treatments-xml/data/03/B7/87/03B787C7FF5876AAFF18D2224F4EF5E0.xml

<document id="2ECCDAB94447B0E8C95D71CE46714997" ID-CLB-Dataset="115951" ID-DOI="10.1093/zoolinnean/zlab033" ID-GBIF-Dataset="75b10efe-a17d-4d10-b46e-094c6a7ebf01" ID-ISSN="0024-4082" ID-Zenodo-Dep="6967774" ID-ZooBank="BE2B3859-8F6A-4543-8A69-91840F82377B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1653991067759" checkinUser="plazi" docAuthor="Lin, Shou-Wang, Lopardo, Lara &amp; Uhl, Gabriele" docDate="2022" docId="03B787C7FF5876AAFF18D2224F4EF5E0" docLanguage="en" docName="zlab033.pdf" docOrigin="Zoological Journal of the Linnean Society 195" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Oedothorax nazareti Scharff 1989, INCERTAE SEDIS" docType="treatment" docVersion="7" lastPageNumber="552" masterDocId="FF8EFFBFFFDD762DFFABD4584A34F51A" masterDocTitle="Evolution of nuptial-gift-related male prosomal structures: taxonomic revision and cladistic analysis of the genus Oedothorax (Araneae: Linyphiidae: Erigoninae)" masterLastPageNumber="584" masterPageNumber="417" pageNumber="550" updateTime="1699338434129" updateUser="ExternalLinkService">
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<mods:title id="659733B40D32390F4AB0E3252A5F5267">Evolution of nuptial-gift-related male prosomal structures: taxonomic revision and cladistic analysis of the genus Oedothorax (Araneae: Linyphiidae: Erigoninae)</mods:title>
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<mods:namePart id="0EA3D7F43F34441334C683D5DFD2534D">Lin, Shou-Wang</mods:namePart>
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<paragraph id="8BA136D1FF5876A8FF18D22248A2F3AA" blockId="133.[179,762,1658,1712]" pageId="133" pageNumber="550">
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<taxonomicName id="4C1E4D52FF5876A8FF18D22248B7F389" ID-CoL="48WV5" authority="SCHARFF, 1989 INCERTAE" authorityName="Scharff" authorityYear="1989" box="[179,643,1658,1683]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="133" pageNumber="550" phylum="Arthropoda" rank="species" species="nazareti" status="INCERTAE SEDIS">
<emphasis id="B96AEAC3FF5876A8FF18D2224BFBF388" box="[179,463,1658,1682]" italics="true" pageId="133" pageNumber="550">‘OEDOTHORAX’ NAZARETI</emphasis>
<bibRefCitation id="EF8F4B20FF5876A8FE7DD22248B7F389" author="Scharff N" box="[470,643,1658,1683]" pageId="133" pageNumber="550" pagination="13 - 20" refId="ref95011" refString="Scharff N, 1989. New species and records of Afrotropical Linyphiidae (Araneae). Bulletin of the British Arachnological Society 8: 13 - 20." type="journal article" year="1989">SCHARFF, 1989</bibRefCitation>
</taxonomicName>
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<taxonomicNameLabel id="A25957B8FF5876A8FD21D2264B6EF3AA" pageId="133" pageNumber="550">INCERTAE SEDIS</taxonomicNameLabel>
;
</emphasis>
NEW FEMALE DESCRIPTION
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C304655AFF5876A8FEF0D2E74865F3C2" box="[347,593,1727,1752]" pageId="133" pageNumber="550" type="description">
<paragraph id="8BA136D1FF5876A8FEF0D2E74865F3C2" blockId="133.[347,593,1727,1752]" box="[347,593,1727,1752]" pageId="133" pageNumber="550">
(
<figureCitation id="13252A54FF5876A8FECFD2E74B83F3CD" box="[356,439,1727,1752]" captionStart="Figure 7" captionStartId="13.[163,240,1315,1337]" captionTargetBox="[166,1440,198,1272]" captionTargetId="figure-25@13.[163,1443,195,1275]" captionTargetPageId="13" captionText="Figure 7. Male prosomal morphology, lateral.A, Emertongone montifera (Emerton, 1882) (Paquin &amp; Dupérré, 2003, fig. 1187). B, Shaanxinus mingchihensis Lin, 2019 (traced from photograph). C, ‘Oedothorax’ paracymbialis Tanasevitch, 2015 (traced from photograph). D, ‘Oe.’ kodaikanal Tanasevitch, 2015 (traced from photograph). E, ‘Oe.’ cunur Tanasevitch, 2015 (traced from photograph). F, ‘Oe.’ stylus Tanasevitch, 2015 (traced from photograph). G, Atypena cirrifrons (Heimer, 1984) (traced from photograph). H, A. formosana (Oi, 1977) (Oi, 1977, fig. 2). I, ‘Oe.’ nazareti Scharff, 1989 (Scharff, 1989: fig. 7). J, Gongylidium rufipes (Linnaeus, 1758) (traced from photograph). K, Ummeliata insecticeps (Bösenberg &amp; Strand, 1906) (Tu &amp; Li 2004: fig. 8A). L, U. esyunini (Zhang, Zhang &amp; Yu, 2003) (Zhang et al., 2003, fig. 2A). M, Hylyphantes graminicola (Sundevall, 1830) (traced from photograph). N, Tmeticus tolli Kulczyński, 1908 (traced from photograph). O, Oedothorax gibbosus (Blackwall, 1841) [Roberts, 1987: fig. 21 (e)]. P, Oe. trilobatus (Banks, 1896) (Bishop &amp; Crosby, 1935: fig. 79). Q, Oe. gibbifer (Kulczyński, 1882) (Heimer &amp; Nentwig, 1991: fig.604.5).R, Oe.apicatus (Blackwall, 1850) [Roberts, 1987: fig.23 (d)].S, Oe. retusus (Westring, 1851) [Roberts, 1987: fig. 23 (c)]. T, Oe. paludigena Simon, 1926 [Millidge, 1975: fig. (5)]. U, Oe. meridionalis Tanasevitch, 1987 (Tanasevitch, 1987: fig. 107).V, Oe. agrestis (Blackwall, 1853) (Roberts, 1987: fig. 23B).W, Oe. fuscus (Blackwall, 1834) (Roberts, 1987: fig. 23A). X, Oe. tingitanus (Simon, 1884) (Denis, 1968: fig. 11)." figureDoi="http://doi.org/10.5281/zenodo.6967802" httpUri="https://zenodo.org/record/6967802/files/figure.png" pageId="133" pageNumber="550">FIGS 7I</figureCitation>
, 
<figureCitation id="13252A54FF5876A8FE6FD2E74BDFF3CD" box="[452,491,1727,1752]" captionStart="Figure 67" captionStartId="110.[144,224,1301,1323]" captionTargetBox="[147,1426,197,1260]" captionTargetId="figure-175@110.[145,1428,195,1261]" captionTargetPageId="110" captionText="Figure 67. Male chelicerae, stridulatory striae on lateral side. A, Jilinus hulongensis (Zhu &amp; Wen, 1980). B, Cornitibia simplicithorax (Tanasevitch, 1998). C, Emertongone montifera (Emerton, 1882). D, ‘Oedothorax’ paracymbialis Tanasevitch, 2015. E, ‘Oe.’ kodaikanal Tanasevitch, 2015. F, ‘Oe.’ cunur Tanasevitch, 2015. G, ‘Oe.’ stylus Tanasevitch, 2015. H, Atypena cirrifrons (Heimer, 1984). I, A. formosana (Oi, 1977). J, ‘Oe.’ nazareti Scharff, 1989. K, Gongylidium rufipes (Linnaeus, 1758). L, Ummeliata insecticeps (Bösenberg &amp; Strand, 1906). M, U. esyunini (Zhang, Zhang &amp; Yu, 2003). N, Hylyphantes graminicola (Sundevall, 1830). O, Tmeticus tolli Kulczyński, 1908. Scale bars 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6967988" httpUri="https://zenodo.org/record/6967988/files/figure.png" pageId="133" pageNumber="550">67J</figureCitation>
, 
<figureCitation id="13252A54FF5876A8FE52D2E74814F3CD" box="[505,544,1727,1752]" captionStart="Figure 68" captionStartId="111.[163,245,1308,1330]" captionTargetBox="[164,1444,196,1268]" captionTargetId="figure-26@111.[163,1446,195,1269]" captionTargetPageId="111" captionText="Figure 68. Male opisthosoma, dorsal view. A, Jilinus hulongensis (Zhu &amp; Wen, 1980). B, Cornitibia simplicithorax (Tanasevitch, 1998). C, Emertongone montifera (Emerton, 1882). D, ‘Oedothorax’ paracymbialis Tanasevitch, 2015. E, ‘Oe.’ kodaikanal Tanasevitch, 2015. F, ‘Oe.’ cunur Tanasevitch, 2015. G, ‘Oe.’ stylus Tanasevitch, 2015. H, Atypena cirrifrons (Heimer, 1984). I, A. formosana (Oi, 1977). J, ‘Oe.’ nazareti Scharff, 1989. K, Gongylidium rufipes (Linnaeus, 1758). L, Ummeliata insecticeps (Bösenberg &amp; Strand, 1906). M, U. esyunini (Zhang, Zhang &amp; Yu, 2003). N, Hylyphantes graminicola (Sundevall, 1830). O, Tmeticus tolli Kulczyński, 1908. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6967990" httpUri="https://zenodo.org/record/6967990/files/figure.png" pageId="133" pageNumber="550">68J</figureCitation>
, 
<figureCitation id="13252A54FF5876A8FD86D2E7487DF3C2" box="[557,585,1727,1752]" captionStart="Figure 78" captionStartId="134.[145,223,1433,1455]" captionTargetBox="[148,1427,198,1382]" captionTargetId="figure-188@134.[145,1431,195,1393]" captionTargetPageId="134" captionText="Figure 78. ‘Oedothorax’ nazareti Scharff, 1989.A–E, male left palp.A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, apical view. F, G, epigyne. F, ventrolateral view. G, external morphology. H, male right spinnerets. I, female left spinnerets. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968022" httpUri="https://zenodo.org/record/6968022/files/figure.png" pageId="133" pageNumber="550">78</figureCitation>
)
</paragraph>
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<paragraph id="8BA136D1FF5876A8FF08D2B0493EF3E4" blockId="133.[163,780,1768,1882]" box="[163,778,1768,1790]" pageId="133" pageNumber="550">
<treatmentCitation id="0ABF10C0FF5876A8FF08D2B04864F3E4" author="Scharff N" box="[163,592,1768,1790]" page="15" pageId="133" pageNumber="550" year="1989">
<taxonomicName id="4C1E4D52FF5876A8FF08D2B04864F3E4" ID-CoL="48WV5" authority="Scharff, 1989: 15" authorityName="Scharff" authorityPageNumber="15" authorityYear="1989" box="[163,592,1768,1790]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="133" pageNumber="550" phylum="Arthropoda" rank="species" species="nazareti">
<emphasis id="B96AEAC3FF5876A8FF08D2B04BBFF3E7" box="[163,395,1768,1789]" italics="true" pageId="133" pageNumber="550">Oedothorax nazareti</emphasis>
<bibRefCitation id="EF8F4B20FF5876A8FE3AD2B04864F3E4" author="Scharff N" box="[401,592,1768,1790]" pageId="133" pageNumber="550" pagination="13 - 20" refId="ref95011" refString="Scharff N, 1989. New species and records of Afrotropical Linyphiidae (Araneae). Bulletin of the British Arachnological Society 8: 13 - 20." type="journal article" year="1989">Scharff, 1989: 15</bibRefCitation>
</taxonomicName>
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, figs 7–12 (Dm).
</paragraph>
<paragraph id="8BA136D1FF5876A8FF08D35E4938F207" blockId="133.[163,780,1768,1882]" box="[163,780,1798,1821]" pageId="133" pageNumber="550">
<treatmentCitation id="0ABF10C0FF5876A8FF08D35E48A1F206" author="Tanasevitch AV" box="[163,661,1798,1820]" page="382" pageId="133" pageNumber="550" year="2015">
<taxonomicName id="4C1E4D52FF5876A8FF08D35E48A1F206" ID-CoL="48WV5" authority="Tanasevitch, 2015: 382" authorityName="Tanasevitch" authorityPageNumber="382" authorityYear="2015" box="[163,661,1798,1820]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="133" pageNumber="550" phylum="Arthropoda" rank="species" species="nazareti">
<emphasis id="B96AEAC3FF5876A8FF08D35E4BBFF206" box="[163,395,1798,1820]" italics="true" pageId="133" pageNumber="550">Oedothorax nazareti</emphasis>
<bibRefCitation id="EF8F4B20FF5876A8FE3BD35E48A1F206" author="Tanasevitch AV" box="[400,661,1798,1820]" pageId="133" pageNumber="550" pagination="381 - 398" refId="ref95840" refString="Tanasevitch AV. 2015. Notes on the spider genus Oedothorax Bertkau, 1883 with description of eleven new species from India (Linyphiidae: Erigoninae). Revue suisse de Zoologie 122: 381 - 398." type="journal article" year="2015">Tanasevitch, 2015: 382</bibRefCitation>
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, fig. 7 (m).
</paragraph>
</subSubSection>
<subSubSection id="C304655AFF5876A8FF08D37E4E72F5E0" pageId="133" pageNumber="550" type="materials_examined">
<paragraph id="8BA136D1FF5876A8FF08D37E4E72F5E0" blockId="133.[163,780,1768,1882]" lastBlockId="133.[827,1442,197,250]" pageId="133" pageNumber="550">
<emphasis id="B96AEAC3FF5876A8FF08D37E4B46F221" box="[163,370,1829,1851]" italics="true" pageId="133" pageNumber="550">Type material:</emphasis>
Holotype: 
<emphasis id="B96AEAC3FF5876A8FD82D37D4883F220" bold="true" box="[553,695,1829,1851]" pageId="133" pageNumber="550">Ethiopia:</emphasis>
Shoa Administrative Province, Nazaret, under stone in cultivated farmland, 
<quantity id="4CE69B34FF5876A8FBEAD49D4EA9F5C1" box="[1089,1181,197,219]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="133" pageNumber="550" unit="m" value="2400.0">2400 m</quantity>
, 
<specimenCount id="9D18FD58FF5876A8FB07D4904E88F5C0" box="[1196,1212,200,218]" pageId="133" pageNumber="550" type="male">♂</specimenCount>
<date id="FFA01011FF5876A8FB6CD49D4F72F5C1" box="[1223,1350,197,219]" pageId="133" pageNumber="550" value="1985-06-22">22.vi.1985</date>
, leg. N, Scharff (not examined).
</paragraph>
</subSubSection>
</treatmentCitationGroup>
<paragraph id="8BA136D1FF5876A8FC90D57A499CF48E" blockId="133.[827,1441,290,404]" pageId="133" pageNumber="550">
<emphasis id="B96AEAC3FF5876A8FC90D57A4E08F42D" box="[827,1084,290,311]" italics="true" pageId="133" pageNumber="550">Examined material:</emphasis>
<materialsCitation id="3B763C8CFF5876A8FBE0D57A4990F48E" ID-GBIF-Occurrence="3880945341" collectingDate="1988-06-11" collectionCode="RMCA" country="Ethiopia" determinerName="A. Russell-Smith" location="Debre Siwa" pageId="133" pageNumber="550" specimenCode="RMCA 224.501" specimenCount="13" specimenCount-female="10" specimenCount-male="3">
<collectingCountry id="F3097641FF5876A8FBE0D57A4E88F422" box="[1099,1212,290,312]" name="Ethiopia" pageId="133" pageNumber="550">Ethiopia</collectingCountry>
, 
<emphasis id="B96AEAC3FF5876A8FB66D57B4EEBF422" box="[1229,1247,291,312]" italics="true" pageId="133" pageNumber="550">c.</emphasis>
<quantity id="4CE69B34FF5876A8FB40D57A4F0FF422" box="[1259,1339,290,312]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="133" pageNumber="550" unit="km" value="15.0">15 km</quantity>
west of 
<location id="8EC1600AFF5876A8FC90D51949FEF44D" LSID="urn:lsid:plazi:treatment:03B787C7FF5876AAFF18D2224F4EF5E0:8EC1600AFF5876A8FC90D51949FEF44D" box="[827,970,321,343]" country="Ethiopia" name="Debre Siwa" pageId="133" pageNumber="550">Debre Siwa</location>
, under stones in overgazed woodland, 
<specimenCount id="9D18FD58FF5876A8FC90D538496EF46F" box="[827,858,352,373]" pageId="133" pageNumber="550" type="male">3♂</specimenCount>
<specimenCount id="9D18FD58FF5876A8FCF7D53849BDF46F" box="[860,905,352,373]" pageId="133" pageNumber="550" type="female">10♀</specimenCount>
<date id="FFA01011FF5876A8FC3FD5384E2EF46C" box="[916,1050,352,374]" pageId="133" pageNumber="550" value="1988-06-11">
<collectingDate id="EFE4E9F9FF5876A8FC3FD5384E2EF46C" box="[916,1050,352,374]" pageId="133" pageNumber="550" value="1988-06-11">11.vi.1988</collectingDate>
</date>
, det. 
<determinerName id="7E7DDA15FF5876A8FBCFD5384F08F46C" box="[1124,1340,352,374]" pageId="133" pageNumber="550">A. Russell-Smith</determinerName>
(
<specimenCode id="DBB89EAAFF5876A8FAE4D53849A9F48E" collectionCode="RMCA" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="133" pageNumber="550" type="Museum">RMCA 224.501</specimenCode>
)
</materialsCitation>
.
</paragraph>
<subSubSection id="C304655AFF5876A8FC90D5E64984F7F3" pageId="133" pageNumber="550" type="diagnosis">
<paragraph id="8BA136D1FF5876A8FC90D5E64986F4C9" blockId="133.[827,946,446,467]" box="[827,946,446,467]" pageId="133" pageNumber="550">
<emphasis id="B96AEAC3FF5876A8FC90D5E64986F4C9" box="[827,946,446,467]" italics="true" pageId="133" pageNumber="550">Diagnosis:</emphasis>
</paragraph>
<paragraph id="8BA136D1FF5876A8FC90D5A44EECF755" blockId="133.[827,1442,508,591]" pageId="133" pageNumber="550">
<emphasis id="B96AEAC3FF5876A8FC90D5A449BCF70B" box="[827,904,508,529]" italics="true" pageId="133" pageNumber="550">Males:</emphasis>
This species can be identified by its unique conical-shaped postocular hump and setae distribution on the hump and interocular region.
</paragraph>
<paragraph id="8BA136D1FF5876A8FC90D6204984F7F3" blockId="133.[827,1443,632,746]" pageId="133" pageNumber="550">
<emphasis id="B96AEAC3FF5876A8FC90D62049ABF797" box="[827,927,632,653]" italics="true" pageId="133" pageNumber="550">Females:</emphasis>
This species has the typical simple epigyne structures as many other 
<taxonomicName id="4C1E4D52FF5876A8FBCFD6CE4EDCF7B1" box="[1124,1256,662,683]" pageId="133" pageNumber="550">
<emphasis id="B96AEAC3FF5876A8FBCFD6CE4EDCF7B1" box="[1124,1256,662,683]" italics="true" pageId="133" pageNumber="550">Oedothorax</emphasis>
</taxonomicName>
species, but can be identified by the trajectory of the copulatory ducts (
<figureCitation id="13252A54FF5876A8FCE8D68C4994F7F3" box="[835,928,724,746]" captionStart="Figure 78" captionStartId="134.[145,223,1433,1455]" captionTargetBox="[148,1427,198,1382]" captionTargetId="figure-188@134.[145,1431,195,1393]" captionTargetPageId="134" captionText="Figure 78. ‘Oedothorax’ nazareti Scharff, 1989.A–E, male left palp.A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, apical view. F, G, epigyne. F, ventrolateral view. G, external morphology. H, male right spinnerets. I, female left spinnerets. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968022" httpUri="https://zenodo.org/record/6968022/files/figure.png" pageId="133" pageNumber="550">Fig. 78F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C304655AFF5876AAFC90D74B48A0F5E0" lastPageId="135" lastPageNumber="552" pageId="133" pageNumber="550" type="description">
<paragraph id="8BA136D1FF5876A8FC90D74B49F7F632" blockId="133.[827,963,787,808]" box="[827,963,787,808]" pageId="133" pageNumber="550">
<emphasis id="B96AEAC3FF5876A8FC90D74B49F7F632" box="[827,963,787,808]" italics="true" pageId="133" pageNumber="550">Description:</emphasis>
</paragraph>
<paragraph id="8BA136D1FF5876A8FC90D7094980F3DB" blockId="133.[827,1444,849,1730]" pageId="133" pageNumber="550">
<emphasis id="B96AEAC3FF5876A8FC90D7094E7DF67C" box="[827,1097,849,870]" italics="true" pageId="133" pageNumber="550">Male (RMCA 224.501):</emphasis>
Total length: 2.29. Prosoma: 1.06 long, 0.74 wide, postocular region conically elevated, covered by long, thick setae; interocular region with dense setae (
<figureCitation id="13252A54FF5876A8FBAED7F54E79F6D9" box="[1029,1101,941,963]" captionStart="Figure 7" captionStartId="13.[163,240,1315,1337]" captionTargetBox="[166,1440,198,1272]" captionTargetId="figure-25@13.[163,1443,195,1275]" captionTargetPageId="13" captionText="Figure 7. Male prosomal morphology, lateral.A, Emertongone montifera (Emerton, 1882) (Paquin &amp; Dupérré, 2003, fig. 1187). B, Shaanxinus mingchihensis Lin, 2019 (traced from photograph). C, ‘Oedothorax’ paracymbialis Tanasevitch, 2015 (traced from photograph). D, ‘Oe.’ kodaikanal Tanasevitch, 2015 (traced from photograph). E, ‘Oe.’ cunur Tanasevitch, 2015 (traced from photograph). F, ‘Oe.’ stylus Tanasevitch, 2015 (traced from photograph). G, Atypena cirrifrons (Heimer, 1984) (traced from photograph). H, A. formosana (Oi, 1977) (Oi, 1977, fig. 2). I, ‘Oe.’ nazareti Scharff, 1989 (Scharff, 1989: fig. 7). J, Gongylidium rufipes (Linnaeus, 1758) (traced from photograph). K, Ummeliata insecticeps (Bösenberg &amp; Strand, 1906) (Tu &amp; Li 2004: fig. 8A). L, U. esyunini (Zhang, Zhang &amp; Yu, 2003) (Zhang et al., 2003, fig. 2A). M, Hylyphantes graminicola (Sundevall, 1830) (traced from photograph). N, Tmeticus tolli Kulczyński, 1908 (traced from photograph). O, Oedothorax gibbosus (Blackwall, 1841) [Roberts, 1987: fig. 21 (e)]. P, Oe. trilobatus (Banks, 1896) (Bishop &amp; Crosby, 1935: fig. 79). Q, Oe. gibbifer (Kulczyński, 1882) (Heimer &amp; Nentwig, 1991: fig.604.5).R, Oe.apicatus (Blackwall, 1850) [Roberts, 1987: fig.23 (d)].S, Oe. retusus (Westring, 1851) [Roberts, 1987: fig. 23 (c)]. T, Oe. paludigena Simon, 1926 [Millidge, 1975: fig. (5)]. U, Oe. meridionalis Tanasevitch, 1987 (Tanasevitch, 1987: fig. 107).V, Oe. agrestis (Blackwall, 1853) (Roberts, 1987: fig. 23B).W, Oe. fuscus (Blackwall, 1834) (Roberts, 1987: fig. 23A). X, Oe. tingitanus (Simon, 1884) (Denis, 1968: fig. 11)." figureDoi="http://doi.org/10.5281/zenodo.6967802" httpUri="https://zenodo.org/record/6967802/files/figure.png" pageId="133" pageNumber="550">Fig. 7I</figureCitation>
). Eyes: AME-AME: 0.04, AME width: 0.05, AME-ALE: 0.05, ALE width: 0.08, ALE- PLE: 0, PLE width: 0.08, PLE-PME: 0.04, PME width: 0.06, PME-PME: 0.19. Clypeus: not hirsute, one sub- AME seta. Sternum: 0.63 long, 0.56 wide. Chelicerae: mastidia absent; stridulatory striae scaly, rows widely and evenly spaced (
<figureCitation id="13252A54FF5876A8FB8CD03D4EB3F161" box="[1063,1159,1125,1147]" captionStart="Figure 67" captionStartId="110.[144,224,1301,1323]" captionTargetBox="[147,1426,197,1260]" captionTargetId="figure-175@110.[145,1428,195,1261]" captionTargetPageId="110" captionText="Figure 67. Male chelicerae, stridulatory striae on lateral side. A, Jilinus hulongensis (Zhu &amp; Wen, 1980). B, Cornitibia simplicithorax (Tanasevitch, 1998). C, Emertongone montifera (Emerton, 1882). D, ‘Oedothorax’ paracymbialis Tanasevitch, 2015. E, ‘Oe.’ kodaikanal Tanasevitch, 2015. F, ‘Oe.’ cunur Tanasevitch, 2015. G, ‘Oe.’ stylus Tanasevitch, 2015. H, Atypena cirrifrons (Heimer, 1984). I, A. formosana (Oi, 1977). J, ‘Oe.’ nazareti Scharff, 1989. K, Gongylidium rufipes (Linnaeus, 1758). L, Ummeliata insecticeps (Bösenberg &amp; Strand, 1906). M, U. esyunini (Zhang, Zhang &amp; Yu, 2003). N, Hylyphantes graminicola (Sundevall, 1830). O, Tmeticus tolli Kulczyński, 1908. Scale bars 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6967988" httpUri="https://zenodo.org/record/6967988/files/figure.png" pageId="133" pageNumber="550">Fig. 67J</figureCitation>
). Legs: dorsal proximal macroseta on tibia I, II, III and IV 0.12, 0.13, 0.91 and 1.46 times diameter of tibia, respectively; Tm I: 0.67. All metatarsi with trichobothrium. Pedipalp: patella prolateral proximal vertical macrosetae absent; tibia with one prolateral, two retrolateral trichobothria; TPS small, ventrally situated at TPA tip; TPA long, base prolateral to palpal tibia prolateral trichobothrium, tip bent retrolaterally (
<figureCitation id="13252A54FF5876A8FBFDD1024E8EF06A" box="[1110,1210,1370,1392]" captionStart="Figure 78" captionStartId="134.[145,223,1433,1455]" captionTargetBox="[148,1427,198,1382]" captionTargetId="figure-188@134.[145,1431,195,1393]" captionTargetPageId="134" captionText="Figure 78. ‘Oedothorax’ nazareti Scharff, 1989.A–E, male left palp.A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, apical view. F, G, epigyne. F, ventrolateral view. G, external morphology. H, male right spinnerets. I, female left spinnerets. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968022" httpUri="https://zenodo.org/record/6968022/files/figure.png" pageId="133" pageNumber="550">Fig. 78C</figureCitation>
); PC median-sized, base not visible from dorsal view, distal setae close to distal clasp, distal clasp without striae, clasp extended apically (
<figureCitation id="13252A54FF5876A8FC0AD1EE49CAF0D6" box="[929,1022,1462,1484]" captionStart="Figure 78" captionStartId="134.[145,223,1433,1455]" captionTargetBox="[148,1427,198,1382]" captionTargetId="figure-188@134.[145,1431,195,1393]" captionTargetPageId="134" captionText="Figure 78. ‘Oedothorax’ nazareti Scharff, 1989.A–E, male left palp.A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, apical view. F, G, epigyne. F, ventrolateral view. G, external morphology. H, male right spinnerets. I, female left spinnerets. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968022" httpUri="https://zenodo.org/record/6968022/files/figure.png" pageId="133" pageNumber="550">Fig. 78A</figureCitation>
); T without papillae; PT long, slender, without papillae; TS long, slender, without papillae; MSA absent; DSA tip straight, simple; EM absent (
<figureCitation id="13252A54FF5876A8FAD3D1AC49B3F332" captionStart="Figure 78" captionStartId="134.[145,223,1433,1455]" captionTargetBox="[148,1427,198,1382]" captionTargetId="figure-188@134.[145,1431,195,1393]" captionTargetPageId="134" captionText="Figure 78. ‘Oedothorax’ nazareti Scharff, 1989.A–E, male left palp.A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, apical view. F, G, epigyne. F, ventrolateral view. G, external morphology. H, male right spinnerets. I, female left spinnerets. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968022" httpUri="https://zenodo.org/record/6968022/files/figure.png" pageId="133" pageNumber="550">Fig. 78A, E</figureCitation>
); ARP with groove fitting E, round and narrow at tip; LER absent; VRP absent; TP wide, round at tip; E retrolaterally spiral at base, prolaterally spiral at tip (
<figureCitation id="13252A54FF5876A8FCC1D23649DCF39E" box="[874,1000,1646,1668]" captionStart="Figure 78" captionStartId="134.[145,223,1433,1455]" captionTargetBox="[148,1427,198,1382]" captionTargetId="figure-188@134.[145,1431,195,1393]" captionTargetPageId="134" captionText="Figure 78. ‘Oedothorax’ nazareti Scharff, 1989.A–E, male left palp.A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, apical view. F, G, epigyne. F, ventrolateral view. G, external morphology. H, male right spinnerets. I, female left spinnerets. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968022" httpUri="https://zenodo.org/record/6968022/files/figure.png" pageId="133" pageNumber="550">Fig. 78B, E</figureCitation>
). Opisthosoma: dorsal pattern see 
<figureCitation id="13252A54FF5876A8FAD3D2364950F3B9" captionStart="Figure 68" captionStartId="111.[163,245,1308,1330]" captionTargetBox="[164,1444,196,1268]" captionTargetId="figure-26@111.[163,1446,195,1269]" captionTargetPageId="111" captionText="Figure 68. Male opisthosoma, dorsal view. A, Jilinus hulongensis (Zhu &amp; Wen, 1980). B, Cornitibia simplicithorax (Tanasevitch, 1998). C, Emertongone montifera (Emerton, 1882). D, ‘Oedothorax’ paracymbialis Tanasevitch, 2015. E, ‘Oe.’ kodaikanal Tanasevitch, 2015. F, ‘Oe.’ cunur Tanasevitch, 2015. G, ‘Oe.’ stylus Tanasevitch, 2015. H, Atypena cirrifrons (Heimer, 1984). I, A. formosana (Oi, 1977). J, ‘Oe.’ nazareti Scharff, 1989. K, Gongylidium rufipes (Linnaeus, 1758). L, Ummeliata insecticeps (Bösenberg &amp; Strand, 1906). M, U. esyunini (Zhang, Zhang &amp; Yu, 2003). N, Hylyphantes graminicola (Sundevall, 1830). O, Tmeticus tolli Kulczyński, 1908. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6967990" httpUri="https://zenodo.org/record/6967990/files/figure.png" pageId="133" pageNumber="550">Fig. 68J</figureCitation>
. PMS with mAP, two AC; PLS with triad, 3+ AC (
<figureCitation id="13252A54FF5876A8FCE8D2F44990F3DB" box="[835,932,1708,1730]" captionStart="Figure 78" captionStartId="134.[145,223,1433,1455]" captionTargetBox="[148,1427,198,1382]" captionTargetId="figure-188@134.[145,1431,195,1393]" captionTargetPageId="134" captionText="Figure 78. ‘Oedothorax’ nazareti Scharff, 1989.A–E, male left palp.A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, apical view. F, G, epigyne. F, ventrolateral view. G, external morphology. H, male right spinnerets. I, female left spinnerets. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968022" httpUri="https://zenodo.org/record/6968022/files/figure.png" pageId="133" pageNumber="550">Fig. 78H</figureCitation>
).
</paragraph>
<paragraph id="8BA136D1FF5876A8FC90D2B24F97F246" blockId="133.[827,1443,1770,1884]" pageId="133" pageNumber="550">
<emphasis id="B96AEAC3FF5876A8FC90D2B24E62F3E5" box="[827,1110,1770,1792]" italics="true" pageId="133" pageNumber="550">Female (RMCA 224.501):</emphasis>
Total length: 2.62. Prosoma: 1.03 long, 0.73 wide. Eyes: AME-AME: 0.02, AME width: 0.05, AME-ALE: 0.05, ALE width: 0.07, ALE- PLE: 0, PLE width: 0.07, PLE-PME: 0.04, PME width:
</paragraph>
<caption id="DF616659FF5B76ABFF3AD1C14BC4F0F0" ID-DOI="http://doi.org/10.5281/zenodo.6968022" ID-Zenodo-Dep="6968022" httpUri="https://zenodo.org/record/6968022/files/figure.png" pageId="134" pageNumber="551" startId="134.[145,223,1433,1455]" targetBox="[148,1427,198,1382]" targetPageId="134">
<paragraph id="8BA136D1FF5B76ABFF3AD1C14BC4F0F0" blockId="134.[145,1425,1433,1514]" pageId="134" pageNumber="551">
<emphasis id="B96AEAC3FF5B76ABFF3AD1C14B31F0B4" bold="true" box="[145,261,1433,1455]" pageId="134" pageNumber="551">Figure 78.</emphasis>
‘ 
<taxonomicName id="4C1E4D52FF5B76ABFEBFD1C14BB8F0B5" authorityName="Lin &amp; Lopardo &amp; Uhl" authorityYear="2022" box="[276,396,1433,1455]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="134" pageNumber="551" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAC3FF5B76ABFEBFD1C14BB8F0B5" box="[276,396,1433,1455]" italics="true" pageId="134" pageNumber="551">Oedothorax</emphasis>
</taxonomicName>
’ 
<taxonomicName id="4C1E4D52FF5B76ABFE3DD1C14848F0B5" authority="Scharff, 1989" authorityName="Scharff" authorityYear="1989" box="[406,636,1433,1455]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="134" pageNumber="551" phylum="Arthropoda" rank="species" species="nazareti">
<emphasis id="B96AEAC3FF5B76ABFE3DD1C14BD8F0B5" box="[406,492,1433,1455]" italics="true" pageId="134" pageNumber="551">nazareti</emphasis>
<bibRefCitation id="EF8F4B20FF5B76ABFE59D1C14848F0B5" author="Scharff N" box="[498,636,1433,1455]" pageId="134" pageNumber="551" pagination="13 - 20" refId="ref95011" refString="Scharff N, 1989. New species and records of Afrotropical Linyphiidae (Araneae). Bulletin of the British Arachnological Society 8: 13 - 20." type="journal article" year="1989">Scharff, 1989</bibRefCitation>
</taxonomicName>
. A–E, male left palp. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, apical view. F, G, epigyne. F, ventrolateral view. G, external morphology. H, male right spinnerets. I, female left spinnerets. Scale bar 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BA136D1FF5B76ABFF3AD24E4AF3F218" blockId="134.[145,762,1558,1794]" pageId="134" pageNumber="551">
0.07, PME-PME: 0.07. Clypeus: not hirsute, one sub- AME seta. Sternum: 0.59 long; 0.54 wide. Legs: dorsal proximal macroseta on tibia I, II, III and IV 1.58, 1.76, 2.26 and 2.52 times diameter of tibia, respectively; Tm I: 0.60. Epigyne: CO posteriorly situated, receptacles long, wide (
<figureCitation id="13252A54FF5B76ABFEB9D2F74BB8F3DE" box="[274,396,1711,1733]" captionStart="Figure 78" captionStartId="134.[145,223,1433,1455]" captionTargetBox="[148,1427,198,1382]" captionTargetId="figure-188@134.[145,1431,195,1393]" captionTargetPageId="134" captionText="Figure 78. ‘Oedothorax’ nazareti Scharff, 1989.A–E, male left palp.A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, apical view. F, G, epigyne. F, ventrolateral view. G, external morphology. H, male right spinnerets. I, female left spinnerets. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968022" httpUri="https://zenodo.org/record/6968022/files/figure.png" pageId="134" pageNumber="551">Fig. 78F, G</figureCitation>
). Opisthosoma: PMS with mAP, two AC, one CY; PLS with triad, two CY, 3+ AC (
<figureCitation id="13252A54FF5B76ABFD66D2964A82F218" captionStart="Figure 78" captionStartId="134.[145,223,1433,1455]" captionTargetBox="[148,1427,198,1382]" captionTargetId="figure-188@134.[145,1431,195,1393]" captionTargetPageId="134" captionText="Figure 78. ‘Oedothorax’ nazareti Scharff, 1989.A–E, male left palp.A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, apical view. F, G, epigyne. F, ventrolateral view. G, external morphology. H, male right spinnerets. I, female left spinnerets. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968022" httpUri="https://zenodo.org/record/6968022/files/figure.png" pageId="134" pageNumber="551">Fig. 78I</figureCitation>
).
</paragraph>
<paragraph id="8BA136D1FF5B76ABFF3AD3744ACEF27A" blockId="134.[145,761,1835,1888]" pageId="134" pageNumber="551">
<emphasis id="B96AEAC3FF5B76ABFF3AD3744B37F25B" box="[145,259,1836,1857]" italics="true" pageId="134" pageNumber="551">Variation:</emphasis>
The measurements are based on examined material.
</paragraph>
<paragraph id="8BA136D1FF5B76ABFC82D24E4F2EF3DF" blockId="134.[809,1425,1558,1733]" pageId="134" pageNumber="551">
<emphasis id="B96AEAC3FF5B76ABFC82D24E49B6F331" box="[809,898,1558,1579]" italics="true" pageId="134" pageNumber="551">Males (</emphasis>
N 
<emphasis id="B96AEAC3FF5B76ABFC0AD24F4EDAF331" box="[929,1262,1558,1580]" italics="true" pageId="134" pageNumber="551">= 3, means in parentheses):</emphasis>
Total length 2.22–2.46 (2.33). Prosoma: 1.06–1.14 (1.09) long, 0.72– 0.74 (0.73) wide. Legs: dorsal proximal macroseta on tibia I, II, III and IV 0.12–0.13 (0.12), 0.13–0.19 (0.16), 0.91–1.68 (1.28) and 1.46–2.28 (1.88) times diameter of tibia, respectively; Tm I: 0.58–0.67 (0.64).
</paragraph>
<paragraph id="8BA136D1FF5B76ABFC82D2B64FA4F27A" blockId="134.[809,1425,1774,1888]" pageId="134" pageNumber="551">
<emphasis id="B96AEAC3FF5B76ABFC82D2B649A2F219" box="[809,918,1774,1795]" italics="true" pageId="134" pageNumber="551">Females (</emphasis>
N 
<emphasis id="B96AEAC3FF5B76ABFC19D2B64ECCF219" box="[946,1272,1774,1795]" italics="true" pageId="134" pageNumber="551">= 10, means in parentheses):</emphasis>
Total length 2.42–3.07 (2.74). Prosoma: 1.03–1.18 (1.13) long, 0.73– 0.86 (0.79) wide. Legs: dorsal proximal macroseta on tibia I, II, III and IV 1.58–1.87 (1.72), 1.63–2.13 (1.90),
</paragraph>
<paragraph id="8BA136D1FF5A76AAFF08D49D48A0F5E0" blockId="135.[163,779,197,250]" pageId="135" pageNumber="552">1.88–2.41 (2.18) and 2.15–2.68 (2.39) times diameter of tibia, respectively; Tm I: 0.60–0.69 (0.64).</paragraph>
</subSubSection>
<subSubSection id="C304655AFF5A76AAFF08D57A4B9FF422" box="[163,427,290,312]" pageId="135" pageNumber="552" type="distribution">
<paragraph id="8BA136D1FF5A76AAFF08D57A4B9FF422" blockId="135.[163,427,290,312]" box="[163,427,290,312]" pageId="135" pageNumber="552">
<emphasis id="B96AEAC3FF5A76AAFF08D57A4B00F42D" box="[163,308,290,311]" italics="true" pageId="135" pageNumber="552">Distribution:</emphasis>
<collectingCountry id="F3097641FF5A76AAFEEAD57A4B93F422" box="[321,423,290,312]" name="Ethiopia" pageId="135" pageNumber="552">Ethiopia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C304655AFF5A76AAFF08D5394B16F48C" pageId="135" pageNumber="552" type="biology_ecology">
<paragraph id="8BA136D1FF5A76AAFF08D5394B16F48C" blockId="135.[163,776,353,406]" pageId="135" pageNumber="552">
<emphasis id="B96AEAC3FF5A76AAFF08D5394B3BF46C" box="[163,271,353,374]" italics="true" pageId="135" pageNumber="552">Habitat:</emphasis>
Under stones in cultivated lands or woodlands.
</paragraph>
</subSubSection>
<subSubSection id="C304655AFF5A76AAFF08D5E64F4EF5E0" pageId="135" pageNumber="552" type="discussion">
<paragraph id="8BA136D1FF5A76AAFF08D5E64F4EF5E0" blockId="135.[163,779,446,591]" lastBlockId="135.[827,1443,197,250]" pageId="135" pageNumber="552">
<emphasis id="B96AEAC3FF5A76AAFF08D5E64B3AF4C9" box="[163,270,446,467]" italics="true" pageId="135" pageNumber="552">Remarks:</emphasis>
According to our phylogenetic analysis, this species lack the synapomorphic traits of 
<emphasis id="B96AEAC3FF5A76AAFD2FD5854AF2F708" italics="true" pageId="135" pageNumber="552">
<taxonomicName id="4C1E4D52FF5A76AAFD2FD585493EF4E8" authorityName="Lin &amp; Lopardo &amp; Uhl" authorityYear="2022" box="[644,778,477,498]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="135" pageNumber="552" phylum="Arthropoda" rank="genus" sensu="stricto">Oedothorax</taxonomicName>
<taxonomicNameLabel id="A25957B8FF5A76AAFF08D5A54AF2F708" box="[163,198,509,530]" pageId="135" pageNumber="552" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
. Its phylogenetic position resulted in Clade 22 sister to Clade 23. However, due to the lack of shared derived characters with other taxa in this study, its taxonomic affinity is undetermined. Therefore, we leave its taxonomic state provisionally unchanged.
</paragraph>
</subSubSection>
</treatment>
</document>