deeply-set, weak parietal tooth.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">Shell of normal size for the genus, slightly higher than wide, body whorl widest from standard apertural view; protoconch consists of 1.25-1.5 whorls, with fine spiral striation; specimens from the type locality with weaker, the shell from site 2020/28 with stronger and denser radial lines; typical shells with ca. 14, the shell from site 2020/28 with 19 spiral striae on body whorl from apertural view; whorls 4, rounded; aperture slightly oblique to shell axis from lateral view; umbilicus wide; aperture large, ovoid (higher than wide) to ovate-subquadrate; peristome slightly expanded, not reflected; parietal callus does not elevate from penultimate whorl; parietal tooth elongated, low, deeply-set (does not reach parietal callus).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Measurements (in mm).</paragraph>
<paragraph pageId="0" pageNumber="1">
H = 0.89-1.0, D = 0.81-0.95, H/D*100 = 105.3-111.6 (
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
= 7), RUD = 27.4-32.3 (
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
= 4).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
Generally larger than
<taxonomicName lsidName="A. elevata" pageId="0" pageNumber="1" rank="species" species="elevata">
<emphasis italics="true" pageId="0" pageNumber="1">A. elevata</emphasis>
</taxonomicName>
, has a rather conical shell (concave-conical in
<taxonomicName lsidName="A. elevata" pageId="0" pageNumber="1" rank="species" species="elevata">
<emphasis italics="true" pageId="0" pageNumber="1">A. elevata</emphasis>
</taxonomicName>
), a less pointed apex, a wider umbilicus, and a comparatively larger, ovate aperture (although
<taxonomicName lsidName="A. elevata" pageId="0" pageNumber="1" rank="species" species="elevata">
<emphasis italics="true" pageId="0" pageNumber="1">A. elevata</emphasis>
</taxonomicName>
samples 2020/18 and 2019/120 have a rather oval aperture) with a weak parietal tooth. In contrast,
<taxonomicName lsidName="A. elevata" pageId="0" pageNumber="1" rank="species" species="elevata">
<emphasis italics="true" pageId="0" pageNumber="1">A. elevata</emphasis>
</taxonomicName>
is always toothless. We note that the tooth-like thickening of the parietal callus in one
<taxonomicName lsidName="A. elevata" pageId="0" pageNumber="1" rank="species" species="elevata">
<emphasis italics="true" pageId="0" pageNumber="1">A. elevata</emphasis>
</taxonomicName>
shell (
<bibRefCitation DOI="https://doi.org/10.17109/AZH.63.3.327.2017" author="Pall-Gergely, B" journalOrPublisher="Acta Zoologica Academiae Scientiarum Hungaricae" pageId="0" pageNumber="1" pagination="143 - 157" refId="B41" refString="Pall-Gergely, B, Jochum, A, Asami, T, 2017. Three new species and a new genus of Hypselostomatidae from Cong Troi Cave, Northern Vietnam (Gastropoda: Pulmonata). Acta Zoologica Academiae Scientiarum Hungaricae 63 (2): 143 - 157, DOI: https://doi.org/10.17109/AZH.63.3.327.2017" title="Three new species and a new genus of Hypselostomatidae from Cong Troi Cave, Northern Vietnam (Gastropoda: Pulmonata)." url="https://doi.org/10.17109/AZH.63.3.327.2017" volume="63" year="2017">
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2017
</bibRefCitation>
: fig. 7F) is not considered to be a tooth. Moreover, although not in all
<taxonomicName lsidName="A. elevata" pageId="0" pageNumber="1" rank="species" species="elevata">
<emphasis italics="true" pageId="0" pageNumber="1">A. elevata</emphasis>
</taxonomicName>
populations, the radial sculpture of that species is more prominent. Populations assigned to
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2015" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila fabella" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="fabella">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila fabella</emphasis>
</taxonomicName>
are larger, possess a stronger parietal tooth, a comparatively smaller aperture and usually, a more pointed apex.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
The specific epithet refers to the ovoid aperture (
<emphasis italics="true" pageId="0" pageNumber="1">ᾠόν</emphasis>
= egg,
<emphasis italics="true" pageId="0" pageNumber="1">
<normalizedToken originalValue="στόμα">στόμα</normalizedToken>
</emphasis>
= mouth).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
This new species is known from two sites: the Cuc Phuong National Park (type locality) and another site ca. 40 km northwest (Fig.
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Distribution of Angustopila species." figureDoi="10.3897/zookeys.1147.93824.figure48" httpUri="https://binary.pensoft.net/fig/808857" pageId="0" pageNumber="1">48</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
The single shell collected at the Cave of Prehistoric Man at Cuc Phuong National Park (JJV 6252) lacks a parietal tooth, has a narrower umbilicus, and weaker sculpture than the other
<taxonomicName lsidName="A. oostoma" pageId="0" pageNumber="1" rank="species" species="oostoma">
<emphasis italics="true" pageId="0" pageNumber="1">A. oostoma</emphasis>
</taxonomicName>
sp. nov. shells. Thus, it may belong to a different species. We refer to it herein as
<taxonomicName lsidName="A. oostoma" pageId="0" pageNumber="1" rank="species" species="oostoma">A. cf. oostoma</taxonomicName>
sp. nov.
</paragraph>
<caption doi="10.3897/zookeys.1147.93824.figure51" httpUri="https://binary.pensoft.net/fig/808860" pageId="0" pageNumber="1" start="Figure 51" startId="F51">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 51.</emphasis>
<taxonomicName authorityName="Pall-Gergely &amp; Vermeulen" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila oostoma" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="oostoma">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila oostoma</emphasis>
</taxonomicName>
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Vermeulen, sp. nov. (holotype, HNHM 105289). Apertural (
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
), apical (
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
), lateral (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
) and ventral (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
) sides of the shell; eroded surface showing some microstructure on the protoconch (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
), aperture (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
), ventral (
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
) and frontal (
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
) surface of the body whorl.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1147.93824.figure52" httpUri="https://binary.pensoft.net/fig/808861" pageId="0" pageNumber="1" start="Figure 52" startId="F52">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 52.</emphasis>
<taxonomicName authorityName="Pall-Gergely &amp; Vermeulen" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila oostoma" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="oostoma">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila oostoma</emphasis>
</taxonomicName>
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Vermeulen, sp. nov. (2020/28). Apertural (
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
), ventral (
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
), apical (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
) and lateral (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
), sides of the shell; aperture (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
), microstructure of the protoconch showing protoconch-teleoconch boundary (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
), ventral (
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
) and frontal (
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
) surface of the body whorl.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>