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<document id="43205819AF8D9152490124EA43E71593" ID-CLB-Dataset="37322" ID-DOI="10.11646/zootaxa.4211.1.1" ID-GBIF-Dataset="7af458af-d56d-44e3-a6f7-af97868175d9" ID-ISSN="1175-5326" ID-Zenodo-Dep="272750" ID-ZooBank="6B86C6BA-6AFE-4AAD-870D-04794C138D47" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1484386885843" checkinUser="plazi" docAuthor="Antić, Dragan Ž. &amp; Makarov, Slobodan E." docDate="2016" docId="03B7878CFF82CD65FF4CEC4AB5AAE82C" docLanguage="en" docName="zootaxa.4211.1.1.pdf.imf" docOrigin="Zootaxa 4211 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Paranotosoma cordatum" docType="treatment" docVersion="8" lastPageNumber="92" masterDocId="FF8EFFF4FFD7CD3EFFDBE875B173E949" masterDocTitle="The Caucasus as a major hotspot of biodiversity: Evidence from the millipede family Anthroleucosomatidae (Diplopoda, Chordeumatida)" masterLastPageNumber="205" masterPageNumber="1" pageNumber="86" updateTime="1698435542289" updateUser="ExternalLinkService">
<mods:mods id="5F2792E135CBB8965ACEBA96BF134B77" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="2EBC91C775554C476812B6321F3A45ED">The Caucasus as a major hotspot of biodiversity: Evidence from the millipede family Anthroleucosomatidae (Diplopoda, Chordeumatida)</mods:title>
</mods:titleInfo>
<mods:name id="0F8AAEF72DC7E494CC8E885511A136F5" type="personal">
<mods:role id="907B57625909DE8A21E18C54463255A8">
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<mods:namePart id="5705877487CB1BA2DE19CDD6DED19C31">Antić, Dragan Ž.</mods:namePart>
</mods:name>
<mods:name id="813BC431AF4AC9018B54817953874C42" type="personal">
<mods:role id="F67733DA2E75DB0DF0D384B1346F4C2F">
<mods:roleTerm id="4F8E4808E76DD09DB8AD011009F58E93">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A034EF56839E53844B9C9072B6B32AA6">Makarov, Slobodan E.</mods:namePart>
</mods:name>
<mods:typeOfResource id="A62D6EF48F6A7FFD08759469BBF27EDF">text</mods:typeOfResource>
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<mods:titleInfo id="555CDEB769C8C16277E7000A6D26C04B">
<mods:title id="BF7C7D63BC71487341F1581D499BA4C2">Zootaxa</mods:title>
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<mods:part id="BC8ECB586F29473AB0B85870B47381DC">
<mods:date id="C4BA8A76A6B56A23B886CC78D0109A74">2016</mods:date>
<mods:detail id="24253C3C04AEC165171EA039BE410F54" type="volume">
<mods:number id="4162D30E7DAD091F4787042E3BB157BC">4211</mods:number>
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<mods:classification id="492AEEA6F1E2D9E9DBD480DD65963960">journal article</mods:classification>
<mods:identifier id="DF54EDA92C9F5CB6F9ED632DA6FC0C35" type="CLB-Dataset">37322</mods:identifier>
<mods:identifier id="1ADCD42D8700CD61719C1694C2E125B9" type="DOI">10.11646/zootaxa.4211.1.1</mods:identifier>
<mods:identifier id="49E945EBA8D49229F11B683C44909F12" type="GBIF-Dataset">7af458af-d56d-44e3-a6f7-af97868175d9</mods:identifier>
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<mods:identifier id="3F1E0DC49EEF3E819F84FB78784880DD" type="Zenodo-Dep">272750</mods:identifier>
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<treatment id="03B7878CFF82CD65FF4CEC4AB5AAE82C" ID-DOI="http://doi.org/10.5281/zenodo.6074417" ID-GBIF-Taxon="127419157" ID-Zenodo-Dep="6074417" LSID="urn:lsid:plazi:treatment:03B7878CFF82CD65FF4CEC4AB5AAE82C" httpUri="http://treatment.plazi.org/id/03B7878CFF82CD65FF4CEC4AB5AAE82C" lastPageId="91" lastPageNumber="92" pageId="85" pageNumber="86">
<subSubSection id="C3046511FF82CD6BFF4CEC4AB067ED32" pageId="85" pageNumber="86" type="nomenclature">
<paragraph id="8BA1369AFF82CD6BFF4CEC4AB30CED13" blockId="85.[151,639,1087,1147]" box="[151,639,1087,1114]" pageId="85" pageNumber="86">
<heading id="D0E981F6FF82CD6BFF4CEC4AB30CED13" bold="true" box="[151,639,1087,1114]" fontSize="11" level="1" pageId="85" pageNumber="86" reason="1">
<emphasis id="B96AEA88FF82CD6BFF4CEC4AB30CED13" bold="true" box="[151,639,1087,1114]" pageId="85" pageNumber="86">
<taxonomicName id="4C1E4D19FF82CD6BFF4CEC4AB0B1ED10" ID-CoL="84C22" box="[151,450,1087,1113]" class="Diplopoda" family="Anthroleucosomatidae" genus="Paranotosoma" kingdom="Animalia" order="Chordeumatida" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="cordatum">
<emphasis id="B96AEA88FF82CD6BFF4CEC4AB0B1ED10" bold="true" box="[151,450,1087,1113]" italics="true" pageId="85" pageNumber="86">Paranotosoma cordatum</emphasis>
</taxonomicName>
gen. et sp. nov.
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1369AFF82CD6BFF4CEC17B067ED32" blockId="85.[151,639,1087,1147]" box="[151,276,1122,1147]" pageId="85" pageNumber="86">
<figureCitation id="13252A1FFF82CD6BFF4CEC17B067ED32" box="[151,276,1122,1147]" captionStart-0="FIGURE 69" captionStart-1="FIGURE 70" captionStart-2="FIGURE 71" captionStart-3="FIGURE 72" captionStart-4="FIGURE 73" captionStartId-0="86.[151,250,1333,1355]" captionStartId-1="87.[151,250,1964,1986]" captionStartId-2="88.[151,250,1852,1874]" captionStartId-3="89.[151,250,1755,1777]" captionStartId-4="90.[151,250,1576,1598]" captionTargetBox-0="[188,1398,445,1312]" captionTargetBox-1="[208,1371,193,1937]" captionTargetBox-2="[189,1398,193,1830]" captionTargetBox-3="[200,1390,223,1668]" captionTargetBox-4="[188,1398,193,1554]" captionTargetId-0="figure@86.[188,1398,445,1312]" captionTargetId-1="figure@87.[208,1371,193,1937]" captionTargetId-2="figure@88.[188,1398,193,1830]" captionTargetId-3="figure@89.[188,1398,193,1734]" captionTargetId-4="figure@90.[188,1398,193,1554]" captionTargetPageId-0="86" captionTargetPageId-1="87" captionTargetPageId-2="88" captionTargetPageId-3="89" captionTargetPageId-4="90" captionText-0="FIGURE 69. Paranotosoma cordatum gen. et sp. nov., paratype male habitus. Scale line = 1 mm." captionText-1="FIGURE 70. Paranotosoma cordatum gen. et sp. nov., paratype male, pleurotergite 15. A. Posterior view. B. Lateral view. C. Prozonite, dorsal view. D. Lateral keel, dorsal view. E. Dorsal view." captionText-2="FIGURE 71. Paranotosoma cordatum gen. et sp. nov., paratype male. A. Leg 3. B. Leg 4. C. Leg 7. Scale lines = 0.2 mm." captionText-3="FIGURE 72. Paranotosoma cordatum gen. et sp. nov., holotype male gonopods. A. Anterior gonopods, anterior view. B. Anterior gonopods, lateral view. C. Anterior gonopods, posterior view. D. Posterior gonopods, posterior view. c = coxite; h = horn; t = telopodite; ah = acuminate horn; ap / cp = anterior coxal process; cv = coxal vesicle; lg = longitudinal groove; lv = posterior lever; pp = posterior coxal process; rs = row of setae; sp = sternal plate; ss = sternal sac; alv = anterior branch of lever; mdl = medial lamella; plv = posterior branch of lever. Scale line = 0.2 mm." captionText-4="FIGURE 73. Paranotosoma cordatum gen. et sp. nov., paratype male anterior gonopods. A. Anterior view. B. Detail, anterior view. C. Lateral view. D. Posterolateral view. E. Posterior view. F. Distal view. h = horn; ah = acuminate horn; ap / cp = anterior coxal process; cv = coxal vesicle; lg = longitudinal groove; lv = posterior lever; pp = posterior coxal process; rs = row of setae; sp = sternal plate; ss = sternal sac; alv = anterior branch of lever; mdl = medial lamella; plv = posterior branch of lever." httpUri-0="https://zenodo.org/record/272818/files/figure.png" httpUri-1="https://zenodo.org/record/272819/files/figure.png" httpUri-2="https://zenodo.org/record/272820/files/figure.png" httpUri-3="https://zenodo.org/record/272821/files/figure.png" httpUri-4="https://zenodo.org/record/272822/files/figure.png" pageId="85" pageNumber="86">Figs 6973</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3046511FF82CD6BFF4CECDFB3DAEC43" pageId="85" pageNumber="86" type="diagnosis">
<paragraph id="8BA1369AFF82CD6BFF4CECDFB3DAEC43" blockId="85.[151,1437,1194,2011]" pageId="85" pageNumber="86">
<emphasis id="B96AEA88FF82CD6BFF4CECDFB062ED8A" bold="true" box="[151,273,1194,1219]" pageId="85" pageNumber="86">Diagnosis.</emphasis>
Differs from the other two species of the genus by the presence of cordiform anterior shields, by the absence of any lateral processes in the posterior part of the anterior gonopods and by the development of rudiments of posterior coxal processes on male leg-pair 7.
</paragraph>
</subSubSection>
<subSubSection id="C3046511FF82CD6BFF1CED63B078EC1A" pageId="85" pageNumber="86" type="etymology">
<paragraph id="8BA1369AFF82CD6BFF1CED63B078EC1A" blockId="85.[151,1437,1194,2011]" pageId="85" pageNumber="86">
<emphasis id="B96AEA88FF82CD6BFF1CED63B03DEC66" bold="true" box="[199,334,1302,1327]" pageId="85" pageNumber="86">Etymology.</emphasis>
An adjective; the new species is named after the cordiform anterior shield of the anterior gonopods.
</paragraph>
</subSubSection>
<subSubSection id="C3046511FF82CD6BFF1CED2BB57CEF96" pageId="85" pageNumber="86" type="materials_examined">
<paragraph id="8BA1369AFF82CD6BFF1CED2BB5DCECD2" blockId="85.[151,1437,1194,2011]" pageId="85" pageNumber="86">
<materialsCitation id="3B763CC7FF82CD6BFF1CED2BB54FEC3E" ID-GBIF-Occurrence="1424567028" box="[199,1084,1374,1399]" country="Georgia" location="GEORGIA" pageId="85" pageNumber="86" specimenCount="22" specimenCount-female="9" specimenCount-male="13" typeStatus="holotype">
<emphasis id="B96AEA88FF82CD6BFF1CED2BB0E2EC3E" bold="true" box="[199,401,1374,1399]" pageId="85" pageNumber="86">Material studied</emphasis>
(total:
<specimenCount id="9D18FD13FF82CD6BFE30ED2AB324EC3E" box="[491,599,1374,1399]" pageId="85" pageNumber="86" type="male">13 males</specimenCount>
,
<specimenCount id="9D18FD13FF82CD6BFDBDED2AB3AFEC3E" box="[614,732,1374,1399]" pageId="85" pageNumber="86" type="female">9 females</specimenCount>
).
<emphasis id="B96AEA88FF82CD6BFD28ED2BB214EC3E" bold="true" box="[755,871,1374,1399]" pageId="85" pageNumber="86">
<typeStatus id="54A58838FF82CD6BFD28ED2BB211EC3E" box="[755,866,1374,1399]" pageId="85" pageNumber="86" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCountry id="F309760AFF82CD6BFCA9ED2AB286EC3E" box="[882,1013,1375,1399]" name="Georgia" pageId="85" pageNumber="86">GEORGIA</collectingCountry>
: male
</materialsCitation>
,
<materialsCitation id="3B763CC7FF82CD6BFB97ED2BB5DCECD2" ID-GBIF-Occurrence="1424567033" collectingDate="1981-10-09" collectionCode="ZMUM" collectorName="S. Golovatch" country="Georgia" elevation="85" location="Batumi Botanical Garden" municipality="Zelenyi Mys" pageId="85" pageNumber="86" specimenCount="1" stateProvince="Ajaria" typeStatus="holotype">
<collectingRegion id="49DAF878FF82CD6BFB97ED2BB5E6EC3E" box="[1100,1173,1374,1399]" country="Georgia" name="Ajaria" pageId="85" pageNumber="86">Ajaria</collectingRegion>
,
<collectingMunicipality id="6BC5ACE0FF82CD6BFB7FED2BB449EC3E" box="[1188,1338,1374,1399]" pageId="85" pageNumber="86">Zelenyi Mys</collectingMunicipality>
,
<location id="8EC16041FF82CD6BFA92ED2BB02CECD2" LSID="urn:lsid:plazi:treatment:03B7878CFF82CD65FF4CEC4AB5AAE82C:8EC16041FF82CD6BFA92ED2BB02CECD2" country="Georgia" municipality="Zelenyi Mys" name="Batumi Botanical Garden" pageId="85" pageNumber="86" stateProvince="Ajaria">Batumi Botanical Garden</location>
, forest litter,
<elevation id="0033D1A9FF82CD6BFE29EDF7B310ECD3" box="[498,611,1410,1435]" metricMagnitude="1" metricUnit="m" metricValue="8.5" metricValueMax="15.0" metricValueMin="2.0" pageId="85" pageNumber="86" unit="m" value="85.0" valueMax="150.0" valueMin="20.0">
<quantity id="4CE69B7FFF82CD6BFE29EDF7B310ECD3" box="[498,611,1410,1435]" metricMagnitude="1" metricUnit="m" metricValue="8.5" metricValueMax="15.0" metricValueMin="2.0" pageId="85" pageNumber="86" unit="m" value="85.0" valueMax="150.0" valueMin="20.0">20150 m</quantity>
</elevation>
asl,
<collectingDate id="EFE4E9B2FF82CD6BFD4CEDF6B26FECD2" box="[663,796,1411,1435]" pageId="85" pageNumber="86" value="1981-10-09">
<date id="FFA0105AFF82CD6BFD4CEDF6B26FECD2" box="[663,796,1411,1435]" pageId="85" pageNumber="86" value="1981-10-09">9 Oct. 1981</date>
</collectingDate>
,
<collectorName id="26EB534CFF82CD6BFCF3EDF6B2C8ECD2" box="[808,955,1410,1435]" pageId="85" pageNumber="86">S. Golovatch</collectorName>
leg. (
<collectionCode id="ED0FAE5FFF82CD6BFC22EDF6B521ECD2" box="[1017,1106,1411,1435]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="85" pageNumber="86">ZMUM</collectionCode>
ρ3311).
</materialsCitation>
</paragraph>
<paragraph id="8BA1369AFF82CD6BFF1CEDD3B36CEF4E" blockId="85.[151,1437,1194,2011]" pageId="85" pageNumber="86">
<emphasis id="B96AEA88FF82CD6BFF1CEDD3B04CECF6" bold="true" box="[199,319,1446,1471]" pageId="85" pageNumber="86">Paratypes</emphasis>
(total: 5 males, 5 females).
<materialsCitation id="3B763CC7FF82CD6BFDA2EDD3B09AECAA" ID-GBIF-Occurrence="1424567027" collectingDate="1978-10-13" collectionCode="ZMUM" country="Georgia" location="All" pageId="85" pageNumber="86" specimenCount="4" specimenCount-female="3" specimenCount-male="1">
<location id="8EC16041FF82CD6BFDA2EDD3B3EEECF6" LSID="urn:lsid:plazi:treatment:03B7878CFF82CD65FF4CEC4AB5AAE82C:8EC16041FF82CD6BFDA2EDD3B3EEECF6" box="[633,669,1446,1471]" country="Georgia" name="All" pageId="85" pageNumber="86">All</location>
from
<collectingCountry id="F309760AFF82CD6BFD39EDD2B216ECF6" box="[738,869,1447,1471]" name="Georgia" pageId="85" pageNumber="86">GEORGIA</collectingCountry>
:
<specimenCount id="9D18FD13FF82CD6BFCB4EDD2B2CDECF6" box="[879,958,1446,1471]" pageId="85" pageNumber="86" type="male">1 male</specimenCount>
,
<specimenCount id="9D18FD13FF82CD6BFC11EDD2B54AECF6" box="[970,1081,1446,1471]" pageId="85" pageNumber="86" type="female">3 females</specimenCount>
, same data as holotype, except:
<collectingDate id="EFE4E9B2FF82CD6BFF4CEDBEB05BECAA" box="[151,296,1483,1507]" pageId="85" pageNumber="86" value="1978-10-13">
<date id="FFA0105AFF82CD6BFF4CEDBEB05BECAA" box="[151,296,1483,1507]" pageId="85" pageNumber="86" value="1978-10-13">13 Oct. 1978</date>
</collectingDate>
(
<collectionCode id="ED0FAE5FFF82CD6BFEECEDBEB0E3ECAA" box="[311,400,1483,1507]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="85" pageNumber="86">ZMUM</collectionCode>
ρ3312)
</materialsCitation>
;
<materialsCitation id="3B763CC7FF82CD6BFE2FEDBEB36CEF4E" ID-GBIF-Occurrence="1424567041" collectingDate="1981-10-10" collectionCode="ZMUM, IZB" collectorName="J. Bohac" country="Georgia" location="GEORGIA" pageId="85" pageNumber="86" specimenCount="6" specimenCount-female="2" specimenCount-male="4">
<specimenCount id="9D18FD13FF82CD6BFE2FEDBEB33EECAA" box="[500,589,1482,1507]" pageId="85" pageNumber="86" type="male">2 males</specimenCount>
,
<specimenCount id="9D18FD13FF82CD6BFD83EDBEB3C9ECAA" box="[600,698,1482,1507]" pageId="85" pageNumber="86" type="female">1 female</specimenCount>
(
<collectionCode id="ED0FAE5FFF82CD6BFD13EDBEB252ECAA" box="[712,801,1483,1507]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="85" pageNumber="86">ZMUM</collectionCode>
ρ3313),
<specimenCount id="9D18FD13FF82CD6BFC5EEDBEB2ADECAA" box="[901,990,1482,1507]" pageId="85" pageNumber="86" type="male">2 males</specimenCount>
,
<specimenCount id="9D18FD13FF82CD6BFC32EDBEB539ECAA" box="[1001,1098,1482,1507]" pageId="85" pageNumber="86" type="female">1 female</specimenCount>
(
<collectionCode id="ED0FAE5FFF82CD6BFB81EDBEB5F8ECAA" box="[1114,1163,1483,1507]" pageId="85" pageNumber="86">IZB</collectionCode>
), same data as holotype, except:
<collectingDate id="EFE4E9B2FF82CD6BFF34ED9AB0F1EF4E" box="[239,386,1519,1543]" pageId="85" pageNumber="86" value="1981-10-10">
<date id="FFA0105AFF82CD6BFF34ED9AB0F1EF4E" box="[239,386,1519,1543]" pageId="85" pageNumber="86" value="1981-10-10">10 Oct. 1981</date>
</collectingDate>
,
<collectorName id="26EB534CFF82CD6BFE55ED9AB09DEF4E" box="[398,494,1518,1543]" pageId="85" pageNumber="86">J. Boháč</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="8BA1369AFF82CD6BFF1CEE67B372EFF2" blockId="85.[151,1437,1194,2011]" pageId="85" pageNumber="86">
<materialsCitation id="3B763CC7FF82CD6BFF1CEE67B3C2EF62" ID-GBIF-Occurrence="1424567032" box="[199,689,1554,1579]" location="Other" pageId="85" pageNumber="86" specimenCount="11" specimenCount-female="4" specimenCount-male="7">
<emphasis id="B96AEA88FF82CD6BFF1CEE67B00FEF62" bold="true" box="[199,380,1554,1579]" pageId="85" pageNumber="86">
<location id="8EC16041FF82CD6BFF1CEE67B07CEF62" LSID="urn:lsid:plazi:treatment:03B7878CFF82CD65FF4CEC4AB5AAE82C:8EC16041FF82CD6BFF1CEE67B07CEF62" box="[199,271,1554,1579]" name="Other" pageId="85" pageNumber="86">Other</location>
material
</emphasis>
(total:
<specimenCount id="9D18FD13FF82CD6BFE14EE66B359EF62" box="[463,554,1554,1579]" pageId="85" pageNumber="86" type="male">7 males</specimenCount>
,
<specimenCount id="9D18FD13FF82CD6BFDEDEE66B3DAEF62" box="[566,681,1554,1579]" pageId="85" pageNumber="86" type="female">4 females</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="3B763CC7FF82CD6BFD66EE67B5F1EF62" ID-GBIF-Occurrence="1424567038" box="[701,1154,1554,1579]" country="Georgia" location="All" pageId="85" pageNumber="86" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<location id="8EC16041FF82CD6BFD66EE67B392EF62" LSID="urn:lsid:plazi:treatment:03B7878CFF82CD65FF4CEC4AB5AAE82C:8EC16041FF82CD6BFD66EE67B392EF62" box="[701,737,1554,1579]" country="Georgia" name="All" pageId="85" pageNumber="86">All</location>
from
<collectingCountry id="F309760AFF82CD6BFCFCEE66B2D9EF62" box="[807,938,1555,1579]" name="Georgia" pageId="85" pageNumber="86">GEORGIA</collectingCountry>
:
<specimenCount id="9D18FD13FF82CD6BFC6DEE66B563EF62" box="[950,1040,1554,1579]" pageId="85" pageNumber="86" type="male">2 males</specimenCount>
,
<specimenCount id="9D18FD13FF82CD6BFBC6EE66B5F1EF62" box="[1053,1154,1554,1579]" pageId="85" pageNumber="86" type="female">1 female</specimenCount>
</materialsCitation>
,
<materialsCitation id="3B763CC7FF82CD6BFB54EE67B021EF3A" ID-GBIF-Occurrence="1424567037" collectingDate="1981-10-13" collectionCode="ZMUM" collectorName="S. Golovatch" country="Georgia" elevation="525" location="Kintrishi Nature Reserve" pageId="85" pageNumber="86" specimenCount="1" stateProvince="Ajaria">
<collectingRegion id="49DAF878FF82CD6BFB54EE67B5ABEF62" box="[1167,1240,1554,1579]" country="Georgia" name="Ajaria" pageId="85" pageNumber="86">Ajaria</collectingRegion>
,
<location id="8EC16041FF82CD6BFB3FEE67B187EF07" LSID="urn:lsid:plazi:treatment:03B7878CFF82CD65FF4CEC4AB5AAE82C:8EC16041FF82CD6BFB3FEE67B187EF07" country="Georgia" name="Kintrishi Nature Reserve" pageId="85" pageNumber="86" stateProvince="Ajaria">Kintrishi Nature Reserve</location>
,
<taxonomicName id="4C1E4D19FF82CD6BFED8EE43B00FEF06" box="[259,380,1590,1615]" class="Diplopoda" family="Anthroleucosomatidae" genus="Dentatosoma" kingdom="Animalia" order="Chordeumatida" pageId="85" pageNumber="202" phylum="Arthropoda" rank="species" species="zeraboseli">Zeraboseli</taxonomicName>
,
<elevation id="0033D1A9FF82CD6BFE57EE43B37CEF07" box="[396,527,1590,1615]" metricMagnitude="2" metricUnit="m" metricValue="5.25" metricValueMax="6.0" metricValueMin="4.5" pageId="85" pageNumber="86" unit="m" value="525.0" valueMax="600.0" valueMin="450.0">
<quantity id="4CE69B7FFF82CD6BFE57EE43B37CEF07" box="[396,527,1590,1615]" metricMagnitude="2" metricUnit="m" metricValue="5.25" metricValueMax="6.0" metricValueMin="4.5" pageId="85" pageNumber="86" unit="m" value="525.0" valueMax="600.0" valueMin="450.0">450600 m</quantity>
</elevation>
asl, deciduous forest, litter and under stones,
<collectingDate id="EFE4E9B2FF82CD6BFBEAEE42B5B1EF06" box="[1073,1218,1591,1615]" pageId="85" pageNumber="86" value="1981-10-13">
<date id="FFA0105AFF82CD6BFBEAEE42B5B1EF06" box="[1073,1218,1591,1615]" pageId="85" pageNumber="86" value="1981-10-13">13 Oct.1981</date>
</collectingDate>
,
<collectorName id="26EB534CFF82CD6BFB0AEE42B41BEF06" box="[1233,1384,1590,1615]" pageId="85" pageNumber="86">S. Golovatch</collectorName>
leg. (
<collectionCode id="ED0FAE5FFF82CD6BFF45EE2EB184EF3A" box="[158,247,1627,1651]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="85" pageNumber="86">ZMUM</collectionCode>
ρ3314)
</materialsCitation>
;
<materialsCitation id="3B763CC7FF82CD6BFE85EE2EB468EF3A" ID-GBIF-Occurrence="1424567036" box="[350,1307,1626,1651]" collectingDate="1981-10-13" collectionCode="ZMUM" collectorName="S. Golovatch" country="Georgia" elevation="800" location="Kintrishi Nature Reserve" pageId="85" pageNumber="86" specimenCount="2" specimenCount-male="2" stateProvince="Ajaria">
<specimenCount id="9D18FD13FF82CD6BFE85EE2EB0CBEF3A" box="[350,440,1626,1651]" pageId="85" pageNumber="86" type="male">2 males</specimenCount>
, same data, except:
<elevation id="0033D1A9FF82CD6BFD46EE2EB396EF3B" box="[669,741,1627,1651]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="85" pageNumber="86" unit="m" value="800.0">
<quantity id="4CE69B7FFF82CD6BFD46EE2EB396EF3B" box="[669,741,1627,1651]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="85" pageNumber="86" unit="m" value="800.0">800 m</quantity>
</elevation>
asl,
<taxonomicName id="4C1E4D19FF82CD6BFCC0EE2FB2B3EF3A" box="[795,960,1626,1651]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="85" pageNumber="86" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEA88FF82CD6BFCC0EE2FB2B3EF3A" box="[795,960,1626,1651]" italics="true" pageId="85" pageNumber="86">Rhododendron</emphasis>
</taxonomicName>
thicket, litter (
<collectionCode id="ED0FAE5FFF82CD6BFBBCEE2EB5B3EF3A" box="[1127,1216,1627,1651]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="85" pageNumber="86">ZMUM</collectionCode>
ρ3315)
</materialsCitation>
;
<materialsCitation id="3B763CC7FF82CD6BFAF3EE2EB372EFF2" ID-GBIF-Occurrence="1424567039" collectingDate="1981-10-26" collectionCode="ZMUM" collectorName="S. Golovatch" country="Georgia" location="Cave Belaya" municipality="Tskhaltubo" pageId="85" pageNumber="86" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Ajaria">
<specimenCount id="9D18FD13FF82CD6BFAF3EE2EB4F1EF3A" box="[1320,1410,1626,1651]" pageId="85" pageNumber="86" type="male">3 males</specimenCount>
,
<specimenCount id="9D18FD13FF82CD6BFA55EE2EB182EFDE" pageId="85" pageNumber="86" type="female">3 females</specimenCount>
,
<collectingMunicipality id="6BC5ACE0FF82CD6BFF27EE0BB008EFDE" box="[252,379,1662,1687]" pageId="85" pageNumber="86">Tskhaltubo</collectingMunicipality>
, near
<location id="8EC16041FF82CD6BFE66EE0AB33FEFDE" LSID="urn:lsid:plazi:treatment:03B7878CFF82CD65FF4CEC4AB5AAE82C:8EC16041FF82CD6BFE66EE0AB33FEFDE" box="[445,588,1662,1687]" country="Georgia" municipality="Tskhaltubo" name="Cave Belaya" pageId="85" pageNumber="86" stateProvince="Ajaria">Cave Belaya</location>
(= Tetri), deciduous forest on rock, litter and under stones,
<collectingDate id="EFE4E9B2FF82CD6BFB3DEE0BB409EFDE" box="[1254,1402,1662,1687]" pageId="85" pageNumber="86" value="1981-10-26">
<date id="FFA0105AFF82CD6BFB3DEE0BB409EFDE" box="[1254,1402,1662,1687]" pageId="85" pageNumber="86" value="1981-10-26">26 Oct. 1981</date>
</collectingDate>
,
<collectorName id="26EB534CFF82CD6BFA5DEE0AB07FEFF2" pageId="85" pageNumber="86">S. Golovatch</collectorName>
leg. (
<collectionCode id="ED0FAE5FFF82CD6BFE91EED6B0D0EFF2" box="[330,419,1699,1723]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="85" pageNumber="86">ZMUM</collectionCode>
ρ3316).
</materialsCitation>
</paragraph>
<paragraph id="8BA1369AFF82CD6BFF1CEEB3B57CEF96" blockId="85.[151,1437,1194,2011]" box="[199,1039,1734,1759]" pageId="85" pageNumber="86">
<materialsCitation id="3B763CC7FF82CD6BFF1CEEB3B09CEF96" ID-GBIF-Occurrence="1424567034" box="[199,495,1734,1759]" country="Georgia" location="GEORGIA" pageId="85" pageNumber="86" specimenCount="1" typeStatus="holotype">
<emphasis id="B96AEA88FF82CD6BFF1CEEB3B016EF96" bold="true" box="[199,357,1734,1759]" pageId="85" pageNumber="86">
<typeStatus id="54A58838FF82CD6BFF1CEEB3B071EF96" box="[199,258,1734,1759]" pageId="85" pageNumber="86">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F309760AFF82CD6BFEB7EEB2B09CEF96" box="[364,495,1735,1759]" name="Georgia" pageId="85" pageNumber="86">GEORGIA</collectingCountry>
</materialsCitation>
:
<materialsCitation id="3B763CC7FF82CD6BFE22EEB3B57CEF96" ID-GBIF-Occurrence="1424567043" box="[505,1039,1734,1759]" country="Georgia" location="Ajaria" pageId="85" pageNumber="86" specimenCount="1" stateProvince="Ajaria" typeStatus="holotype">
<collectingRegion id="49DAF878FF82CD6BFE22EEB3B331EF96" box="[505,578,1734,1759]" country="Georgia" name="Ajaria" pageId="85" pageNumber="86">Ajaria</collectingRegion>
, Zelenyi Mys, Batumi Botanical Garden.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3046511FF82CD64FF1CEE9FB4FBEEFE" lastPageId="90" lastPageNumber="91" pageId="85" pageNumber="86" type="description">
<paragraph id="8BA1369AFF82CD6BFF1CEE9FB2D1EE4A" blockId="85.[151,1437,1194,2011]" box="[199,930,1770,1795]" pageId="85" pageNumber="86">
<emphasis id="B96AEA88FF82CD6BFF1CEE9FB02AEE4A" bold="true" box="[199,345,1770,1795]" pageId="85" pageNumber="86">Description.</emphasis>
Body with 30 segments (including telson) in adults.
</paragraph>
<paragraph id="8BA1369AFF82CD6BFF1CEF7AB2B2EE03" blockId="85.[151,1437,1194,2011]" pageId="85" pageNumber="86">
MEASUREMENTS. Males
<quantity id="4CE69B7FFF82CD6BFE07EF7BB326EE6F" box="[476,597,1806,1831]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="85" pageNumber="86" unit="mm" value="13.5" valueMax="15.0" valueMin="12.0">1215 mm</quantity>
long, vertical diameter of the largest pleurotergite
<quantity id="4CE69B7FFF82CD6BFB55EF7AB470EE6F" box="[1166,1283,1807,1831]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="85" pageNumber="86" unit="mm" value="1.05" valueMax="1.1" valueMin="1.0">11.1 mm</quantity>
. Females
<quantity id="4CE69B7FFF82CD6BFAA9EF7AB188EE03" metricMagnitude="-2" metricUnit="m" metricValue="1.425" metricValueMax="1.65" metricValueMin="1.2" pageId="85" pageNumber="86" unit="mm" value="14.25" valueMax="16.5" valueMin="12.0">12 16.5 mm</quantity>
long, vertical diameter of the largest pleurotergite
<quantity id="4CE69B7FFF82CD6BFCE8EF46B2CEEE03" box="[819,957,1843,1867]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="85" pageNumber="86" unit="mm" value="1.25" valueMax="1.3" valueMin="1.2">1.21.3 mm</quantity>
.
</paragraph>
<paragraph id="8BA1369AFF82CD6BFF1CEF22B390EEDA" blockId="85.[151,1437,1194,2011]" pageId="85" pageNumber="86">
COLORATION (
<figureCitation id="13252A1FFF82CD6BFEB7EF23B0B2EE26" box="[364,449,1878,1903]" captionStart="FIGURE 69" captionStartId="86.[151,250,1333,1355]" captionTargetBox="[188,1398,445,1312]" captionTargetId="figure@86.[188,1398,445,1312]" captionTargetPageId="86" captionText="FIGURE 69. Paranotosoma cordatum gen. et sp. nov., paratype male habitus. Scale line = 1 mm." httpUri="https://zenodo.org/record/272818/files/figure.png" pageId="85" pageNumber="86">Fig. 69</figureCitation>
). Dorsal and dorsolateral sides greyish to brownish with yellowish spots dorsolaterally. Ventrolateral sides yellowish. Antennae dark brown.
</paragraph>
<paragraph id="8BA1369AFF82CD6BFF1CEFEAB4EEEE93" blockId="85.[151,1437,1194,2011]" pageId="85" pageNumber="86">HEAD. Slightly concave in males. Labrum with three medial teeth and 6+6 labral and 2+2 supralabral setae. Promentum triangular, without setae. Lingual plates with 8+8 setae, on each plate arranged in two irregular rows.</paragraph>
<paragraph id="8BA1369AFF81CD68FF4CE8E2B023E809" blockId="86.[151,1437,151,391]" pageId="86" pageNumber="87">
Stipites with 25+25 setae. Antennae
<quantity id="4CE69B7FFF81CD68FDEFE8EDB3EAE9E6" box="[564,665,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="2.07" pageId="86" pageNumber="87" unit="mm" value="2.07">2.07 mm</quantity>
long in
<typeStatus id="54A58838FF81CD68FD2CE8EDB224E9F9" box="[759,855,152,176]" pageId="86" pageNumber="87" type="paratype">paratype</typeStatus>
male. Length of antennomeres (in mm): I (0.11), II (0.2), III (0.58), IV (0.28), V (0.52), VI (0.22), VII (0.14) and VIII (0.02). Length/breadth ratios of antennomeres I VII: I (1), II (1.7), III (5.8), IV (2.8), V (4.3), VI (1.7) and VII (1.4). Antennomeres II, IV, V, VI and VII with one, three, one, four and one sensillum, respectively. Number of ocelli 2228, arranged in 6 rows in males;
<quantity id="4CE69B7FFF81CD68FAF8E971B4F4E854" box="[1315,1415,260,285]" metricMagnitude="-1" metricUnit="m" metricValue="6.604" metricValueMax="6.858" metricValueMin="6.35" pageId="86" pageNumber="87" unit="in" value="26.0" valueMax="27.0" valueMin="25.0">2527 in</quantity>
6 rows in females.
</paragraph>
<paragraph id="8BA1369AFF81CD68FF1CE939B189E8CE" blockId="86.[151,1437,151,391]" pageId="86" pageNumber="87">COLLUM. Narrower than head, with six macrochaetae. Anterior edge semi-circular, posterior margin gently concave.</paragraph>
<caption id="DF616612FF81CD68FF4CED40B50FEC02" box="[151,1148,1333,1356]" httpUri="https://zenodo.org/record/272818/files/figure.png" pageId="86" pageNumber="87" targetBox="[188,1398,445,1312]" targetPageId="86">
<paragraph id="8BA1369AFF81CD68FF4CED40B50FEC02" blockId="86.[151,1148,1333,1356]" box="[151,1148,1333,1356]" pageId="86" pageNumber="87">
<emphasis id="B96AEA88FF81CD68FF4CED40B053EC02" bold="true" box="[151,288,1333,1355]" pageId="86" pageNumber="87">FIGURE 69.</emphasis>
<taxonomicName id="4C1E4D19FF81CD68FEFEED42B352EC02" box="[293,545,1333,1356]" class="Diplopoda" family="Anthroleucosomatidae" genus="Paranotosoma" kingdom="Animalia" order="Chordeumatida" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="cordatum">
<emphasis id="B96AEA88FF81CD68FEFEED42B352EC02" box="[293,545,1333,1356]" italics="true" pageId="86" pageNumber="87">Paranotosoma cordatum</emphasis>
</taxonomicName>
<emphasis id="B96AEA88FF81CD68FDFCED43B3B7EC02" bold="true" box="[551,708,1334,1355]" pageId="86" pageNumber="87">gen. et sp. nov.</emphasis>
, paratype male habitus. Scale line = 1 mm.
</paragraph>
</caption>
<paragraph id="8BA1369AFF81CD68FF1CED01B0D7ECB1" blockId="86.[151,1436,1395,2032]" pageId="86" pageNumber="87">
BODY SEGMENTS (
<figureCitation id="13252A1FFF81CD68FE4EED06B098ECC5" box="[405,491,1395,1420]" captionStart="FIGURE 69" captionStartId="86.[151,250,1333,1355]" captionTargetBox="[188,1398,445,1312]" captionTargetId="figure@86.[188,1398,445,1312]" captionTargetPageId="86" captionText="FIGURE 69. Paranotosoma cordatum gen. et sp. nov., paratype male habitus. Scale line = 1 mm." httpUri="https://zenodo.org/record/272818/files/figure.png" pageId="86" pageNumber="87">Figs 69</figureCitation>
,
<figureCitation id="13252A1FFF81CD68FE2CED06B364ECC2" box="[503,535,1395,1419]" captionStart="FIGURE 70" captionStartId="87.[151,250,1964,1986]" captionTargetBox="[208,1371,193,1937]" captionTargetId="figure@87.[208,1371,193,1937]" captionTargetPageId="87" captionText="FIGURE 70. Paranotosoma cordatum gen. et sp. nov., paratype male, pleurotergite 15. A. Posterior view. B. Lateral view. C. Prozonite, dorsal view. D. Lateral keel, dorsal view. E. Dorsal view." httpUri="https://zenodo.org/record/272819/files/figure.png" pageId="86" pageNumber="87">70</figureCitation>
). Lateral keels developed, but to a lesser degree than in
<taxonomicName id="4C1E4D19FF81CD68FB5FED01B43BECC5" box="[1156,1352,1395,1420]" class="Diplopoda" family="Anthroleucosomatidae" genus="Paranotosoma" kingdom="Animalia" order="Chordeumatida" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="subrotundatum">
<emphasis id="B96AEA88FF81CD68FB5FED01B43BECC5" box="[1156,1352,1395,1420]" italics="true" pageId="86" pageNumber="87">P. subrotundatum</emphasis>
</taxonomicName>
<emphasis id="B96AEA88FF81CD68FA95ED06B186ECF9" bold="true" pageId="86" pageNumber="87">gen. et sp. nov.</emphasis>
, more prominent in males. Inner and median macrochaetae shorter than outer one, medium-sized, all trichoid. CIX (pleurotergite 15) ~ 0.65; MIX (pleurotergite 15) ~ 0.8; PIX (pleurotergite 15) ~ 0.6; MA (pleurotergite 15) ~ 95˚.
</paragraph>
<paragraph id="8BA1369AFF81CD68FF1CEE71B39FEF76" blockId="86.[151,1436,1395,2032]" pageId="86" pageNumber="87">TELSON. Epiproct with a pair of spinnerets and 3+3 setae (1+1 paramedian, 2+2 marginal). Hypoproct with 1+1 apical setae. Paraprocts with 3+3 marginal setae.</paragraph>
<paragraph id="8BA1369AFF81CD68FF1CEE39B295EFCE" blockId="86.[151,1436,1395,2032]" pageId="86" pageNumber="87">WALKING LEGS. In both sexes, leg-pairs 1 and 2 with tarsal combs; prefemora with several long and robust setae; femora and postfemora with a group of several long and robust setae.</paragraph>
<paragraph id="8BA1369AFF81CD68FF1CEEE1B2D5EE51" blockId="86.[151,1436,1395,2032]" pageId="86" pageNumber="87">
MALE SEXUAL CHARACTERS (
<figureCitation id="13252A1FFF81CD68FDC7EEE6B302EFE2" box="[540,625,1683,1708]" captionStart="FIGURE 71" captionStartId="88.[151,250,1852,1874]" captionTargetBox="[189,1398,193,1830]" captionTargetId="figure@88.[188,1398,193,1830]" captionTargetPageId="88" captionText="FIGURE 71. Paranotosoma cordatum gen. et sp. nov., paratype male. A. Leg 3. B. Leg 4. C. Leg 7. Scale lines = 0.2 mm." httpUri="https://zenodo.org/record/272820/files/figure.png" pageId="86" pageNumber="87">Fig. 71</figureCitation>
). Leg-pairs 37 enlarged. Leg-pairs 3 and 4 each with a basal exterior protrusion on prefemur. Leg-pair 5 with a barely visible oral protrusion on prefemur. Leg-pair 6 without peculiarities. Leg-pair 7 only with rudiments of two posterior coxal processes present in
<taxonomicName id="4C1E4D19FF81CD68FB5AEEA9B436EFBD" box="[1153,1349,1755,1780]" class="Diplopoda" family="Anthroleucosomatidae" genus="Paranotosoma" kingdom="Animalia" order="Chordeumatida" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="subrotundatum">
<emphasis id="B96AEA88FF81CD68FB5AEEA9B436EFBD" box="[1153,1349,1755,1780]" italics="true" pageId="86" pageNumber="87">P. subrotundatum</emphasis>
</taxonomicName>
<emphasis id="B96AEA88FF81CD68FA96EEAEB183EE51" bold="true" pageId="86" pageNumber="87">gen. et sp. nov.</emphasis>
Leg-pairs 10 and 11 with coxal glands, no other peculiarities.
</paragraph>
<paragraph id="8BA1369AFF81CD64FF1CEF56B369EE02" blockId="86.[151,1436,1395,2032]" lastBlockId="90.[151,1437,1734,2011]" lastPageId="90" lastPageNumber="91" pageId="86" pageNumber="87">
ANTERIOR GONOPODS (
<figureCitation id="13252A1FFF81CD68FE10EF56B352EE72" box="[459,545,1827,1852]" captionStart="FIGURE 72" captionStartId="89.[151,250,1755,1777]" captionTargetBox="[200,1390,223,1668]" captionTargetId="figure@89.[188,1398,193,1734]" captionTargetPageId="89" captionText="FIGURE 72. Paranotosoma cordatum gen. et sp. nov., holotype male gonopods. A. Anterior gonopods, anterior view. B. Anterior gonopods, lateral view. C. Anterior gonopods, posterior view. D. Posterior gonopods, posterior view. c = coxite; h = horn; t = telopodite; ah = acuminate horn; ap / cp = anterior coxal process; cv = coxal vesicle; lg = longitudinal groove; lv = posterior lever; pp = posterior coxal process; rs = row of setae; sp = sternal plate; ss = sternal sac; alv = anterior branch of lever; mdl = medial lamella; plv = posterior branch of lever. Scale line = 0.2 mm." httpUri="https://zenodo.org/record/272821/files/figure.png" pageId="86" pageNumber="87">Figs 72</figureCitation>
AC, 73). Sternal plate (
<emphasis id="B96AEA88FF81CD68FCE8EF51B23FEE72" box="[819,844,1828,1851]" italics="true" pageId="86" pageNumber="87">sp</emphasis>
) with a medial, anterior, triangular, hairy, sternal sac (
<emphasis id="B96AEA88FF81CD68FF7BEF3DB1C5EE16" box="[160,182,1864,1887]" italics="true" pageId="86" pageNumber="87">ss</emphasis>
). A thin medial lamella (
<emphasis id="B96AEA88FF81CD68FE3BEF32B379EE29" box="[480,522,1863,1888]" italics="true" pageId="86" pageNumber="87">mdl</emphasis>
) present between sternal sac and anterior shield. Anterior coxal processes (
<emphasis id="B96AEA88FF81CD68FAA3EF3DB4E0EE16" box="[1400,1427,1864,1887]" italics="true" pageId="86" pageNumber="87">cp</emphasis>
) wide, distal third divided, remaining parts fused, cordiform, only an anterior longitudinal groove (
<emphasis id="B96AEA88FF81CD68FAD8EF1EB46AEECD" box="[1283,1305,1899,1924]" italics="true" pageId="86" pageNumber="87">lg</emphasis>
) present in basal two-thirds. Distomesal parts of shields denticulate and curved posteriorly. Mesal edges folded inside and forming a projection with two short horns (
<emphasis id="B96AEA88FF81CD68FD5BEFC6B3FDEE85" box="[640,654,1971,1996]" italics="true" pageId="86" pageNumber="87">h</emphasis>
). Anterior and posterior (
<emphasis id="B96AEA88FF81CD68FC69EFC1B2BDEE82" box="[946,974,1972,1995]" italics="true" pageId="86" pageNumber="87">pp</emphasis>
) coxal processes fused basally. Posterior coxal processes (
<emphasis id="B96AEA88FF81CD68FE8DEFADB001EEA6" box="[342,370,2008,2031]" italics="true" pageId="86" pageNumber="87">pp</emphasis>
) consisting of an anterior, relatively long and more or less acuminate horn (
<emphasis id="B96AEA88FF81CD68FB1EEFA2B592EEB9" box="[1221,1249,2007,2032]" italics="true" pageId="86" pageNumber="87">ah</emphasis>
) which abuts to inner side of anterior coxal process, and a posterior lever (
<emphasis id="B96AEA88FF8DCD64FCEDEEB3B239EF96" box="[822,842,1734,1759]" italics="true" pageId="90" pageNumber="91">lv</emphasis>
) with an anterior (
<emphasis id="B96AEA88FF8DCD64FBFFEEB3B535EF96" box="[1060,1094,1734,1759]" italics="true" pageId="90" pageNumber="91">alv</emphasis>
) and a posterior (
<emphasis id="B96AEA88FF8DCD64FACFEEB3B445EF96" box="[1300,1334,1734,1759]" italics="true" pageId="90" pageNumber="91">plv</emphasis>
) branch. Posterior branch curved anteriorly; anterior branch orientated anteriorly. Posterior coxal processes without lateral process. Posteromesal edges of posterior coxal processes with one row of short setae (
<emphasis id="B96AEA88FF8DCD64FB81EF7AB503EE6F" box="[1114,1136,1807,1830]" italics="true" pageId="90" pageNumber="91">rs</emphasis>
). Hairy coxal vesicles (
<emphasis id="B96AEA88FF8DCD64FAA1EF7AB4E0EE6F" box="[1402,1427,1807,1830]" italics="true" pageId="90" pageNumber="91">cv</emphasis>
) present medially on posterior side.
</paragraph>
<caption id="DF616612FF80CD69FF4CEFD9B234EEA8" httpUri="https://zenodo.org/record/272819/files/figure.png" pageId="87" pageNumber="88" targetBox="[208,1371,193,1937]" targetPageId="87">
<paragraph id="8BA1369AFF80CD69FF4CEFD9B234EEA8" blockId="87.[151,1436,1964,2017]" pageId="87" pageNumber="88">
<emphasis id="B96AEA88FF80CD69FF4CEFD9B053EE88" bold="true" box="[151,288,1964,1986]" pageId="87" pageNumber="88">FIGURE 70.</emphasis>
<taxonomicName id="4C1E4D19FF80CD69FEFDEFD8B351EE8B" box="[294,546,1964,1986]" class="Diplopoda" family="Anthroleucosomatidae" genus="Paranotosoma" kingdom="Animalia" order="Chordeumatida" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="cordatum">
<emphasis id="B96AEA88FF80CD69FEFDEFD8B351EE8B" box="[294,546,1964,1986]" italics="true" pageId="87" pageNumber="88">Paranotosoma cordatum</emphasis>
</taxonomicName>
<emphasis id="B96AEA88FF80CD69FDF3EFD8B3B5EE8B" bold="true" box="[552,710,1965,1986]" pageId="87" pageNumber="88">gen. et sp. nov.</emphasis>
, paratype male, pleurotergite 15.
<emphasis id="B96AEA88FF80CD69FBC6EFD9B545EE88" bold="true" box="[1053,1078,1964,1985]" pageId="87" pageNumber="88">A.</emphasis>
Posterior view.
<emphasis id="B96AEA88FF80CD69FB00EFD8B582EE8B" bold="true" box="[1243,1265,1965,1986]" pageId="87" pageNumber="88">B.</emphasis>
Lateral view.
<emphasis id="B96AEA88FF80CD69FA58EFD9B4EFEE8B" bold="true" box="[1411,1436,1964,1986]" pageId="87" pageNumber="88">C.</emphasis>
Prozonite, dorsal view.
<emphasis id="B96AEA88FF80CD69FE5EEFBEB0EDEEA9" bold="true" box="[389,414,1995,2016]" pageId="87" pageNumber="88">D.</emphasis>
Lateral keel, dorsal view.
<emphasis id="B96AEA88FF80CD69FD72EFBEB3B3EEA9" bold="true" box="[681,704,1995,2016]" pageId="87" pageNumber="88">E.</emphasis>
Dorsal view.
</paragraph>
</caption>
<caption id="DF616612FF8FCD66FF4CEF49B40AEE1B" box="[151,1401,1852,1874]" httpUri="https://zenodo.org/record/272820/files/figure.png" pageId="88" pageNumber="89" targetBox="[189,1398,193,1830]" targetPageId="88">
<paragraph id="8BA1369AFF8FCD66FF4CEF49B40AEE1B" blockId="88.[151,1401,1852,1874]" box="[151,1401,1852,1874]" pageId="88" pageNumber="89">
<emphasis id="B96AEA88FF8FCD66FF4CEF49B06CEE18" bold="true" box="[151,287,1852,1874]" pageId="88" pageNumber="89">FIGURE 71.</emphasis>
<taxonomicName id="4C1E4D19FF8FCD66FEFEEF48B352EE1B" box="[293,545,1852,1874]" class="Diplopoda" family="Anthroleucosomatidae" genus="Paranotosoma" kingdom="Animalia" order="Chordeumatida" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="cordatum">
<emphasis id="B96AEA88FF8FCD66FEFEEF48B352EE1B" box="[293,545,1852,1874]" italics="true" pageId="88" pageNumber="89">Paranotosoma cordatum</emphasis>
</taxonomicName>
<emphasis id="B96AEA88FF8FCD66FDFCEF49B3B7EE18" bold="true" box="[551,708,1852,1873]" pageId="88" pageNumber="89">gen. et sp. nov.</emphasis>
, paratype male.
<emphasis id="B96AEA88FF8FCD66FCB6EF49B2F5EE18" bold="true" box="[877,902,1852,1873]" pageId="88" pageNumber="89">A.</emphasis>
Leg 3.
<emphasis id="B96AEA88FF8FCD66FC08EF49B299EE18" bold="true" box="[979,1002,1852,1873]" pageId="88" pageNumber="89">B.</emphasis>
Leg 4.
<emphasis id="B96AEA88FF8FCD66FBECEF49B523EE1B" bold="true" box="[1079,1104,1852,1874]" pageId="88" pageNumber="89">C.</emphasis>
Leg 7. Scale lines = 0.2 mm.
</paragraph>
</caption>
<caption id="DF616612FF8ECD67FF4CEEAEB2FFEE25" httpUri="https://zenodo.org/record/272821/files/figure.png" pageId="89" pageNumber="90" targetBox="[200,1390,223,1668]" targetPageId="89">
<paragraph id="8BA1369AFF8ECD67FF4CEEAEB2FFEE25" blockId="89.[151,1436,1755,1900]" pageId="89" pageNumber="90">
<emphasis id="B96AEA88FF8ECD67FF4CEEAEB050EFB9" bold="true" box="[151,291,1755,1777]" pageId="89" pageNumber="90">FIGURE 72.</emphasis>
<taxonomicName id="4C1E4D19FF8ECD67FEF7EEA9B359EFB8" box="[300,554,1755,1777]" class="Diplopoda" family="Anthroleucosomatidae" genus="Paranotosoma" kingdom="Animalia" order="Chordeumatida" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="cordatum">
<emphasis id="B96AEA88FF8ECD67FEF7EEA9B359EFB8" box="[300,554,1755,1777]" italics="true" pageId="89" pageNumber="90">Paranotosoma cordatum</emphasis>
</taxonomicName>
<emphasis id="B96AEA88FF8ECD67FDEFEEA9B3A9EFB8" bold="true" box="[564,730,1756,1777]" pageId="89" pageNumber="90">gen. et sp. nov.</emphasis>
, holotype male gonopods.
<emphasis id="B96AEA88FF8ECD67FC21EEAEB561EFB9" bold="true" box="[1018,1042,1755,1776]" pageId="89" pageNumber="90">A.</emphasis>
Anterior gonopods, anterior view.
<emphasis id="B96AEA88FF8ECD67FA5FEEA9B4E8EFB8" bold="true" box="[1412,1435,1756,1777]" pageId="89" pageNumber="90">B.</emphasis>
Anterior gonopods, lateral view.
<emphasis id="B96AEA88FF8ECD67FE33EE8FB373EE59" bold="true" box="[488,512,1786,1808]" pageId="89" pageNumber="90">C.</emphasis>
Anterior gonopods, posterior view.
<emphasis id="B96AEA88FF8ECD67FCAFEE8FB2FEEE46" bold="true" box="[884,909,1786,1807]" pageId="89" pageNumber="90">D.</emphasis>
Posterior gonopods, posterior view.
<emphasis id="B96AEA88FF8ECD67FADEEE8EB463EE59" box="[1285,1296,1787,1808]" italics="true" pageId="89" pageNumber="90">c</emphasis>
= coxite;
<emphasis id="B96AEA88FF8ECD67FAA1EE8FB4F4EE59" box="[1402,1415,1786,1808]" italics="true" pageId="89" pageNumber="90">h</emphasis>
= horn;
<emphasis id="B96AEA88FF8ECD67FF0FEF6FB1A8EE66" box="[212,219,1818,1839]" italics="true" pageId="89" pageNumber="90">t</emphasis>
= telopodite;
<emphasis id="B96AEA88FF8ECD67FEB4EF6DB0FBEE67" box="[367,392,1816,1838]" italics="true" pageId="89" pageNumber="90">ah</emphasis>
= acuminate horn;
<emphasis id="B96AEA88FF8ECD67FD8EEF6CB3FDEE67" box="[597,654,1817,1838]" italics="true" pageId="89" pageNumber="90">ap/cp</emphasis>
= anterior coxal process;
<emphasis id="B96AEA88FF8ECD67FC45EF6FB2C7EE66" box="[926,948,1818,1839]" italics="true" pageId="89" pageNumber="90">cv</emphasis>
= coxal vesicle;
<emphasis id="B96AEA88FF8ECD67FBB3EF6DB50FEE67" box="[1128,1148,1816,1838]" italics="true" pageId="89" pageNumber="90">lg</emphasis>
= longitudinal groove;
<emphasis id="B96AEA88FF8ECD67FAA8EF6DB4F5EE67" box="[1395,1414,1816,1838]" italics="true" pageId="89" pageNumber="90">lv</emphasis>
= posterior lever;
<emphasis id="B96AEA88FF8ECD67FEEDEF4DB03CEE04" box="[310,335,1848,1869]" italics="true" pageId="89" pageNumber="90">pp</emphasis>
= posterior coxal process;
<emphasis id="B96AEA88FF8ECD67FD85EF4DB301EE04" box="[606,626,1848,1869]" italics="true" pageId="89" pageNumber="90">rs</emphasis>
= row of setae;
<emphasis id="B96AEA88FF8ECD67FCCFEF4DB258EE04" box="[788,811,1848,1869]" italics="true" pageId="89" pageNumber="90">sp</emphasis>
= sternal plate;
<emphasis id="B96AEA88FF8ECD67FC10EF4DB2ACEE04" box="[971,991,1848,1869]" italics="true" pageId="89" pageNumber="90">ss</emphasis>
= sternal sac;
<emphasis id="B96AEA88FF8ECD67FBB4EF42B5FEEE04" box="[1135,1165,1847,1869]" italics="true" pageId="89" pageNumber="90">alv</emphasis>
= anterior branch of lever;
<emphasis id="B96AEA88FF8ECD67FF4CEF23B1CEEE25" box="[151,189,1878,1900]" italics="true" pageId="89" pageNumber="90">mdl</emphasis>
= medial lamella;
<emphasis id="B96AEA88FF8ECD67FEA0EF23B0EAEE25" box="[379,409,1878,1900]" italics="true" pageId="89" pageNumber="90">plv</emphasis>
= posterior branch of lever. Scale line = 0.2 mm.
</paragraph>
</caption>
<caption id="DF616612FF8DCD64FF4CEE5DB439EFD3" httpUri="https://zenodo.org/record/272822/files/figure.png" pageId="90" pageNumber="91" targetBox="[188,1398,193,1554]" targetPageId="90">
<paragraph id="8BA1369AFF8DCD64FF4CEE5DB439EFD3" blockId="90.[151,1436,1576,1690]" pageId="90" pageNumber="91">
<emphasis id="B96AEA88FF8DCD64FF4CEE5DB053EF74" bold="true" box="[151,288,1576,1598]" pageId="90" pageNumber="91">FIGURE 73.</emphasis>
<taxonomicName id="4C1E4D19FF8DCD64FEFDEE5CB351EF77" box="[294,546,1576,1598]" class="Diplopoda" family="Anthroleucosomatidae" genus="Paranotosoma" kingdom="Animalia" order="Chordeumatida" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="cordatum">
<emphasis id="B96AEA88FF8DCD64FEFDEE5CB351EF77" box="[294,546,1576,1598]" italics="true" pageId="90" pageNumber="91">Paranotosoma cordatum</emphasis>
</taxonomicName>
<emphasis id="B96AEA88FF8DCD64FDF3EE5DB3B5EF74" bold="true" box="[552,710,1576,1597]" pageId="90" pageNumber="91">gen. et sp. nov.</emphasis>
, paratype male anterior gonopods.
<emphasis id="B96AEA88FF8DCD64FBF6EE5DB535EF74" bold="true" box="[1069,1094,1576,1597]" pageId="90" pageNumber="91">A.</emphasis>
Anterior view.
<emphasis id="B96AEA88FF8DCD64FB3EEE5DB58FEF74" bold="true" box="[1253,1276,1576,1597]" pageId="90" pageNumber="91">B.</emphasis>
Detail, anterior view.
<emphasis id="B96AEA88FF8DCD64FF08EE33B198EF15" bold="true" box="[211,235,1606,1628]" pageId="90" pageNumber="91">C.</emphasis>
Lateral view.
<emphasis id="B96AEA88FF8DCD64FEA0EE32B0E0EF15" bold="true" box="[379,403,1607,1628]" pageId="90" pageNumber="91">D.</emphasis>
Posterolateral view.
<emphasis id="B96AEA88FF8DCD64FDBDEE32B30DEF15" bold="true" box="[614,638,1607,1628]" pageId="90" pageNumber="91">E.</emphasis>
Posterior view.
<emphasis id="B96AEA88FF8DCD64FCFAEE32B241EF15" bold="true" box="[801,818,1607,1628]" pageId="90" pageNumber="91">F.</emphasis>
Distal view.
<emphasis id="B96AEA88FF8DCD64FC62EE33B2B6EF15" box="[953,965,1606,1628]" italics="true" pageId="90" pageNumber="91">h</emphasis>
= horn;
<emphasis id="B96AEA88FF8DCD64FBC0EE33B546EF15" box="[1051,1077,1606,1628]" italics="true" pageId="90" pageNumber="91">ah</emphasis>
= acuminate horn;
<emphasis id="B96AEA88FF8DCD64FB23EE32B434EF14" box="[1272,1351,1607,1629]" italics="true" pageId="90" pageNumber="91">ap/cp =</emphasis>
anterior coxal process;
<emphasis id="B96AEA88FF8DCD64FEF6EE13B030EF32" box="[301,323,1638,1659]" italics="true" pageId="90" pageNumber="91">cv</emphasis>
= coxal vesicle;
<emphasis id="B96AEA88FF8DCD64FE34EE10B370EF32" box="[495,515,1637,1659]" italics="true" pageId="90" pageNumber="91">lg</emphasis>
= longitudinal groove;
<emphasis id="B96AEA88FF8DCD64FD29EE10B277EF32" box="[754,772,1637,1659]" italics="true" pageId="90" pageNumber="91">lv</emphasis>
= posterior lever;
<emphasis id="B96AEA88FF8DCD64FC1BEE13B2A9EF32" box="[960,986,1638,1659]" italics="true" pageId="90" pageNumber="91">pp</emphasis>
= posterior coxal process;
<emphasis id="B96AEA88FF8DCD64FB30EE13B58CEF32" box="[1259,1279,1638,1659]" italics="true" pageId="90" pageNumber="91">rs</emphasis>
= row of setae;
<emphasis id="B96AEA88FF8DCD64FF4CEEF0B1DDEFD3" box="[151,174,1669,1690]" italics="true" pageId="90" pageNumber="91">sp</emphasis>
= sternal plate;
<emphasis id="B96AEA88FF8DCD64FE8AEEF0B016EFD3" box="[337,357,1669,1690]" italics="true" pageId="90" pageNumber="91">ss</emphasis>
= sternal sac;
<emphasis id="B96AEA88FF8DCD64FE2CEEF1B366EFD3" box="[503,533,1668,1690]" italics="true" pageId="90" pageNumber="91">alv</emphasis>
= anterior branch of lever;
<emphasis id="B96AEA88FF8DCD64FCF7EEF1B221EFD3" box="[812,850,1668,1690]" italics="true" pageId="90" pageNumber="91">mdl</emphasis>
= medial lamella;
<emphasis id="B96AEA88FF8DCD64FBCBEEF1B55CEFD3" box="[1040,1071,1668,1690]" italics="true" pageId="90" pageNumber="91">plv</emphasis>
= posterior branch of lever.
</paragraph>
</caption>
<paragraph id="8BA1369AFF8DCD64FF1CEF22B4FBEEFE" blockId="90.[151,1437,1734,2011]" pageId="90" pageNumber="91">
POSTERIOR GONOPODS (
<figureCitation id="13252A1FFF8DCD64FE0DEF23B358EE26" box="[470,555,1878,1903]" captionStart="FIGURE 72" captionStartId="89.[151,250,1755,1777]" captionTargetBox="[200,1390,223,1668]" captionTargetId="figure@89.[188,1398,193,1734]" captionTargetPageId="89" captionText="FIGURE 72. Paranotosoma cordatum gen. et sp. nov., holotype male gonopods. A. Anterior gonopods, anterior view. B. Anterior gonopods, lateral view. C. Anterior gonopods, posterior view. D. Posterior gonopods, posterior view. c = coxite; h = horn; t = telopodite; ah = acuminate horn; ap / cp = anterior coxal process; cv = coxal vesicle; lg = longitudinal groove; lv = posterior lever; pp = posterior coxal process; rs = row of setae; sp = sternal plate; ss = sternal sac; alv = anterior branch of lever; mdl = medial lamella; plv = posterior branch of lever. Scale line = 0.2 mm." httpUri="https://zenodo.org/record/272821/files/figure.png" pageId="90" pageNumber="91">Fig. 72</figureCitation>
D). Coxites (
<emphasis id="B96AEA88FF8DCD64FD67EF22B3BBEE27" box="[700,712,1879,1902]" italics="true" pageId="90" pageNumber="91">c</emphasis>
) divided. Telopodites (
<emphasis id="B96AEA88FF8DCD64FC0BEF22B2ABEE27" box="[976,984,1879,1902]" italics="true" pageId="90" pageNumber="91">t</emphasis>
) present on posterolateral side, apically with a small claw and one or two setae; two-segmented. Both coxites with two coxal processes: an anterior narrower and shorter (
<emphasis id="B96AEA88FF8DCD64FE4BEFEAB0DFEEFF" box="[400,428,1951,1974]" italics="true" pageId="90" pageNumber="91">ap</emphasis>
), and a posterior wider and acuminate (
<emphasis id="B96AEA88FF8DCD64FCB0EFEAB2F4EEFF" box="[875,903,1951,1974]" italics="true" pageId="90" pageNumber="91">pp</emphasis>
). Coxal vesicles (
<emphasis id="B96AEA88FF8DCD64FB8AEFEAB519EEFF" box="[1105,1130,1951,1974]" italics="true" pageId="90" pageNumber="91">cv</emphasis>
) present on anterior side.
</paragraph>
</subSubSection>
<subSubSection id="C3046511FF8DCD65FF1CEFB7B5AAE82C" lastPageId="91" lastPageNumber="92" pageId="90" pageNumber="91" type="distribution">
<paragraph id="8BA1369AFF8DCD64FF1CEFB7B3A9EE92" blockId="90.[151,1437,1734,2011]" box="[199,730,1986,2011]" pageId="90" pageNumber="91">
<emphasis id="B96AEA88FF8DCD64FF1CEFB7B013EE92" bold="true" box="[199,352,1986,2011]" pageId="90" pageNumber="91">Distribution.</emphasis>
<collectingRegion id="49DAF878FF8DCD64FEBCEFB7B0B2EE92" box="[359,449,1986,2011]" country="United States of America" name="Georgia" pageId="90" pageNumber="91">
<collectingCountry id="F309760AFF8DCD64FEBCEFB7B0B2EE92" box="[359,449,1986,2011]" name="Georgia" pageId="90" pageNumber="91">Georgia</collectingCountry>
</collectingRegion>
(
<figureCitation id="13252A1FFF8DCD64FE0BEFB7B342EE92" box="[464,561,1986,2011]" captionStart="FIGURE 167" captionStartId="196.[151,250,2007,2029]" captionTargetBox="[188,1398,1136,1986]" captionTargetId="figure@196.[188,1398,1136,1986]" captionTargetPageId="196" captionText="FIGURE 167. Distribution of the Caucasian Anthroleucosomatidae: the Caucaseuma complex." httpUri="https://zenodo.org/record/272916/files/figure.png" pageId="90" pageNumber="91">Fig. 167</figureCitation>
, black square).
</paragraph>
<paragraph id="8BA1369AFF8CCD65FF1CE8E2B5AAE82C" blockId="91.[151,1437,151,357]" pageId="91" pageNumber="92">
<emphasis id="B96AEA88FF8CCD65FF1CE8E2B062E9F9" bold="true" box="[199,273,151,176]" pageId="91" pageNumber="92">Notes.</emphasis>
The population from Tskhaltubo, near Cave Belaya, is characterized by the presence of a lamellar structure between
<emphasis id="B96AEA88FF8CCD65FEB1E8C9B0FFE99C" box="[362,396,188,213]" italics="true" pageId="91" pageNumber="92">alv</emphasis>
and
<emphasis id="B96AEA88FF8CCD65FE12E8C9B09AE99C" box="[457,489,188,213]" italics="true" pageId="91" pageNumber="92">plv</emphasis>
, a condition which can be seen neither in populations from the Batumi Botanical Garden nor from the Kintrish Nature Reserve. Bearing in mind that the Tskhaltubo population is remote enough from the two latter populations, and that such a difference is only minor compared to the differences from
<taxonomicName id="4C1E4D19FF8CCD65FA5CE970B19BE809" class="Diplopoda" family="Anthroleucosomatidae" genus="Paranotosoma" kingdom="Animalia" order="Chordeumatida" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="attemsi">
<emphasis id="B96AEA88FF8CCD65FA5CE970B19BE809" italics="true" pageId="91" pageNumber="92">P. attemsi</emphasis>
</taxonomicName>
<emphasis id="B96AEA88FF8CCD65FF28E952B0DCE809" bold="true" box="[243,431,295,320]" pageId="91" pageNumber="92">gen. et sp. nov.</emphasis>
and
<taxonomicName id="4C1E4D19FF8CCD65FE35E95CB3C5E809" box="[494,694,295,320]" class="Diplopoda" family="Anthroleucosomatidae" genus="Paranotosoma" kingdom="Animalia" order="Chordeumatida" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="subrotundatum">
<emphasis id="B96AEA88FF8CCD65FE35E95CB3C5E809" box="[494,694,295,320]" italics="true" pageId="91" pageNumber="92">P. subrotundatum</emphasis>
</taxonomicName>
<emphasis id="B96AEA88FF8CCD65FD19E952B20DE809" bold="true" box="[706,894,295,320]" pageId="91" pageNumber="92">gen. et sp. nov.</emphasis>
, we assign it to
<taxonomicName id="4C1E4D19FF8CCD65FB91E95CB5A7E809" box="[1098,1236,295,320]" class="Diplopoda" family="Anthroleucosomatidae" genus="Paranotosoma" kingdom="Animalia" order="Chordeumatida" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="cordatum">
<emphasis id="B96AEA88FF8CCD65FB91E95CB5A7E809" box="[1098,1236,295,320]" italics="true" pageId="91" pageNumber="92">P. cordatum</emphasis>
</taxonomicName>
<emphasis id="B96AEA88FF8CCD65FB3BE952B4EFE809" bold="true" box="[1248,1436,295,320]" pageId="91" pageNumber="92">gen. et sp. nov.</emphasis>
Similarly to
<taxonomicName id="4C1E4D19FF8CCD65FEF8E938B353E82C" box="[291,544,332,357]" class="Diplopoda" family="Anthroleucosomatidae" genus="Caucaseuma" kingdom="Animalia" order="Chordeumatida" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="variabile" status="sp. nov.">
<emphasis id="B96AEA88FF8CCD65FEF8E938B353E82C" box="[291,544,332,357]" italics="true" pageId="91" pageNumber="92">Caucaseuma variabile</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25957F3FF8CCD65FDFDE939B3F3E82C" box="[550,640,332,357]" pageId="91" pageNumber="92" rank="species">
<emphasis id="B96AEA88FF8CCD65FDFDE939B3F3E82C" bold="true" box="[550,640,332,357]" pageId="91" pageNumber="92">sp. nov.</emphasis>
</taxonomicNameLabel>
, this taxon is probably in a stage of active speciation.
</paragraph>
</subSubSection>
</treatment>
</document>