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<document id="4D0B6C99FE0D8DE499A977A9F4B5EA3C" ID-DOI="10.1163/18759866-BJA10061" ID-ISSN="1875-9866" ID-Zenodo-Dep="12518921" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1719244263854" checkinUser="felipe" docAuthor="Copilaş-Ciocianu, Denis &amp; Ionesi, Viorel" docDate="2024" docId="6705878F6470FFC1FF1EFA2B2071FBF4" docLanguage="en" docName="ContribZool.93.3.268-288.pdf" docOrigin="Contributions to Zoology 93" docSource="http://dx.doi.org/10.1163/18759866-bja10061" docStyle="DocumentStyle:E1C3C5FD6CDDA08D62BFE928526826AF.3:ContribZool.2019-.journal_article" docStyleId="E1C3C5FD6CDDA08D62BFE928526826AF" docStyleName="ContribZool.2019-.journal_article" docStyleVersion="3" docTitle="Eogmelina Copilaş-Ciocianu &amp; Ionesi, 2024, gen. nov." docType="treatment" docVersion="1" lastPageNumber="276" masterDocId="9B3CFFF76477FFC9FF9FFF932419FFD7" masterDocTitle="New Miocene fossil taxa illuminate the evolution and paleobiogeography of the Ponto-Caspian gammaroid amphipod radiation" masterLastPageNumber="288" masterPageNumber="268" pageNumber="275" updateTime="1729258561795" updateUser="juliana" zenodo-license-document="CC-BY-4.0">
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<mods:title id="2E099C4218F2F0D55799CDB7362F4D01">New Miocene fossil taxa illuminate the evolution and paleobiogeography of the Ponto-Caspian gammaroid amphipod radiation</mods:title>
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<mods:namePart id="1E44B4D85C5AFDD437712816FED77531">Copilaş-Ciocianu, Denis</mods:namePart>
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<mods:affiliation id="A09C8D6DD833201F46A6C29EE501013A">Laboratory of Evolutionary Ecology of Hydrobionts, Nature Research Centre, 08412 Vilnius, Lithuania denis. copilas-ciocianu @ gamtc. lt</mods:affiliation>
<mods:nameIdentifier id="1CF4347DCE8298CA7D91EB478DA3347A" type="email">denis.copilas-ciocianu@gamtc.lt</mods:nameIdentifier>
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<mods:namePart id="184DFABF3A95F8AF8481FD3D52DEE450">Ionesi, Viorel</mods:namePart>
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<mods:affiliation id="B19133090BF4AE70EB8064BB408890AB">Department of Geology, Alexandru Ioan Cuza University of Iași, Iași 700506, Romania RECEIVED 1 FEBRUARY 2024; REVISED 12 MARCH 2024; ACCEPTED 12 MARCH 2024;</mods:affiliation>
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<mods:title id="FC63E4FB8D384F4AFB5519DF8301486F">Contributions to Zoology</mods:title>
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<mods:date id="D9AA98EC4390BE542ACE280A14951F6C">2024</mods:date>
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<paragraph id="EF1336996470FFCEFF1EFA2B25A3FA04" blockId="7.[129,640,1464,1670]" box="[129,442,1464,1491]" pageId="7" pageNumber="275">
Genus †
<taxonomicName id="28AC4D1A6470FFCEFF42FA2B254BFA05" ID-CoL="CCKVL" authorityName="Copilaş-Ciocianu &amp; Ionesi" authorityYear="2024" box="[221,338,1464,1490]" class="Malacostraca" family="Gammaridae" genus="Eogmelina" higherTaxonomySource="CoL" kingdom="Animalia" order="Amphipoda" pageId="7" pageNumber="275" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="DDD8EA8B6470FFCEFF42FA2B254BFA05" box="[221,338,1464,1490]" italics="true" pageId="7" pageNumber="275">Eogmelina</emphasis>
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<taxonomicNameLabel id="C6EB57F06470FFCEFEC7FA2A25A3FA04" box="[344,442,1465,1491]" pageId="7" pageNumber="275" rank="genus">gen. nov.</taxonomicNameLabel>
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Zoobank LSID
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urn:lsid:zoobank.org:act:
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Zoobank publication LSID:
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urn:lsid:zoobank.org:pub:
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species
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. †
<taxonomicName id="28AC4D1A6470FFCEFCD8FB0A202BFB64" authorityName="Copilaş-Ciocianu &amp; Ionesi" authorityYear="2024" box="[839,1074,1177,1203]" class="Malacostraca" family="Gammaridae" genus="Eogmelina" higherTaxonomySource="CoL" kingdom="Animalia" order="Amphipoda" pageId="7" pageNumber="275" phylum="Arthropoda" rank="species" species="moldavica" status="gen. et sp. nov.">
<emphasis id="DDD8EA8B6470FFCEFCD8FB0A202BFB64" box="[839,1074,1177,1203]" italics="true" pageId="7" pageNumber="275">Eogmelina moldavica</emphasis>
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<taxonomicNameLabel id="C6EB57F06470FFCEFBA8FB0A26CBFB00" pageId="7" pageNumber="275" rank="species">gen. et sp. nov.</taxonomicNameLabel>
(
<figureCitation id="77972A1C6470FFCEFD41FB2E270FFB00" box="[734,790,1212,1239]" captionStart="FIGURE 5" captionStartId="9.[129,196,1291,1312]" captionTargetBox="[129,928,164,1263]" captionTargetId="figure-57@9.[129,929,164,1263]" captionTargetPageId="9" captionText="FIGURE 5 †Eogmelina moldavica gen. et sp. nov.On the right are interpretative drawings of the corresponding fossils on the left.(A) Holotype male and paratype female (inventory No. 8236,Vlădiceni quarry), (B) male (inventory No.8237,Vlădiceni quarry), (C) female (inventory No.8238, Iași City), (D) female (inventory No.8239, Vlădiceni quarry).Scale bars = 1 mm. Abbreviations:A = antenna; B = basis; C = coxa; G = gnathopod; H = head; P = pereonite; PL = pleonite; PP = pereopod; U = urosomite; UP = uropod; T = telson." figureDoi="http://doi.org/10.5281/zenodo.12518933" httpUri="https://zenodo.org/record/12518933/files/figure.png" pageId="7" pageNumber="275">fig. 5</figureCitation>
)
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<paragraph id="EF1336996470FFCEFD5DFB722706FAC9" blockId="7.[676,1187,1177,1670]" pageId="7" pageNumber="275">
<emphasis id="DDD8EA8B6470FFCEFD5DFB72274AFB2D" box="[706,851,1249,1275]" italics="true" pageId="7" pageNumber="275">Other species</emphasis>
. †
<taxonomicName id="28AC4D1A6470FFCEFCF0FB722034FB2D" authorityName="Copilaş-Ciocianu &amp; Ionesi" authorityYear="2024" box="[879,1069,1249,1275]" class="Malacostraca" family="Gammaridae" genus="Eogmelina" higherTaxonomySource="CoL" kingdom="Animalia" order="Amphipoda" pageId="7" pageNumber="275" phylum="Arthropoda" rank="species" species="prisca" status="gen. et sp. nov.">
<emphasis id="DDD8EA8B6470FFCEFCF0FB722034FB2D" box="[879,1069,1249,1275]" italics="true" pageId="7" pageNumber="275">Eogmelina prisca</emphasis>
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<taxonomicNameLabel id="C6EB57F06470FFCEFBACFB7226CBFAC8" pageId="7" pageNumber="275" rank="species">gen. et sp. nov.</taxonomicNameLabel>
(
<figureCitation id="77972A1C6470FFCEFD41FA97270EFAC9" box="[734,791,1284,1310]" captionStart="FIGURE 6" captionStartId="10.[98,165,1050,1071]" captionTargetBox="[98,898,164,1022]" captionTargetId="figure-188@10.[98,898,164,1022]" captionTargetPageId="10" captionText="FIGURE 6 †Eogmelina prisca gen.et sp. nov.On the right are interpretative drawings of the corresponding fossils on the left.(A) Holotype male (inventory No. 8241,Vlădiceni quarry), (B) paratype male (inventory No. 8242, Iași City), (C) disarticulated remains of unknown sex (inventory No.8243,Vlădiceni quarry).Scale bars = 1 mm.Abbreviations:A = antenna; B = basis; C = coxa; G = gnathopod; H = head; P = pereonite; PL = pleonite; PP = pereopod; U = urosomite; UP = uropod; T = telson." figureDoi="http://doi.org/10.5281/zenodo.12518935" httpUri="https://zenodo.org/record/12518935/files/figure.png" pageId="7" pageNumber="275">fig. 6</figureCitation>
)
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<subSubSection id="A7B665126470FFC1FD5DFABA2071FBF4" lastPageId="8" lastPageNumber="276" pageId="7" pageNumber="275" type="diagnosis">
<paragraph id="EF1336996470FFC1FD5DFABA24C4FAB1" blockId="7.[676,1187,1177,1670]" lastBlockId="8.[98,610,889,1706]" lastPageId="8" lastPageNumber="276" pageId="7" pageNumber="275">
<emphasis id="DDD8EA8B6470FFCEFD5DFABA2737FA95" box="[706,814,1321,1346]" italics="true" pageId="7" pageNumber="275">Diagnosis</emphasis>
. Body tall and robust (up to
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length) with or without dorsal pleonal projections, lateral pereonal projections absent, coxal plates I-III taller than wide, urosomites free and devoid of humps and tubercles. Antenna 1 peduncle short, not extending beyond the distal end of the 4th peduncular segment of antenna 2. The 3rd peduncular segment of antenna 2 is shorter than half the length of the 4th segment; 4th segment length subequal to head length. Propodi of gnathopods 12 similar in size and shape, rectangular, moderately developed, palm length subequal to posterior margin. Pereopods 37 robust and well developed, dactyli moderately developed and relatively straight. Pereopods 34 carpus longer than wide, not expanded. Length of pereopod 7 exceeds 0.4 of body length. Pereopod 7 basis taller than wide, posterior margin convex, without postero-distal lobe. Epimeral plates 23 with acute postero-inferior corners. Uropod 3 extends beyond the distal end of uropod 1, endopod/exopod ratio is 0.9.
</paragraph>
<caption id="BBD36611647FFFC1FFFDFDD6276BFC92" ID-DOI="http://doi.org/10.5281/zenodo.12518931" ID-Zenodo-Dep="12518931" httpUri="https://zenodo.org/record/12518931/files/figure.png" pageId="8" pageNumber="276" startId="8.[98,165,581,602]" targetBox="[98,961,165,551]" targetPageId="8" targetType="figure">
<paragraph id="EF133699647FFFC1FFFDFDD6276BFC92" blockId="8.[98,933,581,837]" pageId="8" pageNumber="276">FIGURE 4 Morphological diversity of extant and fossil Ponto-Caspian gammaroids based on 43 morphometric measurements. (A) PCA plot depicting the morphospace occupation along the first two axes. Extant non-monotypic genera are shown with dimmed colors (dots or convex hulls if n&gt; 2 species) while monotypic genera are shown with a gray square. Fossil taxa are shown with colored stars that are numbered according to species (see legend on lower left). Extreme morphologies are exemplified by drawings. (B) Biplot of variables along the first two PCA axes. C) A 3D PCA indicating morphospace occupation of fossils (colored triangles) and extant (gray dots) taxa within the first three axes.</paragraph>
</caption>
<paragraph id="EF133699647FFFC1FF1EFAE32656F97E" blockId="8.[98,610,889,1706]" pageId="8" pageNumber="276">
<emphasis id="DDD8EA8B647FFFC1FF1EFAE32574FA5D" box="[129,365,1392,1418]" italics="true" pageId="8" pageNumber="276">Differential diagnosis</emphasis>
.
<taxonomicName id="28AC4D1A647FFFC1FEE2FAE325EBFA5D" authorityName="Copilaş-Ciocianu &amp; Ionesi" authorityYear="2024" box="[381,498,1392,1418]" class="Malacostraca" family="Gammaridae" genus="Eogmelina" higherTaxonomySource="CoL" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="276" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="DDD8EA8B647FFFC1FEE2FAE325EBFA5D" box="[381,498,1392,1418]" italics="true" pageId="8" pageNumber="276">Eogmelina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C6EB57F0647FFFC1FE63FAE2267BFA5C" box="[508,610,1393,1419]" pageId="8" pageNumber="276" rank="genus">gen. nov.</taxonomicNameLabel>
can be readily distinguished from the fossil Ponto-Caspian genera
<taxonomicName id="28AC4D1A647FFFC1FE03FA2B263EFA05" authorityName="Derzhavin" authorityYear="1927" box="[412,551,1464,1490]" class="Malacostraca" family="Pontogammaridae" genus="Praegmelina" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="276" phylum="Arthropoda" rank="genus">
<emphasis id="DDD8EA8B647FFFC1FE03FA2B263EFA05" box="[412,551,1464,1490]" italics="true" pageId="8" pageNumber="276">Praegmelina</emphasis>
</taxonomicName>
and
<taxonomicName id="28AC4D1A647FFFC1FFFDFA4F24F2FA21" authorityName="Derzhavin" authorityYear="1927" box="[98,235,1500,1526]" class="Malacostraca" family="Pontogammaridae" genus="Andrussovia" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="276" phylum="Arthropoda" rank="genus">
<emphasis id="DDD8EA8B647FFFC1FFFDFA4F24F2FA21" box="[98,235,1500,1526]" italics="true" pageId="8" pageNumber="276">Andrussovia</emphasis>
</taxonomicName>
by the following characters: 1) combined length of antenna 2 peduncle segments 4 and 5 longer than head length (vs. shorter), 2) antenna 1 peduncle not reaching the distal end of the 4th peduncular segment of antenna 2 (vs. extending beyond it) (see key).
</paragraph>
<paragraph id="EF133699647FFFC1FD3BFCEA2071FBF4" blockId="8.[645,1157,889,1060]" pageId="8" pageNumber="276">
<emphasis id="DDD8EA8B647FFFC1FD3BFCEA270EFC44" box="[676,791,889,915]" italics="true" pageId="8" pageNumber="276">Etymology</emphasis>
. Derived from the Greek word
<emphasis id="DDD8EA8B647FFFC1FD1AFC0D26B1FC60" box="[645,680,926,951]" italics="true" pageId="8" pageNumber="276">eos</emphasis>
(dawn), referring to its early appearance and plesiomorphic morphology, and the name of the extant genus
<taxonomicName id="28AC4D1A647FFFC1FC31FC762098FC28" authority="Sars, 1894" authorityName="Sars" authorityYear="1894" box="[942,1153,997,1023]" class="Malacostraca" family="Gammaridae" genus="Gmelina" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="276" phylum="Arthropoda" rank="genus">
<emphasis id="DDD8EA8B647FFFC1FC31FC762012FC28" box="[942,1035,997,1023]" italics="true" pageId="8" pageNumber="276">Gmelina</emphasis>
Sars, 1894
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, to which it bares a superficial resemblance.
</paragraph>
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