288 lines
29 KiB
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288 lines
29 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.905.39592" ID-GBIF-Dataset="ee5a0817-7cfc-4c21-ba7d-3beabb06cccd" ID-PMC="PMC7004242" ID-Pensoft-Pub="1313-2970-905-1" ID-Pensoft-UUID="4C7E0CB428DF56BDB15B7AC8D6707961" ID-PubMed="32063726" ID-ZooBank="F4C766E4633A41039FFDE952718F41FB" ModsDocID="1313-2970-905-1" checkinTime="1579539902851" checkinUser="pensoft" docAuthor="Salata, Sebastian & Fisher, Brian L." docDate="2020" docId="A60C37DFF157168749FF30CA92792D92" docLanguage="en" docName="ZooKeys 905: 1-235" docOrigin="ZooKeys 905" docSource="http://dx.doi.org/10.3897/zookeys.905.39592" docTitle="Pheidole ocypodea Salata & Fisher 2020, sp. nov." docType="treatment" docUuid="C9F6D256-6F79-4517-90A2-FFA380105D9D" docUuidSource="ZooBank" docVersion="5" id="4C7E0CB428DF56BDB15B7AC8D6707961" lastPageNumber="1" masterDocId="4C7E0CB428DF56BDB15B7AC8D6707961" masterDocTitle="Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups" masterLastPageNumber="235" masterPageNumber="1" pageNumber="1" updateTime="1668168337901" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Salata, Sebastian</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fisher, Brian L.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>905</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>235</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.905.39592</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.905.39592</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-905-1</mods:identifier>
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<mods:identifier type="ZooBank">F4C766E4633A41039FFDE952718F41FB</mods:identifier>
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<mods:identifier type="Pensoft-UUID">4C7E0CB428DF56BDB15B7AC8D6707961</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="161777027" LSID="urn:lsid:zoobank.org:act:C9F6D256-6F79-4517-90A2-FFA380105D9D" httpUri="http://treatment.plazi.org/id/A60C37DFF157168749FF30CA92792D92" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/C9F6D256-6F79-4517-90A2-FFA380105D9D" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole ocypodea" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ocypodea" status="sp. nov.">Pheidole ocypodea</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Pheidole ocypodea sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923220) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0304390)." figureDoi="10.3897/zookeys.905.39592.figure32" httpUri="https://binary.pensoft.net/fig/373079" pageId="0" pageNumber="1">Figs 32A-F</figureCitation>
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<figureCitation captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Hypostomal teeth. Pheidole madinika sp. nov. (A). P. mahaboensis sp. nov. (B). P. maizina sp. nov. (C). P. makaensis sp. nov. (D). P. makirovana sp. nov. (E). P. manantenensis sp. nov. (F). P. mantadia sp. nov. (G). P. marieannae sp. nov. (H). P. masoala sp. nov. (I). P. mavesatra sp. nov. (J). P. miramila sp. nov. (K). P. moramanaensis sp. nov. (L). P. navoatrensis sp. nov. (M). P. nemoralis Forel (N). P. ocypodea sp. nov. (O). P. parviocula sp. nov. (P). P. petax Forel (Q). P. podargea sp. nov. (R). P. praegrandis sp. nov. (S). P. ranohirensis sp. nov. (T). P. rugocephala sp. nov. (U). P. rugofitarata sp. nov. (V). P. scabrata Forel (W). P. typhlos sp. nov. (X). P. vatovavensis sp. nov. (Y). P. voasara sp. nov. (Z). P. vohemarensis sp. nov. (AA). P. zavamanira sp. nov. (AB)." figureDoi="10.3897/zookeys.905.39592.figure85" httpUri="https://binary.pensoft.net/fig/373132" pageId="0" pageNumber="1">, 85O</figureCitation>
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<figureCitation captionStart="Figure 87" captionStartId="F87" captionText="Figure 87. Distribution. Pheidole lamperos sp. nov. (A). P. longipilosa sp. nov. (B). P. longispinosa Forel (C). P. lutea sp. nov. (D). P. madinika sp. nov. (E). P. mahaboensis sp. nov. (F). P. maizina sp. nov. (G). P. makaensis sp. nov. (H). P. makirovana sp. nov. (I). P. manantenensis sp. nov. (J). P. mantadia sp. nov. (K). P. marieannae sp. nov. (L). P. masoala sp. nov. (M). P. mavesatra sp. nov. (N). P. miramila sp. nov. (O). P. moramanaensis sp. nov. (P). P. navoatrensis sp. nov. (R). P. nemoralis Forel (S). P. ocypodea sp. nov. (T). P. parviocula sp. nov. (U)." figureDoi="10.3897/zookeys.905.39592.figure87" httpUri="https://binary.pensoft.net/fig/373134" pageId="0" pageNumber="1">, 87T</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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.
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</emphasis>
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Madagascar. •1 major worker; Antsiranana; Galoko chain, Mont Galoko;
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<geoCoordinate degrees="13.5888" direction="south" orientation="latitude" precision="5" value="-13.5888">-13.5888</geoCoordinate>
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,
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<geoCoordinate degrees="48.72864" direction="east" orientation="longitude" precision="1" value="48.72864">48.72864</geoCoordinate>
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; alt. 980 m; 20 Feb 2013; B.L. Fisher et al. leg.; BLF30960, CASENT0304390 (CASC).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Paratype</emphasis>
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.
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</emphasis>
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Madagascar. •1 minor worker; same data as for holotype; CASENT0923220 (CASC).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Other material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Madagascar. -
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Antsiranana</emphasis>
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: •8w., 11s.; R.S. Manongarivo, 10.8 km 229°SW Antanambao;
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<geoCoordinate degrees="13.96167" direction="south" orientation="latitude" precision="1" value="-13.96167">-13.96167</geoCoordinate>
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,
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<geoCoordinate degrees="48.43333" direction="east" orientation="longitude" precision="1" value="48.43333">48.43333</geoCoordinate>
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; alt. 400 m; 8 Nov 1998; B.L. Fisher leg.; CASENT0198625, CASENT0198626, CASENT0198834, CASENT0198836, CASENT0846551-CASENT0846562 (CASC). •32w., 24s.; R.S. Manongarivo, 12.8 km 228°SW Antanambao;
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<geoCoordinate degrees="13.97667" direction="south" orientation="latitude" precision="1" value="-13.97667">-13.97667</geoCoordinate>
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,
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<geoCoordinate degrees="48.42333" direction="east" orientation="longitude" precision="1" value="48.42333">48.42333</geoCoordinate>
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; alt. 780 m; 11 Sep 1998; B.L. Fisher leg.; CASENT0196883, CASENT0198627, CASENT0198837, CASENT0846501-CASENT0846550 (CASC).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
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.
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</emphasis>
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Body size moderate: HL: 1.82-2.01 (1.95); HW: 1.52-1.66 (1.6), WL: 1.16-1.31 (1.26); propodeal spines very long (PSL: 0.38-0.45 (0.41)); head in full-face view rectangular, with lateral sides relatively straight, only their posteriormost part slightly convex; sides of the head with sparse, relatively long, erect pilosity; occipital lobes shiny, with sparse and thick rugoreticulation; inner hypostomal teeth distinct, low, closely spaced, triangular, with rounded apex directed outward; outer hypostomal teeth distinct, low, lobe-like, with base wide and tops directed inward; inner and outer teeth closely spaced and connected by concavity.
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
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.
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</emphasis>
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Body size moderate: HL: 0.63-0.72 (0.66); HW: 0.6-0.7 (0.64), WL: 0.8-0.93 (0.85); propodeal spines very long (PSL: 0.26-0.32 (0.28)); scape, when laid back, surpassing posterior head margin by two-fifths of its length; lateral sides of head and malar area smooth and shiny or with indistinct, sparse rugulae, sculpture weakening posteriorly; vertex, genae, and frons smooth; pronotum, dorsal surface of mesonotum, and dorsal and posterior surface of propodeum smooth; katepisternum, anepisternum, and lateral sides of propodeum with thick and sparse rugae.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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Measurements (
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<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 10): HL: 1.82-2.01 (1.95); HW: 1.52-1.66 (1.6); SL: 0.71-0.8 (0.75); EL: 0.15-0.19 (0.17); WL: 1.16-1.31 (1.26); PSL: 0.38-0.45 (0.41); MTL: 0.69-0.79 (0.75); PNW: 0.68-0.78 (0.75); PTW: 0.18-0.25 (0.21); PPW: 0.62-0.73 (0.65); CI: 79.8-84.5 (82.3); SI: 44.5-50.1 (46.8); PSLI: 20.3-22.7 (21.3); PPI: 28.0-37.7 (32.4); PNI: 44.7-48.9 (46.6); MTI: 44.5-48.3 (46.5).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
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.
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</emphasis>
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In full-face view rectangular, with lateral sides relatively straight, only their posteriormost part slightly convex (Fig.
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<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Pheidole ocypodea sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923220) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0304390)." figureDoi="10.3897/zookeys.905.39592.figure32" httpUri="https://binary.pensoft.net/fig/373079" pageId="0" pageNumber="1">32B</figureCitation>
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). In lateral view oval; ventral and dorsal faces convex; inner hypostomal teeth visible. Sides of the head with sparse, relatively long, erect pilosity; whole head with moderately dense, relatively long, erect pilosity. Antennal scrobes indistinct and not delimited by carinulae. Occipital lobes shiny, with sparse and thick rugoreticulation; genae smooth and shiny, only anterior part sometimes with sparse and fine rugulae; frons with longitudinal and sparse rugae, on posterior part of frons rugae directed outward and more irregular, interspaces with sparse to moderately sparse rugose foveolae; malar area and lateral sides of head shiny, with longitudinal rugae, surface between rugae with dense rugose foveolae. Centre of clypeus smooth and shiny, lateral sides with longitudinal rugae; median notch present, moderately wide, and narrow; median longitudinal carina present; lateral longitudinal carinae absent. Scape, when laid back, reaching midlength of head; pilosity suberect to erect (Fig.
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<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Pheidole ocypodea sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923220) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0304390)." figureDoi="10.3897/zookeys.905.39592.figure32" httpUri="https://binary.pensoft.net/fig/373079" pageId="0" pageNumber="1">32B, D</figureCitation>
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). Inner hypostomal teeth distinct, low, closely spaced, triangular, with rounded apex directed outward; outer hypostomal teeth distinct, low, lobe-like, with base wide and tops directed inward; inner and outer teeth closely spaced and connected by concavity (Fig.
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<figureCitation captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Hypostomal teeth. Pheidole madinika sp. nov. (A). P. mahaboensis sp. nov. (B). P. maizina sp. nov. (C). P. makaensis sp. nov. (D). P. makirovana sp. nov. (E). P. manantenensis sp. nov. (F). P. mantadia sp. nov. (G). P. marieannae sp. nov. (H). P. masoala sp. nov. (I). P. mavesatra sp. nov. (J). P. miramila sp. nov. (K). P. moramanaensis sp. nov. (L). P. navoatrensis sp. nov. (M). P. nemoralis Forel (N). P. ocypodea sp. nov. (O). P. parviocula sp. nov. (P). P. petax Forel (Q). P. podargea sp. nov. (R). P. praegrandis sp. nov. (S). P. ranohirensis sp. nov. (T). P. rugocephala sp. nov. (U). P. rugofitarata sp. nov. (V). P. scabrata Forel (W). P. typhlos sp. nov. (X). P. vatovavensis sp. nov. (Y). P. voasara sp. nov. (Z). P. vohemarensis sp. nov. (AA). P. zavamanira sp. nov. (AB)." figureDoi="10.3897/zookeys.905.39592.figure85" httpUri="https://binary.pensoft.net/fig/373132" pageId="0" pageNumber="1">85O</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
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.
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</emphasis>
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In lateral view, promesonotum short, angular, and high, with relatively steep posterior declivity; promesonotal groove absent; metanotal groove absent; propodeal spines very long, narrow, with acute apex; humeral area laterally absent to weakly produced (Fig.
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<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Pheidole ocypodea sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923220) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0304390)." figureDoi="10.3897/zookeys.905.39592.figure32" httpUri="https://binary.pensoft.net/fig/373079" pageId="0" pageNumber="1">32D</figureCitation>
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). Surface shiny, pronotum and dorsal surface of mesonotum smooth or sometimes with indistinct, sparse rugulae; propodeum, katepisternum, and anepisternum with thick and dense rugoreticulation. Pilosity sparse, long, and erect (Fig.
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<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Pheidole ocypodea sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923220) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0304390)." figureDoi="10.3897/zookeys.905.39592.figure32" httpUri="https://binary.pensoft.net/fig/373079" pageId="0" pageNumber="1">32D, F</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
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.
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</emphasis>
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Shagreened; peduncle relatively long, without horizontal lobes on its basal part; node triangular and thin, with rounded apex, in rear view node dorsoventrally slightly depressed; pilosity moderately sparse and erect (Fig.
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<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Pheidole ocypodea sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923220) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0304390)." figureDoi="10.3897/zookeys.905.39592.figure32" httpUri="https://binary.pensoft.net/fig/373079" pageId="0" pageNumber="1">32D, F</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
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.
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</emphasis>
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Shagreened; in dorsal view sides with acute, horn-like, moderately long projections; pilosity long, moderately sparse, and erect (Fig.
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<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Pheidole ocypodea sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923220) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0304390)." figureDoi="10.3897/zookeys.905.39592.figure32" httpUri="https://binary.pensoft.net/fig/373079" pageId="0" pageNumber="1">32D, F</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
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.
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</emphasis>
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First gastral tergite shagreened, at least on its basal part; pilosity moderately dense, long, and erect (Fig.
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<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Pheidole ocypodea sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923220) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0304390)." figureDoi="10.3897/zookeys.905.39592.figure32" httpUri="https://binary.pensoft.net/fig/373079" pageId="0" pageNumber="1">32D, F</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
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.
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</emphasis>
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Unicolourous, ocherous to reddish-brown (Fig.
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<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Pheidole ocypodea sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923220) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0304390)." figureDoi="10.3897/zookeys.905.39592.figure32" httpUri="https://binary.pensoft.net/fig/373079" pageId="0" pageNumber="1">32D, F</figureCitation>
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).
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</paragraph>
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<caption doi="10.3897/zookeys.905.39592.figure32" httpUri="https://binary.pensoft.net/fig/373079" pageId="0" pageNumber="1" start="Figure 32" startId="F32">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 32.</emphasis>
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<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole ocypodea" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ocypodea">
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<emphasis italics="true" pageId="0" pageNumber="1">Pheidole ocypodea</emphasis>
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</taxonomicName>
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sp. nov., full-face view (
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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), profile (
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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), and dorsal view (
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<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
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) of paratype minor worker (CASENT0923220) and full-face view (
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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), profile (
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<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
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), and dorsal view (
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<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
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) of holotype major worker (CASENT0304390).
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</paragraph>
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</caption>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
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Measurements (
|
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<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
|
|
= 10): HL: 0.63-0.72 (0.66); HW: 0.6-0.7 (0.64); SL: 0.63-0.69 (0.66); EL: 0.09-0.13 (0.11); WL: 0.8-0.93 (0.85); PSL: 0.26-0.32 (0.28); MTL: 0.49-0.57 (0.53); PNW: 0.38-0.45 (0.42); PTW: 0.09-0.12 (0.1); PPW: 0.18-0.24 (0.19); CI: 93.6-98.2 (96.3); SI: 98.3-106.9 (102.9); PSLI: 40.4-45.5 (42.6); PPI: 45.1-57.6 (52.4); PNI: 62.5-68.1 (65.2); MTI: 78.9-86.9 (83.3).
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
|
|
.
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|
</emphasis>
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|
In full-face view square, posterior of eyes slightly convex, anterior of eyes relatively straight, occipital margin straight or indistinctly convex; occipital carina indistinct, weakly developed (Fig.
|
|
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Pheidole ocypodea sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923220) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0304390)." figureDoi="10.3897/zookeys.905.39592.figure32" httpUri="https://binary.pensoft.net/fig/373079" pageId="0" pageNumber="1">32A</figureCitation>
|
|
). Pilosity sparse, long, and erect. Lateral sides of head and malar area smooth and shiny or with indistinct, sparse rugulae, sculpture weakening posteriorly; vertex, genae, and frons smooth; antennal sockets with sparse, interrupted carinae curved outward. Clypeus with median longitudinal carina present; two lateral longitudinal carinae present. Scape, when laid back, surpassing posterior head margin by two-fifths of its length; pilosity suberect to erect (Fig.
|
|
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Pheidole ocypodea sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923220) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0304390)." figureDoi="10.3897/zookeys.905.39592.figure32" httpUri="https://binary.pensoft.net/fig/373079" pageId="0" pageNumber="1">32A, C</figureCitation>
|
|
).
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<emphasis bold="true" pageId="0" pageNumber="1">
|
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<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
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|
.
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|
</emphasis>
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|
In lateral view, promesonotum high, short, and convex; promesonotal groove absent; metanotal groove shallow and indistinct; propodeal spines very long, massive basally, with acute apex (Fig.
|
|
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Pheidole ocypodea sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923220) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0304390)." figureDoi="10.3897/zookeys.905.39592.figure32" httpUri="https://binary.pensoft.net/fig/373079" pageId="0" pageNumber="1">32C</figureCitation>
|
|
). Surface shiny, pronotum, dorsal surface of mesonotum, and dorsal and posterior surface of propodeum smooth; katepisternum, anepisternum, and lateral sides of propodeum with thick and sparse rugae. Pilosity sparse, long, and erect (Fig.
|
|
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Pheidole ocypodea sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923220) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0304390)." figureDoi="10.3897/zookeys.905.39592.figure32" httpUri="https://binary.pensoft.net/fig/373079" pageId="0" pageNumber="1">32C, E</figureCitation>
|
|
).
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
|
.
|
|
</emphasis>
|
|
Peduncle moderately long and thin; node low, bulge-like; with few long, erect setae (Fig.
|
|
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Pheidole ocypodea sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923220) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0304390)." figureDoi="10.3897/zookeys.905.39592.figure32" httpUri="https://binary.pensoft.net/fig/373079" pageId="0" pageNumber="1">32C, E</figureCitation>
|
|
).
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
|
|
.
|
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</emphasis>
|
|
Long, low, and slightly convex; with few long, erect setae at the anterior edge (Fig.
|
|
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Pheidole ocypodea sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923220) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0304390)." figureDoi="10.3897/zookeys.905.39592.figure32" httpUri="https://binary.pensoft.net/fig/373079" pageId="0" pageNumber="1">32C, E</figureCitation>
|
|
).
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
|
.
|
|
</emphasis>
|
|
Pilosity sparse and erect (Fig.
|
|
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Pheidole ocypodea sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923220) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0304390)." figureDoi="10.3897/zookeys.905.39592.figure32" httpUri="https://binary.pensoft.net/fig/373079" pageId="0" pageNumber="1">32C, E</figureCitation>
|
|
).
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
|
.
|
|
</emphasis>
|
|
Unicolourous, bright brown to brown (Fig.
|
|
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Pheidole ocypodea sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923220) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0304390)." figureDoi="10.3897/zookeys.905.39592.figure32" httpUri="https://binary.pensoft.net/fig/373079" pageId="0" pageNumber="1">32C, E</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="etymology">
|
|
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">Named after Ocypode, a harpy from Greek mythology, in reference to the long and sharp propodeal spines of minor workers reminiscent of claws.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
|
|
<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">The species was collected between 400-980 m in elevation, in rainforest and montane rainforest. Nests were located in in rotten logs and rotten sticks on ground.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="comments">
|
|
<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
This species is most similar to
|
|
<taxonomicName lsidName="P. aelloea" pageId="0" pageNumber="1" rank="species" species="aelloea">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. aelloea</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. and
|
|
<taxonomicName lsidName="P. podargea" pageId="0" pageNumber="1" rank="species" species="podargea">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. podargea</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
|
|
.
|
|
</emphasis>
|
|
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole ocypodea" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ocypodea">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole ocypodea</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. can be distinguished from
|
|
<taxonomicName lsidName="P. aelloea" pageId="0" pageNumber="1" rank="species" species="aelloea">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. aelloea</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. by steep to relatively steep posterior declivity of promesonotum, inner hypostomal teeth pointed outward and outer hypostomal teeth pointed inward; from
|
|
<taxonomicName lsidName="P. podargea" pageId="0" pageNumber="1" rank="species" species="podargea">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. podargea</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. it differs in longer and sparser pilosity on sides of head, reduced to absent sculpture on genae and propodeum, and inner hypostomal teeth pointed outward.
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
|
|
.
|
|
</emphasis>
|
|
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole ocypodea" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ocypodea">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole ocypodea</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. can be distinguished from
|
|
<taxonomicName lsidName="P. aelloea" pageId="0" pageNumber="1" rank="species" species="aelloea">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. aelloea</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. by smooth lateral sides of head and malar area, presence of metanotal groove, short petiolar peduncle, and long postpetiole which is approximately 1.5 times longer than high; from
|
|
<taxonomicName lsidName="P. podargea" pageId="0" pageNumber="1" rank="species" species="podargea">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. podargea</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. it differs in smooth sculpture of vertex, genae, and area between frontal carinae, absence of promesonotal groove and at least partially smooth surface of pronotum, dorsal surface of mesonotum, and dorsal and posterior surface of propodeum.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |