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<document id="7F680830AED77DEF0481F0AD96A29041" ID-CLB-Dataset="45889" ID-DOI="10.5281/zenodo.279337" ID-GBIF-Dataset="367d3d94-fa26-4f5d-ba39-1fb904401752" ID-ISSN="1175-5326" ID-Zenodo-Dep="279337" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460414521621" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2011" docId="03DF87B01F528E0A86C5FA1B242AFB71" docLanguage="en" docName="zt03113p052.pdf" docOrigin="Zootaxa 3113" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Cephennula sibugana Jałoszyński, 2011, sp. n." docType="treatment" docVersion="8" lastPageNumber="50" masterDocId="FFE6FFC81F5F8E048652FFD7274EFFD1" masterDocTitle="Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia" masterLastPageNumber="52" masterPageNumber="36" pageNumber="49" updateTime="1698328417371" updateUser="ExternalLinkService">
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<mods:title id="37211B0A0AE8C83FCB9E36D304FD4742">Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia</mods:title>
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<mods:namePart id="28A6D6467DFC6081202DDAA987049B3A">Jałoszyński, Paweł</mods:namePart>
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<mods:date id="18DB8DDD380DA2CF03446677EB15A23B">2011</mods:date>
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<treatment id="03DF87B01F528E0A86C5FA1B242AFB71" ID-DOI="http://doi.org/10.5281/zenodo.6185253" ID-GBIF-Taxon="119424603" ID-Zenodo-Dep="6185253" LSID="urn:lsid:plazi:treatment:03DF87B01F528E0A86C5FA1B242AFB71" httpUri="http://treatment.plazi.org/id/03DF87B01F528E0A86C5FA1B242AFB71" lastPageId="14" lastPageNumber="50" pageId="13" pageNumber="49">
<subSubSection id="C36C652D1F528E0986C5FA1B2616F9D7" pageId="13" pageNumber="49" type="nomenclature">
<paragraph id="8BC936A61F528E0986C5FA1B26A4FA34" blockId="13.[151,490,1483,1542]" box="[151,490,1483,1510]" pageId="13" pageNumber="49">
<heading id="D08181CA1F528E0986C5FA1B26A4FA34" bold="true" box="[151,490,1483,1510]" fontSize="11" level="1" pageId="13" pageNumber="49" reason="1">
<emphasis id="B902EAB41F528E0986C5FA1B26A4FA34" bold="true" box="[151,490,1483,1510]" pageId="13" pageNumber="49">
<taxonomicName id="4C764D251F528E0986C5FA1B26EFFA37" ID-CoL="SFQX" box="[151,417,1484,1510]" class="Insecta" family="Staphylinidae" genus="Cephennula" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="49" phylum="Arthropoda" rank="species" species="sibugana" status="sp. nov.">
<emphasis id="B902EAB41F528E0986C5FA1B26EFFA37" bold="true" box="[151,417,1484,1510]" italics="true" pageId="13" pageNumber="49">Cephennula sibugana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157CF1F528E0987FAFA1C26A4FA34" box="[424,490,1483,1509]" pageId="13" pageNumber="49" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC936A61F528E0986C5FA392616F9D7" blockId="13.[151,490,1483,1542]" box="[151,344,1518,1542]" pageId="13" pageNumber="49">
(
<figureCitation id="134D2A231F528E0986CDFA3927B3F9D7" box="[159,253,1518,1542]" captionStart="FIGURES 8 12" captionStartId="9.[151,264,1930,1953]" captionTargetBox="[197,1397,207,1882]" captionTargetId="figure@9.[181,1405,193,1909]" captionTargetPageId="9" captionText="FIGURES 8 12. Habitus of males. 8 ― Cephennula titiwangsana sp. n.; 9 ― Cephennula medioglabra sp. n.; 10 ― Cephennula sibugana sp. n.; 11 ― Cephennula gombakiana sp. n.; 12 ― Cephennula micropunctata sp. n. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/279340/files/figure.png" pageId="13" pageNumber="49">Figs. 10</figureCitation>
,
<figureCitation id="134D2A231F528E09875AFA392601F9D7" box="[264,335,1518,1542]" captionStart="FIGURES 25 32" captionStartId="11.[151,264,1617,1640]" captionTargetBox="[170,1423,185,1557]" captionTargetId="figure@11.[151,1436,157,1596]" captionTargetPageId="11" captionText="FIGURES 25 32. Aedeagus in ventral (25, 27, 29, 31) and lateral (26, 28, 30, 32) views. 25, 26 ― Cephennula medioglabra sp. n.; 27, 28 ― Cephennula sibugana sp. n.; 29, 30 ― Cephennula gombakiana sp. n.; 31, 32 ― Cephennula micropunctata sp. n. Scale bars: 0.05 mm." httpUri="https://zenodo.org/record/279341/files/figure.png" pageId="13" pageNumber="49">27, 28</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36C652D1F528E0986C5F9E0227DF947" pageId="13" pageNumber="49" type="materials_examined">
<paragraph id="8BC936A61F528E0986C5F9E0227DF947" blockId="13.[151,1436,1590,2010]" pageId="13" pageNumber="49">
<emphasis id="B902EAB41F528E0986C5F9E02681F99E" bold="true" box="[151,463,1590,1615]" pageId="13" pageNumber="49">
<typeStatus id="54CD88041F528E0986C5F9E02646F99E" box="[151,264,1591,1615]" pageId="13" pageNumber="49" type="holotype">Holotype</typeStatus>
: E
<collectingCountry id="F36176361F528E09877DF9E12684F99E" box="[303,458,1590,1615]" name="Malaysia" pageId="13" pageNumber="49">MALAYSIA</collectingCountry>
:
</emphasis>
3, two labels: &quot;Borneo Sabah \ Sepilok Sibuga Forest Res. \ nr. Sandakan
<date id="FFC810661F528E098371F9E122C2F99F" box="[1315,1420,1590,1614]" pageId="13" pageNumber="49" value="1968-06-09">9.vi.1968</date>
\ 68.378 \ R.W. Tazlor&quot; [white, partly printed, “Sepilok” crossed out, “Sibuga” and numbers handwritten in black ink]; &quot;
<emphasis id="B902EAB41F528E09868CF9A926DCF947" box="[222,402,1662,1686]" italics="true" pageId="13" pageNumber="49">CEPHENNULA</emphasis>
\
<taxonomicName id="4C764D251F528E0987FAF9A92541F947" box="[424,527,1662,1686]" class="Insecta" family="Staphylinidae" genus="Cephennula" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="49" phylum="Arthropoda" rank="species" species="sibugana">
<emphasis id="B902EAB41F528E0987FAF9A92541F947" box="[424,527,1662,1686]" italics="true" pageId="13" pageNumber="49">sibugana</emphasis>
</taxonomicName>
m. \ det. P. Jałoszyński, 2011 \
<typeStatus id="54CD88041F528E09853DF9A92357F947" box="[879,1049,1662,1686]" pageId="13" pageNumber="49" type="holotype">HOLOTYPUS</typeStatus>
&quot; [red, printed] (
<collectionCode id="ED67AE631F528E098298F9A92268F947" box="[1226,1318,1662,1686]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="13" pageNumber="49">MHNG</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C36C652D1F528E098695F9752619F8BF" pageId="13" pageNumber="49" type="diagnosis">
<paragraph id="8BC936A61F528E098695F9752619F8BF" blockId="13.[151,1436,1590,2010]" pageId="13" pageNumber="49">
<emphasis id="B902EAB41F528E098695F975260FF96A" bold="true" box="[199,321,1698,1723]" pageId="13" pageNumber="49">Diagnosis.</emphasis>
Body length between 0.7 and
<quantity id="4C8E9B431F528E0984F3F97525B5F96B" box="[673,763,1698,1722]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="13" pageNumber="49" unit="mm" value="0.8">0.8 mm</quantity>
; antennal club strongly flattened and compact; head nearly evenly covered with large, deep and dense punctures; punctures on median part of pronotum moderately large, dense; sublateral pronotal carinae as long as pronotum; ante-basal pronotal groove with distinct median fovea; subhumeral carinae nearly as long as 1/
<date id="FFC810661F528E098475F8D9250AF8F7" box="[551,580,1806,1830]" pageId="13" pageNumber="49">2x</date>
EL, humeral carinae less distinct and about half shorter; aedeagus in ventral view bottle-shaped with subtrapezoidal and rounded apex; endophallus composed of lightly pigmented elongate tubular structure.
</paragraph>
</subSubSection>
<subSubSection id="C36C652D1F528E0A8695F8AD26D0FB89" lastPageId="14" lastPageNumber="50" pageId="13" pageNumber="49" type="description">
<paragraph id="8BC936A61F528E098695F8AD26CCF80B" blockId="13.[151,1436,1590,2010]" pageId="13" pageNumber="49">
<emphasis id="B902EAB41F528E098695F8AD2617F842" bold="true" box="[199,345,1914,1939]" pageId="13" pageNumber="49">Description.</emphasis>
Body of male (
<figureCitation id="134D2A231F528E098455F8AD2517F843" box="[519,601,1914,1938]" captionStart="FIGURES 8 12" captionStartId="9.[151,264,1930,1953]" captionTargetBox="[197,1397,207,1882]" captionTargetId="figure@9.[181,1405,193,1909]" captionTargetPageId="9" captionText="FIGURES 8 12. Habitus of males. 8 ― Cephennula titiwangsana sp. n.; 9 ― Cephennula medioglabra sp. n.; 10 ― Cephennula sibugana sp. n.; 11 ― Cephennula gombakiana sp. n.; 12 ― Cephennula micropunctata sp. n. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/279340/files/figure.png" pageId="13" pageNumber="49">Fig. 10</figureCitation>
) strongly convex, elongate, with shallow but distinct constriction between pronotum and elytra, moderately glossy, dark brown, covered with yellowish vestiture, legs and antennae slightly lighter. BL
<quantity id="4C8E9B431F528E098745F8152630F80B" box="[279,382,1986,2010]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="13" pageNumber="49" unit="mm" value="0.75">0.75 mm</quantity>
.
</paragraph>
<paragraph id="8BC936A61F518E0A8695FF40268BFE1E" blockId="14.[151,1437,151,1184]" pageId="14" pageNumber="50">
Head broadest at large, coarsely faceted and strongly convex eyes, HL
<quantity id="4C8E9B431F518E0A85ACFF402328FF7E" box="[1022,1126,151,175]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="50" unit="mm" value="0.1">0.10 mm</quantity>
, HW
<quantity id="4C8E9B431F518E0A82FEFF402258FF7E" box="[1196,1302,151,175]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="14" pageNumber="50" unit="mm" value="0.19">0.19 mm</quantity>
; vertex and frontoclypeal area convex; supraantennal tubercles barely marked. Punctures on head dorsum nearly equally distributed, large, deep and dense; setae very short and sparse, nearly recumbent. Antennae with strongly enlarged, strongly flattened and compactly assembled antennomeres XXI and only slightly enlarged antennomere IX, so that club appears two-segmented, AnL
<quantity id="4C8E9B431F518E0A840EFEF02588FEEE" box="[604,710,295,319]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="14" pageNumber="50" unit="mm" value="0.25">0.25 mm</quantity>
; antennomere I about 1.3x as long as broad; II 2.2x as long as broad; III and IV each about 1.1x as long as broad; V 1.2x as long as broad; VI 1.1x as long as broad; VII 1.2x as long as broad; VIII slightly broader but shorter than VII, as long as broad; IX distinctly broader and slightly longer than VII, distinctly transverse; X much broader and longer than IX, strongly transverse; XI about as broad as X but only 1.1x as long as broad.
</paragraph>
<paragraph id="8BC936A61F518E0A8695FE0A24FCFD3E" blockId="14.[151,1437,151,1184]" pageId="14" pageNumber="50">
Pronotum approximately semielliptical, broadest between middle and anterior 1/3; PL
<quantity id="4C8E9B431F518E0A82DDFE0B23B8FE25" box="[1167,1270,476,500]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="14" pageNumber="50" unit="mm" value="0.25">0.25 mm</quantity>
, PW
<quantity id="4C8E9B431F518E0A8363FE0B22D6FE25" box="[1329,1432,476,500]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="14" pageNumber="50" unit="mm" value="0.31">0.31 mm</quantity>
, anterior margin weakly rounded; lateral margins distinctly microserrate, broadly rounded in anterior 2/3, straight and parallel in posterior 1/4; hind angles sharp and acute; posterior margin very weakly bisinuate; sublateral carinae distinct and as long as pronotum.. Transverse ante-basal groove shallow and indistinct, slightly arcuate and with distinct median foveae; sublateral ante-basal foveae small and indistinct; lateral ante-basal foveae larger and deeper. Punctures on median part of pronotal disc slightly smaller and shallower than those on head but distinct and dense, areas between sublateral carinae and lateral margins of pronotum covered with much larger and denser, coarse punctures; setae sparse, moderately long, only slightly suberect.
</paragraph>
<paragraph id="8BC936A61F518E0A8695FD2B263EFC75" blockId="14.[151,1437,151,1184]" pageId="14" pageNumber="50">
Elytra strongly convex, oval, broadest between middle and anterior 1/3; EL
<quantity id="4C8E9B431F518E0A8274FD2B23C3FCC5" box="[1062,1165,764,788]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="50" unit="mm" value="0.4">0.40 mm</quantity>
, EW
<quantity id="4C8E9B431F518E0A829FFD2B227AFCC5" box="[1229,1332,764,788]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="14" pageNumber="50" unit="mm" value="0.35">0.35 mm</quantity>
, EI 1.14; base of each elytron with rudimentary basal fovea located much closer to scutellum than to base of humeral carina; subhumeral carinae very distinct and nearly as long as 1/
<date id="FFC810661F518E0A8548FC93247BFC8D" box="[794,821,836,860]" pageId="14" pageNumber="50">2x</date>
EL, nearly twice as long as less distinct humeral carinae. Punctures on elytra very fine and very shallow, barely discernible under 100x magnification; setae similar as those on pronotum.
</paragraph>
<paragraph id="8BC936A61F518E0A8695FC66226AFC16" blockId="14.[151,1437,151,1184]" box="[199,1316,943,969]" pageId="14" pageNumber="50">Legs moderately long and slender; pro- and mesotibiae nearly straight, metatibiae distinctly curved.</paragraph>
<paragraph id="8BC936A61F518E0A8695FC032258FBE5" blockId="14.[151,1437,151,1184]" pageId="14" pageNumber="50">
Aedeagus (
<figureCitation id="134D2A231F518E0A8719FC032693FC3D" box="[331,477,980,1004]" captionStart="FIGURES 25 32" captionStartId="11.[151,264,1617,1640]" captionTargetBox="[170,1423,185,1557]" captionTargetId="figure@11.[151,1436,157,1596]" captionTargetPageId="11" captionText="FIGURES 25 32. Aedeagus in ventral (25, 27, 29, 31) and lateral (26, 28, 30, 32) views. 25, 26 ― Cephennula medioglabra sp. n.; 27, 28 ― Cephennula sibugana sp. n.; 29, 30 ― Cephennula gombakiana sp. n.; 31, 32 ― Cephennula micropunctata sp. n. Scale bars: 0.05 mm." httpUri="https://zenodo.org/record/279341/files/figure.png" pageId="14" pageNumber="50">Figs. 27, 28</figureCitation>
) in ventral view bottle-shaped with moderately narrow, subtrapezoidal apex and strongly rounded apical margin; AeL
<quantity id="4C8E9B431F518E0A8403FC2025F2FBC1" box="[593,700,1015,1040]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="14" pageNumber="50" unit="mm" value="0.13">0.13 mm</quantity>
. Endophallus lightly pigmented, with median tubular structure; parameres slender, not exceeding apex of median lobe, each bearing two apical and one subapical seta.
</paragraph>
<paragraph id="8BC936A61F518E0A8695FB9726D0FB89" blockId="14.[151,1437,151,1184]" box="[199,414,1088,1112]" pageId="14" pageNumber="50">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C36C652D1F518E0A8695FBB42582FBAD" box="[199,716,1123,1148]" pageId="14" pageNumber="50" type="distribution">
<paragraph id="8BC936A61F518E0A8695FBB42582FBAD" blockId="14.[151,1437,151,1184]" box="[199,716,1123,1148]" pageId="14" pageNumber="50">
<emphasis id="B902EAB41F518E0A8695FBB4262EFBAD" bold="true" box="[199,352,1123,1148]" pageId="14" pageNumber="50">Distribution.</emphasis>
East
<collectingCountry id="F36176361F518E0A87CCFBB3254BFBAD" box="[414,517,1124,1148]" name="Malaysia" pageId="14" pageNumber="50">Malaysia</collectingCountry>
(Borneo: Sabah).
</paragraph>
</subSubSection>
<subSubSection id="C36C652D1F518E0A8695FB5F242AFB71" box="[199,868,1160,1184]" pageId="14" pageNumber="50" type="etymology">
<paragraph id="8BC936A61F518E0A8695FB5F242AFB71" blockId="14.[151,1437,151,1184]" box="[199,868,1160,1184]" pageId="14" pageNumber="50">
<emphasis id="B902EAB41F518E0A8695FB5F2600FB71" bold="true" box="[199,334,1160,1184]" pageId="14" pageNumber="50">Etymology.</emphasis>
Locotypical, after the Sibuga Forest in Borneo.
</paragraph>
</subSubSection>
</treatment>
</document>