414 lines
56 KiB
XML
414 lines
56 KiB
XML
<document id="1ADD4A97ED7C2EC08C808D04324D878C" ID-DOI="10.12782/specdiv.25.153" ID-ISSN="2189-7301" ID-Zenodo-Dep="5738560" ID-ZooBank="00B865F7-1923-4F75-9075-14CB51A96782" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638229709865" checkinUser="felipe" docAuthor="Ogawa, Akito, Morita, Takami & Fujita, Toshihiko" docDate="2020" docId="03FC87CEAE7CDC1CFEDADB4DFEF6FD3A" docLanguage="en" docName="SpeciesDiversity.25.2.153-162.pdf" docOrigin="Species Diversity 25" docStyle="DocumentStyle:27FECD2C3CD153DA405029F2D33C9514.7:SpeciesDiversity.2012-.journal_article" docStyleId="27FECD2C3CD153DA405029F2D33C9514" docStyleName="SpeciesDiversity.2012-.journal_article" docStyleVersion="7" docTitle="Elpidia soyoae Ogawa & Morita & Fujita 2020, sp. nov." docType="treatment" docUuid="25D0076A-7C3D-468F-AE55-11058945757F" docUuidSource="ZooBank" docVersion="3" lastPageNumber="161" masterDocId="FFC5FFB6AE79DC14FF80D901FFFDFFB9" masterDocTitle="Elpidia soyoae, a New Species of Deep-sea Holothurian (Echinodermata) from the Japan Trench Area" masterLastPageNumber="162" masterPageNumber="153" pageNumber="158" updateTime="1699434177605" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="E2BA92226B73AA7AEA2B97ACAD572467">
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<mods:title id="DEF4F10ABD470310B1012076F698439D">Elpidia soyoae, a New Species of Deep-sea Holothurian (Echinodermata) from the Japan Trench Area</mods:title>
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</mods:titleInfo>
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<mods:name id="E71C5CCDDA22047B1FD623C40469C6AA" type="personal">
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<mods:roleTerm id="E6AA926F964DF14C4FE774D8EFA15C90">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="775024AB3A05F29EF345FBA6256C1F1C">Ogawa, Akito</mods:namePart>
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<mods:affiliation id="CF02AD65272F14F6C69035B71600DB26">Graduate School of Science, The University of Tokyo, 7 - 3 - 1 Hongo, Bunkyo-ku, Tokyo 113 - 0033, Japan E-mail: a-ogawa @ kahaku. go. jp & Department of Zoology, National Museum of Nature and Science, 4 - 1 - 1 Amakubo, Tsukuba, Ibaraki 305 - 0005, Japan & Corresponding author</mods:affiliation>
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<mods:nameIdentifier id="8E6848A0AC62ED209EA70595EA7E9380" type="email">ogawa@kahaku.go.jp</mods:nameIdentifier>
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</mods:name>
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<mods:name id="D8854090C772995DC890B5096541DD08" type="personal">
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<mods:roleTerm id="7650289858E6520DD456AE763CACB1DA">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9B34D57825B36A49E0EBE64A6FF48BA3">Morita, Takami</mods:namePart>
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<mods:affiliation id="6B4DE591FC03B27FB68EED6533EA736C">National Research Institute of Fisheries Science, Japan Fisheries Research and Education Agency, 2 - 12 - 4 Fukuura, Kanazawa-ku, Yokohama, Kanagawa 236 - 8648, Japan</mods:affiliation>
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</mods:name>
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<mods:name id="2A4B264B5FC43C3DA3DA22FD357EF6A7" type="personal">
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<mods:roleTerm id="9C9E21AFB60F245B4ED0CD1943C9BDD3">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="40316BBD41AD46EE431123806F9EA5D4">Fujita, Toshihiko</mods:namePart>
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<mods:affiliation id="72509CAB1C46E6C9F1295BBB4DB634A6">Graduate School of Science, The University of Tokyo, 7 - 3 - 1 Hongo, Bunkyo-ku, Tokyo 113 - 0033, Japan E-mail: a-ogawa @ kahaku. go. jp & Department of Zoology, National Museum of Nature and Science, 4 - 1 - 1 Amakubo, Tsukuba, Ibaraki 305 - 0005, Japan</mods:affiliation>
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<mods:nameIdentifier id="0ED54554DA96D033AA2062D9F722AF73" type="email">ogawa@kahaku.go.jp</mods:nameIdentifier>
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<mods:typeOfResource id="BC93B723050F028982FDA3E9BE8B5E72">text</mods:typeOfResource>
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<mods:titleInfo id="1845B9CE2FDC74A2754FAB4E8A3B7512">
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<mods:title id="CCFF5543E78B77B45F17ECF6C023D405">Species Diversity</mods:title>
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</mods:titleInfo>
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<mods:part id="C32CA403CBFDE145E318D3FCE338C0CA">
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<mods:date id="5F87B57FF62A20C16EAA29A66CF06738">2020</mods:date>
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<mods:number id="E9FE5DCBBE450F06AEC3CB47820D3DC1">2020-08-07</mods:number>
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<mods:detail id="242E0728742DA729C364FDE250B29092" type="volume">
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<mods:number id="781A8D3EE137EF3762083CED8C6A7AE8">25</mods:number>
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</mods:detail>
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<mods:start id="9F8FB4B2D8E9EDBC3FBE520708A69434">153</mods:start>
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<mods:end id="F58FD174464916DF03673677D42C987A">162</mods:end>
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<mods:classification id="6F8F3EAC33F7B872A13DF3E17108A298">journal article</mods:classification>
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<mods:identifier id="68A349D5BB4228C61863A2663E619D44" type="DOI">10.12782/specdiv.25.153</mods:identifier>
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<mods:identifier id="806C5E94689DC9F12DC377C00A88A058" type="ISSN">2189-7301</mods:identifier>
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<mods:identifier id="4EF63C25AAA5DE0DA38C48DB1CA295B6" type="Zenodo-Dep">5738560</mods:identifier>
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<mods:identifier id="746364457763434065ACEA8939D028A0" type="ZooBank">00B865F7-1923-4F75-9075-14CB51A96782</mods:identifier>
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<treatment id="03FC87CEAE7CDC1CFEDADB4DFEF6FD3A" LSID="urn:lsid:zoobank.org:act:25D0076A-7C3D-468F-AE55-11058945757F" httpUri="http://treatment.plazi.org/id/03FC87CEAE7CDC1CFEDADB4DFEF6FD3A" lastPageId="8" lastPageNumber="161" pageId="5" pageNumber="158">
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<subSubSection id="C34F6553AE7CDC11FEDADB4DFDC6FD1F" pageId="5" pageNumber="158" type="nomenclature">
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<paragraph id="8BEA36D8AE7CDC11FEDADB4DFDBFFDDD" blockId="5.[346,578,588,612]" box="[346,578,588,612]" pageId="5" pageNumber="158">
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<heading id="D0A281B4AE7CDC11FEDADB4DFDBFFDDD" box="[346,578,588,612]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="158" reason="2">
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<taxonomicName id="4C554D5BAE7CDC11FEDADB4DFE0CFDDD" ID-CoL="848LM" authority="Ogawa & Morita & Fujita, 2020" authorityName="Ogawa & Morita & Fujita" authorityYear="2020" box="[346,497,588,612]" class="Holothuroidea" family="Elpidiidae" genus="Elpidia" kingdom="Animalia" order="Elasipodida" pageId="5" pageNumber="158" phylum="Echinodermata" rank="species" species="soyoae" status="sp. nov.">
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<emphasis id="B921EACAAE7CDC11FEDADB4DFE0CFDDD" bold="true" box="[346,497,588,612]" italics="true" pageId="5" pageNumber="158">Elpidia soyoae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21257B1AE7CDC11FE76DB4CFDBFFDDD" box="[502,578,589,612]" pageId="5" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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||
</paragraph>
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||
<paragraph id="8BEA36D8AE7CDC11FF73DB6FFDC6FD1F" blockId="5.[243,680,621,678]" pageId="5" pageNumber="158">
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||
[New Japanese name: Soyo-kuma-namako] (
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<figureCitation id="136E2A5DAE7CDC11FEE7DB8FFE42FD1F" box="[359,447,654,678]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStartId-0="1.[156,187,2056,2078]" captionStartId-1="2.[156,187,1972,1994]" captionStartId-2="3.[156,187,2000,2022]" captionStartId-3="5.[156,187,2000,2022]" captionStartId-4="6.[156,187,2028,2050]" captionStartId-5="7.[156,187,2028,2050]" captionTargetBox-0="[189,1398,581,2015]" captionTargetBox-1="[189,1398,445,1931]" captionTargetBox-2="[189,1398,1213,1960]" captionTargetBox-3="[189,1398,984,1960]" captionTargetBox-4="[189,1398,890,1987]" captionTargetBox-5="[189,1398,1063,1987]" captionTargetId-0="figure-162@1.[187,1400,579,2017]" captionTargetId-1="figure-130@2.[187,1400,443,1933]" captionTargetId-2="figure-475@3.[187,1400,1211,1962]" captionTargetId-3="figure-382@5.[187,1400,982,1962]" captionTargetId-4="figure-405@6.[187,1400,888,1989]" captionTargetId-5="figure-472@7.[187,1400,1061,1989]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionText-0="Fig. 1. Fresh specimen of Elpidia soyoae sp. nov. (NSMT E-12635: paratype). A, dorsal view; B, ventral view." captionText-1="Fig. 2. Papillae arrangements of Elpidia soyoae sp. nov. A, left side view; B, right side view; C–F, dorsal side. A–B, holotype; C, NSMT E-12635: paratype; D, NSMT E-12636: paratype; E, NSMT E-12637: paratype; F, NSMT E-12639: paratype. Abbreviations: lp, paired papillae on dorsal left radius; lu, unpaired papillae on dorsal left radius; rp, paired papillae on dorsal right radius; ru, unpaired papillae on dorsal right radius." captionText-2="Fig. 3. Calcareous ring of Elpidia soyoae sp. nov. (holotype). A, entire ring in dorsal view; B, variation of isolated pieces; C, branched inside posterior arm. Black arrowheads show five pieces surrounding pharynx. Abbreviations: ba, branched arm of inside posterior arms; iaa, inside anterior arms; ipa, inside posterior arms; oaa, outside anterior arms; opa, outside posterior arms." captionText-3="Fig. 4. SEM images of ossicles from dorsal body wall of Elpidia soyoae sp. nov. (NSMT E-12635: paratype). A, rods with straight axis, well developed horizontal arms, and vertical apophyses; B, wheel from convex side and obliquely lateral side. Abbreviations: ax, axis; bt, brim teeth; ccp, central connecting portion; cr, central rays; ha, horizontal arms; sp, spokes; va, vertical apophyses." captionText-4="Fig. 5. SEM images of ossicles from ventral body wall of Elpidia soyoae sp. nov. (holotype). Rods with straight axis, well developed horizontal arms, and vertical apophyses. Abbreviations: ax, axis; ha, horizontal arms; va, vertical apophyses." captionText-5="Fig. 6. SEM images of rods from tentacles of Elpidia soyoae sp. nov. (NSMT E-12635: paratype). Rods with arched axis, especially shortened horizontal arms, and vertical apophyses. Abbreviations: ax, axis; ha, horizontal arms; va, vertical apophyses." figureDoi-0="http://doi.org/10.5281/zenodo.5738562" figureDoi-1="http://doi.org/10.5281/zenodo.5738564" figureDoi-2="http://doi.org/10.5281/zenodo.5738566" figureDoi-3="http://doi.org/10.5281/zenodo.5738568" figureDoi-4="http://doi.org/10.5281/zenodo.5738570" figureDoi-5="http://doi.org/10.5281/zenodo.5738572" httpUri-0="https://zenodo.org/record/5738562/files/figure.png" httpUri-1="https://zenodo.org/record/5738564/files/figure.png" httpUri-2="https://zenodo.org/record/5738566/files/figure.png" httpUri-3="https://zenodo.org/record/5738568/files/figure.png" httpUri-4="https://zenodo.org/record/5738570/files/figure.png" httpUri-5="https://zenodo.org/record/5738572/files/figure.png" pageId="5" pageNumber="158">Figs 1–6</figureCitation>
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;
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<tableCitation id="C6D70363AE7CDC11FE49DB8FFDE7FD1F" box="[457,538,654,678]" captionStart="Table 1" captionStartId="4.[156,204,590,613]" captionText="Table 1. Morphological counts and measurements of type specimens of Elpidia soyoae sp.nov." pageId="5" pageNumber="158">Tables 1</tableCitation>
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,
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<tableCitation id="C6D70363AE7CDC11FDA6DB8FFDCEFD1F" box="[550,563,654,678]" captionStart="Table 2" captionStartId="8.[156,204,592,615]" captionText="Table 2. Morphological comparison of Elpidia soyoae sp. nov. with Elpidia species in the slender axis group. DAH and DOL indicate apophyses height and ossicle length of rods in dorsal body wall," pageId="5" pageNumber="158">2</tableCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C34F6553AE7CDC11FF35DBCEFBAFFEA6" pageId="5" pageNumber="158" type="materials_examined">
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<paragraph id="8BEA36D8AE7CDC11FF35DBCEFD02FC30" blockId="5.[156,767,719,906]" pageId="5" pageNumber="158">
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<emphasis id="B921EACAAE7CDC11FF35DBCEFE72FD5E" bold="true" box="[181,399,719,743]" pageId="5" pageNumber="158">Material examined.</emphasis>
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<materialsCitation id="3B3D3C85AE7CDC11FE23DBCEFF04FC91" accessionNumber="LC528684" collectionCode="NSMT, BL, INSD" pageId="5" pageNumber="158" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54EE887AAE7CDC11FE23DBCEFDF9FD5E" box="[419,516,719,743]" pageId="5" pageNumber="158" type="holotype">Holotype</typeStatus>
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||
:
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<collectionCode id="ED44AE1DAE7CDC11FD9ADBCEFDA2FD5E" box="[538,607,719,743]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="5" pageNumber="158">NSMT</collectionCode>
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E-12638 (
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<collectionCode id="ED44AE1DAE7CDC11FD60DBCEFD02FD5E" box="[736,767,719,743]" pageId="5" pageNumber="158">BL</collectionCode>
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<quantity id="4CAD9B3DAE7CDC11FF1CDBEEFF1FFCBE" box="[156,226,751,775]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="158" unit="mm" value="18.0">18 mm</quantity>
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||
; BW
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<quantity id="4CAD9B3DAE7CDC11FEA5DBEEFE93FCBE" box="[293,366,751,775]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="158" unit="mm" value="11.0">11 mm</quantity>
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),
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||
<collectionCode id="ED44AE1DAE7CDC11FE05DBEEFE42FCBE" box="[389,447,751,775]" pageId="5" pageNumber="158">INSD</collectionCode>
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accession number
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||
<accessionNumber id="9406AB3BAE7CDC11FD12DBEEFD06FCBE" box="[658,763,751,775]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC528684" pageId="5" pageNumber="158" type="EnaNcbi">LC528684</accessionNumber>
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||
, 610 bps
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||
</materialsCitation>
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||
.
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<typeStatus id="54EE887AAE7CDC11FE8DDA11FE8CFC91" box="[269,369,784,808]" pageId="5" pageNumber="158" type="paratype">Paratypes</typeStatus>
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:
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<collectionCode id="ED44AE1DAE7CDC11FE07DA11FE31FC91" box="[391,460,784,808]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="5" pageNumber="158">NSMT</collectionCode>
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E-12635 (
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<collectionCode id="ED44AE1DAE7CDC11FDCADA11FD94FC91" box="[586,617,784,808]" pageId="5" pageNumber="158">BL</collectionCode>
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<quantity id="4CAD9B3DAE7CDC11FDFBDA11FD3FFC9E" box="[635,706,784,808]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="158" unit="mm" value="25.0">25 mm</quantity>
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||
; BW
|
||
<quantity id="4CAD9B3DAE7CDC11FF1CDA30FF19FCF0" box="[156,228,817,841]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="158" unit="mm" value="13.0">13 mm</quantity>
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),
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<collectionCode id="ED44AE1DAE7CDC11FE81DA30FEC6FCF0" box="[257,315,817,841]" pageId="5" pageNumber="158">INSD</collectionCode>
|
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accession number
|
||
<accessionNumber id="9406AB3BAE7CDC11FDA1DA31FD77FCF1" box="[545,650,816,840]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC528681" pageId="5" pageNumber="158" type="EnaNcbi">LC528681</accessionNumber>
|
||
, 628 bps;
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||
<collectionCode id="ED44AE1DAE7CDC11FF1CDA50FF1CFCD0" box="[156,225,849,873]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="5" pageNumber="158">NSMT</collectionCode>
|
||
E-12636 (
|
||
<collectionCode id="ED44AE1DAE7CDC11FED0DA50FE92FCD0" box="[336,367,849,873]" pageId="5" pageNumber="158">BL</collectionCode>
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<quantity id="4CAD9B3DAE7CDC11FEF8DA50FE42FCD1" box="[376,447,849,873]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="5" pageNumber="158" unit="mm" value="21.0">21 mm</quantity>
|
||
; BW
|
||
<quantity id="4CAD9B3DAE7CDC11FE7EDA50FDBAFCD0" box="[510,583,849,873]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="158" unit="mm" value="12.0">12 mm</quantity>
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||
),
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||
<collectionCode id="ED44AE1DAE7CDC11FDDCDA50FD6BFCD0" box="[604,662,849,873]" pageId="5" pageNumber="158">INSD</collectionCode>
|
||
accession number
|
||
<accessionNumber id="9406AB3BAE7CDC11FF78DA70FE9CFC30" box="[248,353,881,905]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC528682" pageId="5" pageNumber="158" type="EnaNcbi">LC528682</accessionNumber>
|
||
, 639 bps;
|
||
<collectionCode id="ED44AE1DAE7CDC11FE54DA73FDE4FC33" box="[468,537,882,906]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="5" pageNumber="158">NSMT</collectionCode>
|
||
E-12637 (
|
||
<collectionCode id="ED44AE1DAE7CDC11FD0ADA73FD54FC33" box="[650,681,882,906]" pageId="5" pageNumber="158">BL</collectionCode>
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<quantity id="4CAD9B3DAE7CDC11FD34DA73FD06FC30" box="[692,763,882,906]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="5" pageNumber="158" unit="mm" value="19.0">19 mm</quantity>
|
||
;
|
||
</paragraph>
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<paragraph id="8BEA36D8AE7CDC11FCB5D9C6FBAFFEA6" blockId="5.[821,1432,198,905]" pageId="5" pageNumber="158">
|
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BW
|
||
<quantity id="4CAD9B3DAE7CDC11FCE5D9C7FC53FF67" box="[869,942,198,222]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="158" unit="mm" value="12.0">12 mm</quantity>
|
||
),
|
||
<collectionCode id="ED44AE1DAE7CDC11FC43D9C7FC00FF67" box="[963,1021,198,222]" pageId="5" pageNumber="158">INSD</collectionCode>
|
||
accession number
|
||
<accessionNumber id="9406AB3BAE7CDC11FB49D9C7FACEFF67" box="[1225,1331,198,222]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC528683" pageId="5" pageNumber="158" type="EnaNcbi">LC528683</accessionNumber>
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||
, 636 bps;
|
||
<collectionCode id="ED44AE1DAE7CDC11FCB5D9E6FC87FF46" box="[821,890,231,255]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="5" pageNumber="158">NSMT</collectionCode>
|
||
E-12639 (
|
||
<collectionCode id="ED44AE1DAE7CDC11FC6CD9E6FBF6FF46" box="[1004,1035,231,255]" pageId="5" pageNumber="158">BL</collectionCode>
|
||
<quantity id="4CAD9B3DAE7CDC11FB96D9E6FBA0FF47" box="[1046,1117,231,255]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="158" unit="mm" value="12.0">12 mm</quantity>
|
||
; BW
|
||
<quantity id="4CAD9B3DAE7CDC11FB20D9E6FB21FF46" box="[1184,1244,231,255]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="158" unit="mm" value="8.0">8 mm</quantity>
|
||
),
|
||
<collectionCode id="ED44AE1DAE7CDC11FB73D9E6FAD0FF46" box="[1267,1325,231,255]" pageId="5" pageNumber="158">INSD</collectionCode>
|
||
accession number
|
||
<accessionNumber id="9406AB3BAE7CDC11FC0BD806FC09FEA6" box="[907,1012,263,287]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC528685" pageId="5" pageNumber="158" type="EnaNcbi">LC528685</accessionNumber>
|
||
, 623 bps.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34F6553AE7CDC11FCCED829FB71FDFD" pageId="5" pageNumber="158" type="diagnosis">
|
||
<paragraph id="8BEA36D8AE7CDC11FCCED829FB71FDFD" blockId="5.[821,1432,198,905]" pageId="5" pageNumber="158">
|
||
<emphasis id="B921EACAAE7CDC11FCCED829FC43FEF9" bold="true" box="[846,958,296,320]" pageId="5" pageNumber="158">Diagnosis.</emphasis>
|
||
BL up to
|
||
<quantity id="4CAD9B3DAE7CDC11FBB3D829FB86FE86" box="[1075,1147,296,320]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="158" unit="mm" value="25.0">25 mm</quantity>
|
||
. Dorsal papillae up to five pairs along the entire dorsal radii; second pairs widely separated from third ones. Purple pigmentation spots on dorsal and ventral body walls, tentacles, and tube feet. Ossicles in dorsal body wall mainly rods,
|
||
<taxonomicName id="4C554D5BAE7CDC11FBE7D8ABFB53FE7B" authorityName="Theel" authorityYear="1876" box="[1127,1198,426,450]" class="Holothuroidea" family="Elpidiidae" genus="Elpidia" kingdom="Animalia" order="Elasipodida" pageId="5" pageNumber="158" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B921EACAAE7CDC11FBE7D8ABFB53FE7B" box="[1127,1198,426,450]" italics="true" pageId="5" pageNumber="158">Elpidia</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="54EE887AAE7CDC11FB34D8ABFB1FFE7B" box="[1204,1250,426,450]" pageId="5" pageNumber="158">type</typeStatus>
|
||
, with two pairs of horizontal arms and one pair of vertical apophyses, rod axis up to 1336 µm long, 8.1–33 µm in diameter, length of apophyses 5.1–47% of axis length; tentacle rods with arched axis, and short arms and/or apophyses.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34F6553AE7CDC13FCCEDB4DFE9BFD9D" lastPageId="7" lastPageNumber="160" pageId="5" pageNumber="158" type="description">
|
||
<paragraph id="8BEA36D8AE7CDC12FCCEDB4DFAB8FF46" blockId="5.[821,1432,198,905]" lastBlockId="6.[820,1432,198,841]" lastPageId="6" lastPageNumber="159" pageId="5" pageNumber="158">
|
||
<emphasis id="B921EACAAE7CDC11FCCEDB4DFC2FFDDD" bold="true" box="[846,978,588,612]" pageId="5" pageNumber="158">Description.</emphasis>
|
||
Body semi-ellipsoidal with flat ventrum,
|
||
<quantity id="4CAD9B3DAE7CDC11FCB5DB6CFC63FD3C" box="[821,926,621,645]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.5" metricValueMin="1.2" pageId="5" pageNumber="158" unit="mm" value="18.5" valueMax="25.0" valueMin="12.0">12–25mm</quantity>
|
||
long (5 individuals), 1.5–1.9 times as long as wide (
|
||
<figureCitation id="136E2A5DAE7CDC11FCF3DB8FFC51FD1F" box="[883,940,654,678]" captionStart="Fig" captionStartId="1.[156,187,2056,2078]" captionTargetBox="[189,1398,581,2015]" captionTargetId="figure-162@1.[187,1400,579,2017]" captionTargetPageId="1" captionText="Fig. 1. Fresh specimen of Elpidia soyoae sp. nov. (NSMT E-12635: paratype). A, dorsal view; B, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5738562" httpUri="https://zenodo.org/record/5738562/files/figure.png" pageId="5" pageNumber="158">Fig. 1</figureCitation>
|
||
). Mouth anteroventral. Anus at posterior end of body. Dorsal papillae conical, 0.2–5.0 mm long (N=49 from 5 individuals,
|
||
<tableCitation id="C6D70363AE7CDC11FC45DBCEFBF0FD5E" box="[965,1037,719,743]" captionStart="Table 1" captionStartId="4.[156,204,590,613]" captionText="Table 1. Morphological counts and measurements of type specimens of Elpidia soyoae sp.nov." pageId="5" pageNumber="158">Table 1</tableCitation>
|
||
), four or five paired and 0–3 unpaired papillae present on two dorsal radii (four to seven papillae on each radius,
|
||
<figureCitation id="136E2A5DAE7CDC11FC65DA11FBDBFC91" box="[997,1062,784,808]" captionStart="Fig" captionStartId="2.[156,187,1972,1994]" captionTargetBox="[189,1398,445,1931]" captionTargetId="figure-130@2.[187,1400,443,1933]" captionTargetPageId="2" captionText="Fig. 2. Papillae arrangements of Elpidia soyoae sp. nov. A, left side view; B, right side view; C–F, dorsal side. A–B, holotype; C, NSMT E-12635: paratype; D, NSMT E-12636: paratype; E, NSMT E-12637: paratype; F, NSMT E-12639: paratype. Abbreviations: lp, paired papillae on dorsal left radius; lu, unpaired papillae on dorsal left radius; rp, paired papillae on dorsal right radius; ru, unpaired papillae on dorsal right radius." figureDoi="http://doi.org/10.5281/zenodo.5738564" httpUri="https://zenodo.org/record/5738564/files/figure.png" pageId="5" pageNumber="158">Fig. 2</figureCitation>
|
||
). Paired large papillae 0.3–5.0 mm long (N=42 from 5 individuals), anterior two pairs widely separated from third paired ones. Unpaired minute papillae
|
||
<quantity id="4CAD9B3DAE7CDC11FCDDDA70FC2EFC30" box="[861,979,881,905]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="2.7" metricValueMin="0.2" pageId="5" pageNumber="158" unit="mm" value="1.45" valueMax="2.7" valueMin="0.2">0.2–2.7 mm</quantity>
|
||
long (N=7 from 3 individuals), between second and third paired papillae on
|
||
<typeStatus id="54EE887AAE7FDC12FD90D9C7FD94FF67" box="[528,617,198,222]" pageId="6" pageNumber="159" type="holotype">holotype</typeStatus>
|
||
and
|
||
<specimenCount id="9D53FD51AE7FDC12FD1DD9C6FF35FF46" pageId="6" pageNumber="159" type="generic" typeStatus="paratype">one paratype</specimenCount>
|
||
(NSMT E-12639), and on the posterior dorsum of
|
||
<typeStatus id="54EE887AAE7FDC12FD5BD9E6FF26FEA6" pageId="6" pageNumber="159" type="holotype">holotype</typeStatus>
|
||
and
|
||
<specimenCount id="9D53FD51AE7FDC12FE9AD809FE59FE99" box="[282,420,264,288]" pageId="6" pageNumber="159" type="generic" typeStatus="paratype">one paratype</specimenCount>
|
||
(NSMT E-12635). Ventrolateral radii without conical papillae. Tube feet cylindrical, four on each ventrolateral radius. Mid-ventral radius without tube feet. Tentacles ten, with cylindrical stems, and four conical digits on terminal discs. Papillae, tube feet, and tentacles nonretractile. Calcareous ring comprising five pieces surrounding pharynx (
|
||
<figureCitation id="136E2A5DAE7FDC12FEEAD8CAFE46FE5A" box="[362,443,459,483]" captionStart="Fig" captionStartId="3.[156,187,2000,2022]" captionTargetBox="[189,1398,1213,1960]" captionTargetId="figure-475@3.[187,1400,1211,1962]" captionTargetPageId="3" captionText="Fig. 3. Calcareous ring of Elpidia soyoae sp. nov. (holotype). A, entire ring in dorsal view; B, variation of isolated pieces; C, branched inside posterior arm. Black arrowheads show five pieces surrounding pharynx. Abbreviations: ba, branched arm of inside posterior arms; iaa, inside anterior arms; ipa, inside posterior arms; oaa, outside anterior arms; opa, outside posterior arms." figureDoi="http://doi.org/10.5281/zenodo.5738566" httpUri="https://zenodo.org/record/5738566/files/figure.png" pageId="6" pageNumber="159">Fig. 3A</figureCitation>
|
||
), each piece with four pairs of straight arms (
|
||
<figureCitation id="136E2A5DAE7FDC12FEB6D8EAFE7BFDBA" box="[310,390,491,515]" captionStart="Fig" captionStartId="3.[156,187,2000,2022]" captionTargetBox="[189,1398,1213,1960]" captionTargetId="figure-475@3.[187,1400,1211,1962]" captionTargetPageId="3" captionText="Fig. 3. Calcareous ring of Elpidia soyoae sp. nov. (holotype). A, entire ring in dorsal view; B, variation of isolated pieces; C, branched inside posterior arm. Black arrowheads show five pieces surrounding pharynx. Abbreviations: ba, branched arm of inside posterior arms; iaa, inside anterior arms; ipa, inside posterior arms; oaa, outside anterior arms; opa, outside posterior arms." figureDoi="http://doi.org/10.5281/zenodo.5738566" httpUri="https://zenodo.org/record/5738566/files/figure.png" pageId="6" pageNumber="159">Fig. 3B</figureCitation>
|
||
): inside anterior pair (495–1131 µm long, N=
|
||
<quantity id="4CAD9B3DAE7FDC12FF7DDB0DFECCFD9D" box="[253,305,524,548]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="6" pageNumber="159" unit="in" value="10.0">10 in</quantity>
|
||
<typeStatus id="54EE887AAE7FDC12FEB7DB0DFE69FD9D" box="[311,404,524,548]" pageId="6" pageNumber="159" type="holotype">holotype</typeStatus>
|
||
), inside posterior pair (266–907 µm long, N=
|
||
<quantity id="4CAD9B3DAE7FDC12FE87DB2DFEC4FDFD" box="[263,313,556,580]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="6" pageNumber="159" unit="in" value="8.0">8 in</quantity>
|
||
<typeStatus id="54EE887AAE7FDC12FECADB2DFE5AFDFD" box="[330,423,556,580]" pageId="6" pageNumber="159" type="holotype">holotype</typeStatus>
|
||
), outside anterior pair (1092– 2075 µm long, N=
|
||
<quantity id="4CAD9B3DAE7FDC12FED9DB4CFE7CFDDC" box="[345,385,589,613]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="6" pageNumber="159" unit="in" value="7.0">7 in</quantity>
|
||
<typeStatus id="54EE887AAE7FDC12FE08DB4CFE18FDDC" box="[392,485,589,613]" pageId="6" pageNumber="159" type="holotype">holotype</typeStatus>
|
||
), and outside posterior pair (968–1972 µm long, N=
|
||
<quantity id="4CAD9B3DAE7FDC12FE19DB6CFE3EFD3C" box="[409,451,621,645]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="6" pageNumber="159" unit="in" value="9.0">9 in</quantity>
|
||
<typeStatus id="54EE887AAE7FDC12FE4CDB6CFDD4FD3C" box="[460,553,621,645]" pageId="6" pageNumber="159" type="holotype">holotype</typeStatus>
|
||
). Tips of inside arm pairs branched or broadened in
|
||
<typeStatus id="54EE887AAE7FDC12FE6DDB8FFDBBFD1F" box="[493,582,654,678]" pageId="6" pageNumber="159" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="136E2A5DAE7FDC12FDD6DB8FFD58FD1F" box="[598,677,654,678]" captionStart="Fig" captionStartId="3.[156,187,2000,2022]" captionTargetBox="[189,1398,1213,1960]" captionTargetId="figure-475@3.[187,1400,1211,1962]" captionTargetPageId="3" captionText="Fig. 3. Calcareous ring of Elpidia soyoae sp. nov. (holotype). A, entire ring in dorsal view; B, variation of isolated pieces; C, branched inside posterior arm. Black arrowheads show five pieces surrounding pharynx. Abbreviations: ba, branched arm of inside posterior arms; iaa, inside anterior arms; ipa, inside posterior arms; oaa, outside anterior arms; opa, outside posterior arms." figureDoi="http://doi.org/10.5281/zenodo.5738566" httpUri="https://zenodo.org/record/5738566/files/figure.png" pageId="6" pageNumber="159">Fig. 3C</figureCitation>
|
||
). Gonad white unpaired tuft, comprising central genital ducts and surrounding genital vesicles. Polian vesicle single, fusiform, transparent, whitish in
|
||
<typeStatus id="54EE887AAE7FDC12FE12DBEEFE13FCBE" box="[402,494,751,775]" pageId="6" pageNumber="159" type="holotype">holotype</typeStatus>
|
||
. Body skin whitish, occasionally with purple pigmentation spots composed of small purple particles on dorsal and ventral surfaces. Discs of tube feet and tentacles light yellow, sometimes with purple pigmentation spots composed of small purple particles.
|
||
</paragraph>
|
||
<caption id="DF2A6650AE7CDC11FF1CDED1FB8DF7A7" ID-DOI="http://doi.org/10.5281/zenodo.5738568" ID-Zenodo-Dep="5738568" httpUri="https://zenodo.org/record/5738568/files/figure.png" pageId="5" pageNumber="158" startId="5.[156,187,2000,2022]" targetBox="[189,1398,984,1960]" targetPageId="5">
|
||
<paragraph id="8BEA36D8AE7CDC11FF1CDED1FB8DF7A7" blockId="5.[156,1431,2000,2078]" pageId="5" pageNumber="158">
|
||
Fig. 4. SEM images of ossicles from dorsal body wall of
|
||
<taxonomicName id="4C554D5BAE7CDC11FD2FDED1FCD2F85C" authority="Ogawa & Morita & Fujita, 2020" authorityName="Ogawa & Morita & Fujita" authorityYear="2020" box="[687,815,2000,2021]" class="Holothuroidea" family="Elpidiidae" genus="Elpidia" kingdom="Animalia" order="Elasipodida" pageId="5" pageNumber="158" phylum="Echinodermata" rank="species" species="soyoae" status="sp. nov.">
|
||
<emphasis id="B921EACAAE7CDC11FD2FDED1FCD2F85C" box="[687,815,2000,2021]" italics="true" pageId="5" pageNumber="158">Elpidia soyoae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21257B1AE7CDC11FCB5DED1FC87F85C" box="[821,890,2000,2021]" pageId="5" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(NSMT E-12635: paratype). A, rods with straight axis, well developed horizontal arms, and vertical apophyses; B, wheel from convex side and obliquely lateral side. Abbreviations: ax, axis; bt, brim teeth; ccp, central connecting portion; cr, central rays; ha, horizontal arms; sp, spokes; va, vertical apophyses.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BEA36D8AE7FDC13FCCED806FE9BFD9D" blockId="6.[820,1432,198,841]" lastBlockId="7.[156,768,198,1003]" lastPageId="7" lastPageNumber="160" pageId="6" pageNumber="159">
|
||
<emphasis id="B921EACAAE7FDC12FCCED806FBEAFEA6" bold="true" box="[846,1047,263,287]" pageId="6" pageNumber="159">
|
||
Ossicles (
|
||
<tableCitation id="C6D70363AE7FDC12FC36D806FBF5FEA6" box="[950,1032,263,287]" captionStart="Table 1" captionStartId="4.[156,204,590,613]" captionText="Table 1. Morphological counts and measurements of type specimens of Elpidia soyoae sp.nov." pageId="6" pageNumber="159">Table 1</tableCitation>
|
||
).
|
||
</emphasis>
|
||
Ossicles in dorsal body wall mostly rods,
|
||
<taxonomicName id="4C554D5BAE7FDC12FC0ED829FC28FEF9" authorityName="Theel" authorityYear="1876" box="[910,981,296,320]" class="Holothuroidea" family="Elpidiidae" genus="Elpidia" kingdom="Animalia" order="Elasipodida" pageId="6" pageNumber="159" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B921EACAAE7FDC12FC0ED829FC28FEF9" box="[910,981,296,320]" italics="true" pageId="6" pageNumber="159">Elpidia</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<typeStatus id="54EE887AAE7FDC12FC5CD829FBF6FEF9" box="[988,1035,296,320]" pageId="6" pageNumber="159">type</typeStatus>
|
||
, and rarely wheels (
|
||
<figureCitation id="136E2A5DAE7FDC12FB64D829FAD8FEF9" box="[1252,1317,296,320]" captionStart="Fig" captionStartId="5.[156,187,2000,2022]" captionTargetBox="[189,1398,984,1960]" captionTargetId="figure-382@5.[187,1400,982,1962]" captionTargetPageId="5" captionText="Fig. 4. SEM images of ossicles from dorsal body wall of Elpidia soyoae sp. nov. (NSMT E-12635: paratype). A, rods with straight axis, well developed horizontal arms, and vertical apophyses; B, wheel from convex side and obliquely lateral side. Abbreviations: ax, axis; bt, brim teeth; ccp, central connecting portion; cr, central rays; ha, horizontal arms; sp, spokes; va, vertical apophyses." figureDoi="http://doi.org/10.5281/zenodo.5738568" httpUri="https://zenodo.org/record/5738568/files/figure.png" pageId="6" pageNumber="159">Fig. 4</figureCitation>
|
||
). Rod axis 193–1336 µm in length (DOL, N=104 from 5 individuals), straight, with two paired horizontal arms, and one pair of vertical apophyses. Axis 8.1–33 µm in diameter (N=192 from 5 individuals), with conical tips at both ends. The height of apophyses 17–232 µm (DAH, N=236 from 5 individuals). DAH/DOL ratio between 0.051–0.47 (N=150 from 5 individuals). Wheels rare, 30–33 µm in diameter (N=3 from 3 individuals), central connecting portion 18–21 µm in diameter (N=3 from 3 individuals), with nine spokes (N=3 from 3 individuals), three central rays (N=2 from 2 individuals), and triangular brim teeth (N=3 from 3 individuals) (
|
||
<figureCitation id="136E2A5DAE7FDC12FC53DBAFFBDFFD7F" box="[979,1058,686,710]" captionStart="Fig" captionStartId="5.[156,187,2000,2022]" captionTargetBox="[189,1398,984,1960]" captionTargetId="figure-382@5.[187,1400,982,1962]" captionTargetPageId="5" captionText="Fig. 4. SEM images of ossicles from dorsal body wall of Elpidia soyoae sp. nov. (NSMT E-12635: paratype). A, rods with straight axis, well developed horizontal arms, and vertical apophyses; B, wheel from convex side and obliquely lateral side. Abbreviations: ax, axis; bt, brim teeth; ccp, central connecting portion; cr, central rays; ha, horizontal arms; sp, spokes; va, vertical apophyses." figureDoi="http://doi.org/10.5281/zenodo.5738568" httpUri="https://zenodo.org/record/5738568/files/figure.png" pageId="6" pageNumber="159">Fig. 4B</figureCitation>
|
||
). Ossicles in ventral body wall only rods,
|
||
<taxonomicName id="4C554D5BAE7FDC12FCF1DBCEFC45FD5E" authorityName="Theel" authorityYear="1876" box="[881,952,719,743]" class="Holothuroidea" family="Elpidiidae" genus="Elpidia" kingdom="Animalia" order="Elasipodida" pageId="6" pageNumber="159" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B921EACAAE7FDC12FCF1DBCEFC45FD5E" box="[881,952,719,743]" italics="true" pageId="6" pageNumber="159">Elpidia</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<typeStatus id="54EE887AAE7FDC12FC3EDBCEFC11FD5E" box="[958,1004,719,743]" pageId="6" pageNumber="159">type</typeStatus>
|
||
ossicles (
|
||
<figureCitation id="136E2A5DAE7FDC12FBD1DBCEFB6DFD5E" box="[1105,1168,719,743]" captionStart="Fig" captionStartId="6.[156,187,2028,2050]" captionTargetBox="[189,1398,890,1987]" captionTargetId="figure-405@6.[187,1400,888,1989]" captionTargetPageId="6" captionText="Fig. 5. SEM images of ossicles from ventral body wall of Elpidia soyoae sp. nov. (holotype). Rods with straight axis, well developed horizontal arms, and vertical apophyses. Abbreviations: ax, axis; ha, horizontal arms; va, vertical apophyses." figureDoi="http://doi.org/10.5281/zenodo.5738570" httpUri="https://zenodo.org/record/5738570/files/figure.png" pageId="6" pageNumber="159">Fig. 5</figureCitation>
|
||
). Rod axis 73–557 µm in length (VOL, N=200 from 5 individuals), straight, with two paired horizontal arms, and one pair of vertical apophyses, sometimes shortened. Axis in diameter 4–24µm (N=274 from 5 individuals). The height of apophyses 10–142 µm (VAH, N=438 from 5 individuals), VAH/VOL ratio between 0.071–0.70 (N=332 from 5 individuals). Ossicles in tentacles only rods,
|
||
<taxonomicName id="4C554D5BAE7EDC13FEECD829FE4EFEF9" authorityName="Theel" authorityYear="1876" box="[364,435,296,320]" class="Holothuroidea" family="Elpidiidae" genus="Elpidia" kingdom="Animalia" order="Elasipodida" pageId="7" pageNumber="160" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B921EACAAE7EDC13FEECD829FE4EFEF9" box="[364,435,296,320]" italics="true" pageId="7" pageNumber="160">Elpidia</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<typeStatus id="54EE887AAE7EDC13FE39D829FE1AFEF9" box="[441,487,296,320]" pageId="7" pageNumber="160">type</typeStatus>
|
||
ossicles (
|
||
<figureCitation id="136E2A5DAE7EDC13FDCBD829FD75FEF9" box="[587,648,296,320]" captionStart="Fig" captionStartId="7.[156,187,2028,2050]" captionTargetBox="[189,1398,1063,1987]" captionTargetId="figure-472@7.[187,1400,1061,1989]" captionTargetPageId="7" captionText="Fig. 6. SEM images of rods from tentacles of Elpidia soyoae sp. nov. (NSMT E-12635: paratype). Rods with arched axis, especially shortened horizontal arms, and vertical apophyses. Abbreviations: ax, axis; ha, horizontal arms; va, vertical apophyses." figureDoi="http://doi.org/10.5281/zenodo.5738572" httpUri="https://zenodo.org/record/5738572/files/figure.png" pageId="7" pageNumber="160">Fig. 6</figureCitation>
|
||
). Rods 77– 776 µm in length (TOL, N=223 from 5 individuals), with one arched axis, shortened arms and/or shortened apophyses sometimes completely lacking arms and apophyses. Axis in diameter 4.2–39µm (N=243 from 5 individuals). The height of apophyses 2.7–227 µm (TAH, N=380 from 5 individuals), TAH/TOL ratio between 0.0077–0.58 (N=380 from 5 individuals).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF2A6650AE7FDC12FF1CDEEDFB9AF7A7" ID-DOI="http://doi.org/10.5281/zenodo.5738570" ID-Zenodo-Dep="5738570" httpUri="https://zenodo.org/record/5738570/files/figure.png" pageId="6" pageNumber="159" startId="6.[156,187,2028,2050]" targetBox="[189,1398,890,1987]" targetPageId="6">
|
||
<paragraph id="8BEA36D8AE7FDC12FF1CDEEDFB9AF7A7" blockId="6.[156,1431,2028,2078]" pageId="6" pageNumber="159">
|
||
Fig. 5. SEM images of ossicles from ventral body wall of
|
||
<taxonomicName id="4C554D5BAE7FDC12FD57DEEDFCA6F7B8" authority="Ogawa & Morita & Fujita, 2020" authorityName="Ogawa & Morita & Fujita" authorityYear="2020" box="[727,859,2028,2049]" class="Holothuroidea" family="Elpidiidae" genus="Elpidia" kingdom="Animalia" order="Elasipodida" pageId="6" pageNumber="159" phylum="Echinodermata" rank="species" species="soyoae" status="sp. nov.">
|
||
<emphasis id="B921EACAAE7FDC12FD57DEEDFCA6F7B8" box="[727,859,2028,2049]" italics="true" pageId="6" pageNumber="159">Elpidia soyoae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21257B1AE7FDC12FCE4DEEDFC50F7B8" box="[868,941,2028,2049]" pageId="6" pageNumber="159" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(holotype). Rods with straight axis, well developed horizontal arms, and vertical apophyses. Abbreviations: ax, axis; ha, horizontal arms; va, vertical apophyses.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C34F6553AE7EDC13FF35DB2DFE11FDDD" pageId="7" pageNumber="160" type="distribution">
|
||
<paragraph id="8BEA36D8AE7EDC13FF35DB2DFE11FDDD" blockId="7.[156,768,198,1003]" pageId="7" pageNumber="160">
|
||
<emphasis id="B921EACAAE7EDC13FF35DB2DFEBCFDFD" bold="true" box="[181,321,556,580]" pageId="7" pageNumber="160">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="54EE887AAE7EDC13FDB8DB2CFD99FDFC" box="[568,612,557,581]" pageId="7" pageNumber="160">type</typeStatus>
|
||
locality:
|
||
<collectingCountry id="F3427648AE7EDC13FD46DB2CFD03FDFD" box="[710,766,557,580]" name="Japan" pageId="7" pageNumber="160">Japan</collectingCountry>
|
||
Trench area, depth
|
||
<quantity id="4CAD9B3DAE7EDC13FEDFDB4CFE1AFDDD" box="[351,487,589,613]" metricMagnitude="3" metricUnit="m" metricValue="3.8575000000000004" metricValueMax="4.1450000000000005" metricValueMin="3.5700000000000003" pageId="7" pageNumber="160" unit="m" value="3857.5" valueMax="4145.0" valueMin="3570.0">3570–4145 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34F6553AE7EDC13FF35DB6CFDDBFD1F" pageId="7" pageNumber="160" type="etymology">
|
||
<paragraph id="8BEA36D8AE7EDC13FF35DB6CFDDBFD1F" blockId="7.[156,768,198,1003]" pageId="7" pageNumber="160">
|
||
<emphasis id="B921EACAAE7EDC13FF35DB6CFED0FD3C" bold="true" box="[181,301,621,645]" pageId="7" pageNumber="160">Etymology.</emphasis>
|
||
The species is named after the R/V
|
||
<emphasis id="B921EACAAE7EDC13FD48DB6FFF2EFD1F" italics="true" pageId="7" pageNumber="160">Soyomaru</emphasis>
|
||
which collected these specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34F6553AE7EDC1CFF35DBAFFEF6FD3A" lastPageId="8" lastPageNumber="161" pageId="7" pageNumber="160" type="discussion">
|
||
<paragraph id="8BEA36D8AE7EDC13FF35DBAFFAC9FDBA" blockId="7.[156,768,198,1003]" lastBlockId="7.[820,1432,198,1003]" pageId="7" pageNumber="160">
|
||
<emphasis id="B921EACAAE7EDC13FF35DBAFFEE4FD7F" bold="true" box="[181,281,686,710]" pageId="7" pageNumber="160">Remarks.</emphasis>
|
||
Morphological features of examined specimens were well coincident with five diagnostic characters of the genus
|
||
<taxonomicName id="4C554D5BAE7EDC13FE9EDBEEFE98FCBE" authorityName="Theel" authorityYear="1876" box="[286,357,751,775]" class="Holothuroidea" family="Elpidiidae" genus="Elpidia" kingdom="Animalia" order="Elasipodida" pageId="7" pageNumber="160" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B921EACAAE7EDC13FE9EDBEEFE98FCBE" box="[286,357,751,775]" italics="true" pageId="7" pageNumber="160">Elpidia</emphasis>
|
||
</taxonomicName>
|
||
(see the Diagnosis section for the genus above). The dorsal rods of the examined specimens ranged from 8.1 to 33 µm in axis diameter, indicating
|
||
<taxonomicName id="4C554D5BAE7EDC13FDF0DA31FD03FCF1" authority="Ogawa & Morita & Fujita, 2020" authorityName="Ogawa & Morita & Fujita" authorityYear="2020" box="[624,766,816,840]" class="Holothuroidea" family="Elpidiidae" genus="Elpidia" kingdom="Animalia" order="Elasipodida" pageId="7" pageNumber="160" phylum="Echinodermata" rank="species" species="soyoae" status="sp. nov.">
|
||
<emphasis id="B921EACAAE7EDC13FDF0DA31FD03FCF1" box="[624,766,816,840]" italics="true" pageId="7" pageNumber="160">Elpidia soyoae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21257B1AE7EDC13FF1CDA50FF16FCD1" box="[156,235,849,872]" pageId="7" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
|
||
belong to the slender axis group. A morphological comparison of
|
||
<taxonomicName id="4C554D5BAE7EDC13FEB5DA73FE6DFC30" authority="Ogawa & Morita & Fujita, 2020" authorityName="Ogawa & Morita & Fujita" authorityYear="2020" box="[309,400,882,905]" class="Holothuroidea" family="Elpidiidae" genus="Elpidia" kingdom="Animalia" order="Elasipodida" pageId="7" pageNumber="160" phylum="Echinodermata" rank="species" species="soyoae" status="sp. nov.">
|
||
<emphasis id="B921EACAAE7EDC13FEB5DA73FE6DFC30" box="[309,400,882,905]" italics="true" pageId="7" pageNumber="160">E. soyoae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21257B1AE7EDC13FE17DA73FE19FC30" box="[407,484,882,905]" pageId="7" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
|
||
with six species of the slen- der axis group is provided in
|
||
<tableCitation id="C6D70363AE7EDC13FE48DA93FDEEFC13" box="[456,531,914,938]" captionStart="Table 2" captionStartId="8.[156,204,592,615]" captionText="Table 2. Morphological comparison of Elpidia soyoae sp. nov. with Elpidia species in the slender axis group. DAH and DOL indicate apophyses height and ossicle length of rods in dorsal body wall," pageId="7" pageNumber="160">Table 2</tableCitation>
|
||
.
|
||
<taxonomicName id="4C554D5BAE7EDC13FD9DDA93FD51FC13" authority="Ogawa & Morita & Fujita, 2020" authorityName="Ogawa & Morita & Fujita" authorityYear="2020" box="[541,684,914,938]" class="Holothuroidea" family="Elpidiidae" genus="Elpidia" kingdom="Animalia" order="Elasipodida" pageId="7" pageNumber="160" phylum="Echinodermata" rank="species" species="soyoae" status="sp. nov.">
|
||
<emphasis id="B921EACAAE7EDC13FD9DDA93FD51FC13" box="[541,684,914,938]" italics="true" pageId="7" pageNumber="160">Elpidia soyoae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21257B1AE7EDC13FD33DA93FCFDFC10" box="[691,768,914,937]" pageId="7" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is distinguished from other species by the following four features: i) dorsal papillae on two dorsal radii, four or five paired papillae and unpaired papillae present along the entire dorsal radii (four to seven papillae on each radius), with wide separation between second and third paired papillae, ii) maximum length of dorsal rods exceeds 1000 µm, iii) tentacle rods with arched axis and shortened or occasionally completely reduced arms and apophyses, and iv) purple pigmentation spots composed of small purple particles on dorsal and ventral body walls. In addition, this species is easily distinguished from non-grouped species,
|
||
<taxonomicName id="4C554D5BAE7EDC13FB5ED8CAFAB0FE5B" baseAuthorityName="Perrier" baseAuthorityYear="1896" box="[1246,1357,458,482]" class="Holothuroidea" family="Elpidiidae" genus="Elpidia" kingdom="Animalia" order="Elasipodida" pageId="7" pageNumber="160" phylum="Echinodermata" rank="species" species="echinata">
|
||
<emphasis id="B921EACAAE7EDC13FB5ED8CAFAB0FE5B" box="[1246,1357,458,482]" italics="true" pageId="7" pageNumber="160">E. echinata</emphasis>
|
||
</taxonomicName>
|
||
, which has 3 papillae on each dorsal radius (
|
||
<bibRefCitation id="EFC44B29AE7EDC13FB26D8EAFAD4FDBA" author="Perrier, R." box="[1190,1321,491,515]" pageId="7" pageNumber="160" pagination="900 - 903" refId="ref5866" refString="Perrier, R. 1896. Sur les Elasipodes recueillis par le Travailleur et le Talisman. Comptes Rendus Hebdomadaires des Seances de l'Academie des Sciences 123: 900 - 903." type="journal article" year="1896">Perrier 1896</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BEA36D8AE7EDC13FCCEDB0DFBCEFC52" blockId="7.[820,1432,198,1003]" pageId="7" pageNumber="160">
|
||
<emphasis id="B921EACAAE7EDC13FCCEDB0DFB7BFD9A" bold="true" box="[846,1158,523,548]" pageId="7" pageNumber="160">Molecular data comparison.</emphasis>
|
||
<materialsCitation id="3B3D3C85AE7EDC13FB14DB0DFB93FD1F" accessionNumber="HM196413, HM196420" collectionCode="BLAST, INSD, HM" pageId="7" pageNumber="160" specimenCount="1">
|
||
In
|
||
<collectionCode id="ED44AE1DAE7EDC13FB38DB0DFAFCFD9D" box="[1208,1281,524,548]" pageId="7" pageNumber="160">BLAST</collectionCode>
|
||
searches, the closest hits for the five obtained partial COI gene sequences were those of
|
||
<taxonomicName id="4C554D5BAE7EDC13FC69DB4CFB75FDDC" authorityName="Theel" authorityYear="1876" box="[1001,1160,589,613]" class="Holothuroidea" family="Elpidiidae" genus="Elpidia" kingdom="Animalia" order="Elasipodida" pageId="7" pageNumber="160" phylum="Echinodermata" rank="species" species="glacialis">
|
||
<emphasis id="B921EACAAE7EDC13FC69DB4CFB75FDDC" box="[1001,1160,589,613]" italics="true" pageId="7" pageNumber="160">Elpidia glacialis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<collectionCode id="ED44AE1DAE7EDC13FB19DB4CFB28FDDC" box="[1177,1237,589,613]" pageId="7" pageNumber="160">INSD</collectionCode>
|
||
accession number
|
||
<accessionNumber id="9406AB3BAE7EDC13FCB4DB6CFC55FD3C" box="[820,936,621,645]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/HM196413" pageId="7" pageNumber="160">
|
||
<collectionCode id="ED44AE1DAE7EDC13FCB4DB6CFCA0FD3C" box="[820,861,621,645]" country="0" name="Hastings Museum" pageId="7" pageNumber="160">HM</collectionCode>
|
||
196413
|
||
</accessionNumber>
|
||
or
|
||
<accessionNumber id="9406AB3BAE7EDC13FC51DB6CFBBBFD3C" box="[977,1094,621,645]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/HM196420" pageId="7" pageNumber="160">
|
||
<collectionCode id="ED44AE1DAE7EDC13FC51DB6CFC07FD3C" box="[977,1018,621,645]" country="0" name="Hastings Museum" pageId="7" pageNumber="160">HM</collectionCode>
|
||
196420
|
||
</accessionNumber>
|
||
, with 90.1% and 90.8% similarity in 98% to 100% coverage)
|
||
</materialsCitation>
|
||
. K2P genetic distances were calculated in terms of 603 bps COI partial sequences in the five obtained sequences and 17
|
||
<taxonomicName id="4C554D5BAE7EDC13FB23DBCEFAF3FD5E" authorityName="Theel" authorityYear="1876" box="[1187,1294,719,743]" class="Holothuroidea" family="Elpidiidae" genus="Elpidia" kingdom="Animalia" order="Elasipodida" pageId="7" pageNumber="160" phylum="Echinodermata" rank="species" species="glacialis">
|
||
<emphasis id="B921EACAAE7EDC13FB23DBCEFAF3FD5E" box="[1187,1294,719,743]" italics="true" pageId="7" pageNumber="160">
|
||
E.
|
||
<materialsCitation id="3B3D3C85AE7EDC13FB3FDBCEFAF3FD5E" accessionNumber="HM196413" box="[1215,1294,719,743]" collectionCode="INSD" latitude="0.101" longLatPrecision="78" longitude="-0.12" pageId="7" pageNumber="160" specimenCode="HM196413-196429" specimenCount="1">glacialis</materialsCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
sequences in GenBank (INSD accession numbers
|
||
<specimenCode id="DBF39EA3AE7EDC13FB40DBEEFA73FCBE" box="[1216,1422,751,775]" collectionCode="HM" country="0" name="Hastings Museum" pageId="7" pageNumber="160">
|
||
<accessionNumber id="9406AB3BAE7EDC13FB40DBEEFAC9FCBE" box="[1216,1332,751,775]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/HM196413" pageId="7" pageNumber="160">HM196413</accessionNumber>
|
||
–196429
|
||
</specimenCode>
|
||
): the interspecific K2P distance was
|
||
<geoCoordinate id="EE61501FAE7EDC13FB1CDA11FB2FFC91" box="[1180,1234,784,808]" degrees="0.101" direction="north" orientation="latitude" pageId="7" pageNumber="160" precision="55" value="0.101">0.101</geoCoordinate>
|
||
<geoCoordinate id="EE61501FAE7EDC13FB52DA11FAEBFC91" box="[1234,1302,784,808]" degrees="0.120" direction="west" orientation="longitude" pageId="7" pageNumber="160" precision="55" value="-0.12">–0.120</geoCoordinate>
|
||
between
|
||
<taxonomicName id="4C554D5BAE7EDC13FAFADA11FC4CFCF1" authority="Ogawa & Morita & Fujita, 2020" authorityName="Ogawa & Morita & Fujita" authorityYear="2020" class="Holothuroidea" family="Elpidiidae" genus="Elpidia" kingdom="Animalia" order="Elasipodida" pageId="7" pageNumber="160" phylum="Echinodermata" rank="species" species="soyoae" status="sp. nov.">
|
||
<emphasis id="B921EACAAE7EDC13FAFADA11FC4CFCF1" italics="true" pageId="7" pageNumber="160">Elpidia soyoae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21257B1AE7EDC13FC3ADA30FBF4FCF1" box="[954,1033,817,840]" pageId="7" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C554D5BAE7EDC13FBC0DA30FB51FCF1" authorityName="Theel" authorityYear="1876" box="[1088,1196,816,840]" class="Holothuroidea" family="Elpidiidae" genus="Elpidia" kingdom="Animalia" order="Elasipodida" pageId="7" pageNumber="160" phylum="Echinodermata" rank="species" species="glacialis">
|
||
<emphasis id="B921EACAAE7EDC13FBC0DA30FB51FCF1" box="[1088,1196,816,840]" italics="true" pageId="7" pageNumber="160">E. glacialis</emphasis>
|
||
</taxonomicName>
|
||
, and the intra-specific distance was
|
||
<geoCoordinate id="EE61501FAE7EDC13FC3EDA50FC08FCD0" box="[958,1013,849,873]" degrees="0.005" direction="north" orientation="latitude" pageId="7" pageNumber="160" precision="55" value="0.005">0.005</geoCoordinate>
|
||
<geoCoordinate id="EE61501FAE7EDC13FC75DA50FBC5FCD0" box="[1013,1080,849,873]" degrees="0.017" direction="west" orientation="longitude" pageId="7" pageNumber="160" precision="55" value="-0.017">–0.017</geoCoordinate>
|
||
and
|
||
<quantity id="4CAD9B3DAE7EDC13FBEFDA50FB21FCD0" box="[1135,1244,849,873]" metricMagnitude="-4" metricUnit="m" metricValue="2.413" metricValueMax="4.826" metricValueMin="0.0" pageId="7" pageNumber="160" unit="in" value="0.0095" valueMax="0.019" valueMin="0.0">0–0.019 in</quantity>
|
||
<taxonomicName id="4C554D5BAE7EDC13FB64DA53FABCFCD0" authority="Ogawa & Morita & Fujita, 2020" authorityName="Ogawa & Morita & Fujita" authorityYear="2020" box="[1252,1345,850,873]" class="Holothuroidea" family="Elpidiidae" genus="Elpidia" kingdom="Animalia" order="Elasipodida" pageId="7" pageNumber="160" phylum="Echinodermata" rank="species" species="soyoae" status="sp. nov.">
|
||
<emphasis id="B921EACAAE7EDC13FB64DA53FABCFCD0" box="[1252,1345,850,873]" italics="true" pageId="7" pageNumber="160">E. soyoae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21257B1AE7EDC13FACADA50FA65FCD1" box="[1354,1432,849,872]" pageId="7" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C554D5BAE7EDC13FCE2DA73FC30FC30" authorityName="Theel" authorityYear="1876" box="[866,973,881,905]" class="Holothuroidea" family="Elpidiidae" genus="Elpidia" kingdom="Animalia" order="Elasipodida" pageId="7" pageNumber="160" phylum="Echinodermata" rank="species" species="glacialis">
|
||
<emphasis id="B921EACAAE7EDC13FCE2DA73FC30FC30" box="[866,973,881,905]" italics="true" pageId="7" pageNumber="160">E. glacialis</emphasis>
|
||
</taxonomicName>
|
||
, respectively. The results confirmed that
|
||
<specimenCount id="9D53FD51AE7EDC13FAF4DA70FC60FC13" pageId="7" pageNumber="160" type="generic">five specimens</specimenCount>
|
||
belong to the genus
|
||
<taxonomicName id="4C554D5BAE7EDC13FBFCDA93FB3EFC13" authorityName="Theel" authorityYear="1876" box="[1148,1219,914,938]" class="Holothuroidea" family="Elpidiidae" genus="Elpidia" kingdom="Animalia" order="Elasipodida" pageId="7" pageNumber="160" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B921EACAAE7EDC13FBFCDA93FB3EFC13" box="[1148,1219,914,938]" italics="true" pageId="7" pageNumber="160">Elpidia</emphasis>
|
||
</taxonomicName>
|
||
, and supported that
|
||
<taxonomicName id="4C554D5BAE7EDC13FCB4DAB2FC6EFC73" authority="Ogawa & Morita & Fujita, 2020" authorityName="Ogawa & Morita & Fujita" authorityYear="2020" box="[820,915,947,970]" class="Holothuroidea" family="Elpidiidae" genus="Elpidia" kingdom="Animalia" order="Elasipodida" pageId="7" pageNumber="160" phylum="Echinodermata" rank="species" species="soyoae" status="sp. nov.">
|
||
<emphasis id="B921EACAAE7EDC13FCB4DAB2FC6EFC73" box="[820,915,947,970]" italics="true" pageId="7" pageNumber="160">E. soyoae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21257B1AE7EDC13FC1EDAB2FC13FC73" box="[926,1006,947,970]" pageId="7" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C554D5BAE7EDC13FBA6DAB2FB6EFC73" authorityName="Theel" authorityYear="1876" box="[1062,1171,946,970]" class="Holothuroidea" family="Elpidiidae" genus="Elpidia" kingdom="Animalia" order="Elasipodida" pageId="7" pageNumber="160" phylum="Echinodermata" rank="species" species="glacialis">
|
||
<emphasis id="B921EACAAE7EDC13FBA6DAB2FB6EFC73" box="[1062,1171,946,970]" italics="true" pageId="7" pageNumber="160">E. glacialis</emphasis>
|
||
</taxonomicName>
|
||
are genetically independent and distinct species.
|
||
</paragraph>
|
||
<caption id="DF2A6650AE7EDC13FF1CDEEDFB3AF7A7" ID-DOI="http://doi.org/10.5281/zenodo.5738572" ID-Zenodo-Dep="5738572" httpUri="https://zenodo.org/record/5738572/files/figure.png" pageId="7" pageNumber="160" startId="7.[156,187,2028,2050]" targetBox="[189,1398,1063,1987]" targetPageId="7">
|
||
<paragraph id="8BEA36D8AE7EDC13FF1CDEEDFB3AF7A7" blockId="7.[156,1431,2028,2078]" pageId="7" pageNumber="160">
|
||
Fig. 6. SEM images of rods from tentacles of
|
||
<taxonomicName id="4C554D5BAE7EDC13FDEADEEDFD13F7B8" authority="Ogawa & Morita & Fujita, 2020" authorityName="Ogawa & Morita & Fujita" authorityYear="2020" box="[618,750,2028,2049]" class="Holothuroidea" family="Elpidiidae" genus="Elpidia" kingdom="Animalia" order="Elasipodida" pageId="7" pageNumber="160" phylum="Echinodermata" rank="species" species="soyoae" status="sp. nov.">
|
||
<emphasis id="B921EACAAE7EDC13FDEADEEDFD13F7B8" box="[618,750,2028,2049]" italics="true" pageId="7" pageNumber="160">Elpidia soyoae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21257B1AE7EDC13FD78DEEDFCBFF7B8" box="[760,834,2028,2049]" pageId="7" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(NSMT E-12635: paratype). Rods with arched axis, especially shortened horizontal arms, and vertical apophyses. Abbreviations: ax, axis; ha, horizontal arms; va, vertical apophyses.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2A6650AE71DC1CFF1CDB51F804FDDE" box="[156,2041,592,615]" pageId="8" pageNumber="161" startId="8.[156,204,592,615]">
|
||
<paragraph id="8BEA36D8AE71DC1CFF1CDB51F804FDDE" blockId="8.[156,2041,592,643]" box="[156,2041,592,615]" pageId="8" pageNumber="161">
|
||
Table 2. Morphological comparison of
|
||
<taxonomicName id="4C554D5BAE71DC1CFD9BDB51FD63FDDE" authority="Ogawa & Morita & Fujita, 2020" authorityName="Ogawa & Morita & Fujita" authorityYear="2020" box="[539,670,592,615]" class="Holothuroidea" family="Elpidiidae" genus="Elpidia" kingdom="Animalia" order="Elasipodida" pageId="8" pageNumber="161" phylum="Echinodermata" rank="species" species="soyoae" status="sp. nov.">
|
||
<emphasis id="B921EACAAE71DC1CFD9BDB51FD63FDDE" box="[539,670,592,615]" italics="true" pageId="8" pageNumber="161">Elpidia soyoae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21257B1AE71DC1CFD26DB51FD13FDDE" box="[678,750,592,615]" pageId="8" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
|
||
with
|
||
<taxonomicName id="4C554D5BAE71DC1CFCA6DB51FC53FDDE" box="[806,942,592,615]" class="Holothuroidea" family="Elpidiidae" genus="Elpidia" kingdom="Animalia" order="Elasipodida" pageId="8" pageNumber="161" phylum="Echinodermata" rank="species" species="undetermined">
|
||
<emphasis id="B921EACAAE71DC1CFCA6DB51FC98FDDE" box="[806,869,592,615]" italics="true" pageId="8" pageNumber="161">Elpidia</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
in the slender axis group. DAH and DOL indicate apophyses height and ossicle length of rods in dorsal body wall,
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BEA36D8AE71DC1CFF1CDB6DFEF6FD3A" blockId="8.[156,2041,592,643]" box="[156,267,620,643]" pageId="8" pageNumber="161">respectively.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |