335 lines
18 KiB
XML
335 lines
18 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.100.23152" ID-PMC="PMC6023955" ID-Pensoft-Pub="1314-2003-100-91" ID-Pensoft-UUID="117AFFB5FFE90945FB26EE02FF92FFA4" ID-PubMed="29962891" ID-Zenodo-Dep="1300062" ModsDocID="1314-2003-100-91" checkinTime="1529641986672" checkinUser="pensoft" docAuthor="Martinez-Dominguez, Lili, Nicolalde-Morejon, Fernando, Vergara-Silva, Francisco & Stevenson, Dennis Wm." docDate="2018" docId="F968ECB821AB54CB98BEB9C3479E8FB0" docLanguage="en" docName="PhytoKeys 100: 91-124" docOrigin="PhytoKeys 100" docPubDate="2018-06-21" docSource="http://dx.doi.org/10.3897/phytokeys.100.23152" docTitle="Ceratozamia chamberlainii Mart. - Domínguez, Nic-Mor. & D. W. Stev. Phytotaxa 317 (1): 17 – 28. 2017." docType="treatment" docVersion="3" id="117AFFB5FFE90945FB26EE02FF92FFA4" lastPageNumber="97" masterDocId="117AFFB5FFE90945FB26EE02FF92FFA4" masterDocTitle="Taxonomic review of Ceratozamia (Zamiaceae) in the Sierra Madre Oriental, Mexico" masterLastPageNumber="124" masterPageNumber="91" pageNumber="96" updateTime="1668138916235" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic review of Ceratozamia (Zamiaceae) in the Sierra Madre Oriental, Mexico</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Martinez-Dominguez, Lili</mods:namePart>
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<mods:affiliation>Laboratorio de Taxonomia Integrativa, Instituto de Investigaciones Biologicas, Universidad Veracruzana, Xalapa, 91190, Veracruz, Mexico & Centro de Investigaciones Tropicales, Universidad Veracruzana, Jose Maria Morelos 44, Zona Centro, Xalapa, 91000, Veracruz, Mexico</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Nicolalde-Morejon, Fernando</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1423-7474</mods:nameIdentifier>
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<mods:affiliation>Laboratorio de Taxonomia Integrativa, Instituto de Investigaciones Biologicas, Universidad Veracruzana, Xalapa, 91190, Veracruz, Mexico</mods:affiliation>
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<mods:nameIdentifier type="email">f_nicolalde@yahoo.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vergara-Silva, Francisco</mods:namePart>
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<mods:affiliation>Laboratorio de Sistematica Molecular (Jardin Botanico), Instituto de Biologia, Universidad Nacional Autonoma de Mexico, 3 er Circuito Exterior, Ciudad Universitaria, Coyoacan 04510, Mexico, D. F., Mexico</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Stevenson, Dennis Wm.</mods:namePart>
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<mods:affiliation>The New York Botanical Garden, Bronx, Nueva York, 10458 - 5120, USA</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2018</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2018-06-21</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>100</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>91</mods:start>
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<mods:end>124</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.100.23152</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.100.23152</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-100-91</mods:identifier>
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||
<mods:identifier type="Pensoft-UUID">117AFFB5FFE90945FB26EE02FF92FFA4</mods:identifier>
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<mods:identifier type="Zenodo-Dep">1300062</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="182225056" LSID="urn:lsid:plazi:treatment:F968ECB821AB54CB98BEB9C3479E8FB0" httpUri="http://treatment.plazi.org/id/F968ECB821AB54CB98BEB9C3479E8FB0" lastPageId="6" lastPageNumber="97" pageId="5" pageNumber="96">
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<subSubSection pageId="5" pageNumber="96" type="nomenclature">
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<paragraph pageId="5" pageNumber="96">
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<pageBreakToken pageId="5" pageNumber="96" start="start">2</pageBreakToken>
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.
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<taxonomicName LSID="F968ECB8-21AB-54CB-98BE-B9C3479E8FB0" authority="Mart. - Dominguez, Nic-Mor. & D. W. Stev. Phytotaxa 317 (1): 17 - 28. 2017." authorityName="Mart. - Domínguez, Nic-Mor. & D. W. Stev. Phytotaxa 317 (1): 17 – 28. 2017." class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ceratozamia chamberlainii" order="Cycadales" pageId="5" pageNumber="96" phylum="Tracheophyta" rank="species" species="chamberlainii">
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Ceratozamia chamberlainii Mart.-
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<normalizedToken originalValue="Domínguez">Dominguez</normalizedToken>
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, Nic-Mor. & D.W. Stev. Phytotaxa 317(1): 17-28. 2017.
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</taxonomicName>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Shapes of leaflets. A C. latifolia B C. kuesteriana C C. morettii D C. sabatoi E Ceratozamia brevifrons F C. chamberlainii G C. decumbens H C. fuscoviridis I C. tenuis J C. totonacorum K C. delucana L C. mexicana." figureDoi="10.3897/phytokeys.100.23152.figure3" httpUri="https://binary.pensoft.net/fig/211374" pageId="5" pageNumber="96">Figure 3F</figureCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection pageId="6" pageNumber="97" type="materials_examined">
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<paragraph pageId="6" pageNumber="97">
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<pageBreakToken pageId="6" pageNumber="97" start="start">
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<typeStatus>Type</typeStatus>
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</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="6" pageNumber="97">
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<materialsCitation collectingDate="2016-03-20" collectorName="L. Martinez-Dominguez" country="MEXICO" location="Xilitla" specimenCount="933" specimenCount-female="933" stateProvince="San Luis Potosi" typeStatus="holotype">
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<collectingCountry name="Mexico">MEXICO</collectingCountry>
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.
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<collectingRegion country="Mexico" name="San Luis Potosi">
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San Luis
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<normalizedToken originalValue="Potosí">Potosi</normalizedToken>
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</collectingRegion>
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:
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<location LSID="urn:lsid:plazi:treatment:F968ECB821AB54CB98BEB9C3479E8FB0:FB7712CC96BDB4C84465C581825F67CE" country="MEXICO" name="Xilitla" stateProvince="San Luis Potosi">Xilitla</location>
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,
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<collectingDate value="2016-03-20">20 Mar. 2016</collectingDate>
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,
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<emphasis italics="true" pageId="6" pageNumber="97">
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<collectorName>
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L.
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<normalizedToken originalValue="Martínez-Domínguez">Martinez-Dominguez</normalizedToken>
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</collectorName>
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et al.
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<specimenCount type="female">933 ♀</specimenCount>
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</emphasis>
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(
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<typeStatus>holotype</typeStatus>
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: CIB; isotypes: MEXU, NY)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="97" type="description">
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<paragraph pageId="6" pageNumber="97">Description.</paragraph>
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<paragraph pageId="6" pageNumber="97">
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<emphasis italics="true" pageId="6" pageNumber="97">Stem</emphasis>
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epigeous, erect and decumbent, 20-60 cm in length, 15-30 cm in diameter.
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<emphasis italics="true" pageId="6" pageNumber="97">Cataphylls</emphasis>
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persistent, densely tomentose at emergence, reddish-brown and glabrous at maturity, triangular, apex acuminate, 2.5-5.5
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<normalizedToken originalValue="×">x</normalizedToken>
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1.5-3.5 cm at base.
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<emphasis italics="true" pageId="6" pageNumber="97">Leaves</emphasis>
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5-40, descending, 100-207 cm, reddish-brown at emergence with whitish-grey pubescence, glabrous at maturity.
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<emphasis italics="true" pageId="6" pageNumber="97">Petiole</emphasis>
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terete, straight, 30-69 cm, armed with short and robust prickles, blackish-brown in adult leaves.
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<emphasis italics="true" pageId="6" pageNumber="97">Rachis</emphasis>
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terete, straight, 65-144 cm, armed with prickles, reddish-brown in mature leaves.
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<emphasis italics="true" pageId="6" pageNumber="97">Leaflets</emphasis>
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20-42, oblong, planar and abaxially curved, not basally falcate, coriaceous, flat, opposite to subopposite, plane, dark green, adaxial and abaxial surfaces glabrous, acuminate and asymmetric apex, attenuate at base, with conspicuous and reddish-brown veins; median leaflets 20-37
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<normalizedToken originalValue="×">x</normalizedToken>
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2.3-4 cm, 1.6-3.5 cm between leaflets; articulations generally reddish-brown, 0.5-1.3 cm wide.
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<emphasis italics="true" pageId="6" pageNumber="97">Polliniferous strobilus</emphasis>
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generally solitary (1-2), cylindrical, erect, 26-31 cm in length, 4.8-6 cm in diameter, greenish-brown at emergence, greenish with reddish-brown pubescence at maturity; peduncle tomentose, reddish-brown, 6-10.3 cm in length, 1.6-2 cm in diameter; microsporophylls 1.6-2.3
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<normalizedToken originalValue="×">x</normalizedToken>
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1.1-1.5 cm, not recurved distal face.
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<emphasis italics="true" pageId="6" pageNumber="97">Ovuliferous strobilus</emphasis>
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cylindrical, erect, 28-30 cm in length, 9-10.5 cm in diameter, greyish-green with reddish-brown pubescence at emergence, light greyish-brown with reddish-brown trichomes at maturity, acuminate apex; peduncle tomentose, brown to reddish-brown, 4.5-11 cm in length, 1.3-2.5 cm in diameter; megasporophylls 72-190, 1.5-2.5
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<normalizedToken originalValue="×">x</normalizedToken>
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2.3-3.0 cm, prominent distal face, acute angle between horns.
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<emphasis italics="true" pageId="6" pageNumber="97">Seeds</emphasis>
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ovoid, sarcotesta whitish-red to pink when immature, light brown at maturity, 3.0-4.0 cm in length, 0.6-1.2 cm in diameter.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="97" type="distribution">
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<paragraph pageId="6" pageNumber="97">Distribution and habitat.</paragraph>
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<paragraph pageId="6" pageNumber="97">
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Its range includes the states of San Luis
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<normalizedToken originalValue="Potosí">Potosi</normalizedToken>
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,
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<normalizedToken originalValue="Querétaro">Queretaro</normalizedToken>
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and Hidalgo (Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Distribution map of Ceratozamia sabatoi, C. latifolia, C. zaragozae, C. kuesteriana, C. hildae, C. fuscoviridis and C. chamberlainii." figureDoi="10.3897/phytokeys.100.23152.figure9" httpUri="https://binary.pensoft.net/fig/211380" pageId="6" pageNumber="97">9</figureCitation>
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).
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<taxonomicName class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ceratozamia chamberlainii" order="Cycadales" pageId="6" pageNumber="97" phylum="Tracheophyta" rank="species" species="chamberlainii">
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<emphasis italics="true" pageId="6" pageNumber="97">Ceratozamia chamberlainii</emphasis>
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</taxonomicName>
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occurs in the cloud forest on rocky outcrops at 900-1,200 m.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="97" type="etymology">
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<paragraph pageId="6" pageNumber="97">Etymology.</paragraph>
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<paragraph pageId="6" pageNumber="97">The specific epithet honours Charles Joseph Chamberlain in recognition of his outstanding contributions to knowledge of the biology of the cycads and his fieldwork on Mexican cycads.</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="97" type="distinguishing features">
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<paragraph pageId="6" pageNumber="97">Distinguishing features.</paragraph>
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<paragraph pageId="6" pageNumber="97">This species is distinguished by its petioles and rachis with abundant, short prickles; reddish-brown leaves at emergence with coloured articulations at the leaflet attachment to the rachis in adult leaves. Leaflets are oblong and coriaceous with conspicuous, reddish-brown veins.</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="97" type="materials_examined">
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<paragraph pageId="6" pageNumber="97">Specimens examined.</paragraph>
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<paragraph pageId="6" pageNumber="97">
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||
<materialsCitation collectingDate="2018-01-01" collectingDateMax="2018-12-31" collectingDateMin="2018-01-01" collectorName="F. Nicolalde-Morejon, L. Martinez-Dominguez, La Mision" country="MEXICO" location="Alcantara-Ayala" municipality="Chapulhuacan" specimenCount="890" specimenCount-female="443" specimenCount-male="447" stateProvince="Hidalgo">
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<collectingCountry name="Mexico">MEXICO</collectingCountry>
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.
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<emphasis bold="true" pageId="6" pageNumber="97">
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<collectingRegion country="Mexico" name="Hidalgo">Hidalgo</collectingRegion>
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</emphasis>
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:
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<normalizedToken originalValue="Chapulhuacán">
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<collectingMunicipality>Chapulhuacan</collectingMunicipality>
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</normalizedToken>
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,
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<emphasis italics="true" pageId="6" pageNumber="97">
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<normalizedToken originalValue="Alcántara-Ayala">
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||
<location LSID="urn:lsid:plazi:treatment:F968ECB821AB54CB98BEB9C3479E8FB0:6F48AAD203AAC2AC148E5F0802D20CF7" country="MEXICO" municipality="Chapulhuacan" name="Alcantara-Ayala" stateProvince="Hidalgo">Alcantara-Ayala</location>
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</normalizedToken>
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3650
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||
</emphasis>
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(FCME),
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<emphasis italics="true" pageId="6" pageNumber="97">
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<collectorName>
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F.
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<normalizedToken originalValue="Nicolalde-Morejón">Nicolalde-Morejon</normalizedToken>
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</collectorName>
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et al. 2200-2203
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</emphasis>
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(CIB),
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<emphasis italics="true" pageId="6" pageNumber="97">
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<collectorName>
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L.
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<normalizedToken originalValue="Martínez-Domínguez">Martinez-Dominguez</normalizedToken>
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||
</collectorName>
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||
et al. 429
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||
</emphasis>
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||
-
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||
<emphasis italics="true" pageId="6" pageNumber="97">430</emphasis>
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||
(CIB),
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<emphasis italics="true" pageId="6" pageNumber="97">
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441-
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<specimenCount type="female">443 ♀</specimenCount>
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||
</emphasis>
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||
(CIB),
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||
<specimenCount type="male">
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<emphasis italics="true" pageId="6" pageNumber="97">447</emphasis>
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♂
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</specimenCount>
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(CIB),
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<emphasis italics="true" pageId="6" pageNumber="97">Vite-Reyes et al. 23</emphasis>
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(XAL);
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<collectorName>
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La
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<normalizedToken originalValue="Misión">Mision</normalizedToken>
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</collectorName>
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,
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||
<emphasis italics="true" pageId="6" pageNumber="97">Castro-Castro et al</emphasis>
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.
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<emphasis italics="true" pageId="6" pageNumber="97">1017</emphasis>
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(XAL),
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<emphasis italics="true" pageId="6" pageNumber="97">Vite-Reyes et al. 20</emphasis>
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(XAL)
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||
</materialsCitation>
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||
.
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||
<materialsCitation collectingDate="2018-01-01" collectingDateMax="2018-12-31" collectingDateMin="2018-01-01" collectorName="Landa de Matamoros, A. P, E. Carranza, F. Nicolalde-Morejon, L. Martinez-Dominguez" country="Mexico" location="Landa de Matamoros" specimenCount="833" specimenCount-male="833" stateProvince="Queretaro">
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||
<emphasis bold="true" pageId="6" pageNumber="97">
|
||
<normalizedToken originalValue="Querétaro">
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||
<collectingRegion country="Mexico" name="Queretaro de Arteaga">Queretaro</collectingRegion>
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||
</normalizedToken>
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||
</emphasis>
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||
:
|
||
<collectorName>
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||
<location LSID="urn:lsid:plazi:treatment:F968ECB821AB54CB98BEB9C3479E8FB0:DC6A24B7B07F8D4E1769DC1AF78A9F16" country="Mexico" name="Landa de Matamoros" stateProvince="Queretaro">Landa de Matamoros</location>
|
||
,
|
||
<emphasis italics="true" pageId="6" pageNumber="97">A. P. Vovides 1291</emphasis>
|
||
</collectorName>
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||
(XAL, MEXU),
|
||
<emphasis italics="true" pageId="6" pageNumber="97">1288, 1290</emphasis>
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||
, 2000 (XAL),
|
||
<emphasis italics="true" pageId="6" pageNumber="97">
|
||
<collectorName>E. Carranza</collectorName>
|
||
633
|
||
</emphasis>
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||
(IEB),
|
||
<emphasis italics="true" pageId="6" pageNumber="97">3119</emphasis>
|
||
(MEXU),
|
||
<emphasis italics="true" pageId="6" pageNumber="97">
|
||
<collectorName>
|
||
F.
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||
<normalizedToken originalValue="Nicolalde-Morejón">Nicolalde-Morejon</normalizedToken>
|
||
</collectorName>
|
||
et al. 2192-2199
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||
</emphasis>
|
||
(CIB);
|
||
<emphasis italics="true" pageId="6" pageNumber="97">
|
||
<collectorName>
|
||
L.
|
||
<normalizedToken originalValue="Martínez-Domínguez">Martinez-Dominguez</normalizedToken>
|
||
</collectorName>
|
||
et al. 379,
|
||
<specimenCount type="male">405 ♂</specimenCount>
|
||
</emphasis>
|
||
(CIB),
|
||
<emphasis italics="true" pageId="6" pageNumber="97">380-404</emphasis>
|
||
(CIB);
|
||
<emphasis italics="true" pageId="6" pageNumber="97">
|
||
<collectorName>
|
||
L.
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||
<normalizedToken originalValue="Martínez-Domínguez">Martinez-Dominguez</normalizedToken>
|
||
</collectorName>
|
||
et al. 408-426
|
||
</emphasis>
|
||
(CIB),
|
||
<emphasis italics="true" pageId="6" pageNumber="97">
|
||
427,
|
||
<specimenCount type="male">428 ♂</specimenCount>
|
||
</emphasis>
|
||
(CIB),
|
||
<emphasis italics="true" pageId="6" pageNumber="97">Rubio Hiram 6332</emphasis>
|
||
,
|
||
<emphasis italics="true" pageId="6" pageNumber="97">6334</emphasis>
|
||
(IEB),
|
||
<emphasis italics="true" pageId="6" pageNumber="97">647</emphasis>
|
||
(XAL)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation collectingDate="2018-01-01" collectingDateMax="2018-12-31" collectingDateMin="2018-01-01" collectorName="Tamazunchale, C. L, Xilitla, F, L. Martinez-Dominguez, T. W. Walters" country="Mexico" location="San Luis Potosi" specimenCount="1" stateProvince="San Luis Potosi">
|
||
<emphasis bold="true" pageId="6" pageNumber="97">
|
||
<collectingRegion country="Mexico" name="San Luis Potosi">
|
||
San Luis
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<normalizedToken originalValue="Potosí">Potosi</normalizedToken>
|
||
</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<collectorName>
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||
Tamazunchale,
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||
<emphasis italics="true" pageId="6" pageNumber="97">C. L. Lundell 7235</emphasis>
|
||
</collectorName>
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||
(IEB);
|
||
<collectorName>
|
||
Xilitla,
|
||
<emphasis italics="true" pageId="6" pageNumber="97">
|
||
F.
|
||
<normalizedToken originalValue="Nicolalde-Morejón">Nicolalde-Morejon</normalizedToken>
|
||
et al. 2407-2420
|
||
</emphasis>
|
||
</collectorName>
|
||
(CIB),
|
||
<emphasis italics="true" pageId="6" pageNumber="97">
|
||
<collectorName>
|
||
L.
|
||
<normalizedToken originalValue="Martínez-Domínguez">Martinez-Dominguez</normalizedToken>
|
||
</collectorName>
|
||
et al. 924
|
||
</emphasis>
|
||
-
|
||
<emphasis italics="true" pageId="6" pageNumber="97">932</emphasis>
|
||
(CIB);
|
||
<emphasis italics="true" pageId="6" pageNumber="97">
|
||
<collectorName>T. W. Walters</collectorName>
|
||
, TW-2001-04-A
|
||
</emphasis>
|
||
(XAL)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |