410 lines
52 KiB
XML
410 lines
52 KiB
XML
<document id="4279CF9D8EA6398DD546A3505C9EF87C" ID-DOI="10.5281/zenodo.186322" ID-GBIF-Dataset="287e35b6-1422-48a8-9468-5ff7e7ccfcd6" ID-ISSN="1175-5326" ID-Zenodo-Dep="186322" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461049966650" checkinUser="plazi" docAuthor="Fery, Hans & Erman, Ömer Köksal" docDate="2009" docId="03882B52517FFFAEFF62D7BF50232D86" docLanguage="en" docName="zt02033p012.pdf" docOrigin="Zootaxa 2033" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Hydroporus cagrankaya Fery & Erman, 2009, sp. n." docType="treatment" docVersion="7" lastPageNumber="6" masterDocId="FFB1532A517DFFABFFF5D50E54272940" masterDocTitle="Five new species of the longulus - group of Hydroporus Clairville, 1806 from north-eastern Turkey (Coleoptera: Dytiscidae)" masterLastPageNumber="12" masterPageNumber="1" pageNumber="3" updateTime="1698578434421" updateUser="plazi">
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<mods:mods id="738819364AACF3F084290809923425CA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="9085C2B670C508EB34535E79F2F97D63">
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<mods:title id="B9E37D2613EE9635DFC07A034403F3D1">Five new species of the longulus - group of Hydroporus Clairville, 1806 from north-eastern Turkey (Coleoptera: Dytiscidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="C9382F01E3BAFC199CD4CB5C02149722" type="personal">
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<mods:role id="2D63BE15A5879B060B6E449294DB4914">
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<mods:roleTerm id="08B4221F4F88F81F050AA64F14579286">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7714E0D73688FF3BAA937F38E6F9FBA0">Fery, Hans</mods:namePart>
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</mods:name>
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<mods:name id="F54FC63524B4B51CF19EBC4FF84C74A3" type="personal">
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<mods:role id="1DE2662AFCDB90AF821CC7AAA95A4355">
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<mods:roleTerm id="5309B712C2DEDEA50CD7D8FFE50B997A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="78E4DEA77F4E06C2423449C1F42B70CE">Erman, Ömer Köksal</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="95EE1D2C797246A2380B74D2C5F08968">text</mods:typeOfResource>
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<mods:relatedItem id="0B1E9D18B02779DC5CAAE813630B6F8F" type="host">
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<mods:titleInfo id="1870CF894F65D3B90C35744A2AB13E4F">
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<mods:title id="8DB0E82F1D4F02F67D08F6294A276C71">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="DAB5FD8C7F121435FECD896C98B8518C">
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<mods:date id="2D814CA7C3148C2377623562A322395D">2009</mods:date>
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<mods:detail id="EFD921A1D31C9E729FCD084C3ADFCBAB" type="volume">
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<mods:number id="C15AF0C04E7C1B22C2D105704EDD4742">2033</mods:number>
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</mods:detail>
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<mods:extent id="FE203DD45170A7D0A5E73FC12C7D8EE7" unit="page">
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<mods:start id="FD740A27DC2C1EF0D4ABF98189054F4E">1</mods:start>
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<mods:end id="7724A342E1587B2CBDFDA73FDCBF9561">12</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="E97BD73F80E78960F16E430BAA800473">journal article</mods:classification>
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<mods:identifier id="6CEFC57BD99CB7D352B872D2475C7C28" type="DOI">10.5281/zenodo.186322</mods:identifier>
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<mods:identifier id="1E07BCCF2898BDB64854A33497A8662B" type="GBIF-Dataset">287e35b6-1422-48a8-9468-5ff7e7ccfcd6</mods:identifier>
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<mods:identifier id="290CDABBA3C28204C1675E3EFFA213FE" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="8E85691098FB9D486EABD3E9775D8D7E" type="Zenodo-Dep">186322</mods:identifier>
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</mods:mods>
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<treatment id="03882B52517FFFAEFF62D7BF50232D86" ID-DOI="http://doi.org/10.5281/zenodo.6222789" ID-GBIF-Taxon="119611635" ID-Zenodo-Dep="6222789" LSID="urn:lsid:plazi:treatment:03882B52517FFFAEFF62D7BF50232D86" httpUri="http://treatment.plazi.org/id/03882B52517FFFAEFF62D7BF50232D86" lastPageId="5" lastPageNumber="6" pageId="2" pageNumber="3">
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<subSubSection id="C33BC9CF517FFFA9FF62D7BF551B2BAE" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="8B9E9A44517FFFA9FF62D7BF56212B8A" blockId="2.[151,518,688,750]" box="[151,518,688,715]" pageId="2" pageNumber="3">
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<heading id="D0D62D28517FFFA9FF62D7BF56212B8A" bold="true" box="[151,518,688,715]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
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||
<emphasis id="B9554656517FFFA9FF62D7BF56212B8A" bold="true" box="[151,518,688,715]" pageId="2" pageNumber="3">
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<taxonomicName id="4C21E1C7517FFFA9FF62D7BF559A2B8B" ID-CoL="3N8QS" box="[151,445,689,715]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="cagrankaya" status="sp. nov.">
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<emphasis id="B9554656517FFFA9FF62D7BF559A2B8B" bold="true" box="[151,445,689,715]" italics="true" pageId="2" pageNumber="3">Hydroporus cagrankaya</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A266FB2D517FFFA9FE31D7BE56212B8A" box="[452,518,688,714]" pageId="2" pageNumber="3" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="8B9E9A44517FFFA9FF62D7DA551B2BAE" blockId="2.[151,518,688,750]" box="[151,316,724,750]" pageId="2" pageNumber="3">
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(
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<figureCitation id="131A86C1517FFFA9FF6AD7DA54CE2BAE" box="[159,233,724,750]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,273,1930,1954]" captionTargetBox="[223,1378,304,1892]" captionTargetId="figure@3.[200,1387,283,1906]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Median lobe of aedeagus of: (1) Hydroporus cagrankaya sp. n. (holotype), (2) H. sivrikaya sp. n. (holotype), (3) H. artvinensis sp. n. (holotype), (4) H. toledoi sp. n. (holotype), (5 a, b) H. lundbergi sp. n. (holotype) (the arrow indicates the direction of the " frontal view ", Fig. 5 b)" httpUri="https://zenodo.org/record/186323/files/figure.png" pageId="2" pageNumber="3">Figs 1</figureCitation>
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,
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<figureCitation id="131A86C1517FFFA9FF0DD7DA55142BAE" box="[248,307,724,750]" captionStart-0="FIGURES 6 – 8" captionStart-1="FIGURES 9 – 12" captionStart-2="FIGURE 13" captionStartId-0="4.[151,269,733,757]" captionStartId-1="6.[151,274,1767,1791]" captionStartId-2="9.[151,255,1395,1419]" captionTargetBox-0="[195,1350,222,702]" captionTargetBox-1="[212,1374,194,1743]" captionTargetBox-2="[155,1432,514,1349]" captionTargetId-0="figure@4.[182,1404,194,710]" captionTargetId-1="figure@6.[212,1376,194,1743]" captionTargetId-2="figure@9.[151,1436,506,1368]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionTargetPageId-2="9" captionText-0="FIGURES 6 – 8. Hydroporus cagrankaya sp. n.: (6) paramere, (7) gonocoxae, (8) gonocoxosternum." captionText-1="FIGURES 9 – 12. Hydroporus cagrankaya sp. n.: SEM photos of: (9) parts of venter, (10) left protarsus, (11) left mesotarsus, (12) details of first and second tarsomeres of left protarsus." captionText-2="FIGURE 13. Distribution of Hydroporus lundbergi sp. n. (1 – 3; see text), H. cagrankaya sp. n. (4), H. sivrikaya sp. n. (5), H. toledoi sp. n. (6), H. artvinensis sp. n. (7), and H. erzurumensis (8 – 14). The map was made by using MS ENCARTA 2000." httpUri-0="https://zenodo.org/record/186324/files/figure.png" httpUri-1="https://zenodo.org/record/186325/files/figure.png" httpUri-2="https://zenodo.org/record/186326/files/figure.png" pageId="2" pageNumber="3">6–13</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33BC9CF517FFFA9FF62D62E50982DD7" pageId="2" pageNumber="3" type="materials_examined">
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<paragraph id="8B9E9A44517FFFA9FF62D62E56D72A22" blockId="2.[151,1437,796,2026]" pageId="2" pageNumber="3">
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<emphasis id="B9554656517FFFA9FF62D62E55612A7A" bold="true" box="[151,326,800,826]" pageId="2" pageNumber="3">
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<typeStatus id="549A24E6517FFFA9FF62D62E54F02A7A" box="[151,215,800,826]" pageId="2" pageNumber="3">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="F336DAD4517FFFA9FEBAD62F558F2A7B" box="[335,424,801,827]" name="Turkey" pageId="2" pageNumber="3">Turkey</collectingCountry>
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, Rize province, ca.
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<quantity id="4CD937A1517FFFA9FD6AD62F56F92A7B" box="[671,734,801,827]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="3" unit="km" value="9.0">9 km</quantity>
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due NE İkizdere, near
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<taxonomicName id="4C21E1C7517FFFA9FBF4D61150B42A79" box="[1025,1171,799,825]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="11" phylum="Arthropoda" rank="species" species="cagrankaya" status="sp. nov.">Çaġrankaya</taxonomicName>
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plateau, ca.
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<geoCoordinate id="EE15FC83517FFFA9FAC7D62F51BB2A7B" box="[1330,1436,801,827]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="55" value="40.831">40.831N</geoCoordinate>
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<geoCoordinate id="EE15FC83517FFFA9FF62D64654DB2A22" box="[151,252,840,866]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="55" value="40.633">40.633E</geoCoordinate>
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, altitude ca.
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<quantity id="4CD937A1517FFFA9FE65D64655CE2A22" box="[400,489,840,866]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="2" pageNumber="3" unit="m" value="1900.0">1900 m</quantity>
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(locality
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<quantity id="4CD937A1517FFFA9FDACD64656A12A22" box="[601,646,840,866]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="2" pageNumber="3" unit="in" value="4.0">4 in</quantity>
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<figureCitation id="131A86C1517FFFA9FD78D64656C52A22" box="[653,738,840,866]" captionStart="FIGURE 13" captionStartId="9.[151,255,1395,1419]" captionTargetBox="[155,1432,514,1349]" captionTargetId="figure@9.[151,1436,506,1368]" captionTargetPageId="9" captionText="FIGURE 13. Distribution of Hydroporus lundbergi sp. n. (1 – 3; see text), H. cagrankaya sp. n. (4), H. sivrikaya sp. n. (5), H. toledoi sp. n. (6), H. artvinensis sp. n. (7), and H. erzurumensis (8 – 14). The map was made by using MS ENCARTA 2000." httpUri="https://zenodo.org/record/186326/files/figure.png" pageId="2" pageNumber="3">Fig. 13</figureCitation>
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).
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</paragraph>
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<paragraph id="8B9E9A44517FFFA9FF33D66050982DD7" blockId="2.[151,1437,796,2026]" pageId="2" pageNumber="3">
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<emphasis id="B9554656517FFFA9FF33D66055672AC8" bold="true" box="[198,320,878,904]" pageId="2" pageNumber="3">
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||
<typeStatus id="549A24E6517FFFA9FF33D660551C2AC8" box="[198,315,878,904]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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3, "
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<date id="FF9FBC84517FFFA9FE82D66055D92AC8" box="[375,510,878,904]" pageId="2" pageNumber="3" value="2001-07-29">29.07.2001</date>
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<collectingCountry id="F336DAD4517FFFA9FDFDD66056082AC8" box="[520,559,878,904]" name="Chile" pageId="2" pageNumber="3">TR</collectingCountry>
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–Rize, İkizdere distr., ca.
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<quantity id="4CD937A1517FFFA9FC95D66057BA2AC8" box="[864,925,878,904]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="3" unit="km" value="9.0">9 km</quantity>
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||
NE İkizdere, nr
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<taxonomicName id="4C21E1C7517FFFA9FB9DD66350D32AC7" box="[1128,1268,877,903]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="11" phylum="Arthropoda" rank="species" species="cagrankaya" status="sp. nov.">Çaġrankaya</taxonomicName>
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plateau,
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<collectingCountry id="F336DAD4517FFFA9FA97D660515D2AC8" box="[1378,1402,878,904]" name="Iceland" pageId="2" pageNumber="3">Ö</collectingCountry>
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||
.K. Erman leg." [printed], "ca.
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<quantity id="4CD937A1517FFFA9FE13D69B56612AEF" box="[486,582,917,943]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="2" pageNumber="3" unit="m" value="1900.0">1900 m</quantity>
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, small brooklets without vegetation, ca.
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<geoCoordinate id="EE15FC83517FFFA9FBCED69B50822AEF" box="[1083,1189,917,943]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="55" value="40.831">40.831N</geoCoordinate>
|
||
<geoCoordinate id="EE15FC83517FFFA9FB58D69B513F2AEF" box="[1197,1304,917,943]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="55" value="40.633">40.633E</geoCoordinate>
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" [printed], "
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<typeStatus id="549A24E6517FFFA9FF55D6B2553A2A96" box="[160,285,956,982]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
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,
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<taxonomicName id="4C21E1C7517FFFA9FEDED6B2567C2A96" box="[299,603,956,982]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="cagrankaya" status="sp. nov.">Hydroporus cagrankaya</taxonomicName>
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<taxonomicNameLabel id="A266FB2D517FFFA9FD90D6B556892A95" box="[613,686,955,981]" pageId="2" pageNumber="3" rank="species">
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<emphasis id="B9554656517FFFA9FD90D6B556892A95" bold="true" box="[613,686,955,981]" pageId="2" pageNumber="3">sp. n.</emphasis>
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</taxonomicNameLabel>
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, H. Fery &
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<collectingCountry id="F336DAD4517FFFA9FCB9D6B257412A96" box="[844,870,956,982]" name="Iceland" pageId="2" pageNumber="3">Ö</collectingCountry>
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||
.K. Erman det. 2009" [red, printed] (
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<collectionCode id="ED300281517FFFA9FACCD6B251B72A96" box="[1337,1424,956,982]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="3">NMW</collectionCode>
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||
).
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<emphasis id="B9554656517FFFA9FF62D6EC55032ABC" bold="true" box="[151,292,994,1020]" pageId="2" pageNumber="3">
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||
<typeStatus id="549A24E6517FFFA9FF62D6EC55382ABC" box="[151,287,994,1020]" pageId="2" pageNumber="3" type="paratype">Paratypes</typeStatus>
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||
.
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</emphasis>
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||
27 33, 31 ƤƤ, same collecting data as the
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<typeStatus id="549A24E6517FFFA9FCACD6EC57E02ABC" box="[857,967,994,1020]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
|
||
(
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<collectionCode id="ED300281517FFFA9FC2DD6EC50302ABC" box="[984,1047,994,1020]" pageId="2" pageNumber="3">CHF</collectionCode>
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||
,
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||
<collectionCode id="ED300281517FFFA9FBD0D6EC504F2ABC" LSID="urn:lsid:biocol.org:col:12741" box="[1061,1128,994,1020]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12741" name="Calke Abbey" pageId="2" pageNumber="3">CKE</collectionCode>
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||
). 1 3, idem, but with an additional label "
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<collectionCode id="ED300281517FFFA9FEA9D107559F2D63" LSID="urn:lsid:biocol.org:col:35275" box="[348,440,1033,1059]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="2" pageNumber="3">MNCN</collectionCode>
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||
<collectionCode id="ED300281517FFFA9FE4AD10755CD2D63" box="[447,490,1033,1059]" pageId="2" pageNumber="3">AI-</collectionCode>
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||
62,
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||
<collectionCode id="ED300281517FFFA9FDE1D10756732D63" LSID="urn:lsid:biocol.org:col:15536" box="[532,596,1033,1059]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="2" pageNumber="3">DNA</collectionCode>
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||
specimen"; the
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<collectionCode id="ED300281517FFFA9FCE4D10757762D63" LSID="urn:lsid:biocol.org:col:15536" box="[785,849,1033,1059]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="2" pageNumber="3">DNA</collectionCode>
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extraction of this specimen is stored in the tissue-
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<collectionCode id="ED300281517FFFA9FF62D13E54F02D0A" LSID="urn:lsid:biocol.org:col:15536" box="[151,215,1072,1098]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="2" pageNumber="3">DNA</collectionCode>
|
||
collection of the
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||
<collectionCode id="ED300281517FFFA9FE52D13E55D82D0A" LSID="urn:lsid:biocol.org:col:35275" box="[423,511,1072,1098]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="2" pageNumber="3">MNCN</collectionCode>
|
||
with Ref. No. 23408. Each
|
||
<typeStatus id="549A24E6517FFFA9FCB8D13E57952D0A" box="[845,946,1072,1098]" pageId="2" pageNumber="3" type="paratype">paratype</typeStatus>
|
||
is provided with the respective red label.
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||
<emphasis id="B9554656517FFFA9FF62D15855802D30" bold="true" box="[151,423,1110,1136]" pageId="2" pageNumber="3">Additional specimens</emphasis>
|
||
(not treated as
|
||
<typeStatus id="549A24E6517FFFA9FD97D15856FD2D30" box="[610,730,1110,1136]" pageId="2" pageNumber="3" type="paratype">paratypes</typeStatus>
|
||
). 8 exs., with the same collecting data, but without head or destroyed otherwise; altogether mounted on a common glue card, without red label (
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||
<collectionCode id="ED300281517FFFA9FB86D17350952DD7" box="[1139,1202,1149,1175]" pageId="2" pageNumber="3">CHF</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33BC9CF517FFFAEFF33D1AD508F2AD0" lastPageId="5" lastPageNumber="6" pageId="2" pageNumber="3" type="description">
|
||
<paragraph id="8B9E9A44517FFFA9FF33D1AD51262C4B" blockId="2.[151,1437,796,2026]" pageId="2" pageNumber="3">
|
||
<emphasis id="B9554656517FFFA9FF33D1AD55EB2DFD" bold="true" box="[198,460,1187,1213]" pageId="2" pageNumber="3">Description. Habitus</emphasis>
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||
elongate oval, maximum width situated near middle of total length. Lateral body outline in dorsal view with a very slight, but perceptible discontinuity between pronotum and elytra. Almost entire dorsal and ventral surface dark blackish brown. Upper surface microreticulated, but shiny.
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||
</paragraph>
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<paragraph id="8B9E9A44517FFFA9FF33D01957862CB3" blockId="2.[151,1437,796,2026]" pageId="2" pageNumber="3">
|
||
<emphasis id="B9554656517FFFA9FF33D019552E2C71" bold="true" box="[198,265,1303,1329]" pageId="2" pageNumber="3">Head</emphasis>
|
||
rather broad, interocular distance equalling about half of pronotal width at hind angles. Anterior margin of clypeus in middle and above insertion of antennae shining transparently brownish; vertex with a transverse brown marking. Two grooves behind anterior margin of clypeus. Punctures on head rather small and evenly distributed, however, denser in clypeal grooves, and still smaller and slightly sparser on vertex. Antennae with third article short, fourth still shorter; articles five to ten somewhat flattened, less than 1.5 times as long as wide, resembling almost spherical in proper view.
|
||
</paragraph>
|
||
<paragraph id="8B9E9A44517FFFA9FF33D0F155692E68" blockId="2.[151,1437,796,2026]" pageId="2" pageNumber="3">
|
||
<emphasis id="B9554656517FFFA9FF33D0F155632F59" bold="true" box="[198,324,1535,1561]" pageId="2" pageNumber="3">Pronotum</emphasis>
|
||
with anterior and posterior margins, and lateral beading—in particular posteriorly—brownish transparent. Sides of pronotum curved, stronger anteriorly, indistinctly posteriorly. Rim distinct, but not very broad, more or less of same width over entire length. Maximum width of pronotum between hind angles. Punctation on disc finer and sparser than on clypeus or head, coarser and denser near margins. Some very coarse punctures behind anterior margin, a few also before hind margin. Postero-laterally with an oblique depressed area on each side, here reticulation slightly stronger and punctures in part deformed to weakly impressed short scratches. Centre of disc with a short, but distinct scratch. Lateral parts of pronotum with indistinct setae.
|
||
</paragraph>
|
||
<paragraph id="8B9E9A44517FFFA8FF33D23B55DE29BF" blockId="2.[151,1437,796,2026]" lastBlockId="3.[151,1436,152,255]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
|
||
<emphasis id="B9554656517FFFA9FF33D23B55312E0F" bold="true" box="[198,278,1845,1871]" pageId="2" pageNumber="3">Elytra</emphasis>
|
||
indistinctly dark brown next to suture and near apex. Punctation more or less uniformly distributed. Punctures distinctly coarser than on head and also slightly coarser than on sides of pronotum; near sides and suture somewhat smaller. On disc distance between punctures roughly two times that of their diameter. Distinct puncture lines absent, nevertheless weakly perceptible since normal punctures here somewhat denser, and a very few still coarser punctures interspersed. Setae on disc short and indistinct, laterally and posteriorly longer and more distinct. In lateral view margin of elytra ascending slightly towards humeral angle. Lateral elytral beading distinct, more or less as thick as pronotal beading. Epipleura in lateral view visible to humeral angle.
|
||
</paragraph>
|
||
<caption id="DF5ECACC517EFFA8FF62D284572C2EA7" httpUri="https://zenodo.org/record/186323/files/figure.png" pageId="3" pageNumber="4" targetBox="[223,1378,304,1892]" targetPageId="3">
|
||
<paragraph id="8B9E9A44517EFFA8FF62D284572C2EA7" blockId="3.[151,1436,1930,2023]" pageId="3" pageNumber="4">
|
||
<emphasis id="B9554656517EFFA8FF62D284556D2EE2" bold="true" box="[151,330,1930,1954]" pageId="3" pageNumber="4">FIGURES 1–5.</emphasis>
|
||
Median lobe of aedeagus of: (1)
|
||
<taxonomicName id="4C21E1C7517EFFA8FD34D28557F62EE2" box="[705,977,1931,1954]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="cagrankaya" status="sp. nov.">
|
||
<emphasis id="B9554656517EFFA8FD34D28557F62EE2" box="[705,977,1931,1954]" italics="true" pageId="3" pageNumber="4">Hydroporus cagrankaya</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A266FB2D517EFFA8FC2CD284503E2EE2" box="[985,1049,1930,1954]" pageId="3" pageNumber="4" rank="species">
|
||
<emphasis id="B9554656517EFFA8FC2CD284503E2EE2" bold="true" box="[985,1049,1930,1954]" pageId="3" pageNumber="4">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(holotype), (2)
|
||
<taxonomicName id="4C21E1C7517EFFA8FB39D28551722EE2" box="[1228,1365,1931,1954]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="sivrikaya" status="sp. nov.">
|
||
<emphasis id="B9554656517EFFA8FB39D28551722EE2" box="[1228,1365,1931,1954]" italics="true" pageId="3" pageNumber="4">H. sivrikaya</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A266FB2D517EFFA8FAA9D28451BB2EE2" box="[1372,1436,1930,1954]" pageId="3" pageNumber="4" rank="species">
|
||
<emphasis id="B9554656517EFFA8FAA9D28451BB2EE2" bold="true" box="[1372,1436,1930,1954]" pageId="3" pageNumber="4">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(holotype), (3)
|
||
<taxonomicName id="4C21E1C7517EFFA8FECDD2A055EA2E85" box="[312,461,1966,1989]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="artvinensis" status="sp. nov.">
|
||
<emphasis id="B9554656517EFFA8FECDD2A055EA2E85" box="[312,461,1966,1989]" italics="true" pageId="3" pageNumber="4">H. artvinensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A266FB2D517EFFA8FE26D2A356292E85" box="[467,526,1965,1989]" pageId="3" pageNumber="4" rank="species">
|
||
<emphasis id="B9554656517EFFA8FE26D2A356292E85" bold="true" box="[467,526,1965,1989]" pageId="3" pageNumber="4">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(holotype), (4)
|
||
<taxonomicName id="4C21E1C7517EFFA8FD43D2A057072E85" box="[694,800,1966,1989]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="toledoi" status="sp. nov.">
|
||
<emphasis id="B9554656517EFFA8FD43D2A057072E85" box="[694,800,1966,1989]" italics="true" pageId="3" pageNumber="4">H. toledoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A266FB2D517EFFA8FCD3D2A357462E85" box="[806,865,1965,1989]" pageId="3" pageNumber="4" rank="species">
|
||
<emphasis id="B9554656517EFFA8FCD3D2A357462E85" bold="true" box="[806,865,1965,1989]" pageId="3" pageNumber="4">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(holotype), (5a, b)
|
||
<taxonomicName id="4C21E1C7517EFFA8FBDAD2A050912E85" box="[1071,1206,1966,1989]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="lundbergi" status="sp. nov.">
|
||
<emphasis id="B9554656517EFFA8FBDAD2A050912E85" box="[1071,1206,1966,1989]" italics="true" pageId="3" pageNumber="4">H. lundbergi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A266FB2D517EFFA8FB49D2A350D02E85" box="[1212,1271,1965,1989]" pageId="3" pageNumber="4" rank="species">
|
||
<emphasis id="B9554656517EFFA8FB49D2A350D02E85" bold="true" box="[1212,1271,1965,1989]" pageId="3" pageNumber="4">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(holotype) (the arrow indicates the direction of the "frontal view", Fig. 5b)
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5ECACC5179FFAFFF62D7D350E02BB5" box="[151,1223,733,757]" httpUri="https://zenodo.org/record/186324/files/figure.png" pageId="4" pageNumber="5" targetBox="[195,1350,222,702]" targetPageId="4">
|
||
<paragraph id="8B9E9A445179FFAFFF62D7D350E02BB5" blockId="4.[151,1223,733,757]" box="[151,1223,733,757]" pageId="4" pageNumber="5">
|
||
<emphasis id="B95546565179FFAFFF62D7D355642BB5" bold="true" box="[151,323,733,757]" pageId="4" pageNumber="5">FIGURES 6–8.</emphasis>
|
||
<taxonomicName id="4C21E1C75179FFAFFEBCD7D056692BB5" box="[329,590,734,757]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="cagrankaya" status="sp. nov.">
|
||
<emphasis id="B95546565179FFAFFEBCD7D056692BB5" box="[329,590,734,757]" italics="true" pageId="4" pageNumber="5">Hydroporus cagrankaya</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A266FB2D5179FFAFFDA1D7D356B72BB5" box="[596,656,733,757]" pageId="4" pageNumber="5" rank="species">
|
||
<emphasis id="B95546565179FFAFFDA1D7D356B72BB5" bold="true" box="[596,656,733,757]" pageId="4" pageNumber="5">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
: (6) paramere, (7) gonocoxae, (8) gonocoxosternum.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9E9A445179FFAFFF33D62D569C2DA6" blockId="4.[151,1437,803,2028]" pageId="4" pageNumber="5">
|
||
<emphasis id="B95546565179FFAFFF33D62D55A12A7D" bold="true" box="[198,390,803,829]" pageId="4" pageNumber="5">Ventral surface</emphasis>
|
||
(
|
||
<figureCitation id="131A86C15179FFAFFE63D62D55FA2A7D" box="[406,477,803,829]" captionStart="FIGURES 9 – 12" captionStartId="6.[151,274,1767,1791]" captionTargetBox="[212,1374,194,1743]" captionTargetId="figure@6.[212,1376,194,1743]" captionTargetPageId="6" captionText="FIGURES 9 – 12. Hydroporus cagrankaya sp. n.: SEM photos of: (9) parts of venter, (10) left protarsus, (11) left mesotarsus, (12) details of first and second tarsomeres of left protarsus." httpUri="https://zenodo.org/record/186325/files/figure.png" pageId="4" pageNumber="5">Fig. 9</figureCitation>
|
||
) predominantly black; posterior part of metacoxal processes, hind margins of last abdominal ventrite, an indistinctly limited spot on each side of third, fourth and fifth abdominal ventrite transparently brownish. Gula very slightly more brownish than black genae. Legs, mouthparts, and prosternal process brownish; femora only insignificantly darkened, if at all. Antennae brownish, segments apically progressively but only weakly darkened beginning with fifth. Most parts of venter with weakly impressed microreticulation; gula, metacoxal processes, centres of metaventrite and first abdominal ventrites smooth. Genae smooth close to gula, else reticulated. Punctation on metacoxal plates, sides of metaventrite, and first two abdominal ventrites coarse; on epipleura less coarse, also on rest of abdominal ventrites, here medially sparser, in particular on last ventrite; punctation on centre of metaventrite rather fine and sparse, on metacoxal processes absent. Gula laterally with some punctures. Anterior angles of prosternum with a small, flat, rather densely and roughly punctured triangular area; this area well limited against impunctate area behind. Most punctures on venter with a fine, indistinct seta.
|
||
</paragraph>
|
||
<paragraph id="8B9E9A445179FFAFFF33D1FD56D92C8E" blockId="4.[151,1437,803,2028]" pageId="4" pageNumber="5">
|
||
Prosternal process (
|
||
<figureCitation id="131A86C15179FFAFFE4DD1FD56242C4D" box="[440,515,1267,1293]" captionStart="FIGURES 9 – 12" captionStartId="6.[151,274,1767,1791]" captionTargetBox="[212,1374,194,1743]" captionTargetId="figure@6.[212,1376,194,1743]" captionTargetPageId="6" captionText="FIGURES 9 – 12. Hydroporus cagrankaya sp. n.: SEM photos of: (9) parts of venter, (10) left protarsus, (11) left mesotarsus, (12) details of first and second tarsomeres of left protarsus." httpUri="https://zenodo.org/record/186325/files/figure.png" pageId="4" pageNumber="5">Fig. 9</figureCitation>
|
||
) posteriorly lanceolate, here more or less tectiform in cross-section, sides beaded, provided with long setae. Anteriorly, between procoxae, process with a small transverse ridge, subbasally before this ridge with transverse grooves; process not prolonged anteriorly on to prosternum as narrow convexity; here prosternum medially somewhat flattened and rugosely sculptured. Posterior margins of metacoxal processes sinuate and medially protruded backwards; lines of metacoxal processes diverging anteriorly, not reaching hind margin of metaventrite.
|
||
</paragraph>
|
||
<paragraph id="8B9E9A445179FFAFFF33D0D551642F5C" blockId="4.[151,1437,803,2028]" pageId="4" pageNumber="5">
|
||
<emphasis id="B95546565179FFAFFF33D0D555752CB5" bold="true" box="[198,338,1499,1525]" pageId="4" pageNumber="5">Variability.</emphasis>
|
||
Depending on maturity, several
|
||
<typeStatus id="549A24E65179FFAFFD22D0D557602CB5" box="[727,839,1499,1525]" pageId="4" pageNumber="5" type="paratype">paratypes</typeStatus>
|
||
at least with elytra more brownish, and pronotum slightly darker than elytra and head. Sides of pronotum in some specimens posteriorly almost straight.
|
||
</paragraph>
|
||
<paragraph id="8B9E9A445179FFAFFF33D32655B32E38" blockId="4.[151,1437,803,2028]" pageId="4" pageNumber="5">
|
||
<emphasis id="B95546565179FFAFFF33D326553F2F02" bold="true" box="[198,280,1576,1602]" pageId="4" pageNumber="5">Males.</emphasis>
|
||
First three pro- and mesotarsomeres slightly dilated, ventrally provided with adhesive setae (also known as "small suckers"); first protarsomere additionally with three rather regularly arranged pairs of elongate sucker cups (also called "large suckers", "adhesive discs", "scales" etc.) (
|
||
<figureCitation id="131A86C15179FFAFFB60D37851752FD0" box="[1173,1362,1654,1680]" captionStart="FIGURES 9 – 12" captionStartId="6.[151,274,1767,1791]" captionTargetBox="[212,1374,194,1743]" captionTargetId="figure@6.[212,1376,194,1743]" captionTargetPageId="6" captionText="FIGURES 9 – 12. Hydroporus cagrankaya sp. n.: SEM photos of: (9) parts of venter, (10) left protarsus, (11) left mesotarsus, (12) details of first and second tarsomeres of left protarsus." httpUri="https://zenodo.org/record/186325/files/figure.png" pageId="4" pageNumber="5">Figs 10 and 12</figureCitation>
|
||
), first mesotarsomere with seven irregularly arranged cups (
|
||
<figureCitation id="131A86C15179FFAFFCF9D39257452FF6" box="[780,866,1692,1718]" captionStart="FIGURES 9 – 12" captionStartId="6.[151,274,1767,1791]" captionTargetBox="[212,1374,194,1743]" captionTargetId="figure@6.[212,1376,194,1743]" captionTargetPageId="6" captionText="FIGURES 9 – 12. Hydroporus cagrankaya sp. n.: SEM photos of: (9) parts of venter, (10) left protarsus, (11) left mesotarsus, (12) details of first and second tarsomeres of left protarsus." httpUri="https://zenodo.org/record/186325/files/figure.png" pageId="4" pageNumber="5">Fig. 11</figureCitation>
|
||
); protarsal claws short, in particular anterior one thickened, curved near base and almost straight in distal two thirds (
|
||
<figureCitation id="131A86C15179FFAFFC41D3CD502D2F9D" box="[948,1034,1731,1757]" captionStart="FIGURES 9 – 12" captionStartId="6.[151,274,1767,1791]" captionTargetBox="[212,1374,194,1743]" captionTargetId="figure@6.[212,1376,194,1743]" captionTargetPageId="6" captionText="FIGURES 9 – 12. Hydroporus cagrankaya sp. n.: SEM photos of: (9) parts of venter, (10) left protarsus, (11) left mesotarsus, (12) details of first and second tarsomeres of left protarsus." httpUri="https://zenodo.org/record/186325/files/figure.png" pageId="4" pageNumber="5">Fig. 10</figureCitation>
|
||
); mesotarsal claws evenly curved, longer than protarsal claws (
|
||
<figureCitation id="131A86C15179FFAFFE12D3E456192E44" box="[487,574,1770,1796]" captionStart="FIGURES 9 – 12" captionStartId="6.[151,274,1767,1791]" captionTargetBox="[212,1374,194,1743]" captionTargetId="figure@6.[212,1376,194,1743]" captionTargetPageId="6" captionText="FIGURES 9 – 12. Hydroporus cagrankaya sp. n.: SEM photos of: (9) parts of venter, (10) left protarsus, (11) left mesotarsus, (12) details of first and second tarsomeres of left protarsus." httpUri="https://zenodo.org/record/186325/files/figure.png" pageId="4" pageNumber="5">Fig. 11</figureCitation>
|
||
). Median lobe of aedeagus (
|
||
<figureCitation id="131A86C15179FFAFFC7BD3E457F52E44" box="[910,978,1770,1796]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,273,1930,1954]" captionTargetBox="[223,1378,304,1892]" captionTargetId="figure@3.[200,1387,283,1906]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Median lobe of aedeagus of: (1) Hydroporus cagrankaya sp. n. (holotype), (2) H. sivrikaya sp. n. (holotype), (3) H. artvinensis sp. n. (holotype), (4) H. toledoi sp. n. (holotype), (5 a, b) H. lundbergi sp. n. (holotype) (the arrow indicates the direction of the " frontal view ", Fig. 5 b)" httpUri="https://zenodo.org/record/186323/files/figure.png" pageId="4" pageNumber="5">Fig. 1</figureCitation>
|
||
) asymmetrical, with apex twisted (see frontal view for
|
||
<taxonomicName id="4C21E1C75179FFAFFEA9D21F55D32E6A" box="[348,500,1809,1834]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="lundbergi" status="sp. nov.">
|
||
<emphasis id="B95546565179FFAFFEA9D21F55D32E6A" box="[348,500,1809,1834]" italics="true" pageId="4" pageNumber="5">H. lundbergi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A266FB2D5179FFAFFE08D21E56662E6A" box="[509,577,1808,1834]" pageId="4" pageNumber="5" rank="species">
|
||
<emphasis id="B95546565179FFAFFE08D21E56662E6A" bold="true" box="[509,577,1808,1834]" pageId="4" pageNumber="5">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
in
|
||
<figureCitation id="131A86C15179FFAFFD9CD21F56882E6B" box="[617,687,1809,1835]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,273,1930,1954]" captionTargetBox="[223,1378,304,1892]" captionTargetId="figure@3.[200,1387,283,1906]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Median lobe of aedeagus of: (1) Hydroporus cagrankaya sp. n. (holotype), (2) H. sivrikaya sp. n. (holotype), (3) H. artvinensis sp. n. (holotype), (4) H. toledoi sp. n. (holotype), (5 a, b) H. lundbergi sp. n. (holotype) (the arrow indicates the direction of the " frontal view ", Fig. 5 b)" httpUri="https://zenodo.org/record/186323/files/figure.png" pageId="4" pageNumber="5">Fig. 5</figureCitation>
|
||
b, a species which—due to the broad apex—has this character particularly conspicuous); in ventral view apically converging, apex rounded; in lateral view rather pointed; paramere as in
|
||
<figureCitation id="131A86C15179FFAFFEBCD25055B62E38" box="[329,401,1886,1912]" captionStart="FIGURES 6 – 8" captionStartId="4.[151,269,733,757]" captionTargetBox="[195,1350,222,702]" captionTargetId="figure@4.[182,1404,194,710]" captionTargetPageId="4" captionText="FIGURES 6 – 8. Hydroporus cagrankaya sp. n.: (6) paramere, (7) gonocoxae, (8) gonocoxosternum." httpUri="https://zenodo.org/record/186324/files/figure.png" pageId="4" pageNumber="5">Fig. 6</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9E9A445179FFAEFF33D28A55E828DA" blockId="4.[151,1437,803,2028]" lastBlockId="5.[151,1437,152,1222]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
|
||
<emphasis id="B95546565179FFAFFF33D28A55332EDE" bold="true" box="[198,276,1924,1950]" pageId="4" pageNumber="5">Notes.</emphasis>
|
||
<bibRefCitation id="EFB0E7B55179FFAFFEEED28A56302EDE" author="Larson" box="[283,535,1924,1950]" pageId="4" pageNumber="12" refString="Larson, D. J. (1975) The predaceous water beetles (Coleoptera: Dytiscidae) of Alberta: systematics, natural history and distribution. Quaestiones Entomologicae, 11, 245 - 498." type="journal article" year="1975">Larson (1975: 280 ff)</bibRefCitation>
|
||
in his key of Canadian
|
||
<taxonomicName id="4C21E1C75179FFAFFCDAD28B579C2EDE" box="[815,955,1925,1950]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95546565179FFAFFCDAD28B579C2EDE" box="[815,955,1925,1950]" italics="true" pageId="4" pageNumber="5">Hydroporus</emphasis>
|
||
</taxonomicName>
|
||
(here still including species which today are treated in the genera
|
||
<taxonomicName id="4C21E1C75179FFAFFE33D2A557BB2E85" authority="Roughley & Larson, 2000" authorityName="Roughley & Larson" authorityYear="2000" box="[454,924,1963,1989]" class="Insecta" family="Dytiscidae" genus="Hydrocolus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95546565179FFAFFE33D2A556752E84" box="[454,594,1963,1988]" italics="true" pageId="4" pageNumber="5">Hydrocolus</emphasis>
|
||
Roughley & Larson, 2000
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C21E1C75179FFAFFC5CD2A5513E2E85" authority="Regimbart, 1906" authorityName="Regimbart" authorityYear="1906" box="[937,1305,1963,1989]" class="Insecta" family="Dytiscidae" genus="Nebrioporus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95546565179FFAFFC5CD2A550652E84" box="[937,1090,1963,1988]" italics="true" pageId="4" pageNumber="5">Nebrioporus</emphasis>
|
||
Régimbart, 1906
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C21E1C75179FFAFFAD2D2A555612EAC" authority="Guignot, 1931" authorityName="Guignot" authorityYear="1931" class="Insecta" family="Dytiscidae" genus="Neoporus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95546565179FFAFFAD2D2A551BB2E84" box="[1319,1436,1963,1988]" italics="true" pageId="4" pageNumber="5">Neoporus</emphasis>
|
||
Guignot, 1931
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C21E1C75179FFAFFEA6D2DC565B2EAC" authority="Seidlitz, 1887" authorityName="Seidlitz" authorityYear="1887" box="[339,636,2002,2028]" class="Insecta" family="Dytiscidae" genus="Oreodytes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95546565179FFAFFEA6D2DC55EB2EAB" box="[339,460,2002,2027]" italics="true" pageId="4" pageNumber="5">Oreodytes</emphasis>
|
||
Seidlitz, 1887
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C21E1C75179FFAFFD7CD2DC50072EAC" authority="Franciscolo, 1979" authorityName="Franciscolo" authorityYear="1979" box="[649,1056,2002,2028]" class="Insecta" family="Dytiscidae" genus="Sanfilippodytes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95546565179FFAFFD7CD2DC57672EAB" box="[649,832,2002,2027]" italics="true" pageId="4" pageNumber="5">Sanfilippodytes</emphasis>
|
||
Franciscolo, 1979
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C21E1C75179FFAEFB95D2DC54F129F2" authority="Zimmermann, 1919" authorityName="Zimmermann" authorityYear="1919" class="Insecta" family="Dytiscidae" genus="Stictotarsus" kingdom="Animalia" lastPageId="5" lastPageNumber="6" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95546565179FFAFFB95D2DC50CC2EAB" box="[1120,1259,2002,2027]" italics="true" pageId="4" pageNumber="5">Stictotarsus</emphasis>
|
||
Zimmermann, 1919
|
||
</taxonomicName>
|
||
) introduced the number of male protarsal sucker cups as a differentiating character. Most of those species for which the number of sucker cups is given in that work have two pairs of cups on the ventral side of the first protarsomere, a few other species have more. Recently,
|
||
<bibRefCitation id="EFB0E7B55178FFAEFC7CD5EB504E29BF" box="[905,1129,229,255]" pageId="5" pageNumber="11" refString="Bilton, D. T., Thompson, A. & Foster, G. N. (2008) Inter- and intrasexual dimorphism in the diving beetle Hydroporus memnonius Nicolai (Coleoptera: Dytiscidae). Biological Journal of the Linnean Society, 94, 685 - 697." type="journal article">Bilton et al. (2008)</bibRefCitation>
|
||
published SEM photos of the pro-and mesotarsomeres of
|
||
<taxonomicName id="4C21E1C75178FFAEFDF8D40257F22866" authority="Nicolai, 1822" authorityName="Nicolai" authorityYear="1822" box="[525,981,268,294]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="memnonius">
|
||
<emphasis id="B95546565178FFAEFDF8D402570F2865" box="[525,808,268,293]" italics="true" pageId="5" pageNumber="6">Hydroporus memnonius</emphasis>
|
||
Nicolai, 1822
|
||
</taxonomicName>
|
||
, which show an irregular arrangement of several sucker cups, the number of which varies in different populations. We wonder whether the number and the arrangement of cups is characteristic for the members of the
|
||
<taxonomicName id="4C21E1C75178FFAEFC2BD45750612832" box="[990,1094,345,370]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="longulus">
|
||
<emphasis id="B95546565178FFAEFC2BD45750612832" box="[990,1094,345,370]" italics="true" pageId="5" pageNumber="6">longulus</emphasis>
|
||
</taxonomicName>
|
||
-group and propose to study these also in other species.
|
||
</paragraph>
|
||
<paragraph id="8B9E9A445178FFAEFF33D4A850B12B4E" blockId="5.[151,1437,152,1222]" pageId="5" pageNumber="6">
|
||
<emphasis id="B95546565178FFAEFF33D4A855132880" bold="true" box="[198,308,422,448]" pageId="5" pageNumber="6">Females.</emphasis>
|
||
Without conspicuous external differences to males, except slightly narrower and ventrally simple pro- and mesotarsomeres, and evenly curved protarsal claws; these longer than in males and—as in males—shorter than mesotarsal ones. Gonocoxae and gonocoxosternum as in
|
||
<figureCitation id="131A86C15178FFAEFBDCD4FA50B52B4E" box="[1065,1170,500,526]" captionStart="FIGURES 6 – 8" captionStartId="4.[151,269,733,757]" captionTargetBox="[195,1350,222,702]" captionTargetId="figure@4.[182,1404,194,710]" captionTargetPageId="4" captionText="FIGURES 6 – 8. Hydroporus cagrankaya sp. n.: (6) paramere, (7) gonocoxae, (8) gonocoxosternum." httpUri="https://zenodo.org/record/186324/files/figure.png" pageId="5" pageNumber="6">Figs 7–8</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9E9A445178FFAEFF33D71455D72B1B" blockId="5.[151,1437,152,1222]" pageId="5" pageNumber="6">
|
||
<emphasis id="B95546565178FFAEFF33D71455A22B74" bold="true" box="[198,389,538,564]" pageId="5" pageNumber="6">Measurements.</emphasis>
|
||
TL:
|
||
<quantity id="4CD937A15178FFAEFE27D71456AE2B74" box="[466,649,538,564]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" metricValueMax="3.65" metricValueMin="3.15" pageId="5" pageNumber="6" unit="mm" value="3.4" valueMax="3.65" valueMin="3.15">3.15–3.65 mm</quantity>
|
||
, MW:
|
||
<quantity id="4CD937A15178FFAEFD1ED71457852B74" box="[747,930,538,564]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" metricValueMax="1.85" metricValueMin="1.55" pageId="5" pageNumber="6" unit="mm" value="1.7" valueMax="1.85" valueMin="1.55">1.55–1.85 mm</quantity>
|
||
, TL/MW: 1.94–2.10, IO/MP: 0.46–0.49 (males), 0.45–0.51 (females).
|
||
</paragraph>
|
||
<paragraph id="8B9E9A445178FFAEFF33D769508F2AD0" blockId="5.[151,1437,152,1222]" pageId="5" pageNumber="6">
|
||
<emphasis id="B95546565178FFAEFF33D76955B72BC1" bold="true" box="[198,400,615,641]" pageId="5" pageNumber="6">Molecular data.</emphasis>
|
||
The DNA of one
|
||
<typeStatus id="549A24E65178FFAEFD83D76656FC2BC2" box="[630,731,616,642]" pageId="5" pageNumber="6" type="paratype">paratype</typeStatus>
|
||
was obtained in the laboratory of our colleague I. Ribera (Barcelona,
|
||
<collectingCountry id="F336DAD45178FFAEFEDCD78055492BE8" box="[297,366,654,680]" name="Spain" pageId="5" pageNumber="6">Spain</collectingCountry>
|
||
) using a standard phenol chlorophorm extraction (voucher number MNCN AI62/23408). The sequence of a fragment of 424 bp from the 3’ end of the gene cox1 and a fragment of 751 bp spanning the mitochondrial genes rrnL - tRNAleu - NDA1 are deposited in the EMBL database with accession numbers
|
||
<accessionNumber id="947207A75178FFAEFF62D60C55072A5C" box="[151,288,770,796]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FM946166" pageId="5" pageNumber="6">FM946166</accessionNumber>
|
||
to
|
||
<accessionNumber id="947207A75178FFAEFEBFD60C55F32A5C" box="[330,468,770,796]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FM946169" pageId="5" pageNumber="6">FM946169</accessionNumber>
|
||
(see
|
||
<bibRefCitation id="EFB0E7B55178FFAEFDEDD60C56DD2A5C" author="Ribera" box="[536,762,770,796]" pageId="5" pageNumber="12" refString="Ribera, I., Hernando, C. & Aguilera, P. (2001) Agabus alexandrae sp. n. from Morocco, with a molecular phylogeny of the Western Mediterranean species of the A. guttatus group (Coleoptera: Dytiscidae). Insect Systematics & Evolution, 32, 253 - 262." type="journal article" year="2001">Ribera et al. 2001</bibRefCitation>
|
||
for primers and PCR conditions). The results will be published in a future work, however, we can give already here the information that
|
||
<taxonomicName id="4C21E1C75178FFAEFB6CD62751692A02" box="[1177,1358,809,834]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="cagrankaya" status="sp. nov.">
|
||
<emphasis id="B95546565178FFAEFB6CD62751692A02" box="[1177,1358,809,834]" italics="true" pageId="5" pageNumber="6">H. cagrankaya</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A266FB2D5178FFAEFAA2D62651BB2A02" box="[1367,1436,808,834]" pageId="5" pageNumber="6" rank="species">
|
||
<emphasis id="B95546565178FFAEFAA2D62651BB2A02" bold="true" box="[1367,1436,808,834]" pageId="5" pageNumber="6">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
belongs to a complex of species which comprises
|
||
<taxonomicName id="4C21E1C75178FFAEFCF3D65E50DC2A2A" authority="Ienistea, 1962" authorityName="Ienistea" authorityYear="1962" box="[774,1275,846,874]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="dobrogeanus">
|
||
<emphasis id="B95546565178FFAEFCF3D65E50662A29" box="[774,1089,848,873]" italics="true" pageId="5" pageNumber="6">Hydroporus dobrogeanus</emphasis>
|
||
Ieniştea, 1962
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C21E1C75178FFAEFAFCD65E55C62AD0" authority="Wewalka, 1975" authorityName="Wewalka" authorityYear="1975" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="gueorguievi">
|
||
<emphasis id="B95546565178FFAEFAFCD65E55052AD0" italics="true" pageId="5" pageNumber="6">Hydroporus gueorguievi</emphasis>
|
||
Wewalka, 1975
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C21E1C75178FFAEFE18D67957AB2AD0" authority="Schaum, 1868" authorityName="Schaum" authorityYear="1868" box="[493,908,886,912]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="kraatzii">
|
||
<emphasis id="B95546565178FFAEFE18D67956FD2AD0" box="[493,730,887,912]" italics="true" pageId="5" pageNumber="6">Hydroporus kraatzii</emphasis>
|
||
Schaum, 1868
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C21E1C75178FFAEFC3FD67950862AD0" box="[970,1185,887,912]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="sardomontanus">
|
||
<emphasis id="B95546565178FFAEFC3FD67950862AD0" box="[970,1185,887,912]" italics="true" pageId="5" pageNumber="6">H. sardomontanus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33BC9CF5178FFAEFF33D69356B32A9E" pageId="5" pageNumber="6" type="distribution">
|
||
<paragraph id="8B9E9A445178FFAEFF33D69356B32A9E" blockId="5.[151,1437,152,1222]" pageId="5" pageNumber="6">
|
||
<emphasis id="B95546565178FFAEFF33D69355402AF7" bold="true" box="[198,359,925,951]" pageId="5" pageNumber="6">Distribution.</emphasis>
|
||
So far known only from the
|
||
<typeStatus id="549A24E65178FFAEFD4DD69356CD2AF7" box="[696,746,925,951]" pageId="5" pageNumber="6">type</typeStatus>
|
||
locality, near the
|
||
<taxonomicName id="4C21E1C75178FFAEFC4FD69550612AF5" box="[954,1094,923,949]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="11" phylum="Arthropoda" rank="species" species="cagrankaya" status="sp. nov.">Çaġrankaya</taxonomicName>
|
||
plateau in the Rize province, north-eastern
|
||
<collectingCountry id="F336DAD45178FFAEFECDD6CA55AC2A9E" box="[312,395,964,990]" name="Turkey" pageId="5" pageNumber="6">Turkey</collectingCountry>
|
||
(locality
|
||
<quantity id="4CD937A15178FFAEFE09D6CA560E2A9E" box="[508,553,964,990]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="5" pageNumber="6" unit="in" value="4.0">4 in</quantity>
|
||
<figureCitation id="131A86C15178FFAEFDC5D6CA56A12A9E" box="[560,646,964,990]" captionStart="FIGURE 13" captionStartId="9.[151,255,1395,1419]" captionTargetBox="[155,1432,514,1349]" captionTargetId="figure@9.[151,1436,506,1368]" captionTargetPageId="9" captionText="FIGURE 13. Distribution of Hydroporus lundbergi sp. n. (1 – 3; see text), H. cagrankaya sp. n. (4), H. sivrikaya sp. n. (5), H. toledoi sp. n. (6), H. artvinensis sp. n. (7), and H. erzurumensis (8 – 14). The map was made by using MS ENCARTA 2000." httpUri="https://zenodo.org/record/186326/files/figure.png" pageId="5" pageNumber="6">Fig. 13</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33BC9CF5178FFAEFF33D6E450212D38" pageId="5" pageNumber="6" type="biology_ecology">
|
||
<paragraph id="8B9E9A445178FFAEFF33D6E450212D38" blockId="5.[151,1437,152,1222]" pageId="5" pageNumber="6">
|
||
<emphasis id="B95546565178FFAEFF33D6E4550D2D44" bold="true" box="[198,298,1002,1028]" pageId="5" pageNumber="6">Biology.</emphasis>
|
||
The new species has been found in small brooklets without vegetation. These are fed by melting snow from spring to early summer, are exposed to the sun, and without rain they dry from middle summer. The specimens have been found in association with
|
||
<taxonomicName id="4C21E1C75178FFAEFCFAD13651272D12" authority="Fabricius, 1792" authorityName="Fabricius" authorityYear="1792" box="[783,1280,1080,1106]" class="Insecta" family="Dytiscidae" genus="Hydroglyphus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="geminus">
|
||
<emphasis id="B95546565178FFAEFCFAD13650032D11" box="[783,1060,1080,1105]" italics="true" pageId="5" pageNumber="6">Hydroglyphus geminus</emphasis>
|
||
(Fabricius, 1792)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C21E1C75178FFAEFAF9D13655FB2D38" authority="Mannerheim, 1853" authorityName="Mannerheim" authorityYear="1853" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="nigellus">
|
||
<emphasis id="B95546565178FFAEFAF9D13654D32D38" italics="true" pageId="5" pageNumber="6">Hydroporus nigellus</emphasis>
|
||
Mannerheim, 1853
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C21E1C75178FFAEFDEFD15150252D38" authority="Gyllenhal, 1808" authorityName="Gyllenhal" authorityYear="1808" box="[538,1026,1118,1144]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="pubescens">
|
||
<emphasis id="B95546565178FFAEFDEFD15157012D38" box="[538,806,1119,1144]" italics="true" pageId="5" pageNumber="6">Hydroporus pubescens</emphasis>
|
||
(Gyllenhal, 1808)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33BC9CF5178FFAEFF33D18B50232D86" pageId="5" pageNumber="6" type="etymology">
|
||
<paragraph id="8B9E9A445178FFAEFF33D18B50232D86" blockId="5.[151,1437,152,1222]" pageId="5" pageNumber="6">
|
||
<emphasis id="B95546565178FFAEFF33D18B558B2DDF" bold="true" box="[198,428,1157,1183]" pageId="5" pageNumber="6">Derivatio nominis</emphasis>
|
||
: The specific name
|
||
<taxonomicName id="4C21E1C75178FFAEFD45D18B571C2DDE" box="[688,827,1157,1182]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="cagrankaya">
|
||
<emphasis id="B95546565178FFAEFD45D18B571C2DDE" box="[688,827,1157,1182]" italics="true" pageId="5" pageNumber="6">cagrankaya</emphasis>
|
||
</taxonomicName>
|
||
is a noun in apposition and is derived from the
|
||
<taxonomicName id="4C21E1C75178FFAEFF62D1A455042D84" box="[151,291,1194,1220]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="11" phylum="Arthropoda" rank="species" species="cagrankaya" status="sp. nov.">Çaġrankaya</taxonomicName>
|
||
plateau, which is close to the locus typicus of the new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |