518 lines
74 KiB
XML
518 lines
74 KiB
XML
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<mods:title id="558F3501C74A34B2B048FA9ACC6F3AD4">A new species of Cristarmadillidium from central Spain with remarks on Estenarmadillidium and Iberiarmadillidium (Isopoda: Oniscidea: Armadillidiidae)</mods:title>
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<mods:namePart id="3E498D9640BE8132D0F9B98B2A9D7A25">Garcia, Lluc</mods:namePart>
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<mods:title id="E43AD136468C664B578AFEFEEE497960">Zootaxa</mods:title>
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<mods:part id="447F77650F45EF917EB9CC9B53F8A5D8">
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<mods:date id="C6A967FD94E090EE74BCBA829482DF1A">2023</mods:date>
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<mods:number id="06A7C014C5A19A595A2758CEF3F94B01">2023-08-01</mods:number>
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<mods:identifier id="3C6F10783B5DB853615BE80932180C07" type="DOI">10.11646/zootaxa.5323.2.8</mods:identifier>
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<treatment id="9B0A98020466FFDCFF6F9D2E0D9F4471" ID-DOI="http://doi.org/10.5281/zenodo.8204298" ID-GBIF-Taxon="212424055" ID-Zenodo-Dep="8204298" LSID="urn:lsid:zoobank.org:act:6236E7C0-6334-4091-9893-10C49459F805" httpUri="http://treatment.plazi.org/id/9B0A98020466FFDCFF6F9D2E0D9F4471" lastPageId="6" lastPageNumber="291" pageId="1" pageNumber="286">
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<subSubSection id="5BB97A9F0466FFDBFF6F9D2E0D334377" box="[151,623,793,820]" pageId="1" pageNumber="286" type="nomenclature">
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<heading id="48549E780466FFDBFF6F9D2E0D334377" bold="true" box="[151,623,793,820]" fontSize="11" level="1" pageId="1" pageNumber="286" reason="1">
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<emphasis id="21D7F5060466FFDBFF6F9D2E0D334377" bold="true" box="[151,623,793,820]" pageId="1" pageNumber="286">
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<taxonomicName id="D4A352970466FFDBFF6F9D2E0D564370" ID-CoL="f01420a4-57d0-4c93-9ce9-8efedb361ef3" authority="Garcia, 2023" authorityName="Garcia" authorityYear="2023" box="[151,522,793,820]" class="Malacostraca" family="Armadillidiidae" genus="Cristarmadillidium" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="286" phylum="Arthropoda" rank="species" species="cabanillasi" status="sp. nov.">
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<emphasis id="21D7F5060466FFDBFF6F9D2E0D564370" bold="true" box="[151,522,793,820]" italics="true" pageId="1" pageNumber="286">Cristarmadillidium cabanillasi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="3AE4487D0466FFDBFDE99D2E0D334377" box="[529,623,793,819]" pageId="1" pageNumber="286" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="5BB97A9F0466FFDBFF6F9D0A0C044332" pageId="1" pageNumber="286" type="description">
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<paragraph id="131C29140466FFDBFF6F9D0A0E414313" blockId="1.[151,856,793,886]" box="[151,285,829,855]" pageId="1" pageNumber="286">
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<figureCitation id="8B9835910466FFDBFF6F9D0A0E414313" box="[151,285,829,855]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="1.[152,255,1893,1917]" captionStartId-1="2.[152,255,1908,1932]" captionStartId-2="3.[152,255,1943,1967]" captionStartId-3="4.[152,255,1943,1967]" captionTargetBox-0="[151,1436,1196,1867]" captionTargetBox-1="[208,1378,182,1880]" captionTargetBox-2="[187,1401,185,1914]" captionTargetBox-3="[193,1394,185,1914]" captionTargetId-0="figure-180@1.[151,1436,1196,1867]" captionTargetId-1="figure-29@2.[204,1383,181,1885]" captionTargetId-2="figure-15@3.[182,1405,181,1919]" captionTargetId-3="figure-29@4.[189,1398,181,1919]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionText-0="FIGURE 1. Cristarmadillidium cabanillasi sp. nov. Habitus of an alive specimen. A, conglobate position. B, walking on the ground. Photos: David Cabanillas-Roldán." captionText-1="FIGURE 2. Cristarmadillidium cabanillasi sp. nov. Male paratype (CLLG 1071). A, cephalon and pereonites 1–7, transversal view (from the front). B, left nodulus lateralis, second tubercle of the posterior row of pereonites. C, cephalothorax and first pereon-tergite, dorsal view. D, first antenna. E, second antenna. F, pleon, pleotelson and uropods, dorsal view. G, left uropod, ventral view. Scale bars: C, 1 mm; D, 0.1 mm; E, 0.5 mm; F, 1 mm; G, 0.3 mm." captionText-2="FIGURE 3. Cristarmadillidium cabanillasi sp. nov. Male paratype (CLLG 1071). A, right mandible. B, left mandible. C, first maxilla. D, second maxilla. E, maxilliped; F, pereopod 1; G, pereopod 7. Scale bars: A–B, 0.3 mm; C–E, 0.1 mm; F–G, 0.5 mm." captionText-3="FIGURE 4. Cristarmadillidium cabanillasi sp. nov. Male paratype (CLLG 1071). A, pleopod 1, ventral view. B, pleopod 2, ventral view. C, genital papilla. D, pleopod 3 exopod, ventral view. E, pleopod 4 exopod, ventral view. F, pleopod 5 exopod, ventral view. Scale bars: A–B, 0.3 mm; C, 0.2 mm; D–F, 0.3 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8204286" figureDoi-1="http://doi.org/10.5281/zenodo.8204288" figureDoi-2="http://doi.org/10.5281/zenodo.8204292" figureDoi-3="http://doi.org/10.5281/zenodo.8204294" httpUri-0="https://zenodo.org/record/8204286/files/figure.png" httpUri-1="https://zenodo.org/record/8204288/files/figure.png" httpUri-2="https://zenodo.org/record/8204292/files/figure.png" httpUri-3="https://zenodo.org/record/8204294/files/figure.png" pageId="1" pageNumber="286">Figures 1–4</figureCitation>
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</paragraph>
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<paragraph id="131C29140466FFDBFF6F9D690C044332" blockId="1.[151,856,793,886]" box="[151,856,862,886]" pageId="1" pageNumber="286">
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<uri id="673225160466FFDBFF6F9D690C044332" box="[151,856,862,886]" pageId="1" pageNumber="286">
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urn:lsid:zoobank.org:act:
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<subSubSection id="5BB97A9F0466FFDBFF6F9D930DDA44D3" pageId="1" pageNumber="286" type="materials_examined">
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<paragraph id="131C29140466FFDBFF6F9D930DDA44D3" blockId="1.[151,1436,932,1175]" pageId="1" pageNumber="286">
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<emphasis id="21D7F5060466FFDBFF6F9D930EDD43FA" bold="true" box="[151,385,932,958]" pageId="1" pageNumber="286">Material examined.</emphasis>
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<materialsCitation id="A3CB23490466FFDBFE719D930ED043A6" ID-GBIF-Occurrence="4168023306" collectingDate="2020-12-20" collectionCode="MBCN" collectorName="David Cabanillas-Roldan" country="Spain" location="Madrid" pageId="1" pageNumber="286" specimenCode="MBCN 26801" specimenCount="1" specimenCount-female="1" stateProvince="Madrid" typeStatus="holotype">
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<emphasis id="21D7F5060466FFDBFE719D930EA543FA" box="[393,505,932,958]" italics="true" pageId="1" pageNumber="286">
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<typeStatus id="CC1897B60466FFDBFE719D930EAF43FA" box="[393,499,932,958]" pageId="1" pageNumber="286" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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<specimenCount id="05A5E29D0466FFDBFDF99D920D4A43FB" box="[513,534,933,959]" pageId="1" pageNumber="286" type="female">♀</specimenCount>
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, Cadalso de los Vidrios,
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<collectingRegion id="D167E7F60466FFDBFCC99D930CDA43FA" box="[817,902,932,958]" country="Spain" name="Madrid" pageId="1" pageNumber="286">Madrid</collectingRegion>
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,
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<collectingCountry id="6BB469840466FFDBFC6B9D930C8A43FA" box="[915,982,932,958]" name="Spain" pageId="1" pageNumber="286">Spain</collectingCountry>
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, leg.
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||
<collectorName id="BE564CC20466FFDBFBEA9D930A6D43FA" box="[1042,1329,932,958]" pageId="1" pageNumber="286">David Cabanillas-Roldán</collectorName>
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,
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<date id="671D0FD40466FFDBFAC59D930F9343A6" pageId="1" pageNumber="286" value="2020-12-20">
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||
<collectingDate id="7759F63C0466FFDBFAC59D930F9343A6" pageId="1" pageNumber="286" value="2020-12-20">20–XII– 2020</collectingDate>
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||
</date>
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||
(
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||
<specimenCode id="4305816F0466FFDBFF269DFF0ED843A6" box="[222,388,968,994]" collectionCode="MBCN" pageId="1" pageNumber="286">MBCN 26801</specimenCode>
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||
)
|
||
</materialsCitation>
|
||
.
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||
<materialsCitation id="A3CB23490466FFDBFE6F9DFE0DD84442" ID-GBIF-Occurrence="4168023308" collectingDate="2020-12-20" collectionCode="MNCN" collectorName="David Cabanillas-Roldan" country="Spain" location="Madrid" pageId="1" pageNumber="286" specimenCode="MNCN 20.04" specimenCount="5" specimenCount-female="5" stateProvince="Madrid" typeStatus="paratype">
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||
<emphasis id="21D7F5060466FFDBFE6F9DFE0D4D43A6" box="[407,529,969,994]" italics="true" pageId="1" pageNumber="286">
|
||
<typeStatus id="CC1897B60466FFDBFE6F9DFE0D5043A6" box="[407,524,969,994]" pageId="1" pageNumber="286" type="paratype">Paratypes</typeStatus>
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||
:
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||
</emphasis>
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||
2 ♁,
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||
<specimenCount id="05A5E29D0466FFDBFDA99DFF0D2743A7" box="[593,635,968,995]" pageId="1" pageNumber="286" type="female">5 ♀</specimenCount>
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||
, same locality as the Holotype, leg.
|
||
<collectorName id="BE564CC20466FFDBFBEB9DFF0A6E43A6" box="[1043,1330,968,994]" pageId="1" pageNumber="286">David Cabanillas-Roldán</collectorName>
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||
,
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||
<date id="671D0FD40466FFDBFAC59DFF0F934442" pageId="1" pageNumber="286" value="2020-12-20">
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||
<collectingDate id="7759F63C0466FFDBFAC59DFF0F934442" pageId="1" pageNumber="286" value="2020-12-20">20–XII– 2020</collectingDate>
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||
</date>
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||
(
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||
<specimenCode id="4305816F0466FFDBFF259DDB0E254442" box="[221,377,1004,1030]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="1" pageNumber="286" type="Museum">MNCN 20.04</specimenCode>
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/20710 to 20.04/20716)
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||
</materialsCitation>
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||
.
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||
<materialsCitation id="A3CB23490466FFDBFD759DDB0AC5446E" ID-GBIF-Occurrence="4168023304" collectingDate="2020-12-20" collectionCode="CLLG" collectorName="David Cabanillas-Roldan" country="Spain" location="Madrid" pageId="1" pageNumber="286" specimenCode="CLLG 1071" specimenCount="1" stateProvince="Madrid" typeStatus="paratype">
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1 ♁ (body parts of the dissected specimen, mounted in slides or stored in vials, as appropriate), same locality as the Holotype, leg.
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||
<collectorName id="BE564CC20466FFDBFCD79A270B10446E" box="[815,1100,1040,1066]" pageId="1" pageNumber="286">David Cabanillas-Roldán</collectorName>
|
||
,
|
||
<date id="671D0FD40466FFDBFBAF9A270BAD446E" box="[1111,1265,1040,1066]" pageId="1" pageNumber="286" value="2020-12-20">
|
||
<collectingDate id="7759F63C0466FFDBFBAF9A270BAD446E" box="[1111,1265,1040,1066]" pageId="1" pageNumber="286" value="2020-12-20">20–XII–2020</collectingDate>
|
||
</date>
|
||
, (
|
||
<specimenCode id="4305816F0466FFDBFAFB9A270ACC446E" box="[1283,1424,1040,1066]" collectionCode="CLLG" pageId="1" pageNumber="286">CLLG 1071</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="A3CB23490466FFDBFF6F9A030A37440A" ID-GBIF-Occurrence="4168023305" box="[151,1387,1076,1103]" collectingDate="2020-11-01" collectionCode="MBCN" collectorName="David Cabanillas-Roldan" country="Spain" location="Villanueva del Pardillo" pageId="1" pageNumber="286" specimenCode="MBCN 26802-26803" specimenCount="2" specimenCount-female="2" stateProvince="Madrid" typeStatus="paratype">
|
||
<specimenCount id="05A5E29D0466FFDBFF6F9A030F9C440B" box="[151,192,1076,1103]" pageId="1" pageNumber="286" type="female">2 ♀</specimenCount>
|
||
,
|
||
<location id="167C7FCF0466FFDBFF349A030E8C440A" LSID="urn:lsid:plazi:treatment:9B0A98020466FFDCFF6F9D2E0D9F4471:167C7FCF0466FFDBFF349A030E8C440A" box="[204,464,1076,1102]" country="Spain" name="Villanueva del Pardillo" pageId="1" pageNumber="286" stateProvince="Madrid">Villanueva del Pardillo</location>
|
||
,
|
||
<collectingRegion id="D167E7F60466FFDBFE219A030D72440A" box="[473,558,1076,1102]" country="Spain" name="Madrid" pageId="1" pageNumber="286">Madrid</collectingRegion>
|
||
,
|
||
<collectingCountry id="6BB469840466FFDBFDC19A030D20440A" box="[569,636,1076,1102]" name="Spain" pageId="1" pageNumber="286">Spain</collectingCountry>
|
||
, leg.
|
||
<collectorName id="BE564CC20466FFDBFD4C9A030C8E440A" box="[692,978,1076,1102]" pageId="1" pageNumber="286">David Cabanillas-Roldán</collectorName>
|
||
,
|
||
<date id="671D0FD40466FFDBFC249A030B07440A" box="[988,1115,1076,1102]" pageId="1" pageNumber="286" value="2020-11-01">
|
||
<collectingDate id="7759F63C0466FFDBFC249A030B07440A" box="[988,1115,1076,1102]" pageId="1" pageNumber="286" value="2020-11-01">1–XI–2020</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCode id="4305816F0466FFDBFB919A030A3F440A" box="[1129,1379,1076,1102]" collectionCode="MBCN" pageId="1" pageNumber="286">MBCN 26802–26803</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="A3CB23490466FFDBFA8B9A030EE84436" ID-GBIF-Occurrence="4168023302" collectionCode="MBCN" country="Spain" pageId="1" pageNumber="286" specimenCode="MBCN 26804, 26807" specimenCount="1" stateProvince="Madrid" typeStatus="paratype">
|
||
4 ♁ (
|
||
<specimenCode id="4305816F0466FFDBFF679A6F0E634436" box="[159,319,1112,1138]" collectionCode="MBCN" pageId="1" pageNumber="286">MBCN 26804</specimenCode>
|
||
to
|
||
<specimenCode id="4305816F0466FFDBFE9A9A6F0EF04436" box="[354,428,1112,1138]" collectionCode="MBCN" pageId="1" pageNumber="286">26807</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="A3CB23490466FFDBFE479A6F0DAB4436" ID-GBIF-Occurrence="4168023301" box="[447,759,1112,1139]" collectionCode="MBCN" country="Spain" pageId="1" pageNumber="286" specimenCode="MBCN 26808-26809" specimenCount="2" specimenCount-female="2" stateProvince="Madrid" typeStatus="paratype">
|
||
<specimenCount id="05A5E29D0466FFDBFE479A6F0EB44437" box="[447,488,1112,1139]" pageId="1" pageNumber="286" type="female">2 ♀</specimenCount>
|
||
(
|
||
<specimenCode id="4305816F0466FFDBFE0E9A6F0DB34436" box="[502,751,1112,1138]" collectionCode="MBCN" pageId="1" pageNumber="286">MBCN 26808–26809</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="A3CB23490466FFDBFCF99A6F0B604436" ID-GBIF-Occurrence="4168023303" box="[769,1084,1112,1138]" collectionCode="MBCN" country="Spain" pageId="1" pageNumber="286" specimenCode="MBCN 26810" specimenCount="1" specimenCount-immature="1" stateProvince="Madrid" typeStatus="paratype">
|
||
<specimenCount id="05A5E29D0466FFDBFCF99A6F0CDD4436" box="[769,897,1112,1138]" pageId="1" pageNumber="286" type="immature">1 immature</specimenCount>
|
||
(
|
||
<specimenCode id="4305816F0466FFDBFC779A6F0B6F4436" box="[911,1075,1112,1138]" collectionCode="MBCN" pageId="1" pageNumber="286">MBCN 26810</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="A3CB23490466FFDBFBBE9A6E0DDE44D3" ID-GBIF-Occurrence="4168023307" collectingDate="2020-11-25" collectionCode="MBCN" collectorName="David Cabanillas-Roldan" country="Spain" pageId="1" pageNumber="286" specimenCount="1" stateProvince="Madrid" typeStatus="paratype">
|
||
same locality as the preceding, leg.
|
||
<collectorName id="BE564CC20466FFDBFF3F9A4B0EBA44D2" box="[199,486,1148,1174]" pageId="1" pageNumber="286">David Cabanillas-Roldán</collectorName>
|
||
,
|
||
<date id="671D0FD40466FFDBFE099A4B0DDE44D3" box="[497,642,1148,1175]" pageId="1" pageNumber="286" value="2020-11-25">
|
||
<collectingDate id="7759F63C0466FFDBFE099A4B0DDE44D3" box="[497,642,1148,1175]" pageId="1" pageNumber="286" value="2020-11-25">25–XI–2020</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="47DC799C0466FFDBFF6099520D1A47E5" ID-DOI="http://doi.org/10.5281/zenodo.8204286" ID-Zenodo-Dep="8204286" httpUri="https://zenodo.org/record/8204286/files/figure.png" pageId="1" pageNumber="286" startId="1.[152,255,1893,1917]" targetBox="[151,1436,1196,1867]" targetPageId="1" targetType="figure">
|
||
<paragraph id="131C29140466FFDBFF6099520D1A47E5" blockId="1.[151,1436,1893,1953]" pageId="1" pageNumber="286">
|
||
<emphasis id="21D7F5060466FFDBFF6099520E454739" bold="true" box="[152,281,1893,1917]" pageId="1" pageNumber="286">FIGURE 1.</emphasis>
|
||
<taxonomicName id="D4A352970466FFDBFED899520D004739" authority="Garcia, 2023" authorityName="Garcia" authorityYear="2023" box="[288,604,1893,1917]" class="Malacostraca" family="Armadillidiidae" genus="Cristarmadillidium" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="286" phylum="Arthropoda" rank="species" species="cabanillasi" status="sp. nov.">
|
||
<emphasis id="21D7F5060466FFDBFED899520D004739" box="[288,604,1893,1917]" italics="true" pageId="1" pageNumber="286">Cristarmadillidium cabanillasi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="21D7F5060466FFDBFD9B99520DE84739" bold="true" box="[611,692,1893,1917]" pageId="1" pageNumber="286">
|
||
<taxonomicNameLabel id="3AE4487D0466FFDBFD9B99520DE84739" box="[611,692,1893,1917]" pageId="1" pageNumber="286" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Habitus of an alive specimen. A, conglobate position. B, walking on the ground. Photos: David Cabanillas-Roldán.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="47DC799C0465FFD8FF6099430CE347BC" ID-DOI="http://doi.org/10.5281/zenodo.8204288" ID-Zenodo-Dep="8204288" httpUri="https://zenodo.org/record/8204288/files/figure.png" pageId="2" pageNumber="287" startId="2.[152,255,1908,1932]" targetBox="[208,1378,182,1880]" targetPageId="2" targetType="figure">
|
||
<paragraph id="131C29140465FFD8FF6099430CE347BC" blockId="2.[151,1436,1908,2041]" pageId="2" pageNumber="287">
|
||
<emphasis id="21D7F5060465FFD8FF6099430E4447C8" bold="true" box="[152,280,1908,1932]" pageId="2" pageNumber="287">FIGURE 2.</emphasis>
|
||
<taxonomicName id="D4A352970465FFD8FEE699420D0547C9" authority="Garcia, 2023" authorityName="Garcia" authorityYear="2023" box="[286,601,1909,1933]" class="Malacostraca" family="Armadillidiidae" genus="Cristarmadillidium" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="287" phylum="Arthropoda" rank="species" species="cabanillasi" status="sp. nov.">
|
||
<emphasis id="21D7F5060465FFD8FEE699420D0547C9" box="[286,601,1909,1933]" italics="true" pageId="2" pageNumber="287">Cristarmadillidium cabanillasi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="21D7F5060465FFD8FDA699420DF347C9" bold="true" box="[606,687,1909,1933]" pageId="2" pageNumber="287">
|
||
<taxonomicNameLabel id="3AE4487D0465FFD8FDA699420DF347C9" box="[606,687,1909,1933]" pageId="2" pageNumber="287" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Male paratype (CLLG 1071). A, cephalon and pereonites 1–7, transversal view (from the front). B, left
|
||
<emphasis id="21D7F5060465FFD8FE3099AE0D2847F5" box="[456,628,1945,1969]" italics="true" pageId="2" pageNumber="287">nodulus lateralis</emphasis>
|
||
, second tubercle of the posterior row of pereonites. C, cephalothorax and first pereon-tergite, dorsal view. D, first antenna. E, second antenna. F, pleon, pleotelson and uropods, dorsal view. G, left uropod, ventral view. Scale bars: C, 1 mm; D, 0.1 mm; E, 0.5 mm; F, 1 mm; G, 0.3 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="47DC799C0464FFD9FF6099A00F9A47B3" ID-DOI="http://doi.org/10.5281/zenodo.8204292" ID-Zenodo-Dep="8204292" httpUri="https://zenodo.org/record/8204292/files/figure.png" pageId="3" pageNumber="288" startId="3.[152,255,1943,1967]" targetBox="[187,1401,185,1914]" targetPageId="3" targetType="figure">
|
||
<paragraph id="131C29140464FFD9FF6099A00F9A47B3" blockId="3.[151,1436,1943,2039]" pageId="3" pageNumber="288">
|
||
<emphasis id="21D7F5060464FFD9FF6099A00E4447EB" bold="true" box="[152,280,1943,1967]" pageId="3" pageNumber="288">FIGURE 3.</emphasis>
|
||
<taxonomicName id="D4A352970464FFD9FEE799A00D0647EB" authority="Garcia, 2023" authorityName="Garcia" authorityYear="2023" box="[287,602,1943,1967]" class="Malacostraca" family="Armadillidiidae" genus="Cristarmadillidium" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="288" phylum="Arthropoda" rank="species" species="cabanillasi" status="sp. nov.">
|
||
<emphasis id="21D7F5060464FFD9FEE799A00D0647EB" box="[287,602,1943,1967]" italics="true" pageId="3" pageNumber="288">Cristarmadillidium cabanillasi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="21D7F5060464FFD9FDA799AF0DEC47F4" bold="true" box="[607,688,1944,1968]" pageId="3" pageNumber="288">
|
||
<taxonomicNameLabel id="3AE4487D0464FFD9FDA799AF0DEC47F4" box="[607,688,1944,1968]" pageId="3" pageNumber="288" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Male paratype (CLLG 1071). A, right mandible. B, left mandible. C, first maxilla. D, second maxilla. E, maxilliped; F, pereopod 1; G, pereopod 7. Scale bars: A–B, 0.3 mm; C–E, 0.1 mm; F–G, 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="47DC799C0463FFDEFF6099A00C7347B3" ID-DOI="http://doi.org/10.5281/zenodo.8204294" ID-Zenodo-Dep="8204294" httpUri="https://zenodo.org/record/8204294/files/figure.png" pageId="4" pageNumber="289" startId="4.[152,255,1943,1967]" targetBox="[193,1394,185,1914]" targetPageId="4" targetType="figure">
|
||
<paragraph id="131C29140463FFDEFF6099A00C7347B3" blockId="4.[151,1437,1943,2039]" pageId="4" pageNumber="289">
|
||
<emphasis id="21D7F5060463FFDEFF6099A00E4547EB" bold="true" box="[152,281,1943,1967]" pageId="4" pageNumber="289">FIGURE 4.</emphasis>
|
||
<taxonomicName id="D4A352970463FFDEFED999A00D0147EB" authority="Garcia, 2023" authorityName="Garcia" authorityYear="2023" box="[289,605,1943,1967]" class="Malacostraca" family="Armadillidiidae" genus="Cristarmadillidium" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="289" phylum="Arthropoda" rank="species" species="cabanillasi" status="sp. nov.">
|
||
<emphasis id="21D7F5060463FFDEFED999A00D0147EB" box="[289,605,1943,1967]" italics="true" pageId="4" pageNumber="289">Cristarmadillidium cabanillasi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="21D7F5060463FFDEFD9C99A00DEA47EB" bold="true" box="[612,694,1943,1967]" pageId="4" pageNumber="289">
|
||
<taxonomicNameLabel id="3AE4487D0463FFDEFD9C99A00DEA47EB" box="[612,694,1943,1967]" pageId="4" pageNumber="289" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Male paratype (CLLG 1071). A, pleopod 1, ventral view. B, pleopod 2, ventral view. C, genital papilla. D, pleopod 3 exopod, ventral view. E, pleopod 4 exopod, ventral view. F, pleopod 5 exopod, ventral view. Scale bars: A–B, 0.3 mm; C, 0.2 mm; D–F, 0.3 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="5BB97A9F0462FFDFFF3F9EA10DB34125" pageId="5" pageNumber="290" type="diagnosis">
|
||
<paragraph id="131C29140462FFDFFF3F9EA10DB34125" blockId="5.[151,1437,150,2043]" pageId="5" pageNumber="290">
|
||
<emphasis id="21D7F5060462FFDFFF3F9EA10E1840F4" bold="true" box="[199,324,150,176]" pageId="5" pageNumber="290">Diagnosis:</emphasis>
|
||
Cephalon
|
||
<taxonomicName id="D4A352970462FFDFFE329EA00D4F40F5" authorityName="Budde-Lund" authorityYear="1885" box="[458,531,151,177]" class="Malacostraca" family="Armadillidiidae" genus="Eluma" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="290" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="21D7F5060462FFDFFE329EA00D4F40F5" box="[458,531,151,177]" italics="true" pageId="5" pageNumber="290">Eluma</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<typeStatus id="CC1897B60462FFDFFDE39EA00D1340F5" box="[539,591,151,177]" pageId="5" pageNumber="290">type</typeStatus>
|
||
, with strong
|
||
<emphasis id="21D7F5060462FFDFFD0C9EA00CCB40F5" box="[756,919,151,177]" italics="true" pageId="5" pageNumber="290">linea frontalis</emphasis>
|
||
, without post-scutellar ridge; frontal shield separated from vertex and protruding over it. Epimera slightly extended laterally. Back with hypertrophic triangular tubercles; first pereon tergite with six transversal rows of tubercles; pereon tergites 2–7 with two main transversal rows of tubercles; pleon and pleotelson with one row of tubercles; posterior row of pereon-tergites with 14 regularly aligned tubercles; rows of pleonites 1–4 with 10 tubercles each; pleonite 5 with 8 tubercles; pleotelson triangular, as wide as long, with rounded tip and 2 dorsal tubercles.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5BB97A9F0462FFDFFF3F9F5D0B9446DF" pageId="5" pageNumber="290" type="description">
|
||
<paragraph id="131C29140462FFDFFF3F9F5D0B7A41C0" blockId="5.[151,1437,150,2043]" box="[199,1062,362,388]" pageId="5" pageNumber="290">
|
||
<emphasis id="21D7F5060462FFDFFF3F9F5D0E0741C0" bold="true" box="[199,347,362,388]" pageId="5" pageNumber="290">Description:</emphasis>
|
||
Maximum body length: male, 7.2 x
|
||
<quantity id="D45B84F10462FFDFFD0D9F5D0C1241C0" box="[757,846,362,388]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="5" pageNumber="290" unit="mm" value="3.2">3.2 mm</quantity>
|
||
; female, 9 x
|
||
<quantity id="D45B84F10462FFDFFC259F5D0B7D41C0" box="[989,1057,362,388]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="290" unit="mm" value="4.0">4 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="131C29140462FFDFFF3F9FBA0EF5418E" blockId="5.[151,1437,150,2043]" pageId="5" pageNumber="290">
|
||
<emphasis id="21D7F5060462FFDFFF3F9FBA0E7C41E3" box="[199,288,397,423]" italics="true" pageId="5" pageNumber="290">Habitus</emphasis>
|
||
and general body form, as in
|
||
<figureCitation id="8B9835910462FFDFFDA79FBA0D8641E3" box="[607,730,397,423]" captionStart="FIGURE 1" captionStartId="1.[152,255,1893,1917]" captionTargetBox="[151,1436,1196,1867]" captionTargetId="figure-180@1.[151,1436,1196,1867]" captionTargetPageId="1" captionText="FIGURE 1. Cristarmadillidium cabanillasi sp. nov. Habitus of an alive specimen. A, conglobate position. B, walking on the ground. Photos: David Cabanillas-Roldán." figureDoi="http://doi.org/10.5281/zenodo.8204286" httpUri="https://zenodo.org/record/8204286/files/figure.png" pageId="5" pageNumber="290">Figs 1A–B</figureCitation>
|
||
and
|
||
<figureCitation id="8B9835910462FFDFFCF49FBA0C0D41E3" box="[780,849,397,423]" captionStart="FIGURE 2" captionStartId="2.[152,255,1908,1932]" captionTargetBox="[208,1378,182,1880]" captionTargetId="figure-29@2.[204,1383,181,1885]" captionTargetPageId="2" captionText="FIGURE 2. Cristarmadillidium cabanillasi sp. nov. Male paratype (CLLG 1071). A, cephalon and pereonites 1–7, transversal view (from the front). B, left nodulus lateralis, second tubercle of the posterior row of pereonites. C, cephalothorax and first pereon-tergite, dorsal view. D, first antenna. E, second antenna. F, pleon, pleotelson and uropods, dorsal view. G, left uropod, ventral view. Scale bars: C, 1 mm; D, 0.1 mm; E, 0.5 mm; F, 1 mm; G, 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.8204288" httpUri="https://zenodo.org/record/8204288/files/figure.png" pageId="5" pageNumber="290">2 C–F</figureCitation>
|
||
.Animals with perfect endo-conglobation (eusphaeric conglobation) (
|
||
<figureCitation id="8B9835910462FFDFFEBA9F870EC7418E" box="[322,411,432,458]" captionStart="FIGURE 1" captionStartId="1.[152,255,1893,1917]" captionTargetBox="[151,1436,1196,1867]" captionTargetId="figure-180@1.[151,1436,1196,1867]" captionTargetPageId="1" captionText="FIGURE 1. Cristarmadillidium cabanillasi sp. nov. Habitus of an alive specimen. A, conglobate position. B, walking on the ground. Photos: David Cabanillas-Roldán." figureDoi="http://doi.org/10.5281/zenodo.8204286" httpUri="https://zenodo.org/record/8204286/files/figure.png" pageId="5" pageNumber="290">Fig. 1A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="131C29140462FFDFFF3F9FE40A0141AA" blockId="5.[151,1437,150,2043]" box="[199,1373,467,494]" pageId="5" pageNumber="290">
|
||
<emphasis id="21D7F5060462FFDFFF3F9FE40E4A41A9" box="[199,278,467,493]" italics="true" pageId="5" pageNumber="290">Colour</emphasis>
|
||
light brown; edges of epimera, antennae, pereopods and pleopods without pigments (
|
||
<figureCitation id="8B9835910462FFDFFB2D9FE40A1341AA" box="[1237,1359,467,494]" captionStart="FIGURE 1" captionStartId="1.[152,255,1893,1917]" captionTargetBox="[151,1436,1196,1867]" captionTargetId="figure-180@1.[151,1436,1196,1867]" captionTargetPageId="1" captionText="FIGURE 1. Cristarmadillidium cabanillasi sp. nov. Habitus of an alive specimen. A, conglobate position. B, walking on the ground. Photos: David Cabanillas-Roldán." figureDoi="http://doi.org/10.5281/zenodo.8204286" httpUri="https://zenodo.org/record/8204286/files/figure.png" pageId="5" pageNumber="290">Fig. 1A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="131C29140462FFDFFF3F9FC00B48439E" blockId="5.[151,1437,150,2043]" pageId="5" pageNumber="290">
|
||
<emphasis id="21D7F5060462FFDFFF3F9FC00E5D4255" box="[199,257,503,529]" italics="true" pageId="5" pageNumber="290">Body</emphasis>
|
||
(
|
||
<figureCitation id="8B9835910462FFDFFEEB9FC00E2E4255" box="[275,370,503,529]" captionStart="FIGURE 1" captionStartId="1.[152,255,1893,1917]" captionTargetBox="[151,1436,1196,1867]" captionTargetId="figure-180@1.[151,1436,1196,1867]" captionTargetPageId="1" captionText="FIGURE 1. Cristarmadillidium cabanillasi sp. nov. Habitus of an alive specimen. A, conglobate position. B, walking on the ground. Photos: David Cabanillas-Roldán." figureDoi="http://doi.org/10.5281/zenodo.8204286" httpUri="https://zenodo.org/record/8204286/files/figure.png" pageId="5" pageNumber="290">Figs 1B</figureCitation>
|
||
,
|
||
<figureCitation id="8B9835910462FFDFFE7A9FC00EB74255" box="[386,491,503,529]" captionStart="FIGURE 2" captionStartId="2.[152,255,1908,1932]" captionTargetBox="[208,1378,182,1880]" captionTargetId="figure-29@2.[204,1383,181,1885]" captionTargetPageId="2" captionText="FIGURE 2. Cristarmadillidium cabanillasi sp. nov. Male paratype (CLLG 1071). A, cephalon and pereonites 1–7, transversal view (from the front). B, left nodulus lateralis, second tubercle of the posterior row of pereonites. C, cephalothorax and first pereon-tergite, dorsal view. D, first antenna. E, second antenna. F, pleon, pleotelson and uropods, dorsal view. G, left uropod, ventral view. Scale bars: C, 1 mm; D, 0.1 mm; E, 0.5 mm; F, 1 mm; G, 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.8204288" httpUri="https://zenodo.org/record/8204288/files/figure.png" pageId="5" pageNumber="290">2A, C, F</figureCitation>
|
||
) convex with epimera slightly extended laterally, more on pleon than on pereon. Dorsum with strong triangular tubercles compressed laterally; scale-setae triangular, tricorn-type, with rounded tip; integument covered with tiny round raised papillae.
|
||
<emphasis id="21D7F5060462FFDFFCEC9C0A0C964213" box="[788,970,573,599]" italics="true" pageId="5" pageNumber="290">Noduli laterales</emphasis>
|
||
(
|
||
<figureCitation id="8B9835910462FFDFFC229C0A0B6D4213" box="[986,1073,573,599]" captionStart="FIGURE 2" captionStartId="2.[152,255,1908,1932]" captionTargetBox="[208,1378,182,1880]" captionTargetId="figure-29@2.[204,1383,181,1885]" captionTargetPageId="2" captionText="FIGURE 2. Cristarmadillidium cabanillasi sp. nov. Male paratype (CLLG 1071). A, cephalon and pereonites 1–7, transversal view (from the front). B, left nodulus lateralis, second tubercle of the posterior row of pereonites. C, cephalothorax and first pereon-tergite, dorsal view. D, first antenna. E, second antenna. F, pleon, pleotelson and uropods, dorsal view. G, left uropod, ventral view. Scale bars: C, 1 mm; D, 0.1 mm; E, 0.5 mm; F, 1 mm; G, 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.8204288" httpUri="https://zenodo.org/record/8204288/files/figure.png" pageId="5" pageNumber="290">Fig. 2B</figureCitation>
|
||
) small (
|
||
<quantity id="D45B84F10462FFDFFB769C0A0A644213" box="[1166,1336,573,600]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="5" pageNumber="290" unit="mm" value="0.04" valueMax="0.05" valueMin="0.03">0.03–0.05 mm</quantity>
|
||
), located medially on concave outer face of second tubercle (posterior row) of pereonites 1–7, not visible in dorsal view. No visible glandular pores. Dorsal tubercles arranged as follows (
|
||
<figureCitation id="8B9835910462FFDFFC879CB40B4642DA" box="[895,1050,643,670]" captionStart="FIGURE 2" captionStartId="2.[152,255,1908,1932]" captionTargetBox="[208,1378,182,1880]" captionTargetId="figure-29@2.[204,1383,181,1885]" captionTargetPageId="2" captionText="FIGURE 2. Cristarmadillidium cabanillasi sp. nov. Male paratype (CLLG 1071). A, cephalon and pereonites 1–7, transversal view (from the front). B, left nodulus lateralis, second tubercle of the posterior row of pereonites. C, cephalothorax and first pereon-tergite, dorsal view. D, first antenna. E, second antenna. F, pleon, pleotelson and uropods, dorsal view. G, left uropod, ventral view. Scale bars: C, 1 mm; D, 0.1 mm; E, 0.5 mm; F, 1 mm; G, 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.8204288" httpUri="https://zenodo.org/record/8204288/files/figure.png" pageId="5" pageNumber="290">Fig. 2A, C, F</figureCitation>
|
||
): cephalothorax with four rows of tubercles irregularly distributed, except rear row, consisting of 14 tubercles, larger than anteriors; pereon-tergite 1 with six rows of tubercles; pereon-tergites 2–7 with two rows and some smaller tubercles between them; posterior row of all pereon-tergites with 14 aligned tubercles; pleon-tergites 1–4 with one row of ten tubercles; pleon-tergite 5 with one row of eight tubercles; pleotelson with two tubercles.
|
||
<emphasis id="21D7F5060462FFDFFC8D9D270B41436E" box="[885,1053,784,810]" italics="true" pageId="5" pageNumber="290">Cephalothorax</emphasis>
|
||
(
|
||
<figureCitation id="8B9835910462FFDFFBD49D270BFA436E" box="[1068,1190,784,810]" captionStart="FIGURE 2" captionStartId="2.[152,255,1908,1932]" captionTargetBox="[208,1378,182,1880]" captionTargetId="figure-29@2.[204,1383,181,1885]" captionTargetPageId="2" captionText="FIGURE 2. Cristarmadillidium cabanillasi sp. nov. Male paratype (CLLG 1071). A, cephalon and pereonites 1–7, transversal view (from the front). B, left nodulus lateralis, second tubercle of the posterior row of pereonites. C, cephalothorax and first pereon-tergite, dorsal view. D, first antenna. E, second antenna. F, pleon, pleotelson and uropods, dorsal view. G, left uropod, ventral view. Scale bars: C, 1 mm; D, 0.1 mm; E, 0.5 mm; F, 1 mm; G, 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.8204288" httpUri="https://zenodo.org/record/8204288/files/figure.png" pageId="5" pageNumber="290">Fig. 2A, C</figureCitation>
|
||
)
|
||
<taxonomicName id="D4A352970462FFDFFB4E9D270BA3436E" authorityName="Budde-Lund" authorityYear="1885" box="[1206,1279,784,810]" class="Malacostraca" family="Armadillidiidae" genus="Eluma" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="290" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="21D7F5060462FFDFFB4E9D270BA3436E" box="[1206,1279,784,810]" italics="true" pageId="5" pageNumber="290">Eluma</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<typeStatus id="CC1897B60462FFDFFAFE9D260A66436F" box="[1286,1338,785,811]" pageId="5" pageNumber="290">type</typeStatus>
|
||
, without postscutellar line; scutellar ridge delimitied by
|
||
<emphasis id="21D7F5060462FFDFFD619D040C684309" box="[665,820,819,845]" italics="true" pageId="5" pageNumber="290">linea frontalis</emphasis>
|
||
on upper margin, convex superior margin; frontal shield triangular, concave lateral margins and excavated anterior surface, distinctly separated from vertex and protruding over it; antennary lobes well developed, obliques and directed frontwards; eyes small, with 6–10 ommatidia.
|
||
<emphasis id="21D7F5060462FFDFFA9E9D4C0E6943F3" italics="true" pageId="5" pageNumber="290">First pereon-tergite</emphasis>
|
||
(
|
||
<figureCitation id="8B9835910462FFDFFEBC9DAA0EFE43F3" box="[324,418,925,951]" captionStart="FIGURE 2" captionStartId="2.[152,255,1908,1932]" captionTargetBox="[208,1378,182,1880]" captionTargetId="figure-29@2.[204,1383,181,1885]" captionTargetPageId="2" captionText="FIGURE 2. Cristarmadillidium cabanillasi sp. nov. Male paratype (CLLG 1071). A, cephalon and pereonites 1–7, transversal view (from the front). B, left nodulus lateralis, second tubercle of the posterior row of pereonites. C, cephalothorax and first pereon-tergite, dorsal view. D, first antenna. E, second antenna. F, pleon, pleotelson and uropods, dorsal view. G, left uropod, ventral view. Scale bars: C, 1 mm; D, 0.1 mm; E, 0.5 mm; F, 1 mm; G, 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.8204288" httpUri="https://zenodo.org/record/8204288/files/figure.png" pageId="5" pageNumber="290">Fig. 2 C</figureCitation>
|
||
) without
|
||
<emphasis id="21D7F5060462FFDFFDF59DAA0D3B43F3" box="[525,615,925,951]" italics="true" pageId="5" pageNumber="290">schisma</emphasis>
|
||
, with angled posterolateral edge and acute anterior tip, extending beyond eyes position.
|
||
<emphasis id="21D7F5060462FFDFFEC09DF70EF2439E" box="[312,430,960,986]" italics="true" pageId="5" pageNumber="290">Pleotelson</emphasis>
|
||
triangular, as wide as long, with rounded tip (
|
||
<figureCitation id="8B9835910462FFDFFC499DF70B5B439E" box="[945,1031,960,986]" captionStart="FIGURE 2" captionStartId="2.[152,255,1908,1932]" captionTargetBox="[208,1378,182,1880]" captionTargetId="figure-29@2.[204,1383,181,1885]" captionTargetPageId="2" captionText="FIGURE 2. Cristarmadillidium cabanillasi sp. nov. Male paratype (CLLG 1071). A, cephalon and pereonites 1–7, transversal view (from the front). B, left nodulus lateralis, second tubercle of the posterior row of pereonites. C, cephalothorax and first pereon-tergite, dorsal view. D, first antenna. E, second antenna. F, pleon, pleotelson and uropods, dorsal view. G, left uropod, ventral view. Scale bars: C, 1 mm; D, 0.1 mm; E, 0.5 mm; F, 1 mm; G, 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.8204288" httpUri="https://zenodo.org/record/8204288/files/figure.png" pageId="5" pageNumber="290">Fig. 2F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="131C29140462FFDFFF3F9DD30E904400" blockId="5.[151,1437,150,2043]" pageId="5" pageNumber="290">
|
||
<emphasis id="21D7F5060462FFDFFF3F9DD30E0043B9" box="[199,348,996,1021]" italics="true" pageId="5" pageNumber="290">First antenna</emphasis>
|
||
(
|
||
<figureCitation id="8B9835910462FFDFFE919DD40EE343BA" box="[361,447,995,1022]" captionStart="FIGURE 2" captionStartId="2.[152,255,1908,1932]" captionTargetBox="[208,1378,182,1880]" captionTargetId="figure-29@2.[204,1383,181,1885]" captionTargetPageId="2" captionText="FIGURE 2. Cristarmadillidium cabanillasi sp. nov. Male paratype (CLLG 1071). A, cephalon and pereonites 1–7, transversal view (from the front). B, left nodulus lateralis, second tubercle of the posterior row of pereonites. C, cephalothorax and first pereon-tergite, dorsal view. D, first antenna. E, second antenna. F, pleon, pleotelson and uropods, dorsal view. G, left uropod, ventral view. Scale bars: C, 1 mm; D, 0.1 mm; E, 0.5 mm; F, 1 mm; G, 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.8204288" httpUri="https://zenodo.org/record/8204288/files/figure.png" pageId="5" pageNumber="290">Fig. 2D</figureCitation>
|
||
) basal article longest, medial article short, distal article with 7–8 aesthetascs and triangular lobe.
|
||
<emphasis id="21D7F5060462FFDFFF2E9A300ED54464" box="[214,393,1031,1057]" italics="true" pageId="5" pageNumber="290">Second antenna</emphasis>
|
||
(
|
||
<figureCitation id="8B9835910462FFDFFE619A300EB24465" box="[409,494,1031,1057]" captionStart="FIGURE 2" captionStartId="2.[152,255,1908,1932]" captionTargetBox="[208,1378,182,1880]" captionTargetId="figure-29@2.[204,1383,181,1885]" captionTargetPageId="2" captionText="FIGURE 2. Cristarmadillidium cabanillasi sp. nov. Male paratype (CLLG 1071). A, cephalon and pereonites 1–7, transversal view (from the front). B, left nodulus lateralis, second tubercle of the posterior row of pereonites. C, cephalothorax and first pereon-tergite, dorsal view. D, first antenna. E, second antenna. F, pleon, pleotelson and uropods, dorsal view. G, left uropod, ventral view. Scale bars: C, 1 mm; D, 0.1 mm; E, 0.5 mm; F, 1 mm; G, 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.8204288" httpUri="https://zenodo.org/record/8204288/files/figure.png" pageId="5" pageNumber="290">Fig. 2E</figureCitation>
|
||
) short, just reaching first pereon tergite posterior margin; distal article of flagellum 2.7 times longer than basal.
|
||
</paragraph>
|
||
<paragraph id="131C29140462FFDFFF3F9A7A0ADB4553" blockId="5.[151,1437,150,2043]" pageId="5" pageNumber="290">
|
||
Mandibles with dichotomized molar process, about 15 branches;
|
||
<emphasis id="21D7F5060462FFDFFC5D9A7A0B164423" box="[933,1098,1101,1127]" italics="true" pageId="5" pageNumber="290">right mandible</emphasis>
|
||
(
|
||
<figureCitation id="8B9835910462FFDFFBA29A7A0BEF4423" box="[1114,1203,1101,1127]" captionStart="FIGURE 3" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[187,1401,185,1914]" captionTargetId="figure-15@3.[182,1405,181,1919]" captionTargetPageId="3" captionText="FIGURE 3. Cristarmadillidium cabanillasi sp. nov. Male paratype (CLLG 1071). A, right mandible. B, left mandible. C, first maxilla. D, second maxilla. E, maxilliped; F, pereopod 1; G, pereopod 7. Scale bars: A–B, 0.3 mm; C–E, 0.1 mm; F–G, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8204292" httpUri="https://zenodo.org/record/8204292/files/figure.png" pageId="5" pageNumber="290">Fig. 3A</figureCitation>
|
||
) with 2 + 3 penicils;
|
||
<emphasis id="21D7F5060462FFDFFF6F9A470E7744CE" box="[151,299,1136,1162]" italics="true" pageId="5" pageNumber="290">left mandible</emphasis>
|
||
: (
|
||
<figureCitation id="8B9835910462FFDFFEBB9A470EC644CE" box="[323,410,1136,1162]" captionStart="FIGURE 3" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[187,1401,185,1914]" captionTargetId="figure-15@3.[182,1405,181,1919]" captionTargetPageId="3" captionText="FIGURE 3. Cristarmadillidium cabanillasi sp. nov. Male paratype (CLLG 1071). A, right mandible. B, left mandible. C, first maxilla. D, second maxilla. E, maxilliped; F, pereopod 1; G, pereopod 7. Scale bars: A–B, 0.3 mm; C–E, 0.1 mm; F–G, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8204292" httpUri="https://zenodo.org/record/8204292/files/figure.png" pageId="5" pageNumber="290">Fig. 3B</figureCitation>
|
||
) 1 + 3 penicils.
|
||
<emphasis id="21D7F5060462FFDFFDAC9A470DE344CE" box="[596,703,1136,1162]" italics="true" pageId="5" pageNumber="290">Maxillule</emphasis>
|
||
(
|
||
<figureCitation id="8B9835910462FFDFFD379A470C7A44CE" box="[719,806,1136,1162]" captionStart="FIGURE 3" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[187,1401,185,1914]" captionTargetId="figure-15@3.[182,1405,181,1919]" captionTargetPageId="3" captionText="FIGURE 3. Cristarmadillidium cabanillasi sp. nov. Male paratype (CLLG 1071). A, right mandible. B, left mandible. C, first maxilla. D, second maxilla. E, maxilliped; F, pereopod 1; G, pereopod 7. Scale bars: A–B, 0.3 mm; C–E, 0.1 mm; F–G, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8204292" httpUri="https://zenodo.org/record/8204292/files/figure.png" pageId="5" pageNumber="290">Fig. 3C</figureCitation>
|
||
) external branch with 4 + 5 teeth, three of them cleft at apex; internal branch with two thick penicils and wide triangular posterior corner.
|
||
<emphasis id="21D7F5060462FFDFFBCA9AA40BD544E9" box="[1074,1161,1171,1197]" italics="true" pageId="5" pageNumber="290">Maxilla</emphasis>
|
||
(Fig, 3D) with subequal lobes, external lobe bearing three thick setae.
|
||
<emphasis id="21D7F5060462FFDFFD6A9A800C574495" box="[658,779,1207,1233]" italics="true" pageId="5" pageNumber="290">Maxilliped</emphasis>
|
||
(
|
||
<figureCitation id="8B9835910462FFDFFCE29A800C324495" box="[794,878,1207,1233]" captionStart="FIGURE 3" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[187,1401,185,1914]" captionTargetId="figure-15@3.[182,1405,181,1919]" captionTargetPageId="3" captionText="FIGURE 3. Cristarmadillidium cabanillasi sp. nov. Male paratype (CLLG 1071). A, right mandible. B, left mandible. C, first maxilla. D, second maxilla. E, maxilliped; F, pereopod 1; G, pereopod 7. Scale bars: A–B, 0.3 mm; C–E, 0.1 mm; F–G, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8204292" httpUri="https://zenodo.org/record/8204292/files/figure.png" pageId="5" pageNumber="290">Fig. 3E</figureCitation>
|
||
) palp with two setae on first article, second article with two long setae and few small setae, distal article with tuft of short setae and two long setae on external margin; endite with rounded external margin, two triangular teeth on anterior edge and strong medial seta on caudal face.
|
||
</paragraph>
|
||
<paragraph id="131C29140462FFDFFF3F9B170D8C4519" blockId="5.[151,1437,150,2043]" pageId="5" pageNumber="290">
|
||
<emphasis id="21D7F5060462FFDFFF3F9B170E62457E" box="[199,318,1312,1338]" italics="true" pageId="5" pageNumber="290">Pereopods</emphasis>
|
||
with more ventral setae in males than in females; dactyli with inner claw shorter than outer one. Pereopod 1 with oblique antennal grooming brush.
|
||
</paragraph>
|
||
<paragraph id="131C29140462FFDFFF3F9B500CC045C5" blockId="5.[151,1437,150,2043]" box="[199,924,1383,1409]" pageId="5" pageNumber="290">
|
||
<emphasis id="21D7F5060462FFDFFF3F9B500E7345C5" box="[199,303,1383,1409]" italics="true" pageId="5" pageNumber="290">Pleopods</emphasis>
|
||
1 and 2 exopods with monospiracular pleopodal lungs.
|
||
</paragraph>
|
||
<paragraph id="131C29140462FFDFFF3F9BBD0C804583" blockId="5.[151,1437,150,2043]" pageId="5" pageNumber="290">
|
||
<emphasis id="21D7F5060462FFDFFF3F9BBD0E7445E0" box="[199,296,1418,1444]" italics="true" pageId="5" pageNumber="290">Uropods</emphasis>
|
||
with trapezoidal, plate-like, exopod about twice wide than long; endopod inserted proximally on medial margin of protopod, not surpassing the posterior margin of exopod.
|
||
</paragraph>
|
||
<paragraph id="131C29140462FFDFFF3F9BE70B9446DF" blockId="5.[151,1437,150,2043]" pageId="5" pageNumber="290">
|
||
<emphasis id="21D7F5060462FFDFFF3F9BE70EE545AE" box="[199,441,1488,1514]" italics="true" pageId="5" pageNumber="290">Male. Pereopods 1–6</emphasis>
|
||
(
|
||
<figureCitation id="8B9835910462FFDFFE329BE70D4345AE" box="[458,543,1488,1514]" captionStart="FIGURE 3" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[187,1401,185,1914]" captionTargetId="figure-15@3.[182,1405,181,1919]" captionTargetPageId="3" captionText="FIGURE 3. Cristarmadillidium cabanillasi sp. nov. Male paratype (CLLG 1071). A, right mandible. B, left mandible. C, first maxilla. D, second maxilla. E, maxilliped; F, pereopod 1; G, pereopod 7. Scale bars: A–B, 0.3 mm; C–E, 0.1 mm; F–G, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8204292" httpUri="https://zenodo.org/record/8204292/files/figure.png" pageId="5" pageNumber="290">Fig. 3F</figureCitation>
|
||
) with ventral brush of setae on meropod and carpopod.
|
||
<emphasis id="21D7F5060462FFDFFB679BE70A7E45AF" box="[1183,1314,1488,1515]" italics="true" pageId="5" pageNumber="290">Pereopod 7</emphasis>
|
||
(
|
||
<figureCitation id="8B9835910462FFDFFACA9BE70AC845AE" box="[1330,1428,1488,1514]" captionStart="FIGURE 3" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[187,1401,185,1914]" captionTargetId="figure-15@3.[182,1405,181,1919]" captionTargetPageId="3" captionText="FIGURE 3. Cristarmadillidium cabanillasi sp. nov. Male paratype (CLLG 1071). A, right mandible. B, left mandible. C, first maxilla. D, second maxilla. E, maxilliped; F, pereopod 1; G, pereopod 7. Scale bars: A–B, 0.3 mm; C–E, 0.1 mm; F–G, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8204292" httpUri="https://zenodo.org/record/8204292/files/figure.png" pageId="5" pageNumber="290">Fig. 3 G</figureCitation>
|
||
) ischiopod with straight ventral margin.
|
||
<emphasis id="21D7F5060462FFDFFDA89BC40DA64649" box="[592,762,1523,1549]" italics="true" pageId="5" pageNumber="290">Genital papilla</emphasis>
|
||
as in
|
||
<figureCitation id="8B9835910462FFDFFCC49BC40CCF464A" box="[828,915,1523,1550]" captionStart="FIGURE 4" captionStartId="4.[152,255,1943,1967]" captionTargetBox="[193,1394,185,1914]" captionTargetId="figure-29@4.[189,1398,181,1919]" captionTargetPageId="4" captionText="FIGURE 4. Cristarmadillidium cabanillasi sp. nov. Male paratype (CLLG 1071). A, pleopod 1, ventral view. B, pleopod 2, ventral view. C, genital papilla. D, pleopod 3 exopod, ventral view. E, pleopod 4 exopod, ventral view. F, pleopod 5 exopod, ventral view. Scale bars: A–B, 0.3 mm; C, 0.2 mm; D–F, 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.8204294" httpUri="https://zenodo.org/record/8204294/files/figure.png" pageId="5" pageNumber="290">Fig. 4C</figureCitation>
|
||
.
|
||
<emphasis id="21D7F5060462FFDFFC669BC40B4C464A" box="[926,1040,1523,1550]" italics="true" pageId="5" pageNumber="290">Pleopod 1</emphasis>
|
||
(
|
||
<figureCitation id="8B9835910462FFDFFBE79BC40B21464A" box="[1055,1149,1523,1550]" captionStart="FIGURE 4" captionStartId="4.[152,255,1943,1967]" captionTargetBox="[193,1394,185,1914]" captionTargetId="figure-29@4.[189,1398,181,1919]" captionTargetPageId="4" captionText="FIGURE 4. Cristarmadillidium cabanillasi sp. nov. Male paratype (CLLG 1071). A, pleopod 1, ventral view. B, pleopod 2, ventral view. C, genital papilla. D, pleopod 3 exopod, ventral view. E, pleopod 4 exopod, ventral view. F, pleopod 5 exopod, ventral view. Scale bars: A–B, 0.3 mm; C, 0.2 mm; D–F, 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.8204294" httpUri="https://zenodo.org/record/8204294/files/figure.png" pageId="5" pageNumber="290">Fig. 4 A</figureCitation>
|
||
) exopod wider than long; posterior lobe subtriangular with rounded tip; respiratory field oblique, with protruding edge on its ventral face partially covering spiraculum; endopod straight, twice as long as exopod. Pleopod 2 (
|
||
<figureCitation id="8B9835910462FFDFFBA7980D0BE24610" box="[1119,1214,1594,1620]" captionStart="FIGURE 4" captionStartId="4.[152,255,1943,1967]" captionTargetBox="[193,1394,185,1914]" captionTargetId="figure-29@4.[189,1398,181,1919]" captionTargetPageId="4" captionText="FIGURE 4. Cristarmadillidium cabanillasi sp. nov. Male paratype (CLLG 1071). A, pleopod 1, ventral view. B, pleopod 2, ventral view. C, genital papilla. D, pleopod 3 exopod, ventral view. E, pleopod 4 exopod, ventral view. F, pleopod 5 exopod, ventral view. Scale bars: A–B, 0.3 mm; C, 0.2 mm; D–F, 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.8204294" httpUri="https://zenodo.org/record/8204294/files/figure.png" pageId="5" pageNumber="290">Fig. 4 B</figureCitation>
|
||
) exopod triangular, longer than wide, bearing posterior lobe; respiratory field oblique, with protruding edge on its ventral face partially covering spiraculum; endopod flagelliform, longer than exopod.
|
||
<emphasis id="21D7F5060462FFDFFC9598B70B5A46DE" box="[877,1030,1664,1690]" italics="true" pageId="5" pageNumber="290">Pleopods 3–5</emphasis>
|
||
as in
|
||
<figureCitation id="8B9835910462FFDFFBB098B70B9F46DF" box="[1096,1219,1664,1691]" captionStart="FIGURE 4" captionStartId="4.[152,255,1943,1967]" captionTargetBox="[193,1394,185,1914]" captionTargetId="figure-29@4.[189,1398,181,1919]" captionTargetPageId="4" captionText="FIGURE 4. Cristarmadillidium cabanillasi sp. nov. Male paratype (CLLG 1071). A, pleopod 1, ventral view. B, pleopod 2, ventral view. C, genital papilla. D, pleopod 3 exopod, ventral view. E, pleopod 4 exopod, ventral view. F, pleopod 5 exopod, ventral view. Scale bars: A–B, 0.3 mm; C, 0.2 mm; D–F, 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.8204294" httpUri="https://zenodo.org/record/8204294/files/figure.png" pageId="5" pageNumber="290">Fig. 4 D–F</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5BB97A9F0462FFDFFF3F98930D0A46A5" pageId="5" pageNumber="290" type="etymology">
|
||
<paragraph id="131C29140462FFDFFF3F98930D0A46A5" blockId="5.[151,1437,150,2043]" pageId="5" pageNumber="290">
|
||
<emphasis id="21D7F5060462FFDFFF3F98930E0E46FA" bold="true" box="[199,338,1700,1726]" pageId="5" pageNumber="290">Etymology:</emphasis>
|
||
The new species is named after David Cabanillas-Roldán, a myriapodologist from
|
||
<collectingRegion id="D167E7F60462FFDFFB1298940A6046F9" box="[1258,1340,1699,1725]" country="Spain" name="Madrid" pageId="5" pageNumber="290">Madrid</collectingRegion>
|
||
(
|
||
<collectingCountry id="6BB469840462FFDFFAB398940ACC46F9" box="[1355,1424,1699,1725]" name="Spain" pageId="5" pageNumber="290">Spain</collectingCountry>
|
||
), who collected the material studied here.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5BB97A9F0462FFDCFF3F98DD0C1A438D" lastPageId="6" lastPageNumber="291" pageId="5" pageNumber="290" type="discussion">
|
||
<paragraph id="131C29140462FFDFFF3F98DD0D1C47BE" blockId="5.[151,1437,150,2043]" pageId="5" pageNumber="290">
|
||
<emphasis id="21D7F5060462FFDFFF3F98DD0E614740" bold="true" box="[199,317,1770,1796]" pageId="5" pageNumber="290">Remarks:</emphasis>
|
||
According to the original diagnosis of
|
||
<taxonomicName id="D4A352970462FFDFFCE598DD0CA94740" authorityName="Arcangeli" authorityYear="1935" box="[797,1013,1770,1796]" class="Malacostraca" family="Armadillidiidae" genus="Cristarmadillidium" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="290" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="21D7F5060462FFDFFCE598DD0CA94740" box="[797,1013,1770,1796]" italics="true" pageId="5" pageNumber="290">Cristarmadillidium</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="773254E50462FFDFFBF298DD0B9E4740" author="Arcangeli, A." box="[1034,1218,1770,1796]" pageId="5" pageNumber="290" pagination="133 - 151" refId="ref3376" refString="Arcangeli, A. (1936) Sopra alcuni sottogeneri di Armadillidium Br. (Isopodi terrestri). Bollettino dei Musei di Zoologia e di Anatomia comparata della R. Universita di Torino, Serie 3, 45 (62), 133 - 151, pls. 1 - 4." type="journal article" year="1936">Arcangeli 1936</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="773254E50462FFDFFB2B98DD0A354741" author="Vandel, A." box="[1235,1385,1770,1797]" pageId="5" pageNumber="290" pagination="61 - 66" refId="ref4089" refString="Vandel, A. (1954) Une nouvelle espece cavernicole appartenant a la famille des Armadillidiidae et provenant de l'Espagne orientale (Crustaces; Isopodes terrestres). Notes biospeologiques, 9, 61 - 66." type="journal article" year="1954">Vandel 1954</bibRefCitation>
|
||
),
|
||
<taxonomicName id="D4A352970462FFDFFA7B98DD0E4F4763" authority="Garcia, 2023" authorityName="Garcia" authorityYear="2023" class="Malacostraca" family="Armadillidiidae" genus="Cristarmadillidium" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="290" phylum="Arthropoda" rank="species" species="cabanillasi" status="sp. nov.">
|
||
<emphasis id="21D7F5060462FFDFFA7B98DD0E4F4763" italics="true" pageId="5" pageNumber="290">C. cabanillasi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="21D7F5060462FFDFFEE3993A0E2A4763" bold="true" box="[283,374,1805,1831]" pageId="5" pageNumber="290">
|
||
<taxonomicNameLabel id="3AE4487D0462FFDFFEE3993A0E2A4763" box="[283,374,1805,1831]" pageId="5" pageNumber="290" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be accommodated in this genus because it presents the following morphological features:
|
||
<taxonomicName id="D4A352970462FFDFFF6F99070FBC470E" authorityName="Budde-Lund" authorityYear="1885" box="[151,224,1840,1866]" class="Malacostraca" family="Armadillidiidae" genus="Eluma" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="290" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="21D7F5060462FFDFFF6F99070FBC470E" box="[151,224,1840,1866]" italics="true" pageId="5" pageNumber="290">Eluma</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<typeStatus id="CC1897B60462FFDFFF1099060E46470F" box="[232,282,1841,1867]" pageId="5" pageNumber="290">type</typeStatus>
|
||
cephalon; eyes reduced; a lack of schisma on the first pereon-tergite, pereon-epimera 6–7 and pleon-epimera 3,4 and 5 expanded horizontally, and male pereopod 7 not specially modified.
|
||
<bibRefCitation id="773254E50462FFDFFB9B99630A0D472A" author="Recuero, E. & Rodriguez-Flores, P. & Garcia-Paris, M." box="[1123,1361,1875,1902]" pageId="5" pageNumber="290" pagination="1312 - 1340" refId="ref3814" refString="Recuero, E., Rodriguez-Flores, P. & Garcia-Paris, M. (2021) Homoplasy and morphological stasis revealed through multilocus phylogeny of new myrmecophilous species in Armadillidiidae (Isopoda: Oniscidea). Zoological Journal of the Linnean Society, 194 (4), 1312 - 1340. https: // doi. org / 10.1093 / zoolinnean / zlab 066" type="journal article" year="2021">Recuero et al. (2021)</bibRefCitation>
|
||
, based on morphological and molecular data, amend the diagnosis by adding, among other characteristics, the presence of a small schisma on the posterior edge of the first pereonite (only in
|
||
<taxonomicName id="D4A352970462FFDFFC7499AD0ED44793" authority="Recuero, Rodriguez-Flores & Garcia-Paris, 2021" authorityName="Recuero, Rodriguez-Flores & Garcia-Paris" authorityYear="2021" class="Malacostraca" family="Armadillidiidae" genus="Cristarmadillidium" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="290" phylum="Arthropoda" rank="species" species="myrmecophylum">
|
||
<emphasis id="21D7F5060462FFDFFC7499AD0CC347F0" box="[908,927,1946,1972]" italics="true" pageId="5" pageNumber="290">C</emphasis>
|
||
.
|
||
<emphasis id="21D7F5060462FFDFFC5599AD0B3A47F0" box="[941,1126,1946,1972]" italics="true" pageId="5" pageNumber="290">myrmecophylum</emphasis>
|
||
<bibRefCitation id="773254E50462FFDFFB9599AD0ED44793" author="Recuero, E. & Rodriguez-Flores, P. & Garcia-Paris, M." pageId="5" pageNumber="290" pagination="1312 - 1340" refId="ref3814" refString="Recuero, E., Rodriguez-Flores, P. & Garcia-Paris, M. (2021) Homoplasy and morphological stasis revealed through multilocus phylogeny of new myrmecophilous species in Armadillidiidae (Isopoda: Oniscidea). Zoological Journal of the Linnean Society, 194 (4), 1312 - 1340. https: // doi. org / 10.1093 / zoolinnean / zlab 066" type="journal article" year="2021">Recuero, Rodríguez-Flores & García-Paris, 2021</bibRefCitation>
|
||
</taxonomicName>
|
||
); external margin of pereon-tergite 2 epimera straight; and distal part of pereonite 4-5 epimera narrow and rounded. (see discussion).
|
||
</paragraph>
|
||
<paragraph id="131C29140461FFDCFF3F9EA00BF542ED" blockId="6.[151,1437,151,1077]" pageId="6" pageNumber="291">
|
||
In having the dorsal surface tuberculate,
|
||
<taxonomicName id="D4A352970461FFDCFD509EA00C1540F5" authority="Garcia, 2023" authorityName="Garcia" authorityYear="2023" box="[680,841,151,177]" class="Malacostraca" family="Armadillidiidae" genus="Cristarmadillidium" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="291" phylum="Arthropoda" rank="species" species="cabanillasi" status="sp. nov.">
|
||
<emphasis id="21D7F5060461FFDCFD509EA00C1540F5" box="[680,841,151,177]" italics="true" pageId="6" pageNumber="291">C. cabanillasi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="21D7F5060461FFDCFCAD9EA00CEE40F5" bold="true" box="[853,946,151,177]" pageId="6" pageNumber="291">
|
||
<taxonomicNameLabel id="3AE4487D0461FFDCFCAD9EA00CEE40F5" box="[853,946,151,177]" pageId="6" pageNumber="291" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to
|
||
<taxonomicName id="D4A352970461FFDCFBAD9EA00F874091" authority="(Budde-Lund, 1885)" baseAuthorityName="Budde-Lund" baseAuthorityYear="1885" class="Malacostraca" family="Armadillidiidae" genus="Armadillidium" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="291" phylum="Arthropoda" rank="species" species="muricatum">
|
||
<emphasis id="21D7F5060461FFDCFBAD9EA00BAF40F4" box="[1109,1267,151,177]" italics="true" pageId="6" pageNumber="291">C. muricatum</emphasis>
|
||
(Budde-Lund, 1885)
|
||
</taxonomicName>
|
||
,
|
||
<typeStatus id="CC1897B60461FFDCFF109E8C0E444091" box="[232,280,187,213]" pageId="6" pageNumber="291">type</typeStatus>
|
||
species of the genus; however, it differs in having the tubercles numerous, and less developed. Also, it differs from
|
||
<taxonomicName id="D4A352970461FFDCFEB99EE80E8140BC" baseAuthorityName="Budde-Lund" baseAuthorityYear="1885" box="[321,477,223,249]" class="Malacostraca" family="Armadillidiidae" genus="Armadillidium" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="291" phylum="Arthropoda" rank="species" species="muricatum">
|
||
<emphasis id="21D7F5060461FFDCFEB99EE80E8140BC" box="[321,477,223,249]" italics="true" pageId="6" pageNumber="291">C. muricatum</emphasis>
|
||
</taxonomicName>
|
||
by the following characteristics: pereon and pleon-epimera less expanded laterally; integument with round raised papillae (diameter =
|
||
<quantity id="D45B84F10461FFDCFD319F340C6C4159" box="[713,816,259,285]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="291" unit="mm" value="0.02">0.02 mm</quantity>
|
||
), instead of larger oval cavities (diameters = 0.04 x
|
||
<quantity id="D45B84F10461FFDCFA939F340F9B4105" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="291" unit="mm" value="0.06">0.06 mm</quantity>
|
||
); pleotelson triangular with straight sides and broadly rounded tip, instead of slightly concave lateral edges and pointed tip; anterolateral tip of first pereon-tergite triangular and acute, forming an angle with the anterior edge, instead of less prominent and rounded, without angular transition to anterior edge; epimera of pereon-tergite 5 broad and rounded instead of narrow and almost triangular; exopod of male first pleopod 1.2 times as wide as long instead of 1.4 times as wide as long; and uropod exopods (
|
||
<figureCitation id="8B9835910461FFDCFC919F800C954195" box="[873,969,439,465]" captionStart="FIGURE 2" captionStartId="2.[152,255,1908,1932]" captionTargetBox="[208,1378,182,1880]" captionTargetId="figure-29@2.[204,1383,181,1885]" captionTargetPageId="2" captionText="FIGURE 2. Cristarmadillidium cabanillasi sp. nov. Male paratype (CLLG 1071). A, cephalon and pereonites 1–7, transversal view (from the front). B, left nodulus lateralis, second tubercle of the posterior row of pereonites. C, cephalothorax and first pereon-tergite, dorsal view. D, first antenna. E, second antenna. F, pleon, pleotelson and uropods, dorsal view. G, left uropod, ventral view. Scale bars: C, 1 mm; D, 0.1 mm; E, 0.5 mm; F, 1 mm; G, 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.8204288" httpUri="https://zenodo.org/record/8204288/files/figure.png" pageId="6" pageNumber="291">Fig. 2 G</figureCitation>
|
||
) wider than long (ratio w/l=1.56) instead of as wide as long (ratio w/l=1), among other characteristics. The morphological differences with the remaining
|
||
<taxonomicName id="D4A352970461FFDCFF6F9FC80E97425D" box="[151,459,511,537]" pageId="6" pageNumber="291">
|
||
<emphasis id="21D7F5060461FFDCFF6F9FC80E33425D" box="[151,367,511,537]" italics="true" pageId="6" pageNumber="291">Cristarmadillidium</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
are even greater. From
|
||
<taxonomicName id="D4A352970461FFDCFD119FC80CA2425D" authority="Vandel, 1954" authorityName="Vandel" authorityYear="1954" box="[745,1022,510,537]" class="Malacostraca" family="Armadillidiidae" genus="Cristarmadillidium" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="291" phylum="Arthropoda" rank="species" species="breuili">
|
||
<emphasis id="21D7F5060461FFDCFD119FC80C04425D" box="[745,856,511,537]" italics="true" pageId="6" pageNumber="291">C. breuili</emphasis>
|
||
<bibRefCitation id="773254E50461FFDCFC9B9FC80CA2425D" author="Vandel, A." box="[867,1022,510,537]" pageId="6" pageNumber="291" pagination="61 - 66" refId="ref4089" refString="Vandel, A. (1954) Une nouvelle espece cavernicole appartenant a la famille des Armadillidiidae et provenant de l'Espagne orientale (Crustaces; Isopodes terrestres). Notes biospeologiques, 9, 61 - 66." type="journal article" year="1954">Vandel, 1954</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="D4A352970461FFDCFBB89FC80A22425D" authority="Cifuentes, 2021" authorityName="Cifuentes" authorityYear="2021" box="[1088,1406,511,537]" class="Malacostraca" family="Armadillidiidae" genus="Cristarmadillidium" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="291" phylum="Arthropoda" rank="species" species="alticola">
|
||
<emphasis id="21D7F5060461FFDCFBB89FC80BE7425D" box="[1088,1211,511,537]" italics="true" pageId="6" pageNumber="291">C. alticola</emphasis>
|
||
Cifuentes, 2021
|
||
</taxonomicName>
|
||
, it differs mainly by the hypertrophic tubercles on the body. From
|
||
<taxonomicName id="D4A352970461FFDCFC8A9C140A704279" authority="Cifuentes & Prieto, 2020" authorityName="Cifuentes & Prieto" authorityYear="2020" box="[882,1324,547,573]" class="Malacostraca" family="Armadillidiidae" genus="Cristarmadillidium" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="291" phylum="Arthropoda" rank="species" species="zaragozai">
|
||
<emphasis id="21D7F5060461FFDCFC8A9C140B5E4278" box="[882,1026,547,573]" italics="true" pageId="6" pageNumber="291">C. zaragozai</emphasis>
|
||
<bibRefCitation id="773254E50461FFDCFBF49C140A704279" author="Cifuentes, J. & Prieto, C. E." box="[1036,1324,547,573]" pageId="6" pageNumber="291" pagination="111" refId="ref3636" refString="Cifuentes, J. & Prieto, C. E. (2020) Cristarmadillidium zaragozai n. sp. (Oniscidea, Armadillidiidae), un nuevo isopodo terrestre de la peninsula iberica. Graellsia, 76 (2), e 111. https: // doi. org / 10.3989 / graellsia. 2020. v 76.266" type="journal article" year="2020">Cifuentes & Prieto, 2020</bibRefCitation>
|
||
</taxonomicName>
|
||
, it can be distinguished mainly by the shape and structure of the tergal tubercles, which are well differentiated and raised in
|
||
<taxonomicName id="D4A352970461FFDCFF6F9C5C0E6842C1" authority="Garcia, 2023" authorityName="Garcia" authorityYear="2023" box="[151,308,619,645]" class="Malacostraca" family="Armadillidiidae" genus="Cristarmadillidium" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="291" phylum="Arthropoda" rank="species" species="cabanillasi" status="sp. nov.">
|
||
<emphasis id="21D7F5060461FFDCFF6F9C5C0E6842C1" box="[151,308,619,645]" italics="true" pageId="6" pageNumber="291">C. cabanillasi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="21D7F5060461FFDCFEC39C5C0EC942C1" bold="true" box="[315,405,619,645]" pageId="6" pageNumber="291">
|
||
<taxonomicNameLabel id="3AE4487D0461FFDCFEC39C5C0EC942C1" box="[315,405,619,645]" pageId="6" pageNumber="291" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and most flattened, longitudinally keeled, and with a different arrangement in
|
||
<taxonomicName id="D4A352970461FFDCFAFE9C5C0AC942C0" authorityName="Cifuentes & Prieto" authorityYear="2020" box="[1286,1429,619,645]" class="Malacostraca" family="Armadillidiidae" genus="Cristarmadillidium" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="291" phylum="Arthropoda" rank="species" species="zaragozai">
|
||
<emphasis id="21D7F5060461FFDCFAFE9C5C0AC942C0" box="[1286,1429,619,645]" italics="true" pageId="6" pageNumber="291">C. zaragozai</emphasis>
|
||
</taxonomicName>
|
||
. All of these species differ from
|
||
<taxonomicName id="D4A352970461FFDCFE039CB80D8842ED" authorityName="Recuero, Rodriguez-Flores & Garcia-Paris" authorityYear="2021" box="[507,724,655,681]" class="Malacostraca" family="Armadillidiidae" genus="Cristarmadillidium" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="291" phylum="Arthropoda" rank="species" species="myrmecophylum">
|
||
<emphasis id="21D7F5060461FFDCFE039CB80D8842ED" box="[507,724,655,681]" italics="true" pageId="6" pageNumber="291">C. myrmecophylum</emphasis>
|
||
</taxonomicName>
|
||
by lacking schisma on the first pereonite.
|
||
</paragraph>
|
||
<paragraph id="131C29140461FFDCFF3F9C840C1A438D" blockId="6.[151,1437,151,1077]" pageId="6" pageNumber="291">
|
||
<taxonomicName id="D4A352970461FFDCFF3F9C840D784289" authority="Garcia, 2023" authorityName="Garcia" authorityYear="2023" box="[199,548,691,717]" class="Malacostraca" family="Armadillidiidae" genus="Cristarmadillidium" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="291" phylum="Arthropoda" rank="species" species="cabanillasi" status="sp. nov.">
|
||
<emphasis id="21D7F5060461FFDCFF3F9C840D784289" box="[199,548,691,717]" italics="true" pageId="6" pageNumber="291">Cristarmadillidium cabanillasi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="21D7F5060461FFDCFDD59C840DD44289" bold="true" box="[557,648,691,717]" pageId="6" pageNumber="291">
|
||
<taxonomicNameLabel id="3AE4487D0461FFDCFDD59C840DD44289" box="[557,648,691,717]" pageId="6" pageNumber="291" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
also shows some morphological characteristics that bring it closer to the genera
|
||
<taxonomicName id="D4A352970461FFDCFEF79CE00DFA42B5" authority="Cifuentes, 2021" authorityName="Cifuentes" authorityYear="2021" box="[271,678,727,753]" class="Malacostraca" family="Armadillidiidae" genus="Estenarmadillidium" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="291" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="21D7F5060461FFDCFEF79CE00EB142B5" box="[271,493,727,753]" italics="true" pageId="6" pageNumber="291">Estenarmadillidium</emphasis>
|
||
Cifuentes, 2021
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="D4A352970461FFDCFD269CE00F8E4351" authority="Recuero, Rodriguez-Flores & Garcia-Paris, 2021" authorityName="Recuero, Rodriguez-Flores & Garcia-Paris" authorityYear="2021" class="Malacostraca" family="Armadillidiidae" genus="Iberiarmadillidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="291" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="21D7F5060461FFDCFD269CE00CE942B5" box="[734,949,727,753]" italics="true" pageId="6" pageNumber="291">Iberiarmadillidium</emphasis>
|
||
<bibRefCitation id="773254E50461FFDCFC429CE00F8E4351" author="Recuero, E. & Rodriguez-Flores, P. & Garcia-Paris, M." pageId="6" pageNumber="291" pagination="1312 - 1340" refId="ref3814" refString="Recuero, E., Rodriguez-Flores, P. & Garcia-Paris, M. (2021) Homoplasy and morphological stasis revealed through multilocus phylogeny of new myrmecophilous species in Armadillidiidae (Isopoda: Oniscidea). Zoological Journal of the Linnean Society, 194 (4), 1312 - 1340. https: // doi. org / 10.1093 / zoolinnean / zlab 066" type="journal article" year="2021">Recuero, Rodríguez-Flores & García-París, 2021</bibRefCitation>
|
||
</taxonomicName>
|
||
. These similarities are as follows: epimera of pereon-tergites less expanded outwards; integument covered with tiny round raised structures (papillae); lateral edge of the pereonite 5 straight with rounded angles and anterolateral tip of the first tergite extended forwards, forming an angle with the corresponding anterior edge (
|
||
<figureCitation id="8B9835910461FFDCFB0A9D740A6A4319" box="[1266,1334,835,861]" captionStart="FIGURE 1" captionStartId="1.[152,255,1893,1917]" captionTargetBox="[151,1436,1196,1867]" captionTargetId="figure-180@1.[151,1436,1196,1867]" captionTargetPageId="1" captionText="FIGURE 1. Cristarmadillidium cabanillasi sp. nov. Habitus of an alive specimen. A, conglobate position. B, walking on the ground. Photos: David Cabanillas-Roldán." figureDoi="http://doi.org/10.5281/zenodo.8204286" httpUri="https://zenodo.org/record/8204286/files/figure.png" pageId="6" pageNumber="291">Fig. 1</figureCitation>
|
||
), among other features (see Cifuentes, 2021;
|
||
<bibRefCitation id="773254E50461FFDCFDCC9D500C4A43C5" author="Recuero, E. & Rodriguez-Flores, P. & Garcia-Paris, M." box="[564,790,871,897]" pageId="6" pageNumber="291" pagination="1312 - 1340" refId="ref3814" refString="Recuero, E., Rodriguez-Flores, P. & Garcia-Paris, M. (2021) Homoplasy and morphological stasis revealed through multilocus phylogeny of new myrmecophilous species in Armadillidiidae (Isopoda: Oniscidea). Zoological Journal of the Linnean Society, 194 (4), 1312 - 1340. https: // doi. org / 10.1093 / zoolinnean / zlab 066" type="journal article" year="2021">Recuero et al. 2021</bibRefCitation>
|
||
). However, all species of these two genera have the first tergite with a schisma. Therefore, until a systematic review of these genera can be made, we believe it is convenient to keep the new species within the genus
|
||
<taxonomicName id="D4A352970461FFDCFD9F9D980C63438D" authorityName="Arcangeli" authorityYear="1935" box="[615,831,943,969]" class="Malacostraca" family="Armadillidiidae" genus="Cristarmadillidium" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="291" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="21D7F5060461FFDCFD9F9D980C63438D" box="[615,831,943,969]" italics="true" pageId="6" pageNumber="291">Cristarmadillidium</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5BB97A9F0461FFDCFF3F9DE40D9F4471" pageId="6" pageNumber="291" type="biology_ecology">
|
||
<paragraph id="131C29140461FFDCFF3F9DE40D9F4471" blockId="6.[151,1437,151,1077]" pageId="6" pageNumber="291">
|
||
<emphasis id="21D7F5060461FFDCFF3F9DE40E7A43A9" bold="true" box="[199,294,979,1005]" pageId="6" pageNumber="291">Ecology</emphasis>
|
||
: The specimens from the
|
||
<typeStatus id="CC1897B60461FFDCFDAD9DE40DD943A9" box="[597,645,979,1005]" pageId="6" pageNumber="291">type</typeStatus>
|
||
locality, Villanueva del Pardillo, were collected from a pastured oak woodland. The specimens from Cadalso de los Vidrios were collected in a rocky clearing inside a forest of
|
||
<taxonomicName id="D4A352970461FFDCFAA49DC00FA34471" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="291" phylum="Tracheophyta" rank="species" species="sylvestris">
|
||
<emphasis id="21D7F5060461FFDCFAA49DC00FA34471" italics="true" pageId="6" pageNumber="291">Pinus sylvestris</emphasis>
|
||
</taxonomicName>
|
||
L., under stones, with unidentified ants.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |