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<document id="CB4951A9B364E1FD8D9387CEC6D4F446" ID-CLB-Dataset="32076" ID-DOI="10.11646/zootaxa.4320.3.6" ID-GBIF-Dataset="e80fc464-0c2a-4003-a867-ab8b5b3b399e" ID-ISSN="1175-5326" ID-Zenodo-Dep="1051951" ID-ZooBank="Ca50E5A5-B417-4000-B67D-5F6028Da9154" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1505712517045" checkinUser="plazi" docAuthor="Bocak, Ladislav" docDate="2017" docId="A312190FEC1DFF8A509F38DABFEBFE79" docLanguage="en" docName="zootaxa.4320.3.6.pdf" docOrigin="Zootaxa 4320 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cladophorus mindikensis Bocak 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="514" masterDocId="5F2B6177EC15FF8350083D11BB5CFFD4" masterDocTitle="Species delimitation of colour polymorphic Cladophorus (Coleoptera: Lycidae) from New Guinea" masterLastPageNumber="522" masterPageNumber="505" pageNumber="513" updateTime="1698468596363" updateUser="ExternalLinkService">
<mods:mods id="BD5D66D25D94073DCE2D87AC5F7694EC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="20E03053CD051F9EE3332458CAC6FF31">
<mods:title id="5628191B84E4F205B1E4C2A2F4D59A38">Species delimitation of colour polymorphic Cladophorus (Coleoptera: Lycidae) from New Guinea</mods:title>
</mods:titleInfo>
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</mods:role>
<mods:namePart id="DE0FB76795E4B396BA03792A5D9FC9B8">Bocak, Ladislav</mods:namePart>
</mods:name>
<mods:typeOfResource id="C32D570E61798FA49607689549894B80">text</mods:typeOfResource>
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<mods:title id="C5109CF02EE4042C60C51A694B930525">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="4CE128E08CF2F15FB4F6FD2AE907AAEC">
<mods:date id="89516EE5186D86339457608CA3FF3E76">2017</mods:date>
<mods:detail id="6F900456306BFA7DDB5DAB9B2DFBA209" type="pubDate">
<mods:number id="1FC2BE482E36E600B16554DFF3A0D0B1">2017-09-18</mods:number>
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<mods:number id="EE61AF5147EFFCB41070A2893E592418">4320</mods:number>
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<mods:classification id="82E09154E63DC84D5B239740866D32FA">journal article</mods:classification>
<mods:identifier id="B6E6B328E95159F99CB8103C289DD2F4" type="CLB-Dataset">32076</mods:identifier>
<mods:identifier id="EB5C1445CD9BDB1B2DF39ED66A59CE6F" type="DOI">10.11646/zootaxa.4320.3.6</mods:identifier>
<mods:identifier id="DB3713B293079B6657C68ED952CD5D6E" type="GBIF-Dataset">e80fc464-0c2a-4003-a867-ab8b5b3b399e</mods:identifier>
<mods:identifier id="D0241E70FAF95B1363664778CC6836D7" type="ISSN">1175-5326</mods:identifier>
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<treatment id="A312190FEC1DFF8A509F38DABFEBFE79" ID-DOI="http://doi.org/10.5281/zenodo.6018063" ID-GBIF-Taxon="134090572" ID-Zenodo-Dep="6018063" LSID="urn:lsid:plazi:treatment:A312190FEC1DFF8A509F38DABFEBFE79" httpUri="http://treatment.plazi.org/id/A312190FEC1DFF8A509F38DABFEBFE79" lastPageId="9" lastPageNumber="514" pageId="8" pageNumber="513">
<subSubSection id="63A1FB92EC1DFF8B509F38DABAF0F9D2" pageId="8" pageNumber="513" type="nomenclature">
<paragraph id="2B04A819EC1DFF8B509F38DAB960FA31" blockId="8.[151,572,1483,1542]" box="[151,572,1483,1509]" pageId="8" pageNumber="513">
<heading id="704C1F75EC1DFF8B509F38DAB960FA31" bold="true" box="[151,572,1483,1509]" fontSize="11" level="1" pageId="8" pageNumber="513" reason="1">
<emphasis id="19CF740BEC1DFF8B509F38DAB960FA31" bold="true" box="[151,572,1483,1509]" pageId="8" pageNumber="513">
<taxonomicName id="ECBBD39AEC1DFF8B509F38DABA8BFA31" authority="Bocak, 2017" authorityName="Bocak" authorityYear="2017" box="[151,471,1483,1509]" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="513" phylum="Arthropoda" rank="species" species="mindikensis" status="sp. nov.">Cladophorus mindikensis</taxonomicName>
<taxonomicNameLabel id="02FCC970EC1DFF8B51D638DAB960FA31" box="[478,572,1483,1509]" pageId="8" pageNumber="513" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="2B04A819EC1DFF8B509F38FCBAF0F9D2" blockId="8.[151,572,1483,1542]" box="[151,428,1517,1542]" pageId="8" pageNumber="513">
(
<figureCitation id="B380B49CEC1DFF8B509738FCBA7AF9D2" box="[159,294,1517,1542]" captionStart="FIGURES 6 – 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 – 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="8" pageNumber="513">Figs. 13, 27</figureCitation>
,
<figureCitation id="B380B49CEC1DFF8B513A38FFBA0DF9D2" box="[306,337,1518,1542]" captionStart="FIGURES 33 – 52" captionStartId="12.[151,264,1876,1899]" captionTargetBox="[174,1412,193,1843]" captionTargetId="figure@12.[174,1418,193,1871]" captionTargetPageId="12" captionText="FIGURES 33 – 52. Elytron. 33 – 36, Cladophorus bicolor Kleine; 37, C. boceki sp. nov.; 38, C. craterensis sp. nov.; 39, C. haiaensis sp. nov.; 40, C. humeralis Kleine; 41, C. kailakiensis sp. nov.; 42, C. manokwarensis sp. nov.; 43, C. mindikensis sp. nov.; 44 – 45, C. motykai sp. nov.; 46, C. pallescens sp. nov.; 47, C. praecipuus Kleine; 48, C. riedeli sp. nov.; 49, C. wasiorensis sp. nov. Structure of elytral costae in C. bicolor Kleine. 50, humeral part of the elytron; 51, transition between the dark and light colored part of the elytron; 52, apical part of the elytron. Scales: elytron 1.5 mm; elytra detail 0.2 mm." httpUri="https://zenodo.org/record/1051959/files/figure.png" pageId="8" pageNumber="513">43</figureCitation>
,
<figureCitation id="B380B49CEC1DFF8B515538FCBAF8F9D2" box="[349,420,1517,1542]" captionStart="FIGURES 53 – 78" captionStartId="13.[151,264,1895,1918]" captionTargetBox="[174,1418,196,1884]" captionTargetId="figure@13.[171,1423,193,1887]" captionTargetPageId="13" captionText="FIGURES 53 – 78. Male genitalia ventrally and laterally. 53 – 54, Cladophorus bicolor Kleine; 55 – 56, C. boceki sp. nov.; 57 – 58, C. craterensis sp. nov.; 59 – 60, C. haiaensis sp. nov.; 61 – 62, C. humeralis Kleine; 63 – 64, C. kailakiensis sp. nov.; 65 – 66, C. manokwarensis sp. nov.; 67 – 68, C. mindikensis sp. nov.; 69 – 70, C. motykai sp. nov.; 71 – 72, C. pallescens sp. nov.; 73 – 74, C. praecipuus Kleine; 75 – 76, C. riedeli sp. nov.; 77 – 78, C. wasiorensis sp. nov. Scales 0.5 mm." httpUri="https://zenodo.org/record/1051961/files/figure.png" pageId="8" pageNumber="513">67–68</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="63A1FB92EC1DFF8B509F3B24B9B2F9A6" pageId="8" pageNumber="513" type="materials_examined">
<paragraph id="2B04A819EC1DFF8B509F3B24B9B2F9A6" blockId="8.[151,1437,1589,2010]" pageId="8" pageNumber="513">
<materialsCitation id="9BD3A244EC1DFF8B509F3B24BEC4F99A" ID-GBIF-Occurrence="1638540614" box="[151,1432,1589,1614]" collectionCode="LMBC" country="Papua New Guinea" elevation="1575" latitude="-6.45" location="Mindik" longLatPrecision="1304" longitude="147.41667" pageId="8" pageNumber="513" specimenCount="2" typeStatus="holotype">
<emphasis id="19CF740BEC1DFF8B509F3B24BA18F99A" bold="true" box="[151,324,1589,1614]" pageId="8" pageNumber="513">
<typeStatus id="F40016BBEC1DFF8B509F3B24BB8EF99A" box="[151,210,1589,1614]" pageId="8" pageNumber="513">Type</typeStatus>
material.
</emphasis>
<typeStatus id="F40016BBEC1DFF8B51423B24BAE9F99A" box="[330,437,1589,1614]" pageId="8" pageNumber="513" type="holotype">Holotype</typeStatus>
, male.
<collectingCountry id="53ACE889EC1DFF8B520C3B27B982F99A" box="[516,734,1589,1614]" name="Papua New Guinea" pageId="8" pageNumber="513">Papua New Guinea</collectingCountry>
,
<location id="2E64FEC2EC1DFF8B52E03B24B862F99A" LSID="urn:lsid:plazi:treatment:A312190FEC1DFF8A509F38DABFEBFE79:2E64FEC2EC1DFF8B52E03B24B862F99A" box="[744,830,1589,1614]" country="Papua New Guinea" latitude="-6.45" longLatPrecision="1304" longitude="147.41667" name="Mindik" pageId="8" pageNumber="513">Mindik</location>
,
<geoCoordinate id="4E8FCEDEEC1DFF8B53403B24B8F8F99A" box="[840,932,1589,1614]" direction="south" orientation="latitude" pageId="8" pageNumber="513" precision="925" value="-6.45">06°27´S</geoCoordinate>
<geoCoordinate id="4E8FCEDEEC1DFF8B53A23B24BF44F99A" box="[938,1048,1589,1614]" direction="east" orientation="longitude" pageId="8" pageNumber="513" precision="925" value="147.41667">147°25´E</geoCoordinate>
,
<elevation id="A0964F2AEC1DFF8B542B3B24BF9CF999" box="[1059,1216,1589,1614]" metricMagnitude="3" metricUnit="m" metricValue="1.575" metricValueMax="1.65" metricValueMin="1.5" pageId="8" pageNumber="513" unit="m" value="1575.0" valueMax="1650.0" valueMin="1500.0">
<quantity id="EC4305FCEC1DFF8B542B3B24BF9CF999" box="[1059,1216,1589,1614]" metricMagnitude="3" metricUnit="m" metricValue="1.575" metricValueMax="1.65" metricValueMin="1.5" pageId="8" pageNumber="513" unit="m" value="1575.0" valueMax="1650.0" valueMin="1500.0">1500–1650 m</quantity>
</elevation>
, (A00181,
<collectionCode id="4DAA30DCEC1DFF8B55343B27BECCF99A" box="[1340,1424,1590,1614]" pageId="8" pageNumber="513">LMBC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="9BD3A244EC1DFF8B509F3B4BB9B6F9A6" ID-GBIF-Occurrence="1638540603" box="[151,746,1625,1650]" collectionCode="LMBC" country="Papua New Guinea" elevation="1575" latitude="-6.45" location="Mindik" longLatPrecision="1304" longitude="147.41667" pageId="8" pageNumber="513" specimenCount="1" typeStatus="paratype">
<typeStatus id="F40016BBEC1DFF8B509F3B4BBBA0F9A6" box="[151,252,1626,1650]" pageId="8" pageNumber="513" type="paratype">Paratype</typeStatus>
, male, same locality data (A00186,
<collectionCode id="4DAA30DCEC1DFF8B52863B4BB9BEF9A6" box="[654,738,1626,1650]" pageId="8" pageNumber="513">LMBC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="63A1FB92EC1DFF8B50CF3B6CBF23F8D6" pageId="8" pageNumber="513" type="diagnosis">
<paragraph id="2B04A819EC1DFF8B50CF3B6CBF23F8D6" blockId="8.[151,1437,1589,2010]" pageId="8" pageNumber="513">
<emphasis id="19CF740BEC1DFF8B50CF3B6CBA1DF942" bold="true" box="[199,321,1661,1686]" pageId="8" pageNumber="513">Diagnosis.</emphasis>
<taxonomicName id="ECBBD39AEC1DFF8B51403B6FBAB2F941" authorityName="Bocak" authorityYear="2017" box="[328,494,1661,1685]" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="513" phylum="Arthropoda" rank="species" species="mindikensis">
<emphasis id="19CF740BEC1DFF8B51403B6FBAB2F941" box="[328,494,1661,1685]" italics="true" pageId="8" pageNumber="513">C. mindikensis</emphasis>
</taxonomicName>
has uniquely coloured and wide elytra (
<figureCitation id="B380B49CEC1DFF8B53B83B6CBF59F942" box="[944,1029,1661,1686]" captionStart="FIGURES 33 – 52" captionStartId="12.[151,264,1876,1899]" captionTargetBox="[174,1412,193,1843]" captionTargetId="figure@12.[174,1418,193,1871]" captionTargetPageId="12" captionText="FIGURES 33 – 52. Elytron. 33 – 36, Cladophorus bicolor Kleine; 37, C. boceki sp. nov.; 38, C. craterensis sp. nov.; 39, C. haiaensis sp. nov.; 40, C. humeralis Kleine; 41, C. kailakiensis sp. nov.; 42, C. manokwarensis sp. nov.; 43, C. mindikensis sp. nov.; 44 – 45, C. motykai sp. nov.; 46, C. pallescens sp. nov.; 47, C. praecipuus Kleine; 48, C. riedeli sp. nov.; 49, C. wasiorensis sp. nov. Structure of elytral costae in C. bicolor Kleine. 50, humeral part of the elytron; 51, transition between the dark and light colored part of the elytron; 52, apical part of the elytron. Scales: elytron 1.5 mm; elytra detail 0.2 mm." httpUri="https://zenodo.org/record/1051959/files/figure.png" pageId="8" pageNumber="513">Fig. 43</figureCitation>
). Each elytron has an extensive pale yellow patch in the apical part and the margin of the elytron is completely dark coloured (
<figureCitation id="B380B49CEC1DFF8B54863BB0BFBFF96E" box="[1166,1251,1697,1722]" captionStart="FIGURES 33 – 52" captionStartId="12.[151,264,1876,1899]" captionTargetBox="[174,1412,193,1843]" captionTargetId="figure@12.[174,1418,193,1871]" captionTargetPageId="12" captionText="FIGURES 33 – 52. Elytron. 33 – 36, Cladophorus bicolor Kleine; 37, C. boceki sp. nov.; 38, C. craterensis sp. nov.; 39, C. haiaensis sp. nov.; 40, C. humeralis Kleine; 41, C. kailakiensis sp. nov.; 42, C. manokwarensis sp. nov.; 43, C. mindikensis sp. nov.; 44 – 45, C. motykai sp. nov.; 46, C. pallescens sp. nov.; 47, C. praecipuus Kleine; 48, C. riedeli sp. nov.; 49, C. wasiorensis sp. nov. Structure of elytral costae in C. bicolor Kleine. 50, humeral part of the elytron; 51, transition between the dark and light colored part of the elytron; 52, apical part of the elytron. Scales: elytron 1.5 mm; elytra detail 0.2 mm." httpUri="https://zenodo.org/record/1051959/files/figure.png" pageId="8" pageNumber="513">Fig. 43</figureCitation>
).
<taxonomicName id="ECBBD39AEC1DFF8B54FE3BB3BEC0F96D" authorityName="Bocak" authorityYear="2017" box="[1270,1436,1697,1721]" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="513" phylum="Arthropoda" rank="species" species="kailakiensis">
<emphasis id="19CF740BEC1DFF8B54FE3BB3BEC0F96D" box="[1270,1436,1697,1721]" italics="true" pageId="8" pageNumber="513">C. kailakiensis</emphasis>
</taxonomicName>
is its sister species (
<figureCitation id="B380B49CEC1DFF8B51883BD4BA9AF90A" box="[384,454,1733,1758]" captionStart="FIGURES 4 – 5. 4" captionStartId="3.[151,264,1220,1243]" captionTargetBox="[159,1417,255,1198]" captionTargetId="figure@3.[151,1436,255,1210]" captionTargetPageId="3" captionText="FIGURES 4 – 5. 4, Phylogenetic hypothesis of Papuan Cladophorus inferred from the cox 1 mtDNA dataset using maximum likelihood criterion. 5, Cladophorus sp., general appearance." httpUri="https://zenodo.org/record/1051955/files/figure.png" pageId="8" pageNumber="513">Fig. 4</figureCitation>
) and differs in the lamella of antennomere 3 (
<figureCitation id="B380B49CEC1DFF8B53E93BD4BF37F90A" box="[993,1131,1733,1758]" captionStart="FIGURES 6 – 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 – 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="8" pageNumber="513">Figs. 11, 13</figureCitation>
) and the coloration of the pronotum (
<figureCitation id="B380B49CEC1DFF8B511B3BF8BAC0F8D6" box="[275,412,1769,1794]" captionStart="FIGURES 6 – 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 – 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="8" pageNumber="513">Figs. 25, 27</figureCitation>
). Male genitalia of these species are similar (
<figureCitation id="B380B49CEC1DFF8B539D3BF8BF2EF8D6" box="[917,1138,1769,1794]" captionStart="FIGURES 53 – 78" captionStartId="13.[151,264,1895,1918]" captionTargetBox="[174,1418,196,1884]" captionTargetId="figure@13.[171,1423,193,1887]" captionTargetPageId="13" captionText="FIGURES 53 – 78. Male genitalia ventrally and laterally. 53 – 54, Cladophorus bicolor Kleine; 55 – 56, C. boceki sp. nov.; 57 – 58, C. craterensis sp. nov.; 59 – 60, C. haiaensis sp. nov.; 61 – 62, C. humeralis Kleine; 63 – 64, C. kailakiensis sp. nov.; 65 – 66, C. manokwarensis sp. nov.; 67 – 68, C. mindikensis sp. nov.; 69 – 70, C. motykai sp. nov.; 71 – 72, C. pallescens sp. nov.; 73 – 74, C. praecipuus Kleine; 75 – 76, C. riedeli sp. nov.; 77 – 78, C. wasiorensis sp. nov. Scales 0.5 mm." httpUri="https://zenodo.org/record/1051961/files/figure.png" pageId="8" pageNumber="513">Figs. 63–64, 67–68</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="63A1FB92EC1DFF8A50CF3A1CB8B7FEB0" lastPageId="9" lastPageNumber="514" pageId="8" pageNumber="513" type="description">
<paragraph id="2B04A819EC1DFF8A50CF3A1CB927FEC9" blockId="8.[151,1437,1589,2010]" lastBlockId="9.[151,1436,151,429]" lastPageId="9" lastPageNumber="514" pageId="8" pageNumber="513">
<emphasis id="19CF740BEC1DFF8B50CF3A1CBA05F8F2" bold="true" box="[199,345,1805,1830]" pageId="8" pageNumber="513">Description.</emphasis>
Male. Body large,
<quantity id="EC4305FCEC1DFF8B523D3A1FB986F8F1" box="[565,730,1806,1830]" metricMagnitude="-2" metricUnit="m" metricValue="1.3800000000000001" metricValueMax="1.3900000000000001" metricValueMin="1.3699999999999999" pageId="8" pageNumber="513" unit="mm" value="13.8" valueMax="13.9" valueMin="13.7">13.7–13.9 mm</quantity>
long, body dark brown to black, pronotum and most of elytra black, light brown patch in apical third of elytra, whole elytral margin black, transition between light and black parts of elytra clearly delimited (
<figureCitation id="B380B49CEC1DFF8B521E3A44B92BF8BA" box="[534,631,1877,1902]" captionStart="FIGURES 6 – 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 – 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="8" pageNumber="513">Figs. 27</figureCitation>
,
<figureCitation id="B380B49CEC1DFF8B528E3A47B9FAF8BA" box="[646,678,1878,1902]" captionStart="FIGURES 33 – 52" captionStartId="12.[151,264,1876,1899]" captionTargetBox="[174,1412,193,1843]" captionTargetId="figure@12.[174,1418,193,1871]" captionTargetPageId="12" captionText="FIGURES 33 – 52. Elytron. 33 – 36, Cladophorus bicolor Kleine; 37, C. boceki sp. nov.; 38, C. craterensis sp. nov.; 39, C. haiaensis sp. nov.; 40, C. humeralis Kleine; 41, C. kailakiensis sp. nov.; 42, C. manokwarensis sp. nov.; 43, C. mindikensis sp. nov.; 44 – 45, C. motykai sp. nov.; 46, C. pallescens sp. nov.; 47, C. praecipuus Kleine; 48, C. riedeli sp. nov.; 49, C. wasiorensis sp. nov. Structure of elytral costae in C. bicolor Kleine. 50, humeral part of the elytron; 51, transition between the dark and light colored part of the elytron; 52, apical part of the elytron. Scales: elytron 1.5 mm; elytra detail 0.2 mm." httpUri="https://zenodo.org/record/1051959/files/figure.png" pageId="8" pageNumber="513">43</figureCitation>
). Head small, with small hemispherically prominent eyes, frontal interocular distance 1.8 times maximum diameter; antennae flabellate, long, reaching behind middle of elytral length, antennomere 3 with lamella 1.7 times longer than stem of antennomere, lamella attached to apical part of antennomere (
<figureCitation id="B380B49CEC1DFF8B513E3AD0BAD5F80E" box="[310,393,1985,2010]" captionStart="FIGURES 6 – 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 – 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="8" pageNumber="513">Fig. 13</figureCitation>
). Pronotum transverse, 1.5 times wider than long at midline, lateral margins slightly elevated, frontal margin slightly anteriorly projected, frontal angles inconspicuous, lateral margins straight to slightly concave, posterior angles acutely projected, all carinae well-developed (
<figureCitation id="B380B49CEC1CFF8A53EB3DADBF60FF00" box="[995,1084,188,213]" captionStart="FIGURES 6 – 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 – 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="9" pageNumber="514">Fig. 27</figureCitation>
). Elytra flat, wide posteriorly, transverse costae well-developed, dense, regular, elytral areoles transverse (
<figureCitation id="B380B49CEC1CFF8A53EF3DCEBF60FF2C" box="[999,1084,223,248]" captionStart="FIGURES 33 – 52" captionStartId="12.[151,264,1876,1899]" captionTargetBox="[174,1412,193,1843]" captionTargetId="figure@12.[174,1418,193,1871]" captionTargetPageId="12" captionText="FIGURES 33 – 52. Elytron. 33 – 36, Cladophorus bicolor Kleine; 37, C. boceki sp. nov.; 38, C. craterensis sp. nov.; 39, C. haiaensis sp. nov.; 40, C. humeralis Kleine; 41, C. kailakiensis sp. nov.; 42, C. manokwarensis sp. nov.; 43, C. mindikensis sp. nov.; 44 – 45, C. motykai sp. nov.; 46, C. pallescens sp. nov.; 47, C. praecipuus Kleine; 48, C. riedeli sp. nov.; 49, C. wasiorensis sp. nov. Structure of elytral costae in C. bicolor Kleine. 50, humeral part of the elytron; 51, transition between the dark and light colored part of the elytron; 52, apical part of the elytron. Scales: elytron 1.5 mm; elytra detail 0.2 mm." httpUri="https://zenodo.org/record/1051959/files/figure.png" pageId="9" pageNumber="514">Fig. 42</figureCitation>
). Phallus robust, parallel-sided, with long slender apical part (
<figureCitation id="B380B49CEC1CFF8A51ED3C15B933FEC9" box="[485,623,260,285]" captionStart="FIGURES 53 – 78" captionStartId="13.[151,264,1895,1918]" captionTargetBox="[174,1418,196,1884]" captionTargetId="figure@13.[171,1423,193,1887]" captionTargetPageId="13" captionText="FIGURES 53 – 78. Male genitalia ventrally and laterally. 53 – 54, Cladophorus bicolor Kleine; 55 – 56, C. boceki sp. nov.; 57 – 58, C. craterensis sp. nov.; 59 – 60, C. haiaensis sp. nov.; 61 – 62, C. humeralis Kleine; 63 – 64, C. kailakiensis sp. nov.; 65 – 66, C. manokwarensis sp. nov.; 67 – 68, C. mindikensis sp. nov.; 69 – 70, C. motykai sp. nov.; 71 – 72, C. pallescens sp. nov.; 73 – 74, C. praecipuus Kleine; 75 – 76, C. riedeli sp. nov.; 77 – 78, C. wasiorensis sp. nov. Scales 0.5 mm." httpUri="https://zenodo.org/record/1051961/files/figure.png" pageId="9" pageNumber="514">Figs. 67–68</figureCitation>
).
</paragraph>
<paragraph id="2B04A819EC1CFF8A50CF3C36B8B7FEB0" blockId="9.[151,1436,151,429]" pageId="9" pageNumber="514">
<emphasis id="19CF740BEC1CFF8A50CF3C36BA21FE94" bold="true" box="[199,381,295,320]" pageId="9" pageNumber="514">Measurements.</emphasis>
BL
<quantity id="EC4305FCEC1CFF8A51BA3C39B905FEEB" box="[434,601,296,320]" metricMagnitude="-2" metricUnit="m" metricValue="1.3800000000000001" metricValueMax="1.3900000000000001" metricValueMin="1.3699999999999999" pageId="9" pageNumber="514" unit="mm" value="13.8" valueMax="13.9" valueMin="13.7">13.7–13.9 mm</quantity>
, PL
<quantity id="EC4305FCEC1CFF8A52983C36B86BFEEB" box="[656,823,295,320]" metricMagnitude="-3" metricUnit="m" metricValue="1.765" metricValueMax="1.78" metricValueMin="1.75" pageId="9" pageNumber="514" unit="mm" value="1.765" valueMax="1.78" valueMin="1.75">1.75–1.78 mm</quantity>
, PW
<quantity id="EC4305FCEC1CFF8A53703C36B8BCFEEB" box="[888,992,295,320]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" pageId="9" pageNumber="514" unit="mm" value="2.75">2.75 mm</quantity>
, WH
<quantity id="EC4305FCEC1CFF8A542D3C36BFD1FEEB" box="[1061,1165,295,320]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="9" pageNumber="514" unit="mm" value="1.6">1.60 mm</quantity>
, Edist
<quantity id="EC4305FCEC1CFF8A54D53C39BE19FEEB" box="[1245,1349,296,320]" metricMagnitude="-3" metricUnit="m" metricValue="1.08" pageId="9" pageNumber="514" unit="mm" value="1.08">1.08 mm</quantity>
, Ediam
<quantity id="EC4305FCEC1CFF8A509F3C5DBBA1FEB0" box="[151,253,332,357]" metricMagnitude="-4" metricUnit="m" metricValue="5.8999999999999995" pageId="9" pageNumber="514" unit="mm" value="0.59">0.59 mm</quantity>
, LE
<quantity id="EC4305FCEC1CFF8A513A3C5DBA88FEB0" box="[306,468,332,356]" metricMagnitude="-2" metricUnit="m" metricValue="1.1350000000000002" metricValueMax="1.1400000000000001" metricValueMin="1.1300000000000001" pageId="9" pageNumber="514" unit="mm" value="11.35" valueMax="11.4" valueMin="11.3">11.3–11.4 mm</quantity>
, antennomere 3: stem
<quantity id="EC4305FCEC1CFF8A52D93C5DB874FEB0" box="[721,808,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="9" pageNumber="514" unit="mm" value="1.1">1.1 mm</quantity>
, lamella
<quantity id="EC4305FCEC1CFF8A53853C5DB8BBFEB0" box="[909,999,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="9" pageNumber="514" unit="mm" value="1.9">1.9 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="63A1FB92EC1CFF8A50CF3C7EB8DCFE5C" box="[199,896,367,392]" pageId="9" pageNumber="514" type="distribution">
<paragraph id="2B04A819EC1CFF8A50CF3C7EB8DCFE5C" blockId="9.[151,1436,151,429]" box="[199,896,367,392]" pageId="9" pageNumber="514">
<emphasis id="19CF740BEC1CFF8A50CF3C7EBA3CFE5C" bold="true" box="[199,352,367,392]" pageId="9" pageNumber="514">Distribution.</emphasis>
Eastern New
<collectingCountry id="53ACE889EC1CFF8A51F63C7EB90EFE5C" box="[510,594,367,392]" name="Guinea-Bissau" pageId="9" pageNumber="514">Guinea</collectingCountry>
,
<collectingRegion id="E97F66FBEC1CFF8A52553C7EB842FE5C" box="[605,798,367,392]" country="Papua New Guinea" name="Morobe" pageId="9" pageNumber="514">Morobe province</collectingRegion>
(
<figureCitation id="B380B49CEC1CFF8A53263C7EB82EFE5C" box="[814,882,367,392]" captionStart="FIGURES 1 – 3. 1 – 2" captionStartId="2.[151,264,1396,1419]" captionTargetBox="[151,1436,339,1375]" captionTargetId="figure@2.[151,1436,339,1375]" captionTargetPageId="2" captionText="FIGURES 1 – 3. 1 – 2, General appearance of Cladophorus bicolor in the Central mountains of New Guinea. 3, Sampled localities and the distribution of Cladophorus species" httpUri="https://zenodo.org/record/1051953/files/figure.png" pageId="9" pageNumber="514">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="63A1FB92EC1CFF8A50CF3C85BFEBFE79" box="[199,1207,404,429]" pageId="9" pageNumber="514" type="etymology">
<paragraph id="2B04A819EC1CFF8A50CF3C85BFEBFE79" blockId="9.[151,1436,151,429]" box="[199,1207,404,429]" pageId="9" pageNumber="514">
<emphasis id="19CF740BEC1CFF8A50CF3C85BA12FE79" bold="true" box="[199,334,404,429]" pageId="9" pageNumber="514">Etymology.</emphasis>
The epithet refers to the local name ‘Mindik’, the
<typeStatus id="F40016BBEC1CFF8A538C3C84B8E8FE79" box="[900,948,405,429]" pageId="9" pageNumber="514">type</typeStatus>
locality of the species.
</paragraph>
</subSubSection>
</treatment>
</document>
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