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<mods:title id="BB9699728BE1788CA6B778D9050E565C">Two new species of Aegopsis Burmeister, 1847 (Coleoptera: Scarabaeidae: Dynastinae) from the central Brazilian Cerrado</mods:title>
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<mods:namePart id="B4C2D98C84B1E782333C2C2DBF459BF3">De Morais, José W.</mods:namePart>
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<treatment id="03FC87E9FF95062AFF4CCCC8FE97C07C" ID-DOI="http://doi.org/10.5281/zenodo.5976897" ID-GBIF-Taxon="154507623" ID-Zenodo-Dep="5976897" LSID="urn:lsid:plazi:treatment:03FC87E9FF95062AFF4CCCC8FE97C07C" httpUri="http://treatment.plazi.org/id/03FC87E9FF95062AFF4CCCC8FE97C07C" lastPageId="18" lastPageNumber="193" pageId="17" pageNumber="192">
<subSubSection id="C34F6574FF950629FF4CCCC8F9FBC5C5" pageId="17" pageNumber="192" type="nomenclature">
<paragraph id="8BEA36FFFF950629FF4CCCC8F9BEC5A4" blockId="17.[151,752,1159,1219]" box="[151,693,1159,1186]" pageId="17" pageNumber="192">
<heading id="D0A28193FF950629FF4CCCC8F9BEC5A4" bold="true" box="[151,693,1159,1186]" fontSize="11" level="1" pageId="17" pageNumber="192" reason="1">
<emphasis id="B921EAEDFF950629FF4CCCC8F9BEC5A4" bold="true" box="[151,693,1159,1186]" pageId="17" pageNumber="192">
Key to species of
<taxonomicName id="4C554D7CFF950629FEB6CCC7F9BEC5A4" ID-CoL="PKT" authority="Burmeister, 1847" authorityName="Burmeister" authorityYear="1847" box="[365,693,1159,1186]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="192" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAEDFF950629FEB6CCC7FAD3C5A7" bold="true" box="[365,472,1159,1185]" italics="true" pageId="17" pageNumber="192">Aegopsis</emphasis>
Burmeister, 1847
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA36FFFF950629FF4CCCEAF9FBC5C5" blockId="17.[151,752,1159,1219]" box="[151,752,1194,1219]" pageId="17" pageNumber="192">
(Male key modified from
<bibRefCitation id="EFC44B0EFF950629FE61CCEAF9ECC5C5" author="Dechambre, R. - P. &amp; Grossi, E. J." box="[442,743,1194,1219]" pageId="17" pageNumber="192" pagination="237 - 240" refId="ref8835" refString="Dechambre, R. - P. &amp; Grossi, E. J. (1990) Agaocephala bolbocerida Thomson: nouvelles captures et mise au point systematique. Bulletin de la Societe entomologique de France, 95, 237 - 240." type="journal article" year="1990">Dechambre &amp; Grossi 1990</bibRefCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34F6574FF95062AFF4CCCB1FE97C07C" lastPageId="18" lastPageNumber="193" pageId="17" pageNumber="192" type="key">
<keyStep id="30A12E5AFF950629FF4CCCB1FE90C422" pageId="17" pageNumber="192">
<paragraph id="8BEA36FFFF950629FF4CCCB1FE90C401" blockId="17.[151,1436,1265,2021]" box="[151,1435,1265,1287]" pageId="17" pageNumber="192">
<keyLead id="30A495CAFF950629FF4CCCB1FE90C401" box="[151,1435,1265,1287]" pageId="17" pageNumber="192">
1. Head and pronotum with horns, pronotal horn reduced in minor specimens (
<emphasis id="B921EAEDFF950629FC61CCB2F8D6C401" box="[954,989,1266,1287]" italics="true" pageId="17" pageNumber="192">e.g.</emphasis>
,
<figureCitation id="136E2A7AFF950629FC32CCB1FF2BC401" box="[1001,1056,1265,1287]" captionStart="FIGURE 1" captionStartId="3.[151,250,1977,1999]" captionTargetBox="[184,1388,203,1935]" captionTargetId="figure@3.[184,1388,203,1935]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Aegopsis bolboceridus. A, Major male in dorsal view; B, minor male in dorsal view; C, female in dorsal view. Scale bars: 3 mm." httpUri="https://zenodo.org/record/2611532/files/figure.png" pageId="17" pageNumber="192">Figs 1</figureCitation>
AB, 9AB, 10AB) (males)..... 2
</keyLead>
</paragraph>
<paragraph id="8BEA36FFFF950629FF4CCD4FFE90C422" blockId="17.[151,1436,1265,2021]" box="[151,1435,1295,1317]" pageId="17" pageNumber="192">
<keyLead id="30A495CAFF950629FF4CCD4FFE90C422" box="[151,1435,1295,1317]" pageId="17" pageNumber="192">
- Head and pronotum without horns (
<emphasis id="B921EAEDFF950629FDE3CD4FF950C422" box="[568,603,1295,1316]" italics="true" pageId="17" pageNumber="192">e.g.</emphasis>
,
<figureCitation id="136E2A7AFF950629FDBDCD4FF9B8C423" box="[614,691,1295,1317]" captionStart="FIGURE 1" captionStartId="3.[151,250,1977,1999]" captionTargetBox="[184,1388,203,1935]" captionTargetId="figure@3.[184,1388,203,1935]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Aegopsis bolboceridus. A, Major male in dorsal view; B, minor male in dorsal view; C, female in dorsal view. Scale bars: 3 mm." httpUri="https://zenodo.org/record/2611532/files/figure.png" pageId="17" pageNumber="192">Figs 1C</figureCitation>
,
<figureCitation id="136E2A7AFF950629FD66CD4FF9D7C423" box="[701,732,1295,1317]" captionStart="FIGURE 9" captionStartId="12.[151,250,1976,1998]" captionTargetBox="[185,1375,193,1943]" captionTargetId="figure@12.[185,1375,193,1943]" captionTargetPageId="12" captionText="FIGURE 9. Habitus of Aegopsis diceratops. A, Major male in dorsal view; B, minor male in dorsal view; C, female in dorsal view. Scale bars: 3 mm." httpUri="https://zenodo.org/record/2611550/files/figure.png" pageId="17" pageNumber="192">9C</figureCitation>
,
<figureCitation id="136E2A7AFF950629FD3ECD4FF805C423" box="[741,782,1295,1317]" captionStart="FIGURE 10" captionStartId="13.[151,250,1970,1992]" captionTargetBox="[194,1312,193,1920]" captionTargetId="figure@13.[194,1388,193,1949]" captionTargetPageId="13" captionText="FIGURE 10. Habitus of Aegopsis vazdemelloi. A, Major male in dorsal view; B, minor male in dorsal view; C, female in dorsal view. Scale bars: 3 mm." httpUri="https://zenodo.org/record/2611553/files/figure.png" pageId="17" pageNumber="192">10C</figureCitation>
) (females)............................................ 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A12E5AFF950629FF4CCD6CFE90C47B" pageId="17" pageNumber="192">
<paragraph id="8BEA36FFFF950629FF4CCD6CFE97C459" blockId="17.[151,1436,1265,2021]" pageId="17" pageNumber="192">
<keyLead id="30A495CAFF950629FF4CCD6CFE97C459" pageId="17" pageNumber="192">
2. Protibia with four external teeth, basal tooth sometimes reduced. Pronotum with disc strongly convex in major males.................................................................................
<taxonomicName id="4C554D7CFF950629FB87CD0AFE97C459" ID-CoL="6596P" authority="Burmeister, 1847" authorityName="Burmeister" authorityYear="1847" box="[1116,1436,1354,1376]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="192" phylum="Arthropoda" rank="species" species="curvicornis">
<emphasis id="B921EAEDFF950629FB87CD0AFE97C459" bold="true" box="[1116,1436,1354,1376]" pageId="17" pageNumber="192">
<emphasis id="B921EAEDFF950629FB87CD0AFFE3C466" bold="true" box="[1116,1256,1354,1376]" italics="true" pageId="17" pageNumber="192">A. curvicornis</emphasis>
Burmeister, 1847
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEA36FFFF950629FF4CCD27FE90C47B" blockId="17.[151,1436,1265,2021]" box="[151,1435,1383,1405]" pageId="17" pageNumber="192">
<keyLead id="30A495CAFF950629FF4CCD27FE90C47B" box="[151,1435,1383,1405]" pageId="17" pageNumber="192">- Protibia with three external teeth. Pronotum with disc slightly convex............................................ 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A12E5AFF950629FF4CCDC4FE90C4D3" pageId="17" pageNumber="192">
<paragraph id="8BEA36FFFF950629FF4CCDC4FE97C4B1" blockId="17.[151,1436,1265,2021]" pageId="17" pageNumber="192">
<keyLead id="30A495CAFF950629FF4CCDC4FE97C4B1" pageId="17" pageNumber="192">
3. Base of cephalic horns connected by a curved carina. Lateral carina of parameres not reaching apex of parameres in lateral view.........................................................................
<taxonomicName id="4C554D7CFF950629FB56CDE2FE97C4B1" ID-CoL="64X74" authority="Arrow, 1941" authorityName="Arrow" authorityYear="1941" box="[1165,1436,1442,1464]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="192" phylum="Arthropoda" rank="species" species="peruvianus">
<emphasis id="B921EAEDFF950629FB56CDE2FE97C4B1" bold="true" box="[1165,1436,1442,1464]" pageId="17" pageNumber="192">
<emphasis id="B921EAEDFF950629FB56CDE2FE13C4BE" bold="true" box="[1165,1304,1442,1464]" italics="true" pageId="17" pageNumber="192">A. peruvianus</emphasis>
Arrow, 1941
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEA36FFFF950629FF4CCDFFFE90C4D3" blockId="17.[151,1436,1265,2021]" box="[151,1435,1471,1493]" pageId="17" pageNumber="192">
<keyLead id="30A495CAFF950629FF4CCDFFFE90C4D3" box="[151,1435,1471,1493]" pageId="17" pageNumber="192">- Base of cephalic horns with no connecting carina. Lateral carina of parameres reaching apex of parameres in lateral view.. 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A12E5AFF950629FF4CCD9CFE90C74C" pageId="17" pageNumber="192">
<paragraph id="8BEA36FFFF950629FF4CCD9CFE90C709" blockId="17.[151,1436,1265,2021]" pageId="17" pageNumber="192">
<keyLead id="30A495CAFF950629FF4CCD9CFE90C709" pageId="17" pageNumber="192">
4. Apex of parameres less than one time longer than base of parameres (
<figureCitation id="136E2A7AFF950629FCA7CD9CF8DDC4F4" box="[892,982,1500,1522]" captionStart="FIGURE 12" captionStartId="15.[151,250,901,923]" captionTargetBox="[172,1416,193,879]" captionTargetId="figure@15.[172,1416,193,879]" captionTargetPageId="15" captionText="FIGURE 12. A, Parameres of Aegopsis chaminadei in dorsal view; B, mentum of Aegopsis chaminadei in ventral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611557/files/figure.png" pageId="17" pageNumber="192">Fig. 12A</figureCitation>
). Mentum round (
<figureCitation id="136E2A7AFF950629FB59CD9CFFDCC4F4" box="[1154,1239,1500,1522]" captionStart="FIGURE 12" captionStartId="15.[151,250,901,923]" captionTargetBox="[172,1416,193,879]" captionTargetId="figure@15.[172,1416,193,879]" captionTargetPageId="15" captionText="FIGURE 12. A, Parameres of Aegopsis chaminadei in dorsal view; B, mentum of Aegopsis chaminadei in ventral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611557/files/figure.png" pageId="17" pageNumber="192">Fig. 12B</figureCitation>
)........................................................................................
<taxonomicName id="4C554D7CFF950629FB8FCDBAFE90C709" ID-CoL="64X6P" authority="Dechambre, 1999" authorityName="Dechambre" authorityYear="1999" box="[1108,1435,1530,1552]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="192" phylum="Arthropoda" rank="species" species="chaminadei">
<emphasis id="B921EAEDFF950629FB8FCDBAFE90C709" bold="true" box="[1108,1435,1530,1552]" pageId="17" pageNumber="192">
<emphasis id="B921EAEDFF950629FB8FCDBAFFEFC716" bold="true" box="[1108,1252,1530,1552]" italics="true" pageId="17" pageNumber="192">A. chaminadei</emphasis>
<bibRefCitation id="EFC44B0EFF950629FB32CDBAFE90C709" author="Dechambre, R. - P." box="[1257,1435,1530,1551]" pageId="17" pageNumber="192" pagination="173 - 174" refId="ref8804" refString="Dechambre, R. - P. (1999) Une nouvelle espece d' Aegopsis Burmeister, 1847. Revue Francaise d'Entomologie, Nouvelle-Serie, 21, 173 - 174." type="journal article" year="1999">Dechambre, 1999</bibRefCitation>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEA36FFFF950629FF4CCE57FE90C74C" blockId="17.[151,1436,1265,2021]" pageId="17" pageNumber="192">
<keyLead id="30A495CAFF950629FF4CCE57FE90C74C" pageId="17" pageNumber="192">
- Apex of parameres about two times longer than base of parameres (
<figureCitation id="136E2A7AFF950629FC5ACE57F8F5C72A" box="[897,1022,1559,1581]" captionStart="FIGURE 6" captionStartId="9.[151,250,1775,1797]" captionTargetBox="[152,1421,205,1743]" captionTargetId="figure@9.[151,1436,193,1754]" captionTargetPageId="9" captionText="FIGURE 6. A, Aedeagus of Aegopsis bolboceridus, in lateral view; B, parameres of Aegopsis bolboceridus, in ventral view; C, parameres of Aegopsis bolboceridus, in dorsal view; D, aedeagus of Aegopsis diceratops in lateral view; E, parameres of Aegopsis diceratops in ventral view; F, parameres of Aegopsis diceratops in dorsal view; G, aedeagus of Aegopsis vazdemelloi in lateral view; H, parameres of Aegopsis vazdemelloi in ventral view; I, parameres of Aegopsis vazdemelloi in dorsal view. White arrows in the left indicating ventral carina of parameres in B, E, H; white arrows upward indicating inferior edge of parameres in B, E, H. Scale bars: A, D, G, 1 mm; B, C, E, F, H, I, 0.5 mm." httpUri="https://zenodo.org/record/2611544/files/figure.png" pageId="17" pageNumber="192">Fig. 6C, F, I</figureCitation>
). Mentum subrectangular or subtriangular (
<figureCitation id="136E2A7AFF950629FF33CE74FA6CC74C" box="[232,359,1588,1610]" captionStart="FIGURE 5" captionStartId="7.[151,250,1292,1314]" captionTargetBox="[151,1436,193,1270]" captionTargetId="figure@7.[151,1436,193,1270]" captionTargetPageId="7" captionText="FIGURE 5. A, Mandible of Aegopsis bolboceridus male, in ventral view; B, mandible of Aegopsis diceratops male, in ventral view; C, mandible of Aegopsis vazdemelloi male, in ventral view; D, mentum of Aegopsis bolboceridus male, in ventral view; E, mentum of Aegopsis diceratops male, in ventral view; F, mentum of Aegopsis vazdemelloi male, in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; Arrows indicating the internal lobe in B, C; arrow indicating the mentum carina in E. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611541/files/figure.png" pageId="17" pageNumber="192">Fig. 5D, E, F</figureCitation>
)....................................................................................... 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A12E5AFF950629FF4CCE11FE90C7B9" pageId="17" pageNumber="192">
<paragraph id="8BEA36FFFF950629FF4CCE11FE90C782" blockId="17.[151,1436,1265,2021]" pageId="17" pageNumber="192">
<keyLead id="30A495CAFF950629FF4CCE11FE90C782" pageId="17" pageNumber="192">
5. Clypeus with round apex (
<figureCitation id="136E2A7AFF950629FE07CE11F923C761" box="[476,552,1617,1639]" captionStart="FIGURE 2" captionStartId="4.[151,250,1882,1904]" captionTargetBox="[200,1377,193,1853]" captionTargetId="figure@4.[200,1377,193,1853]" captionTargetPageId="4" captionText="FIGURE 2. A, Head of Aegopsis bolboceridus male, in dorsal view; B, pronotum of Aegopsis bolboceridus male, in dorsal view; C, head of Aegopsis diceratops male, in dorsal view; D, pronotum of Aegopsis diceratops male, in dorsal view; E, head of Aegopsis vazdemelloi male, in dorsal view; F, pronotum of Aegopsis vazdemelloi male, in dorsal view. Black arrow indicating emarginate apex of clypeus in A, E; white arrow indicating micropunctures on pronotum in F. Scale bars: A, C, F, 2 mm; B, D, E, 3 mm." httpUri="https://zenodo.org/record/2611534/files/figure.png" pageId="17" pageNumber="192">Fig. 2C</figureCitation>
). Ocular canthi protruding anteriorly (
<figureCitation id="136E2A7AFF950629FC4FCE11F8D4C761" box="[916,991,1617,1639]" captionStart="FIGURE 3" captionStartId="5.[151,250,1876,1898]" captionTargetBox="[231,1354,193,1844]" captionTargetId="figure@5.[231,1354,193,1844]" captionTargetPageId="5" captionText="FIGURE 3. A, Canthus of Aegopsis bolboceridus male, in dorsal view; B, canthus of Aegopsis bolboceridus female, in dorsal view; C, canthus of Aegopsis diceratops male, in dorsal view; D, canthus of Aegopsis diceratops female, in dorsal view; E, canthus of Aegopsis vazdemelloi male, in dorsal view; F, canthus of Aegopsis vazdemelloi female, in dorsal view. White arrow indicating apex inconspicuous of canthi carina in A; black arrows indicating canthi carina split in C; black arrow indicating apex conspicuous of canthi carina in E; black arrows indicating external edge of canthi in B, D, F. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611536/files/figure.png" pageId="17" pageNumber="192">Fig. 3C</figureCitation>
). Apical corners of mentum carinate (
<figureCitation id="136E2A7AFF950629FA9CCE11FE86C761" box="[1351,1421,1617,1639]" captionStart="FIGURE 5" captionStartId="7.[151,250,1292,1314]" captionTargetBox="[151,1436,193,1270]" captionTargetId="figure@7.[151,1436,193,1270]" captionTargetPageId="7" captionText="FIGURE 5. A, Mandible of Aegopsis bolboceridus male, in ventral view; B, mandible of Aegopsis diceratops male, in ventral view; C, mandible of Aegopsis vazdemelloi male, in ventral view; D, mentum of Aegopsis bolboceridus male, in ventral view; E, mentum of Aegopsis diceratops male, in ventral view; F, mentum of Aegopsis vazdemelloi male, in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; Arrows indicating the internal lobe in B, C; arrow indicating the mentum carina in E. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611541/files/figure.png" pageId="17" pageNumber="192">Fig. 5E</figureCitation>
).................................................................
<emphasis id="B921EAEDFF950629FC37CE2FFE90C782" bold="true" box="[1004,1435,1647,1669]" pageId="17" pageNumber="192">
<taxonomicName id="4C554D7CFF950629FC37CE2FFE17C782" ID-CoL="64WV4" authority="Sobral &amp; Grossi" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[1004,1308,1647,1669]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="192" phylum="Arthropoda" rank="species" species="diceratops" status="sp. nov.">
<emphasis id="B921EAEDFF950629FC37CE2FFF66C783" bold="true" box="[1004,1133,1647,1669]" italics="true" pageId="17" pageNumber="192">A. diceratops</emphasis>
Sobral &amp; Grossi
</taxonomicName>
,
<taxonomicNameLabel id="A2125796FF950629FAFDCE2FFE90C782" box="[1318,1435,1647,1668]" pageId="17" pageNumber="192" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BEA36FFFF950629FF4CCECDFE90C7B9" blockId="17.[151,1436,1265,2021]" pageId="17" pageNumber="192">
<keyLead id="30A495CAFF950629FF4CCECDFE90C7B9" pageId="17" pageNumber="192">
- Clypeus apex truncate or slightly emarginate (
<figureCitation id="136E2A7AFF950629FD7ACECCF800C7A4" box="[673,779,1676,1698]" captionStart="FIGURE 2" captionStartId="4.[151,250,1882,1904]" captionTargetBox="[200,1377,193,1853]" captionTargetId="figure@4.[200,1377,193,1853]" captionTargetPageId="4" captionText="FIGURE 2. A, Head of Aegopsis bolboceridus male, in dorsal view; B, pronotum of Aegopsis bolboceridus male, in dorsal view; C, head of Aegopsis diceratops male, in dorsal view; D, pronotum of Aegopsis diceratops male, in dorsal view; E, head of Aegopsis vazdemelloi male, in dorsal view; F, pronotum of Aegopsis vazdemelloi male, in dorsal view. Black arrow indicating emarginate apex of clypeus in A, E; white arrow indicating micropunctures on pronotum in F. Scale bars: A, C, F, 2 mm; B, D, E, 3 mm." httpUri="https://zenodo.org/record/2611534/files/figure.png" pageId="17" pageNumber="192">Fig. 2A, E</figureCitation>
). Ocular canthi not protruding (
<figureCitation id="136E2A7AFF950629FBE4CECCFFA1C7A4" box="[1087,1194,1676,1698]" captionStart="FIGURE 3" captionStartId="5.[151,250,1876,1898]" captionTargetBox="[231,1354,193,1844]" captionTargetId="figure@5.[231,1354,193,1844]" captionTargetPageId="5" captionText="FIGURE 3. A, Canthus of Aegopsis bolboceridus male, in dorsal view; B, canthus of Aegopsis bolboceridus female, in dorsal view; C, canthus of Aegopsis diceratops male, in dorsal view; D, canthus of Aegopsis diceratops female, in dorsal view; E, canthus of Aegopsis vazdemelloi male, in dorsal view; F, canthus of Aegopsis vazdemelloi female, in dorsal view. White arrow indicating apex inconspicuous of canthi carina in A; black arrows indicating canthi carina split in C; black arrow indicating apex conspicuous of canthi carina in E; black arrows indicating external edge of canthi in B, D, F. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611536/files/figure.png" pageId="17" pageNumber="192">Fig. 3A, E</figureCitation>
). Mentum lacking carina (
<figureCitation id="136E2A7AFF950629FF33CEE9FA47C7B9" box="[232,332,1705,1727]" captionStart="FIGURE 5" captionStartId="7.[151,250,1292,1314]" captionTargetBox="[151,1436,193,1270]" captionTargetId="figure@7.[151,1436,193,1270]" captionTargetPageId="7" captionText="FIGURE 5. A, Mandible of Aegopsis bolboceridus male, in ventral view; B, mandible of Aegopsis diceratops male, in ventral view; C, mandible of Aegopsis vazdemelloi male, in ventral view; D, mentum of Aegopsis bolboceridus male, in ventral view; E, mentum of Aegopsis diceratops male, in ventral view; F, mentum of Aegopsis vazdemelloi male, in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; Arrows indicating the internal lobe in B, C; arrow indicating the mentum carina in E. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611541/files/figure.png" pageId="17" pageNumber="192">Fig. 5D, F</figureCitation>
).......................................................................................... 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A12E5AFF950629FF4CCE87FE90C669" pageId="17" pageNumber="192">
<paragraph id="8BEA36FFFF950629FF4CCE87FE97C611" blockId="17.[151,1436,1265,2021]" pageId="17" pageNumber="192">
<keyLead id="30A495CAFF950629FF4CCE87FE97C611" pageId="17" pageNumber="192">
6. Punctures on posterior angles of pronotum with no micropunctures between them (
<figureCitation id="136E2A7AFF950629FBDDCE87FF59C7DB" box="[1030,1106,1735,1757]" captionStart="FIGURE 2" captionStartId="4.[151,250,1882,1904]" captionTargetBox="[200,1377,193,1853]" captionTargetId="figure@4.[200,1377,193,1853]" captionTargetPageId="4" captionText="FIGURE 2. A, Head of Aegopsis bolboceridus male, in dorsal view; B, pronotum of Aegopsis bolboceridus male, in dorsal view; C, head of Aegopsis diceratops male, in dorsal view; D, pronotum of Aegopsis diceratops male, in dorsal view; E, head of Aegopsis vazdemelloi male, in dorsal view; F, pronotum of Aegopsis vazdemelloi male, in dorsal view. Black arrow indicating emarginate apex of clypeus in A, E; white arrow indicating micropunctures on pronotum in F. Scale bars: A, C, F, 2 mm; B, D, E, 3 mm." httpUri="https://zenodo.org/record/2611534/files/figure.png" pageId="17" pageNumber="192">Fig. 2B</figureCitation>
). Mandibles with apical protuberance near external carina (
<figureCitation id="136E2A7AFF950629FE06CEA4F920C7FC" box="[477,555,1764,1786]" captionStart="FIGURE 5" captionStartId="7.[151,250,1292,1314]" captionTargetBox="[151,1436,193,1270]" captionTargetId="figure@7.[151,1436,193,1270]" captionTargetPageId="7" captionText="FIGURE 5. A, Mandible of Aegopsis bolboceridus male, in ventral view; B, mandible of Aegopsis diceratops male, in ventral view; C, mandible of Aegopsis vazdemelloi male, in ventral view; D, mentum of Aegopsis bolboceridus male, in ventral view; E, mentum of Aegopsis diceratops male, in ventral view; F, mentum of Aegopsis vazdemelloi male, in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; Arrows indicating the internal lobe in B, C; arrow indicating the mentum carina in E. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611541/files/figure.png" pageId="17" pageNumber="192">Fig. 5A</figureCitation>
). Mentum subrectangular (
<figureCitation id="136E2A7AFF950629FCF5CEA4F870C7FC" box="[814,891,1764,1786]" captionStart="FIGURE 5" captionStartId="7.[151,250,1292,1314]" captionTargetBox="[151,1436,193,1270]" captionTargetId="figure@7.[151,1436,193,1270]" captionTargetPageId="7" captionText="FIGURE 5. A, Mandible of Aegopsis bolboceridus male, in ventral view; B, mandible of Aegopsis diceratops male, in ventral view; C, mandible of Aegopsis vazdemelloi male, in ventral view; D, mentum of Aegopsis bolboceridus male, in ventral view; E, mentum of Aegopsis diceratops male, in ventral view; F, mentum of Aegopsis vazdemelloi male, in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; Arrows indicating the internal lobe in B, C; arrow indicating the mentum carina in E. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611541/files/figure.png" pageId="17" pageNumber="192">Fig. 5D</figureCitation>
). Parameres, in lateral view, narrower than apex of phallobase (
<figureCitation id="136E2A7AFF950629FEF0CF41FA7CC611" box="[299,375,1793,1815]" captionStart="FIGURE 6" captionStartId="9.[151,250,1775,1797]" captionTargetBox="[152,1421,205,1743]" captionTargetId="figure@9.[151,1436,193,1754]" captionTargetPageId="9" captionText="FIGURE 6. A, Aedeagus of Aegopsis bolboceridus, in lateral view; B, parameres of Aegopsis bolboceridus, in ventral view; C, parameres of Aegopsis bolboceridus, in dorsal view; D, aedeagus of Aegopsis diceratops in lateral view; E, parameres of Aegopsis diceratops in ventral view; F, parameres of Aegopsis diceratops in dorsal view; G, aedeagus of Aegopsis vazdemelloi in lateral view; H, parameres of Aegopsis vazdemelloi in ventral view; I, parameres of Aegopsis vazdemelloi in dorsal view. White arrows in the left indicating ventral carina of parameres in B, E, H; white arrows upward indicating inferior edge of parameres in B, E, H. Scale bars: A, D, G, 1 mm; B, C, E, F, H, I, 0.5 mm." httpUri="https://zenodo.org/record/2611544/files/figure.png" pageId="17" pageNumber="192">Fig. 6A</figureCitation>
)...........................................................
<taxonomicName id="4C554D7CFF950629FB94CF42FE97C611" ID-CoL="65972" authority="(Thomson, 1860)" baseAuthorityName="Thomson" baseAuthorityYear="1860" box="[1103,1436,1794,1816]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="192" phylum="Arthropoda" rank="species" species="bolboceridus">
<emphasis id="B921EAEDFF950629FB94CF42FE97C611" bold="true" box="[1103,1436,1794,1816]" pageId="17" pageNumber="192">
<emphasis id="B921EAEDFF950629FB94CF42FFE2C61E" bold="true" box="[1103,1257,1794,1816]" italics="true" pageId="17" pageNumber="192">A. bolboceridus</emphasis>
(
<bibRefCitation id="EFC44B0EFF950629FB2DCF42FE9EC611" author="Thomson, J." box="[1270,1429,1794,1815]" pageId="17" pageNumber="192" refId="ref10005" refString="Thomson, J. (1860) Musee Scientifique ou recueil d'histoire naturelle. Societe Entomologique de France, Paris, 96 pp." type="book" year="1860">Thomson, 1860</bibRefCitation>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEA36FFFF950629FF4CCF5FFE90C669" blockId="17.[151,1436,1265,2021]" pageId="17" pageNumber="192">
<keyLead id="30A495CAFF950629FF4CCF5FFE90C669" pageId="17" pageNumber="192">
- Punctures on posterior angles of pronotum with micropunctures between them (
<figureCitation id="136E2A7AFF950629FC32CF5FFF3FC633" box="[1001,1076,1823,1845]" captionStart="FIGURE 2" captionStartId="4.[151,250,1882,1904]" captionTargetBox="[200,1377,193,1853]" captionTargetId="figure@4.[200,1377,193,1853]" captionTargetPageId="4" captionText="FIGURE 2. A, Head of Aegopsis bolboceridus male, in dorsal view; B, pronotum of Aegopsis bolboceridus male, in dorsal view; C, head of Aegopsis diceratops male, in dorsal view; D, pronotum of Aegopsis diceratops male, in dorsal view; E, head of Aegopsis vazdemelloi male, in dorsal view; F, pronotum of Aegopsis vazdemelloi male, in dorsal view. Black arrow indicating emarginate apex of clypeus in A, E; white arrow indicating micropunctures on pronotum in F. Scale bars: A, C, F, 2 mm; B, D, E, 3 mm." httpUri="https://zenodo.org/record/2611534/files/figure.png" pageId="17" pageNumber="192">Fig. 2F</figureCitation>
). Mandibles with no apical protuberance (
<figureCitation id="136E2A7AFF950629FEC1CF7CFA6EC654" box="[282,357,1852,1874]" captionStart="FIGURE 5" captionStartId="7.[151,250,1292,1314]" captionTargetBox="[151,1436,193,1270]" captionTargetId="figure@7.[151,1436,193,1270]" captionTargetPageId="7" captionText="FIGURE 5. A, Mandible of Aegopsis bolboceridus male, in ventral view; B, mandible of Aegopsis diceratops male, in ventral view; C, mandible of Aegopsis vazdemelloi male, in ventral view; D, mentum of Aegopsis bolboceridus male, in ventral view; E, mentum of Aegopsis diceratops male, in ventral view; F, mentum of Aegopsis vazdemelloi male, in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; Arrows indicating the internal lobe in B, C; arrow indicating the mentum carina in E. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611541/files/figure.png" pageId="17" pageNumber="192">Fig. 5C</figureCitation>
). Mentum subtriangular (
<figureCitation id="136E2A7AFF950629FD82CF7CF9A9C654" box="[601,674,1852,1874]" captionStart="FIGURE 5" captionStartId="7.[151,250,1292,1314]" captionTargetBox="[151,1436,193,1270]" captionTargetId="figure@7.[151,1436,193,1270]" captionTargetPageId="7" captionText="FIGURE 5. A, Mandible of Aegopsis bolboceridus male, in ventral view; B, mandible of Aegopsis diceratops male, in ventral view; C, mandible of Aegopsis vazdemelloi male, in ventral view; D, mentum of Aegopsis bolboceridus male, in ventral view; E, mentum of Aegopsis diceratops male, in ventral view; F, mentum of Aegopsis vazdemelloi male, in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; Arrows indicating the internal lobe in B, C; arrow indicating the mentum carina in E. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611541/files/figure.png" pageId="17" pageNumber="192">Fig. 5F</figureCitation>
). Parameres, in lateral view, almost as thick as apex of phallobase (
<figureCitation id="136E2A7AFF950629FAFCCF7CFE78C654" box="[1319,1395,1852,1874]" captionStart="FIGURE 6" captionStartId="9.[151,250,1775,1797]" captionTargetBox="[152,1421,205,1743]" captionTargetId="figure@9.[151,1436,193,1754]" captionTargetPageId="9" captionText="FIGURE 6. A, Aedeagus of Aegopsis bolboceridus, in lateral view; B, parameres of Aegopsis bolboceridus, in ventral view; C, parameres of Aegopsis bolboceridus, in dorsal view; D, aedeagus of Aegopsis diceratops in lateral view; E, parameres of Aegopsis diceratops in ventral view; F, parameres of Aegopsis diceratops in dorsal view; G, aedeagus of Aegopsis vazdemelloi in lateral view; H, parameres of Aegopsis vazdemelloi in ventral view; I, parameres of Aegopsis vazdemelloi in dorsal view. White arrows in the left indicating ventral carina of parameres in B, E, H; white arrows upward indicating inferior edge of parameres in B, E, H. Scale bars: A, D, G, 1 mm; B, C, E, F, H, I, 0.5 mm." httpUri="https://zenodo.org/record/2611544/files/figure.png" pageId="17" pageNumber="192">Fig. 6G</figureCitation>
)................................................................
<emphasis id="B921EAEDFF950629FC05CF1AFE90C669" bold="true" box="[990,1435,1882,1904]" pageId="17" pageNumber="192">
<taxonomicName id="4C554D7CFF950629FC05CF1AFE17C669" ID-CoL="65974" authority="Sobral &amp; Grossi" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[990,1308,1882,1904]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="192" phylum="Arthropoda" rank="species" species="vazdemelloi" status="sp. nov.">
<emphasis id="B921EAEDFF950629FC05CF1AFF65C676" bold="true" box="[990,1134,1882,1904]" italics="true" pageId="17" pageNumber="192">A. vazdemelloi</emphasis>
Sobral &amp; Grossi
</taxonomicName>
,
<taxonomicNameLabel id="A2125796FF950629FAFDCF1AFE90C669" box="[1318,1435,1882,1903]" pageId="17" pageNumber="192" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A12E5AFF950629FF4CCF37FE90C6AC" pageId="17" pageNumber="192">
<paragraph id="8BEA36FFFF950629FF4CCF37FE97C68A" blockId="17.[151,1436,1265,2021]" box="[151,1436,1911,1933]" pageId="17" pageNumber="192">
<keyLead id="30A495CAFF950629FF4CCF37FE97C68A" box="[151,1436,1911,1933]" pageId="17" pageNumber="192">
<emphasis id="B921EAEDFF950629FF4CCF37FBA8C68A" bold="true" box="[151,163,1911,1932]" pageId="17" pageNumber="192">7</emphasis>
. Protibia with four external teeth, basal tooth sometimes reduced......................
<taxonomicName id="4C554D7CFF950629FB87CF37FE97C68A" ID-CoL="6596P" authority="Burmeister, 1847" authorityName="Burmeister" authorityYear="1847" box="[1116,1436,1911,1933]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="192" phylum="Arthropoda" rank="species" species="curvicornis">
<emphasis id="B921EAEDFF950629FB87CF37FE97C68A" bold="true" box="[1116,1436,1911,1933]" pageId="17" pageNumber="192">
<emphasis id="B921EAEDFF950629FB87CF37FFE3C68B" bold="true" box="[1116,1256,1911,1933]" italics="true" pageId="17" pageNumber="192">A. curvicornis</emphasis>
Burmeister, 1847
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEA36FFFF950629FF4CCFD5FE90C6AC" blockId="17.[151,1436,1265,2021]" box="[151,1435,1940,1962]" pageId="17" pageNumber="192">
<keyLead id="30A495CAFF950629FF4CCFD5FE90C6AC" box="[151,1435,1940,1962]" pageId="17" pageNumber="192">- Protibia with three external teeth......................................................................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A12E5AFF950629FF4CCFF2FE90C6E3" pageId="17" pageNumber="192">
<paragraph id="8BEA36FFFF950629FF4CCFF2FE97C6C1" blockId="17.[151,1436,1265,2021]" box="[151,1436,1970,1992]" pageId="17" pageNumber="192">
<keyLead id="30A495CAFF950629FF4CCFF2FE97C6C1" box="[151,1436,1970,1992]" pageId="17" pageNumber="192">
8. Ocular canthi round, canthi apex posteriorly almost indistinct. Galea triangular, apex recurved..
<taxonomicName id="4C554D7CFF950629FB56CFF2FE97C6C1" ID-CoL="64X74" authority="Arrow, 1941" authorityName="Arrow" authorityYear="1941" box="[1165,1436,1970,1992]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="192" phylum="Arthropoda" rank="species" species="peruvianus">
<emphasis id="B921EAEDFF950629FB56CFF2FE97C6C1" bold="true" box="[1165,1436,1970,1992]" pageId="17" pageNumber="192">
<emphasis id="B921EAEDFF950629FB56CFF2FE13C6CE" bold="true" box="[1165,1304,1970,1992]" italics="true" pageId="17" pageNumber="192">A. peruvianus</emphasis>
Arrow, 1941
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEA36FFFF950629FF4CCF8FFE90C6E3" blockId="17.[151,1436,1265,2021]" box="[151,1435,1999,2021]" pageId="17" pageNumber="192">
<keyLead id="30A495CAFF950629FF4CCF8FFE90C6E3" box="[151,1435,1999,2021]" pageId="17" pageNumber="192">- Ocular canthi convex, canthi apex posteriorly distinct. Galea trapezoidal, apex straight.............................. 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A12E5AFF96062AFF4CC8D7FE90C003" pageId="18" pageNumber="193">
<paragraph id="8BEA36FFFF96062AFF4CC8D7FE90C1CC" blockId="18.[151,1437,151,378]" pageId="18" pageNumber="193">
<keyLead id="30A495CAFF96062AFF4CC8D7FE90C1CC" pageId="18" pageNumber="193">
9. Clypeus trapezoidal (
<figureCitation id="136E2A7AFF96062AFE76C8D7FAF2C1AB" box="[429,505,151,173]" captionStart="FIGURE 7" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[185,1389,202,1867]" captionTargetId="figure@10.[181,1389,193,1871]" captionTargetPageId="10" captionText="FIGURE 7. A, Head of Aegopsis bolboceridus female in dorsal view; B, head of Aegopsis bolboceridus female in lateral view; C, head of Aegopsis diceratops female in dorsal view; D, head of Aegopsis diceratops female in lateral view; E, head of Aegopsis vazdemelloi female in dorsal view; F, head of Aegopsis vazdemelloi female in lateral view. Black arrow indicating head punctures in A; black arrow indicating clypeus trapezoidal in C; white arrow indicating vertex in C; black arrow indicating frontal tubercles in D; white arrow indicating head punctures in E. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2611546/files/figure.png" pageId="18" pageNumber="193">Fig. 7C</figureCitation>
). Frons with two strong tubercles (
<figureCitation id="136E2A7AFF96062AFC9DC8D7F89FC1AA" box="[838,916,151,173]" captionStart="FIGURE 7" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[185,1389,202,1867]" captionTargetId="figure@10.[181,1389,193,1871]" captionTargetPageId="10" captionText="FIGURE 7. A, Head of Aegopsis bolboceridus female in dorsal view; B, head of Aegopsis bolboceridus female in lateral view; C, head of Aegopsis diceratops female in dorsal view; D, head of Aegopsis diceratops female in lateral view; E, head of Aegopsis vazdemelloi female in dorsal view; F, head of Aegopsis vazdemelloi female in lateral view. Black arrow indicating head punctures in A; black arrow indicating clypeus trapezoidal in C; white arrow indicating vertex in C; black arrow indicating frontal tubercles in D; white arrow indicating head punctures in E. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2611546/files/figure.png" pageId="18" pageNumber="193">Fig. 7D</figureCitation>
). Punctures absent on disc of vertex (
<figureCitation id="136E2A7AFF96062AFB22C8D7FE48C1AB" box="[1273,1347,151,173]" captionStart="FIGURE 7" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[185,1389,202,1867]" captionTargetId="figure@10.[181,1389,193,1871]" captionTargetPageId="10" captionText="FIGURE 7. A, Head of Aegopsis bolboceridus female in dorsal view; B, head of Aegopsis bolboceridus female in lateral view; C, head of Aegopsis diceratops female in dorsal view; D, head of Aegopsis diceratops female in lateral view; E, head of Aegopsis vazdemelloi female in dorsal view; F, head of Aegopsis vazdemelloi female in lateral view. Black arrow indicating head punctures in A; black arrow indicating clypeus trapezoidal in C; white arrow indicating vertex in C; black arrow indicating frontal tubercles in D; white arrow indicating head punctures in E. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2611546/files/figure.png" pageId="18" pageNumber="193">Fig. 7C</figureCitation>
).......................................................................
<emphasis id="B921EAEDFF96062AFC37C8F4FE90C1CC" bold="true" box="[1004,1435,180,202]" pageId="18" pageNumber="193">
<taxonomicName id="4C554D7CFF96062AFC37C8F4FE17C1CC" ID-CoL="64WV4" authority="Sobral &amp; Grossi" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[1004,1308,180,202]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="193" phylum="Arthropoda" rank="species" species="diceratops" status="sp. nov.">
<emphasis id="B921EAEDFF96062AFC37C8F4FF66C1CC" bold="true" box="[1004,1133,180,202]" italics="true" pageId="18" pageNumber="193">A. diceratops</emphasis>
Sobral &amp; Grossi
</taxonomicName>
,
<taxonomicNameLabel id="A2125796FF96062AFAFDC8F5FE90C1CC" box="[1318,1435,181,202]" pageId="18" pageNumber="193" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BEA36FFFF96062AFF4CC892FE90C003" blockId="18.[151,1437,151,378]" pageId="18" pageNumber="193">
<keyLead id="30A495CAFF96062AFF4CC892FE90C003" pageId="18" pageNumber="193">
- Clypeus round (
<figureCitation id="136E2A7AFF96062AFEA1C891FAE9C1E1" box="[378,482,209,231]" captionStart="FIGURE 7" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[185,1389,202,1867]" captionTargetId="figure@10.[181,1389,193,1871]" captionTargetPageId="10" captionText="FIGURE 7. A, Head of Aegopsis bolboceridus female in dorsal view; B, head of Aegopsis bolboceridus female in lateral view; C, head of Aegopsis diceratops female in dorsal view; D, head of Aegopsis diceratops female in lateral view; E, head of Aegopsis vazdemelloi female in dorsal view; F, head of Aegopsis vazdemelloi female in lateral view. Black arrow indicating head punctures in A; black arrow indicating clypeus trapezoidal in C; white arrow indicating vertex in C; black arrow indicating frontal tubercles in D; white arrow indicating head punctures in E. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2611546/files/figure.png" pageId="18" pageNumber="193">Fig. 7A, E</figureCitation>
). Frons with two inconspicuous tubercles (
<figureCitation id="136E2A7AFF96062AFCA1C891F8D4C1E1" box="[890,991,209,231]" captionStart="FIGURE 7" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[185,1389,202,1867]" captionTargetId="figure@10.[181,1389,193,1871]" captionTargetPageId="10" captionText="FIGURE 7. A, Head of Aegopsis bolboceridus female in dorsal view; B, head of Aegopsis bolboceridus female in lateral view; C, head of Aegopsis diceratops female in dorsal view; D, head of Aegopsis diceratops female in lateral view; E, head of Aegopsis vazdemelloi female in dorsal view; F, head of Aegopsis vazdemelloi female in lateral view. Black arrow indicating head punctures in A; black arrow indicating clypeus trapezoidal in C; white arrow indicating vertex in C; black arrow indicating frontal tubercles in D; white arrow indicating head punctures in E. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2611546/files/figure.png" pageId="18" pageNumber="193">Fig. 7B, F</figureCitation>
). Punctures present on disc of vertex (
<figureCitation id="136E2A7AFF96062AFA97C891FBFAC002" captionStart="FIGURE 7" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[185,1389,202,1867]" captionTargetId="figure@10.[181,1389,193,1871]" captionTargetPageId="10" captionText="FIGURE 7. A, Head of Aegopsis bolboceridus female in dorsal view; B, head of Aegopsis bolboceridus female in lateral view; C, head of Aegopsis diceratops female in dorsal view; D, head of Aegopsis diceratops female in lateral view; E, head of Aegopsis vazdemelloi female in dorsal view; F, head of Aegopsis vazdemelloi female in lateral view. Black arrow indicating head punctures in A; black arrow indicating clypeus trapezoidal in C; white arrow indicating vertex in C; black arrow indicating frontal tubercles in D; white arrow indicating head punctures in E. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2611546/files/figure.png" pageId="18" pageNumber="193">Fig. 7A, E</figureCitation>
)................................................................................................ 10
</keyLead>
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<keyStep id="30A12E5AFF96062AFF4CC94CFE97C07C" pageId="18" pageNumber="193">
<paragraph id="8BEA36FFFF96062AFF4CC94CFE90C039" blockId="18.[151,1437,151,378]" pageId="18" pageNumber="193">
<keyLead id="30A495CAFF96062AFF4CC94CFE90C039" pageId="18" pageNumber="193">
10. Vertex with deeper and coalescent punctures (
<figureCitation id="136E2A7AFF96062AFD4DC94CF9EBC024" box="[662,736,268,290]" captionStart="FIGURE 7" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[185,1389,202,1867]" captionTargetId="figure@10.[181,1389,193,1871]" captionTargetPageId="10" captionText="FIGURE 7. A, Head of Aegopsis bolboceridus female in dorsal view; B, head of Aegopsis bolboceridus female in lateral view; C, head of Aegopsis diceratops female in dorsal view; D, head of Aegopsis diceratops female in lateral view; E, head of Aegopsis vazdemelloi female in dorsal view; F, head of Aegopsis vazdemelloi female in lateral view. Black arrow indicating head punctures in A; black arrow indicating clypeus trapezoidal in C; white arrow indicating vertex in C; black arrow indicating frontal tubercles in D; white arrow indicating head punctures in E. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2611546/files/figure.png" pageId="18" pageNumber="193">Fig. 7E</figureCitation>
). Distance from canthi carina to canthi apex longer than canthi thickness. Sensorial area of maxillary palpi rugose (
<figureCitation id="136E2A7AFF96062AFDB7C969F9A5C039" box="[620,686,297,319]" captionStart="FIGURE 4" captionStartId="6.[151,250,1693,1715]" captionTargetBox="[151,1415,201,1637]" captionTargetId="figure@6.[151,1436,193,1672]" captionTargetPageId="6" captionText="FIGURE 4. A, Labrum of Aegopsis bolboceridus male, in dorsal view; B, maxillary palpomere IV of Aegopsis bolboceridus male, in dorsal view; C, maxillary palpomere IV of Aegopsis bolboceridus female, in dorsal view; D, labrum of Aegopsis diceratops male, in dorsal view; E, maxillary palpomere IV of Aegopsis diceratops male, in dorsal view; F, maxillary palpomere IV of Aegopsis diceratops female, in dorsal view; G, labrum of Aegopsis vazdemelloi male, in dorsal view; H, maxillary palpomere IV of Aegopsis vazdemelloi male, in dorsal view; I, maxillary palpomere IV of Aegopsis vazdemelloi female, in dorsal view. Arrows indicating the middle of the labrum in A, G; arrows indicating sensorial area in C, I. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611538/files/figure.png" pageId="18" pageNumber="193">
Fig.
<date id="FFEB103FFF96062AFD42C969F9A5C039" box="[665,686,297,319]" pageId="18" pageNumber="193">4I</date>
</figureCitation>
)........................
<emphasis id="B921EAEDFF96062AFC05C96AFE90C039" bold="true" box="[990,1435,298,320]" pageId="18" pageNumber="193">
<taxonomicName id="4C554D7CFF96062AFC05C96AFE17C039" ID-CoL="65974" authority="Sobral &amp; Grossi" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[990,1308,298,320]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="193" phylum="Arthropoda" rank="species" species="vazdemelloi" status="sp. nov.">
<emphasis id="B921EAEDFF96062AFC05C96AFF65C046" bold="true" box="[990,1134,298,320]" italics="true" pageId="18" pageNumber="193">A. vazdemelloi</emphasis>
Sobral &amp; Grossi
</taxonomicName>
,
<taxonomicNameLabel id="A2125796FF96062AFAFDC96AFE90C039" box="[1318,1435,298,319]" pageId="18" pageNumber="193" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BEA36FFFF96062AFF4CC907FE97C07C" blockId="18.[151,1437,151,378]" pageId="18" pageNumber="193">
<keyLead id="30A495CAFF96062AFF4CC907FE97C07C" pageId="18" pageNumber="193">
- Vertex with shallower and annular punctures (
<figureCitation id="136E2A7AFF96062AFD78C907F9F9C05B" box="[675,754,327,349]" captionStart="FIGURE 7" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[185,1389,202,1867]" captionTargetId="figure@10.[181,1389,193,1871]" captionTargetPageId="10" captionText="FIGURE 7. A, Head of Aegopsis bolboceridus female in dorsal view; B, head of Aegopsis bolboceridus female in lateral view; C, head of Aegopsis diceratops female in dorsal view; D, head of Aegopsis diceratops female in lateral view; E, head of Aegopsis vazdemelloi female in dorsal view; F, head of Aegopsis vazdemelloi female in lateral view. Black arrow indicating head punctures in A; black arrow indicating clypeus trapezoidal in C; white arrow indicating vertex in C; black arrow indicating frontal tubercles in D; white arrow indicating head punctures in E. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2611546/files/figure.png" pageId="18" pageNumber="193">Fig. 7A</figureCitation>
). Distance from canthi carina to canthi apex subequal to canthi thick- ness. Sensorial area of maxillary palpi striate (
<figureCitation id="136E2A7AFF96062AFD47C924F9ECC07C" box="[668,743,356,378]" captionStart="FIGURE 4" captionStartId="6.[151,250,1693,1715]" captionTargetBox="[151,1415,201,1637]" captionTargetId="figure@6.[151,1436,193,1672]" captionTargetPageId="6" captionText="FIGURE 4. A, Labrum of Aegopsis bolboceridus male, in dorsal view; B, maxillary palpomere IV of Aegopsis bolboceridus male, in dorsal view; C, maxillary palpomere IV of Aegopsis bolboceridus female, in dorsal view; D, labrum of Aegopsis diceratops male, in dorsal view; E, maxillary palpomere IV of Aegopsis diceratops male, in dorsal view; F, maxillary palpomere IV of Aegopsis diceratops female, in dorsal view; G, labrum of Aegopsis vazdemelloi male, in dorsal view; H, maxillary palpomere IV of Aegopsis vazdemelloi male, in dorsal view; I, maxillary palpomere IV of Aegopsis vazdemelloi female, in dorsal view. Arrows indicating the middle of the labrum in A, G; arrows indicating sensorial area in C, I. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611538/files/figure.png" pageId="18" pageNumber="193">Fig. 4C</figureCitation>
)............................
<taxonomicName id="4C554D7CFF96062AFB94C924FE97C07C" ID-CoL="65972" authority="(Thomson, 1860)" baseAuthorityName="Thomson" baseAuthorityYear="1860" box="[1103,1436,356,378]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="193" phylum="Arthropoda" rank="species" species="bolboceridus">
<emphasis id="B921EAEDFF96062AFB94C924FE97C07C" bold="true" box="[1103,1436,356,378]" pageId="18" pageNumber="193">
<emphasis id="B921EAEDFF96062AFB94C924FFE2C07C" bold="true" box="[1103,1257,356,378]" italics="true" pageId="18" pageNumber="193">A. bolboceridus</emphasis>
(
<bibRefCitation id="EFC44B0EFF96062AFB2DC925FE9EC07C" author="Thomson, J." box="[1270,1429,357,378]" pageId="18" pageNumber="193" refId="ref10005" refString="Thomson, J. (1860) Musee Scientifique ou recueil d'histoire naturelle. Societe Entomologique de France, Paris, 96 pp." type="book" year="1860">Thomson, 1860</bibRefCitation>
)
</emphasis>
</taxonomicName>
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</paragraph>
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</subSubSection>
</treatment>
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