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<document id="2A3F2E207058AB1C085721A3A40FF406" ID-CLB-Dataset="299803" ID-DOI="10.3897/zookeys.1207.120068" ID-GBIF-Dataset="d7935e1b-a3af-427e-afd7-88ee385dc1e7" ID-ZooBank="8B2E2B5D-CA1D-4B61-8736-ECF013EDC384" ID-publisher-id="120068" URI-arpha="1314A2C0-E9C6-52F9-821B-A77D9D85F5DA" URI-zoobank="http://zoobank.org/8B2E2B5D-CA1D-4B61-8736-ECF013EDC384" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1720801545516" checkinUser="pensoft" docAuthor="Porto, Willians, Derkarabetian, Shahan, Giribet, Gonzalo &amp; Pérez-González, Abel" docDate="2024" docId="F8C8242EC12E5790AD4270EBFC0EC275" docLanguage="en" docName="ZooKeys 1207: 1-149" docOrigin="ZooKeys 1207" docSource="https://zookeys.pensoft.net/article/120068/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Nerudiella caramavida Porto, Derkarabetian, Giribet &amp; Pérez-González, 2024, sp. nov." docType="treatment" docUuid="45A9D45E-90CE-43D9-9223-BD691DA31146" docUuidSource="ZooBank" docVersion="5" dtd-version="3.0" lastPageNumber="149" masterDocId="1314A2C0E9C652F9821BA77D9D85F5DA" masterDocTitle="Systematic revision of the South American “ Nuncia ” (Opiliones, Laniatores, Triaenonychidae)" masterLastPageNumber="149" masterPageNumber="1" pageNumber="1" updateTime="1720815513282" updateUser="ExternalLinkService">
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<mods:title id="F38C61BF8853C6ED5111F407CD0A061C">Systematic revision of the South American “ Nuncia ” (Opiliones, Laniatores, Triaenonychidae)</mods:title>
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<mods:namePart id="2AF0685CAD7E163CA668EFC4F338AE6C">Porto, Willians</mods:namePart>
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<mods:affiliation id="54B95D568BBE059D02F0B4CC50DC3E70">División Aracnología, Museo Argentino de Ciencias Naturales CONICET, Av. Ángel Gallardo 470, C 1405 DJR Buenos Aires, Argentina &amp; Sección Aracnología y Miriapodología, Universidad Nacional de La Plata, Facultad de Ciencias Naturales y Museo, Paseo del Bosque s / n, (1900) La Plata, Argentina</mods:affiliation>
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<mods:namePart id="764FDB9BD439EC03E70E47B2C22537C5">Derkarabetian, Shahan</mods:namePart>
<mods:nameIdentifier id="035B6F621408D08049608C6A16EB0A75" type="ORCID">0000-0002-9163-9277</mods:nameIdentifier>
<mods:affiliation id="75060EA8AE4DAB5B7839B6251ADE0C0E">Museum of Comparative Zoology, Department of Organismic and Evolutionary Biology, Harvard University, Cambridge, MA 02138, USA &amp; San Diego Natural History Museum, Department of Entomology, San Diego, CA 92101, USA</mods:affiliation>
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<mods:namePart id="6E649510BEECC62ECE049F80B758524B">Giribet, Gonzalo</mods:namePart>
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<mods:affiliation id="EC0724031560E5C408B4151A994615C9">Museum of Comparative Zoology, Department of Organismic and Evolutionary Biology, Harvard University, Cambridge, MA 02138, USA</mods:affiliation>
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<mods:namePart id="387BA75F831C9CAAB573F00B8B726548">Pérez-González, Abel</mods:namePart>
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<mods:affiliation id="B00318F89BFCA4EF91270F7D0C57B414">División Aracnología, Museo Argentino de Ciencias Naturales CONICET, Av. Ángel Gallardo 470, C 1405 DJR Buenos Aires, Argentina</mods:affiliation>
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<mods:date id="1337DC3A3E667341B57BF06E111C4957">2024</mods:date>
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<treatment id="F8C8242EC12E5790AD4270EBFC0EC275" ID-DOI="http://doi.org/10.5281/zenodo.12734512" ID-GBIF-Taxon="235091451" ID-Zenodo-Dep="12734512" ID-arpha="F8C8242E-C12E-5790-AD42-70EBFC0EC275" ID-zoobank="https://zoobank.org/45A9D45E-90CE-43D9-9223-BD691DA31146" LSID="urn:lsid:zoobank.org:act:45A9D45E-90CE-43D9-9223-BD691DA31146" LSID-ZBK="urn:lsid:zoobank.org:act:45A9D45E-90CE-43D9-9223-BD691DA31146" httpUri="http://treatment.plazi.org/id/F8C8242EC12E5790AD4270EBFC0EC275">
<subSubSection id="73798B7B9313CE0B0F9B2CD4EE815CF4" type="nomenclature">
<paragraph id="11B7808414D614AD728623C682586568">
<taxonomicName id="033FAEDA08BE57CB4164776E3976889B" authorityName="Porto &amp; Derkarabetian &amp; Giribet &amp; Pérez-González" authorityYear="2024" class="Arachnida" family="Triaenonychidae" genus="Nerudiella" kingdom="Animalia" order="Opiliones" phylum="Arthropoda" rank="species" species="caramavida" status="sp. nov.">
<emphasis id="D9F10F8DAD82ED7C88DCB4124B4E1B34" italics="true">Nerudiella caramavida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3C48ABC1777C0F61AC3B03CD88236386" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="50DAF42D7967C26FA020A0E6B8046C3B" type="description">
<paragraph id="3EB49CD93E6C52D5288C49E40124983E">
<figureCitation id="8F973F851BEAB972BA100D2C0302E613" captionStart="Figure 80" captionStartId="F80" captionText="Figure 80. Nerudiella caramavida sp. nov. habitus, male A dorsal view C lateral view E ventral view. Female B dorsal view D lateral view F ventral view. Scale bars: 1 mm. Species of Clade C, see Fig. 3." figureDoi="10.3897/zookeys.1207.120068.figure80" httpUri="https://binary.pensoft.net/fig/1090693">Figs 80</figureCitation>
,
<figureCitation id="4CBFD353DEB5FD6BE62698C3D0B85D9C" captionStart="Figure 81" captionStartId="F81" captionText="Figure 81. Nerudiella caramavida sp. nov. male, SEM images of habitus A dorsal view B lateral view C, D ventral view. Scale bars: 1 mm (A, B, C); 500 µm (D)." figureDoi="10.3897/zookeys.1207.120068.figure81" httpUri="https://binary.pensoft.net/fig/1090694">81</figureCitation>
,
<figureCitation id="13A36A3912A2C5A52482DA14CFD69D00" captionStart="Figure 82" captionStartId="F82" captionText="Figure 82. Nerudiella caramavida sp. nov. chelicerae: mesal A ectal B pedipalps: mesal C ectal D. Scale bars: 200 µm (A, B); 500 µm (C, D)." figureDoi="10.3897/zookeys.1207.120068.figure82" httpUri="https://binary.pensoft.net/fig/1090695">82</figureCitation>
,
<figureCitation id="FEADDF5B1D9B2FCBFC7E1B241CE3B921" captionStart="Figure 83" captionStartId="F83" captionText="Figure 83. Nerudiella caramavida sp. nov. legs I A II B III C IV D. Scale bars: 500 µm (A, B, D); 200 µm (C)." figureDoi="10.3897/zookeys.1207.120068.figure83" httpUri="https://binary.pensoft.net/fig/1090696">83</figureCitation>
,
<figureCitation id="55E838EC981970387F54CEE715884E6E" captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Nerudiella caramavida sp. nov. penis: ventral A, B apical C lateral D, E dorsal F, G. Scale bars: 200 µm (A, D, F); 50 µm (B, C, E, G)." figureDoi="10.3897/zookeys.1207.120068.figure84" httpUri="https://binary.pensoft.net/fig/1090697">84</figureCitation>
,
<figureCitation id="4B1402307E63CD078E157A2209AFD22A" captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Nerudiella caramavida sp. nov. penis: ventral A apical B lateral C dorsal D. Colors: ventral plate (yellow), capsula externa (blue), capsula interna (red). Scale bars: 50 µm (A C); 200 µm (D)." figureDoi="10.3897/zookeys.1207.120068.figure85" httpUri="https://binary.pensoft.net/fig/1090698">85</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELUCK" type="material">
<paragraph id="C879E8AEC561FA74BF215A2A7AF973FE">
<heading id="7800427B1B9C2EEA726EA2F194174A5F" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="4FBBA193B4E33962C6C49ABE2DC3F472">
<materialsCitation id="E4C81AF84E47264B125B5580C9FF38B6" ID-GBIF-Occurrence="4905322343" collectingDate="2005-02-12" collectionCode="MNHNCL" collectorName="M. Ramirez &amp; F. Labarque" country="Chile" location="P. N. Nahuelbuta" specimenCount="1" specimenCount-male="1" stateProvince="Malleco" typeStatus="holotype">
<emphasis id="C5769252274FE441CF4B1A034B17D9B2" bold="true">
<emphasis id="2B5FB4EDF495034C80D293DCF38F3FC7" bold="true" italics="true">
<typeStatus id="F8334B92584B23308CCAA9C40322F285" type="holotype">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<specimenCount id="C9AD98FFE9BA563BEE92E039E3F3A262" count="1" type="male"></specimenCount>
<emphasis id="A9FB261FC19D251A55BCBBA87A53DAC7" bold="true">
<collectingCountry id="D8874F82B1D842091EE9B4FF5F9A900C" name="Chile">Chile</collectingCountry>
.
</emphasis>
<collectingRegion id="C7DF126157D367AA77848C8515281E6A">Malleco</collectingRegion>
:
<location id="1FEE7BC43FE61A2BDA4EB44F02C549C7" LSID="urn:lsid:plazi:treatment:F8C8242EC12E5790AD4270EBFC0EC275:1FEE7BC43FE61A2BDA4EB44F02C549C7" country="Chile" name="P. N. Nahuelbuta" stateProvince="Malleco">P. N. Nahuelbuta</location>
,
<collectorName id="B552EB33B316B3314C7E8BE5A68FD12F">M. Ramírez</collectorName>
,
<collectorName id="0C47931E1D1E6E69A15D939359965837">F. Labarque</collectorName>
,
<date id="D41DBB08C18365B323B84C92E2F55F27" value="2005-02-12">
<collectingDate id="A0B0A4C68ED6BE91F4FCD5BE8FA358C2" value="2005-02-12">12. II. 2005</collectingDate>
</date>
(
<named-content id="A88CC47C6E7905284AC1F85E626598A7" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-nacional-de-historia-natural-santiago" xlink_title="Museo Nacional de Historia Natural, Santiago de Chile">
<collectionCode id="D74C3B3C1D17D034DC2F0FE8D634F08B">MNHNCL</collectionCode>
</named-content>
)
</materialsCitation>
.
<materialsCitation id="1C52026F2ABEF9DB51C998403907B9DA" ID-GBIF-Occurrence="4905322323" collectingDate="2005-02-12" collectionCode="MACN" collectorName="M. Ramirez &amp; F. Labarque" country="Chile" location="P. N. Nahuelbuta" specimenCount="19" specimenCount-female="7" specimenCount-male="9" stateProvince="Malleco" typeStatus="paratype">
<emphasis id="A3517A84AB659FF7BD769FFE70A5915A" bold="true">
<emphasis id="14C5DCA04D1AF973B699FAE5871D5F2E" bold="true" italics="true">
<typeStatus id="D363C7F710771002405BA8324DA18310" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<collectingCountry id="69FD0A37ED4085C6D1C1BF8471857D4A" name="Chile">Chile</collectingCountry>
.
</emphasis>
<collectingRegion id="12E38483A250A42B5D3CC541DCB7189D">Malleco</collectingRegion>
:
<location id="AD177AF73B58E20F51EA3807EA51096A" LSID="urn:lsid:plazi:treatment:F8C8242EC12E5790AD4270EBFC0EC275:AD177AF73B58E20F51EA3807EA51096A" country="Chile" name="P. N. Nahuelbuta" stateProvince="Malleco">P. N. Nahuelbuta</location>
,
<collectorName id="63209A01904C83DD073B80CC23B31C47">M. Ramírez</collectorName>
,
<collectorName id="CD15C02F460B1162C0BC2A23D00BC60B">F. Labarque</collectorName>
,
<date id="998B0AA24E47384DB10B20F713F6C999" value="2005-02-12">
<collectingDate id="66CE691692AB4DA2C6899F7B6514D8E8" value="2005-02-12">12. II. 2005</collectingDate>
</date>
,
<specimenCount id="54F5208756F687A656CB2F9D063EB7D6" count="9" type="male">9 ♂</specimenCount>
<specimenCount id="EB4B6466E1E520233B2A4E97DDEE0372" count="7" type="female">7 ♀</specimenCount>
<specimenCount id="91670E65B2FCD9BFBA9DD1EB6137D2FE" count="3" type="generic">3 imm.</specimenCount>
(
<named-content id="1B65BFBE8FF88B06A2181116BC9C166E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-argentino-de-ciencias-naturales-bernardino-rivadavia" xlink_title="Museo Argentino de Ciencias Naturales">
<collectionCode id="417F6AD0FDA2A9B6A525AFEE992953F5" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" type="Museum">MACN</collectionCode>
</named-content>
)
</materialsCitation>
,
<materialsCitation id="0684BFFAA6CBF6D52EADD6BFDB193A9F" ID-GBIF-Occurrence="4905322332" collectingDate="1981-11-19" collectionCode="AMNH" collectorName="N. Platnick &amp; R. Schuh" elevation="1250" location="P. N. Nahuelbuta" specimenCount="9" specimenCount-female="2" specimenCount-male="1" stateProvince="Malleco">
<collectingRegion id="9DA42695573089DD54A486AC645AB5EE">Malleco</collectingRegion>
:
<location id="3E815A0E5B15FA36FA2B19786A2D0726" LSID="urn:lsid:plazi:treatment:F8C8242EC12E5790AD4270EBFC0EC275:3E815A0E5B15FA36FA2B19786A2D0726" name="P. N. Nahuelbuta" stateProvince="Malleco">P. N. Nahuelbuta</location>
,
<quantity id="EB25DE2ECECAD85235FB49D79224C9BB" metricMagnitude="3" metricUnit="m" metricValue="1.25" unit="m" value="1250.0">
<elevation id="D8F88A59CFB8AAF7597B40AD43022311" metricMagnitude="3" metricUnit="m" metricValue="1.25" unit="m" value="1250.0">1250 m</elevation>
</quantity>
,
<collectorName id="E6BE8782B4CCF529F1B472EA148CFAFF">N. Platnick</collectorName>
,
<collectorName id="B2BEE5200F7451B7F89973FE344843BF">R. Schuh</collectorName>
.
<date id="89F12EC66567466527B281ADA00390E6" value="1981-11-19">
<collectingDate id="6610878AC0996A201FDB3C11ED352EC2" value="1981-11-19">19. XI. 1981</collectingDate>
</date>
,
<specimenCount id="C559DCD3E88A35E514D64C6A931AE01D" count="1" type="male">1 ♂</specimenCount>
<specimenCount id="99697108BF299EECC6B99E21B936D5E0" count="2" type="female">2 ♀</specimenCount>
<specimenCount id="AA44A9AD8F8EA0C16419E71757E10657" count="6" type="generic">6 imm.</specimenCount>
(
<named-content id="9DD29DFC56169CCD48F981C316A4B1E2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/american-museum-natural-history" xlink_title="American Museum of Natural History">
<collectionCode id="9E7A7F068D1971CAA2C7375E75C4534C" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" type="">AMNH</collectionCode>
</named-content>
)
</materialsCitation>
.
<materialsCitation id="836D548F65F9282A297A01681DE371AB" ID-GBIF-Occurrence="4905322320" collectingDate="1989-01-09" collectionCode="MACN" collectorName="E. Maury" location="Las Trancas" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Nuble">
<collectingRegion id="293B6215150614FE927E0D47FB08B955">Ñuble</collectingRegion>
:
<locationDeviation id="94C2DFBCADDA030CEDF58EE1AEFB21C1" bearing="90.0" distance="2000.0">
<quantity id="62BA4DC0D45DA78E3049E4B42AA9BBF3" metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="km" value="2.0">2 km</quantity>
E
</locationDeviation>
de
<location id="EAD0FFD7A5353C32D1459F0DC2F2FEFE" LSID="urn:lsid:plazi:treatment:F8C8242EC12E5790AD4270EBFC0EC275:EAD0FFD7A5353C32D1459F0DC2F2FEFE" name="Las Trancas" stateProvince="Nuble">Las Trancas</location>
,
<collectorName id="FEB6DB59E65DDE8F8C6F2F09826E5C69">E. Maury</collectorName>
.
<date id="0AF5CDFC6ED8C854942E1E67066AB37D" value="1989-01-09">
<collectingDate id="4558CDD3ED4E19722504E4E6BD61E0DD" value="1989-01-09">09. I. 1989</collectingDate>
</date>
,
<specimenCount id="5E15994A5C149D18E676B17F97B62282" count="1" type="male">1 ♂</specimenCount>
<specimenCount id="62C410A38DAAA1F69842ECCEB4EF2B2F" count="1" type="female">1 ♀</specimenCount>
(
<named-content id="E2098918155D1F179626627585546A29" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-argentino-de-ciencias-naturales-bernardino-rivadavia" xlink_title="Museo Argentino de Ciencias Naturales">
<collectionCode id="41F3F0921CA1F2BB6E87DED17DD27056" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" type="Museum">MACN</collectionCode>
</named-content>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQVCK" type="material">
<paragraph id="55F80CF0C95381AB4652ACD5239B60E9">
<heading id="5E5B018934DE4738FACD593868490DB6" reason="title">Additional material.</heading>
</paragraph>
<paragraph id="90EBCFE2B9037493714A1DF7C7D7506C">
<collectingCountry id="3EC54BC0BE3E1A73B76522020D0E4204" name="Chile">Chile</collectingCountry>
.
<collectingRegion id="9FE35DB4F5FA5813B404009BF25BF2D7" country="Chile" name="Araucania">Arauco</collectingRegion>
. Hualpén, Univ. Concepción Terrestrial Biology Station,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EZVCK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-73.163070,-36.798210]}">
<geoCoordinate id="05E315FE7DBD70AC720AA6FDEB877E2E" degrees="36.79821" direction="south" orientation="latitude" precision="1" value="-36.79821">36.79821 ° S</geoCoordinate>
,
<geoCoordinate id="1526CF68E065606F5E0B335068BB3947" degrees="73.16307" direction="west" orientation="longitude" precision="1" value="-73.16307">73.16307 ° W</geoCoordinate>
</named-content>
,
<quantity id="1A0F60B90DA0878891028E8F63AE81CD" metricMagnitude="1" metricUnit="m" metricValue="5.2" unit="m" value="52.0">52 m</quantity>
, A. Ojanguren, A. Pérez-González, M. Ramírez, G. Azevedo, W. Porto coll.,
<date id="0C483A43E09B59C1EBF0167FC527B5FC" value="2018-01-14">14. I. 2018</date>
,
<specimenCount id="379FC9DEC433C59DAA05A5FE1687ABBC" count="1" type="male">1 ♂</specimenCount>
(
<named-content id="FB3CC135570305B1726634892C8C317B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-argentino-de-ciencias-naturales-bernardino-rivadavia" xlink_title="Museo Argentino de Ciencias Naturales">
<collectionCode id="8E6C46B36504AD85A82670D1E012DEA8" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" type="Museum">MACN</collectionCode>
</named-content>
). Caramávida, San Alfonso, Quebrada Caramávida,
<collectingRegion id="F6AD42D8082D369064FBDCBE9751E258" country="Chile" name="Araucania">Arauco</collectingRegion>
Reserve,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EGWCK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-73.171070,-37.709420]}">
<geoCoordinate id="60BB6766C23CF56BA2FE227165829083" degrees="37.70942" direction="south" orientation="latitude" precision="1" value="-37.70942">37.70942 ° S</geoCoordinate>
,
<geoCoordinate id="96F08B82CFF0E3C38F0F0B3D0147504A" degrees="73.17107" direction="west" orientation="longitude" precision="1" value="-73.17107">73.17107 ° W</geoCoordinate>
</named-content>
,
<quantity id="A4F2833CFE65032CE22626B041BDD434" metricMagnitude="2" metricUnit="m" metricValue="7.5" unit="m" value="750.0">750 m</quantity>
, A. Ojanguren, A. Pérez-González, M. Ramírez, G. Azevedo, W. Porto coll.,
<quantity id="77C18017E1C75D0647CA92AD23C1A217" metricMagnitude="-3" metricUnit="m" metricValue="3.0" unit="mm" value="3.0">3 mm</quantity>
.
<date id="6C7C801015F0296716A7DC8A6D915CEB" value="2018-01-14">14. I. 2018</date>
,
<specimenCount id="F92FCD70D107A17D4D81FD337DD60C96" count="1" type="female">1 ♀</specimenCount>
,
<specimenCount id="1788FD4C23278799716118B9576F47C3" count="1" type="male">1 ♂</specimenCount>
3.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELWCK" type="etymology">
<paragraph id="9BE2BF446872595ED9AE5FC6FDF7F532">
<heading id="5E825F9B7396F78C2178E8BA7D87EB56" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="C40BD56BB6957593C0A92B3A6BD05463">
The specific epithet
<taxonomicName id="EE60B9CE056C672260BD5E4AF02A20A4" authorityName="Porto &amp; Derkarabetian &amp; Giribet &amp; Pérez-González" authorityYear="2024" class="Arachnida" family="Triaenonychidae" genus="Nerudiella" kingdom="Animalia" order="Opiliones" phylum="Arthropoda" rank="species" species="caramavida">
<emphasis id="D72C56CF2F0B64B80F3CFC217AA5DE0F" italics="true">caramavida</emphasis>
</taxonomicName>
” is derived from the type locality of the species, Quebrada Caramávida, located on the western slope of the Cordillera de Nahuelbuta in the
<collectingRegion id="9E8BCEF3E607F4D8E5EA3F950B65E655" country="Chile" name="Biobio">Bío-Bío Region</collectingRegion>
. The choice of this name, a noun in apposition, is based on the geographic location where the species was originally discovered.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E2WCK" type="diagnosis">
<paragraph id="337FED2933A6456B5B81E1FAAF978CF1">
<heading id="43E0EFB8F00079BECC3CEAA7E7E44ACC" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="FA5D47642C1785E89FC0B5B9EC574281">This species can be easily distinguished from its congeners by the capsula interna of the male genitalia, which has two long lateral processes.</paragraph>
</subSubSection>
<subSubSection id="SECID0EAXCK" type="distribution">
<paragraph id="29C86394EF122EADDBFC20AB45A08316">
<heading id="1A3F46B576FCF4E9B18809CAC6FFAB59" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="6786BC83780F1095E38F60875E297719">
<collectingCountry id="00CE81D3ECEE86DDDB455DF4FCA86620" name="Chile">Chile</collectingCountry>
:
<collectingRegion id="01B5A0B069D5D03311FC3F35E0A2043F" country="Chile" name="Biobio">Bío-Bío Region</collectingRegion>
(Fig.
<figureCitation id="7D43227F45D87D206F71C5DA16AE35BA" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Maps showing the distribution of the species used in this work. South America (Chile and western Argentina inset). Chile in green, Argentina / South America in grey. Terrestrial ecoregions (following Olson et al. 2001) highlighted in yellow (Chilean Matorral) and orange (Valdivian temperate forests)." figureDoi="10.3897/zookeys.1207.120068.figure4" httpUri="https://binary.pensoft.net/fig/1090610">4 F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EKXCK" type="description">
<paragraph id="C27DC9E6B799183FAF820D9BBF4238AF">
<heading id="B2227514B2E47B3544185D39D56ABD67" reason="title">
Description of male
<typeStatus id="72BACBB39D66D71AF6DF75BB5192A695" type="holotype">holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="358B5EA4EE88C448B9FD84A0C468C03F">Measurements: Total length 2.74, carapace length 1.1, dorsal scutum length 2.05, carapace max. width 1.37, mesotergum max. width 1.96. Appendage measurements: Pedipalps. Trochanter length 0.20, femora length 0.92, patella length 0.40, tibia length 0.62, tarsus length 0.83. Leg I: trochanter (tr) 0.20, femora (fe) 0.85, patella (pa) 0.46, tibia (ti) 0.62, metatarsus (mt) 0.78, tarsus (ta) 0.65. II: tr 0.25, fe 1.84, pa 0.57, ti 0.90, mt 1.31, ta 1.29. III: tr 0.31, fe 0.79, pa 0.38, ti 0.64, mt 0.66, ta 0.60. IV: tr 0.31, fe 1.08, pa 0.63, ti 0.88, mt 1.04, ta 0.82.</paragraph>
<paragraph id="819D082BD5F8009F40C4A96172783E4F">
Dorsum (Fig.
<figureCitation id="89AE846F5F9D4C0F077FB677F620D56C" captionStart="Figure 80" captionStartId="F80" captionText="Figure 80. Nerudiella caramavida sp. nov. habitus, male A dorsal view C lateral view E ventral view. Female B dorsal view D lateral view F ventral view. Scale bars: 1 mm. Species of Clade C, see Fig. 3." figureDoi="10.3897/zookeys.1207.120068.figure80" httpUri="https://binary.pensoft.net/fig/1090693">80</figureCitation>
,
<figureCitation id="92F8A0BEF9E39585A938851FD6AF5BB7" captionStart="Figure 81" captionStartId="F81" captionText="Figure 81. Nerudiella caramavida sp. nov. male, SEM images of habitus A dorsal view B lateral view C, D ventral view. Scale bars: 1 mm (A, B, C); 500 µm (D)." figureDoi="10.3897/zookeys.1207.120068.figure81" httpUri="https://binary.pensoft.net/fig/1090694">81</figureCitation>
). Eta (η) hourglass-shaped dorsal scutum. Ocularium low and rounded, pointing a bit forward. Dorsal scutum microgranulate. Mesotergal areas of dorsal scutum not well delimited; with small setiferous tubercles. Free tergites with two rows of small setiferous tubercles.
</paragraph>
<caption id="62AA43615790A8F858343DCD12713D5B" ID-DOI="10.3897/zookeys.1207.120068.figure80" ID-arpha="8D2C9DD8-B2EC-5ECD-ADC0-F8909DC705B8" httpUri="https://binary.pensoft.net/fig/1090693" startId="F80">
<paragraph id="F50AD1B1BCA8D2931D43214A8FE39A69">
<label id="8B9602B882CC8D6945CDAA78B6B074B2">Figure 80.</label>
</paragraph>
<paragraph id="B6984C46D2465E9B61656106A43F318D">
<taxonomicName id="01782BB661741F86EFD4C4C6BBD08E74" authorityName="Porto &amp; Derkarabetian &amp; Giribet &amp; Pérez-González" authorityYear="2024" class="Arachnida" family="Triaenonychidae" genus="Nerudiella" kingdom="Animalia" order="Opiliones" phylum="Arthropoda" rank="species" species="caramavida" status="sp. nov.">
<emphasis id="71D3A141E84E5703AB5FEA9641191F68" italics="true">Nerudiella caramavida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EB2986E59A6243D2D6975641937D81D8" rank="species">sp. nov.</taxonomicNameLabel>
habitus, male
<emphasis id="18A51AAB3CA1EC2206DB95EFA875DA4D" bold="true">A</emphasis>
dorsal view
<emphasis id="566B831A2B02C6A7A78466C242E98FEE" bold="true">C</emphasis>
lateral view
<emphasis id="2AC4D8C55A31A3400F96ABD558A722BD" bold="true">E</emphasis>
ventral view. Female
<emphasis id="0D8C8EBFF5C80CBEF0BFA363D5733A95" bold="true">B</emphasis>
dorsal view
<emphasis id="6C063D4B26FC1B012CE670876FA33211" bold="true">D</emphasis>
lateral view
<emphasis id="3D7E3F3139F38F53368EB05EBC677236" bold="true">F</emphasis>
ventral view. Scale bars: 1 mm. Species of Clade C, see Fig.
<figureCitation id="49961EE5D5F7C25CAF19735D7C1B5FF1" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phylogenetics relationships of Triaenonychidae, with clade names as discussed in the main text. Phylogeny is derived from the IQ-TREE analysis. Nodes without circles have &lt; 95 % bootstrap support to AH-aLR and UFBoot." figureDoi="10.3897/zookeys.1207.120068.figure3" httpUri="https://binary.pensoft.net/fig/1090609">3</figureCitation>
.
</paragraph>
</caption>
<caption id="B1253307B26716AE287E360521A230DE" ID-DOI="10.3897/zookeys.1207.120068.figure81" ID-arpha="F2D8D4BE-3C77-5969-9EB8-F3CEE6A8B5E8" httpUri="https://binary.pensoft.net/fig/1090694" startId="F81">
<paragraph id="7EFF99CAF0C0B2595FD71EAE6A05FD25">
<label id="57E399394FBA95B9A436B9D5F0A459FE">Figure 81.</label>
</paragraph>
<paragraph id="81B9559D1E349809ACC580D0E67BE54C">
<taxonomicName id="216E2403778C0BB80D6F52D5BBD907CA" authorityName="Porto &amp; Derkarabetian &amp; Giribet &amp; Pérez-González" authorityYear="2024" class="Arachnida" family="Triaenonychidae" genus="Nerudiella" kingdom="Animalia" order="Opiliones" phylum="Arthropoda" rank="species" species="caramavida" status="sp. nov.">
<emphasis id="4D22F70D768556749668CBFF43B56F11" italics="true">Nerudiella caramavida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F4C1A30F3325368637547C9256B2D925" rank="species">sp. nov.</taxonomicNameLabel>
male,
<abbrev content-type="institution" id="ABBRID0EUZCK" xlink_title="scanning electron microscopy">SEM</abbrev>
images of habitus
<emphasis id="4C8757B831D7678104DA70C333743546" bold="true">A</emphasis>
dorsal view
<emphasis id="3B7E0CA1858E746B8D4F21BC12E36631" bold="true">B</emphasis>
lateral view
<emphasis id="06A549755A1463F8A28A276F2AA9571E" bold="true">C, D</emphasis>
ventral view. Scale bars: 1 mm (
<emphasis id="034BBEC7A135BA799CF9905685CB40C7" bold="true">A, B, C</emphasis>
); 500 µm (
<emphasis id="2685C5089C59962E6365C265DD93D5DB" bold="true">D</emphasis>
).
</paragraph>
</caption>
<paragraph id="8720A869639484BAE8D9E805300B2695">
Chelicerae (Fig.
<figureCitation id="E3C5C51A0BDF9E69998588036CE14B0C" captionStart="Figure 82" captionStartId="F82" captionText="Figure 82. Nerudiella caramavida sp. nov. chelicerae: mesal A ectal B pedipalps: mesal C ectal D. Scale bars: 200 µm (A, B); 500 µm (C, D)." figureDoi="10.3897/zookeys.1207.120068.figure82" httpUri="https://binary.pensoft.net/fig/1090695">82 A, B</figureCitation>
). Segment I with an acute tubercle on the dorso-distal surface. Segment II with two mesal setiferous tubercles and 78 small setiferous tubercles, with one triangular tubercle more prominent than others in frontal view.
</paragraph>
<caption id="8FA34B951BD6056DF9BF4137128533A9" ID-DOI="10.3897/zookeys.1207.120068.figure82" ID-arpha="0E311B24-6A2D-56E7-AF6B-8B96892BA8A6" httpUri="https://binary.pensoft.net/fig/1090695" startId="F82">
<paragraph id="481761F0BA2550082855087FEC08A847">
<label id="7D524F2A19E5CB289904615D89F83D02">Figure 82.</label>
</paragraph>
<paragraph id="C2EFF36EE10958EF67BCAE34A5E6EC41">
<taxonomicName id="D8D78FBFCE51A4D80B5B6FE6A43CF0A8" authorityName="Porto &amp; Derkarabetian &amp; Giribet &amp; Pérez-González" authorityYear="2024" class="Arachnida" family="Triaenonychidae" genus="Nerudiella" kingdom="Animalia" order="Opiliones" phylum="Arthropoda" rank="species" species="caramavida" status="sp. nov.">
<emphasis id="13D3EAE4EBB847BB84F8D8E347A954A0" italics="true">Nerudiella caramavida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7244C0428FAE63AB01A2E2CFDB5B38E" rank="species">sp. nov.</taxonomicNameLabel>
chelicerae: mesal
<emphasis id="08D77ADAE79A50094307386C67C8E11E" bold="true">A</emphasis>
ectal
<emphasis id="95EE28613A13F9BBA192E4166E21C0E6" bold="true">B</emphasis>
pedipalps: mesal
<emphasis id="9BC3788D52AAACFCCA7D8B43F8C66612" bold="true">C</emphasis>
ectal
<emphasis id="CF4D898BE5A89BBAB2900BF7D90C5EEB" bold="true">D</emphasis>
. Scale bars: 200 µm (
<emphasis id="9A0D94B73F9314DFE3D8DF934C2CB095" bold="true">A, B</emphasis>
); 500 µm (
<emphasis id="2D05D7FA315FFADB734F091FC5F1E98D" bold="true">C, D</emphasis>
).
</paragraph>
</caption>
<paragraph id="DAC0B7D429B2CC4E5F9916D68F5C49F4">
Pedipalps (Fig.
<figureCitation id="C5B5BE21EF4E3944158F1DB55F1FB8B2" captionStart="Figure 82" captionStartId="F82" captionText="Figure 82. Nerudiella caramavida sp. nov. chelicerae: mesal A ectal B pedipalps: mesal C ectal D. Scale bars: 200 µm (A, B); 500 µm (C, D)." figureDoi="10.3897/zookeys.1207.120068.figure82" httpUri="https://binary.pensoft.net/fig/1090695">82 C, D</figureCitation>
). Trochanter with tiny dorsal and ventral tubercles. Femora with two robust ventral-proximal tubercles and four minor ventral-distal tubercles. Additionally, with a row of setiferous tubercles along dorsal surface of femora, with the three largest tubercles located in the proximal section. Patella with two small ventral-ectal tubercles and two small ventral-mesal tubercles. Tibia with two rows of weaker apical tubercles and two rows of minor ventral tubercles. Tarsus characterized by three mesal and ectal spines; Also, with a few setae and subdistal setae in this region.
</paragraph>
<paragraph id="3677F65FBB91BF991263BE64BC7A9CD0">
Legs (Fig.
<figureCitation id="81E56C284240722DF9173767A7212E99" captionStart="Figure 83" captionStartId="F83" captionText="Figure 83. Nerudiella caramavida sp. nov. legs I A II B III C IV D. Scale bars: 500 µm (A, B, D); 200 µm (C)." figureDoi="10.3897/zookeys.1207.120068.figure83" httpUri="https://binary.pensoft.net/fig/1090696">83</figureCitation>
). Segment I with nine or ten setiferous tubercles; segment II with 1820 setiferous tubercles; segment IV with five or six small tubercles connected to the opisthosoma. Spiracles not obstructed by bridges. Smooth areas with ~ 1 / 3 of leg II smooth, with two or three small tubercles on each side featuring subdistal setae. The smooth portion extends to ¾ of leg III and &lt;1 / 3 of leg IV. Sternum arrow-shaped. Legs smooth, with notch in tarsus I. Tarsal count: 36 / 7 4 4.
</paragraph>
<caption id="2F43AB0E6372125AF944ADE6A0684DFD" ID-DOI="10.3897/zookeys.1207.120068.figure83" ID-arpha="05365973-0DC2-544C-A23F-7B3269E39ADE" httpUri="https://binary.pensoft.net/fig/1090696" startId="F83">
<paragraph id="49E9B44C1A0AE9CEB02622219B2244A1">
<label id="E471DAC95945235BFBCFBA9EFCCC33C1">Figure 83.</label>
</paragraph>
<paragraph id="C724BEA1F347B0082E6C2E8CC4FC8701">
<taxonomicName id="C129E68E5A085970143DC815E8AAA1BA" authorityName="Porto &amp; Derkarabetian &amp; Giribet &amp; Pérez-González" authorityYear="2024" class="Arachnida" family="Triaenonychidae" genus="Nerudiella" kingdom="Animalia" order="Opiliones" phylum="Arthropoda" rank="species" species="caramavida" status="sp. nov.">
<emphasis id="E69FF081CFB92D63F46FE3A1FFF07FE5" italics="true">Nerudiella caramavida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1CEAC0E2E98811890003D5B22C04EC0C" rank="species">sp. nov.</taxonomicNameLabel>
legs I
<emphasis id="0A08ABA3AB1592C8A034698A421C9584" bold="true">A</emphasis>
II
<emphasis id="D3000D3036618CEE17ECCCD55B3C14F5" bold="true">B</emphasis>
III
<emphasis id="F900BB6F0B46E60CEB8FDAC4765F6E2B" bold="true">C</emphasis>
IV
<emphasis id="D23E9A9DE9F63483BE217801C62A0BDB" bold="true">D</emphasis>
. Scale bars: 500 µm (
<emphasis id="813C49D537C02D3D9105DAD06F4CD5B3" bold="true">A, B, D</emphasis>
); 200 µm (
<emphasis id="AABAAB49A6B2D8511D6C257086B0609B" bold="true">C</emphasis>
).
</paragraph>
</caption>
<paragraph id="7149CCC8CEFE91C43EDD89E4A12E1F6E">
Penis (Figs
<figureCitation id="31E8BE8DAA2191C076F693AFCC44625D" captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Nerudiella caramavida sp. nov. penis: ventral A, B apical C lateral D, E dorsal F, G. Scale bars: 200 µm (A, D, F); 50 µm (B, C, E, G)." figureDoi="10.3897/zookeys.1207.120068.figure84" httpUri="https://binary.pensoft.net/fig/1090697">84</figureCitation>
,
<figureCitation id="44A252EF838399655DD3E1AD71C9444A" captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Nerudiella caramavida sp. nov. penis: ventral A apical B lateral C dorsal D. Colors: ventral plate (yellow), capsula externa (blue), capsula interna (red). Scale bars: 50 µm (A C); 200 µm (D)." figureDoi="10.3897/zookeys.1207.120068.figure85" httpUri="https://binary.pensoft.net/fig/1090698">85</figureCitation>
). Pars distalis with a large ventral plate bearing a cleft that divides the plate into two halves. Each half with three pointed macrosetae on the ventral surface and one macroseta on the dorsal surface. Capsula externa covering dorsal and lateral surface, its apical region U-shaped. With a dorsolateral plate attached to the pars basalis. Capsula interna longer than the capsula externa, which has a long lateral process, perpendicular to the axis of the genitalia, with a visible stylus in its apical portion.
</paragraph>
<caption id="54D4D463EB18D808F4378581596E2BE7" ID-DOI="10.3897/zookeys.1207.120068.figure84" ID-arpha="2F224DC8-1DDF-59E2-859B-82648342F5EB" httpUri="https://binary.pensoft.net/fig/1090697" startId="F84">
<paragraph id="D8AFF1013DD95D654281760A76EF0068">
<label id="EE9B3AF2A40CE2C7B6096A911E598CEF">Figure 84.</label>
</paragraph>
<paragraph id="40965F705A0AB8D0B3B65AFA38438284">
<taxonomicName id="E612EB31C8A41D5B7C06630EA9DA6AEB" authorityName="Porto &amp; Derkarabetian &amp; Giribet &amp; Pérez-González" authorityYear="2024" class="Arachnida" family="Triaenonychidae" genus="Nerudiella" kingdom="Animalia" order="Opiliones" phylum="Arthropoda" rank="species" species="caramavida" status="sp. nov.">
<emphasis id="9FF2604D1FF99C1037E5C083FFA56D5A" italics="true">Nerudiella caramavida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F738E491A4ECF716C47CEE37BA54671D" rank="species">sp. nov.</taxonomicNameLabel>
penis: ventral
<emphasis id="A2AA280B95CBDDF6B0D3553FBE5A4479" bold="true">A, B</emphasis>
apical
<emphasis id="BF57DF8921FE3FE8187FAB590B556CA7" bold="true">C</emphasis>
lateral
<emphasis id="F529C700FB5B7FE3B4C7CB7958115216" bold="true">D, E</emphasis>
dorsal
<emphasis id="B40977E9951A6E92E8AB5B388742F101" bold="true">F, G</emphasis>
. Scale bars: 200 µm (
<emphasis id="E1D952A91C0DC5DF30C637C7218E1856" bold="true">A, D, F</emphasis>
); 50 µm (
<emphasis id="352FBE85BD34D94E62B7CF56BA3A80F5" bold="true">B, C, E, G</emphasis>
).
</paragraph>
</caption>
<caption id="0B5119FBBC8FDF70FFB149E09C9AC133" ID-DOI="10.3897/zookeys.1207.120068.figure85" ID-arpha="108A2A67-BD3C-5D4D-BDA6-EABC9AE1D44F" httpUri="https://binary.pensoft.net/fig/1090698" startId="F85">
<paragraph id="27DCD524E7A96ED00D0D53644A1BE947">
<label id="1A2E007180B9CB4584C3EEC3E0E2CB99">Figure 85.</label>
</paragraph>
<paragraph id="988258D8D819C92EB3429A0596F8C418">
<taxonomicName id="EAE6EEB32ECE4FBF2B77CAA3D316B4DA" authorityName="Porto &amp; Derkarabetian &amp; Giribet &amp; Pérez-González" authorityYear="2024" class="Arachnida" family="Triaenonychidae" genus="Nerudiella" kingdom="Animalia" order="Opiliones" phylum="Arthropoda" rank="species" species="caramavida" status="sp. nov.">
<emphasis id="DD100DBA3CA88D583DA72A86A83E9139" italics="true">Nerudiella caramavida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F0034EE50432C9B342EFB0576B9A55B7" rank="species">sp. nov.</taxonomicNameLabel>
penis: ventral
<emphasis id="EB2266B8C1FEFF20C8BC38F353AED038" bold="true">A</emphasis>
apical
<emphasis id="8ABFC17D3EE3FC9A000DDC09E515D6CB" bold="true">B</emphasis>
lateral
<emphasis id="D304530E69707C086E6E66AF0FE436DA" bold="true">C</emphasis>
dorsal
<emphasis id="77D3E870BE2AB204AF41CC90DEFE8F25" bold="true">D</emphasis>
. Colors: ventral plate (yellow), capsula externa (blue), capsula interna (red). Scale bars: 50 µm (
<emphasis id="B8F6A7C0F80D1C3D849CC07BA5E2CE81" bold="true">A C</emphasis>
); 200 µm (
<emphasis id="DCC60AAA56B924472FF46B45425A9DC0" bold="true">D</emphasis>
).
</paragraph>
</caption>
<paragraph id="904E5524E3232A00E682215DF6B81676">
<emphasis id="243D07C69C362EC7B05C1A60B9C5944F" bold="true">Female.</emphasis>
Similar to male, with shorter pedipalpal femora.
</paragraph>
<paragraph id="C6EC04A4365C87B6DBC5D015AC920F14">Female measurements. Total length 2.6, carapace length 1.13, dorsal scutum length 2.1, carapace max. width 1.39, mesotergum max. width 1.98. Appendage measurements: Pedipalps. Trochanter length 0.26, femora length 0.99, patella length 0.46, tibia length 0.70, tarsus length 0.87. Leg I: trochanter (tr) 0.24, femora (fe) 1.04, patella (pa) 0.54, tibia (ti) 0.79, metatarsus (mt) 0.94, tarsus (ta) 0.78. II: tr 0.26, fe 1.51, pa 0.64, ti 1.08, mt 1.26, ta 1.44. III: tr 0.31, fe 0.93, pa 0.41, ti 0.69, mt 0.78, ta 0.78. IV: tr 0.32, fe 1.26, pa 0.61, ti 1.09, mt 1.28, ta 0.97.</paragraph>
</subSubSection>
</treatment>
</document>