454 lines
58 KiB
XML
454 lines
58 KiB
XML
<document id="C2E9AB1A161B875EC0C2E37BF0879F29" ID-CLB-Dataset="29979" ID-DOI="10.11646/zootaxa.4426.1.1" ID-GBIF-Dataset="cdd567ed-ebd8-4801-a6a4-af6c9fb964fd" ID-ISSN="1175-5326" ID-Zenodo-Dep="1271239" ID-ZooBank="18929E20-5296-4458-8A8A-4F5316A290FD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1528147699440" checkinUser="plazi" docAuthor="Van, Rob W. M. & De, Nicole J." docDate="2018" docId="386CC616DC53A512FF678A2EFDA0F86A" docLanguage="en" docName="zootaxa.4426.1.1.pdf.imf" docOrigin="Zootaxa 4426 (1)" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Borojevia pirella Van & De 2018, sp.nov." docType="treatment" docVersion="7" lastPageNumber="36" masterDocId="C455BE6EDC72A531FFF0880AFFCEFF99" masterDocTitle="Calcareous sponges of the Western Indian Ocean and Red Sea" masterLastPageNumber="160" masterPageNumber="1" pageNumber="34" updateTime="1698547079698" updateUser="ExternalLinkService">
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<mods:mods id="5739645F7F505662DFD6C08053252CBB" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="064561995EDB0248AA18D01667D79369">
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<mods:title id="1707835EEA558DD12E842AF112790D5A">Calcareous sponges of the Western Indian Ocean and Red Sea</mods:title>
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</mods:titleInfo>
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<mods:name id="8A5681BE4E487DAB3EB15BABE4D32D97" type="personal">
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<mods:roleTerm id="A6BDEB595F805E64D623207662ACA03D">Author</mods:roleTerm>
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<mods:namePart id="F83E83A6EDE869A84A2C3453A63EAFD7">Van, Rob W. M.</mods:namePart>
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<mods:name id="03A1BFB1A50639DACD9196FA88F64E38" type="personal">
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<mods:roleTerm id="A65A09BF6DAF7821775CE5609FA8C364">Author</mods:roleTerm>
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<mods:namePart id="C6E316FC905CC19B9B4F38D4F6A33E59">De, Nicole J.</mods:namePart>
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<mods:typeOfResource id="B2053D5C40A7019B89FE9003B7656D08">text</mods:typeOfResource>
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<mods:title id="8F7D5576E3B7C6EBA4E3CA7F051CC963">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="FCD1601F2D6C03C3B3D2F58FA01A12E9">
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<mods:date id="AE5C7A19782B765731AB2B3503307C4E">2018</mods:date>
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<mods:number id="6A9F457FEF7654038D0ED586FCE350B9">2018-06-01</mods:number>
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<mods:detail id="905599F0418205C3F3ACC250B235B838" type="volume">
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<mods:number id="11AD0C2DA5AF4F580A5746B814B4BFB9">4426</mods:number>
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<mods:detail id="DFBFD83115787846D991E311097EB3F2" type="issue">
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<mods:number id="AF942A3B1C3FE6A4906832C1604FEEAF">1</mods:number>
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<mods:classification id="C408AAEC04A19B17E24EA333EB2D147A">journal article</mods:classification>
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<mods:identifier id="8C3D265BA4AD07BC6019457C1758D66A" type="CLB-Dataset">29979</mods:identifier>
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<mods:identifier id="15DB275819D30E4CA853A21191A8076F" type="DOI">10.11646/zootaxa.4426.1.1</mods:identifier>
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<mods:identifier id="0543EF7B11BC9E044FAA45C38F63EBF9" type="GBIF-Dataset">cdd567ed-ebd8-4801-a6a4-af6c9fb964fd</mods:identifier>
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<mods:identifier id="08B1534E2AC9A8647A24B488AA1F5783" type="ISSN">1175-5326</mods:identifier>
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<treatment id="386CC616DC53A512FF678A2EFDA0F86A" ID-DOI="http://doi.org/10.5281/zenodo.5966696" ID-GBIF-Taxon="144239510" ID-Zenodo-Dep="5966696" LSID="urn:lsid:plazi:treatment:386CC616DC53A512FF678A2EFDA0F86A" httpUri="http://treatment.plazi.org/id/386CC616DC53A512FF678A2EFDA0F86A" lastPageId="35" lastPageNumber="36" pageId="33" pageNumber="34">
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<subSubSection id="F8DF248BDC53A510FF678A2EFE46FDC6" pageId="33" pageNumber="34" type="nomenclature">
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<paragraph id="B07A7700DC53A510FF678A2EFE70FDA7" blockId="33.[151,446,548,607]" box="[151,446,548,574]" pageId="33" pageNumber="34">
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<heading id="EB32C06CDC53A510FF678A2EFE70FDA7" bold="true" box="[151,446,548,574]" fontSize="11" level="1" pageId="33" pageNumber="34" reason="1">
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<emphasis id="82B1AB12DC53A510FF678A2EFE70FDA7" bold="true" box="[151,446,548,574]" pageId="33" pageNumber="34">
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<taxonomicName id="77C50C83DC53A510FF678A2EFEAFFDA7" ID-CoL="MKQ3" authority="Van & De, 2018" authorityName="Van & De" authorityYear="2018" box="[151,353,548,574]" class="Calcarea" family="Clathrinidae" genus="Borojevia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clathrinida" pageId="33" pageNumber="34" phylum="Porifera" rank="species" species="pirella" status="sp. nov.">
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<emphasis id="82B1AB12DC53A510FF678A2EFEAFFDA7" bold="true" box="[151,353,548,574]" italics="true" pageId="33" pageNumber="34">Borojevia pirella</emphasis>
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||
</taxonomicName>
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<taxonomicNameLabel id="99821669DC53A510FE988A2EFE70FDA7" box="[360,446,548,574]" pageId="33" pageNumber="34" rank="species">sp.nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
<paragraph id="B07A7700DC53A510FF678A4CFE46FDC6" blockId="33.[151,446,548,607]" box="[151,392,582,607]" pageId="33" pageNumber="34">
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<figureCitation id="28FE6B85DC53A510FF678A4CFEF7FDC6" box="[151,313,582,607]" captionStart="FIGURE 12" captionStartId="34.[151,249,1884,1906]" captionTargetBox="[151,1436,445,1863]" captionTargetId="figure@34.[151,1436,445,1863]" captionTargetPageId="34" captionText="FIGURE 12. Borojevia pirella sp.nov., a, habitus in situ of holotype RMNH Por. 11622, from Rodrigues, the easternmost of the Mascarene Islands (photo N.J. de Voogd), a1, preserved holotype (scale bar = 1 cm), b, habitus in situ of paratype RMNH Por. 11622b (photo N.J. de Voogd), b1, preserved paratype." httpUri="https://zenodo.org/record/1271289/files/figure.png" pageId="33" pageNumber="34">Figures 12a–b</figureCitation>
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,
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<figureCitation id="28FE6B85DC53A510FEB48A4DFE46FDC6" box="[324,392,583,607]" captionStart="FIGURE 13" captionStartId="35.[151,249,1605,1627]" captionTargetBox="[177,1421,199,1578]" captionTargetId="figure@35.[177,1421,199,1578]" captionTargetPageId="35" captionText="FIGURE 13. Borojevia pirella sp.nov., holotype RMNH Por. 11622, from Rodrigues, a, light microscopic cross section of oscular region showing pseudoatrium and surrounding tubuli, b–e, SEM images of the spicules, b, regular triactine, c, rare sagittal irregular triactine, d, tetractines, e, details of apical actines of the tetractines showing characteristic verticil of three conical spines." httpUri="https://zenodo.org/record/1271295/files/figure.png" pageId="33" pageNumber="34">13a–e</figureCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="F8DF248BDC53A510FF678A84FC90FC8A" pageId="33" pageNumber="34" type="materials_examined">
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||
<paragraph id="B07A7700DC53A510FF678A84FBB3FD52" blockId="33.[151,1437,654,2011]" pageId="33" pageNumber="34">
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<materialsCitation id="00AD7D5DDC53A510FF678A84FBB3FD52" ID-GBIF-Occurrence="1848256243" collectingDate="2016-10-16" collectionCode="RMNH" collectorName="N. J. de Voogd" country="Mauritius" elevation="12" latitude="-19.6683" location="Passe Baladirou" longLatPrecision="5" longitude="63.46307" municipality="Material" pageId="33" pageNumber="34" specimenCount="1" stateProvince="Rodrigues" typeStatus="holotype">
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||
<emphasis id="82B1AB12DC53A510FF678A84FE43FD3E" bold="true" box="[151,397,654,679]" pageId="33" pageNumber="34">
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||
<collectingMunicipality id="501EED7ADC53A510FF678A84FF31FD3E" box="[151,255,654,679]" pageId="33" pageNumber="34">Material</collectingMunicipality>
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||
examined.
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||
</emphasis>
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||
<typeStatus id="6F7EC9A2DC53A510FE508A84FDC5FD3E" box="[416,523,654,679]" pageId="33" pageNumber="34" type="holotype">Holotype</typeStatus>
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||
,
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||
<collectionCode id="D6D4EFC5DC53A510FDD38A85FDB9FD3F" box="[547,631,655,678]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="33" pageNumber="34">RMNH</collectionCode>
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Por. 11622a,
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<collectingCountry id="C8D23790DC53A510FCC68A84FC68FD3E" box="[822,934,654,679]" name="Mauritius" pageId="33" pageNumber="34">Mauritius</collectingCountry>
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||
,
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||
<collectingRegion id="7201B9E2DC53A510FC4E8A84FBFAFD3E" box="[958,1076,654,679]" country="Mauritius" name="Rodrigues" pageId="33" pageNumber="34">Rodrigues</collectingRegion>
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,
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||
<location id="B51A21DBDC53A510FBBC8A85FAC0FD3E" LSID="urn:lsid:plazi:treatment:386CC616DC53A512FF678A2EFDA0F86A:B51A21DBDC53A510FBBC8A85FAC0FD3E" box="[1100,1294,654,679]" country="Mauritius" latitude="-19.6683" longLatPrecision="5" longitude="63.46307" municipality="Material" name="Passe Baladirou" pageId="33" pageNumber="34" stateProvince="Rodrigues">Passe Baladirou</location>
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||
,
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||
<geoCoordinate id="D5F111C7DC53A510FAD68A84FA52FD3E" box="[1318,1436,654,679]" direction="south" orientation="latitude" pageId="33" pageNumber="34" precision="5" value="-19.6683">19.6683°S</geoCoordinate>
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<geoCoordinate id="D5F111C7DC53A510FF678AB8FED1FD52" box="[151,287,690,715]" direction="east" orientation="longitude" pageId="33" pageNumber="34" precision="1" value="63.46307">63.46307°E</geoCoordinate>
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||
, depth
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||
<quantity id="773DDAE5DC53A510FE9F8AB9FE64FD53" box="[367,426,691,715]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="33" pageNumber="34" unit="m" value="12.0">
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||
<elevation id="3BE89033DC53A510FE9F8AB9FE64FD53" box="[367,426,691,715]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="33" pageNumber="34" unit="m" value="12.0">12 m</elevation>
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||
</quantity>
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||
, scuba, coll.
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||
<collectorName id="1D3012D6DC53A510FDCC8AB9FD13FD52" box="[572,733,690,715]" pageId="33" pageNumber="34">N.J. de Voogd</collectorName>
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||
, field nr. ROG023,
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||
<date id="C47B51C0DC53A510FC498AB8FBB7FD52" box="[953,1145,690,715]" pageId="33" pageNumber="34" value="2016-10-16">
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||
<collectingDate id="D43FA828DC53A510FC498AB8FBB7FD52" box="[953,1145,690,715]" pageId="33" pageNumber="34" value="2016-10-16">16 October 2016</collectingDate>
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||
</date>
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||
.
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="B07A7700DC53A510FF378ADDFC90FC8A" blockId="33.[151,1437,654,2011]" pageId="33" pageNumber="34">
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||
<materialsCitation id="00AD7D5DDC53A510FF378ADDFC90FC8A" ID-GBIF-Occurrence="1848256138" collectingDate="2016-10-16" collectionCode="RMNH" collectorName="N. J. de Voogd" country="Mauritius" elevation="12" latitude="-19.6683" location="Passe Baladirou" longLatPrecision="5" longitude="63.46307" pageId="33" pageNumber="34" specimenCount="1" stateProvince="Rodrigues" typeStatus="paratype">
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||
<typeStatus id="6F7EC9A2DC53A510FF378ADDFEE2FD76" box="[199,300,727,751]" pageId="33" pageNumber="34" type="paratype">Paratype</typeStatus>
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||
,
|
||
<collectionCode id="D6D4EFC5DC53A510FECA8ADDFE40FD77" box="[314,398,727,750]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="33" pageNumber="34">RMNH</collectionCode>
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Por. 11622b,
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||
<collectingCountry id="C8D23790DC53A510FDC28ADCFD6CFD76" box="[562,674,726,751]" name="Mauritius" pageId="33" pageNumber="34">Mauritius</collectingCountry>
|
||
,
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||
<collectingRegion id="7201B9E2DC53A510FD408ADCFCE8FD76" box="[688,806,726,751]" country="Mauritius" name="Rodrigues" pageId="33" pageNumber="34">Rodrigues</collectingRegion>
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||
,
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||
<location id="B51A21DBDC53A510FCC48ADDFC23FD76" LSID="urn:lsid:plazi:treatment:386CC616DC53A512FF678A2EFDA0F86A:B51A21DBDC53A510FCC48ADDFC23FD76" box="[820,1005,726,751]" country="Mauritius" latitude="-19.6683" longLatPrecision="5" longitude="63.46307" name="Passe Baladirou" pageId="33" pageNumber="34" stateProvince="Rodrigues">Passe Baladirou</location>
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||
,
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||
<geoCoordinate id="D5F111C7DC53A510FC0B8ADCFBBFFD76" box="[1019,1137,726,751]" direction="south" orientation="latitude" pageId="33" pageNumber="34" precision="5" value="-19.6683">19.6683°S</geoCoordinate>
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||
<geoCoordinate id="D5F111C7DC53A510FB8B8ADCFACAFD76" box="[1147,1284,726,751]" direction="east" orientation="longitude" pageId="33" pageNumber="34" precision="1" value="63.46307">63.46307°E</geoCoordinate>
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||
, depth
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||
<quantity id="773DDAE5DC53A510FAAA8ADDFA59FD77" box="[1370,1431,727,751]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="33" pageNumber="34" unit="m" value="12.0">
|
||
<elevation id="3BE89033DC53A510FAAA8ADDFA59FD77" box="[1370,1431,727,751]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="33" pageNumber="34" unit="m" value="12.0">12 m</elevation>
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||
</quantity>
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||
, scuba, coll.
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||
<collectorName id="1D3012D6DC53A510FEED8AF1FE70FC8A" box="[285,446,762,787]" pageId="33" pageNumber="34">N.J. de Voogd</collectorName>
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||
, field nr. ROG023,
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||
<date id="C47B51C0DC53A510FD6A8AF0FC94FC8A" box="[666,858,762,787]" pageId="33" pageNumber="34" value="2016-10-16">
|
||
<collectingDate id="D43FA828DC53A510FD6A8AF0FC94FC8A" box="[666,858,762,787]" pageId="33" pageNumber="34" value="2016-10-16">16 October 2016</collectingDate>
|
||
</date>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="F8DF248BDC53A510FF378B14FA43FA02" pageId="33" pageNumber="34" type="description">
|
||
<paragraph id="B07A7700DC53A510FF378B14FEEEFC72" blockId="33.[151,1437,654,2011]" pageId="33" pageNumber="34">
|
||
<emphasis id="82B1AB12DC53A510FF378B14FE97FCAE" bold="true" box="[199,345,798,823]" pageId="33" pageNumber="34">Description.</emphasis>
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||
Small pear-shaped cormus (
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||
<figureCitation id="28FE6B85DC53A510FD6A8B14FCD7FCAE" box="[666,793,798,823]" captionStart="FIGURE 12" captionStartId="34.[151,249,1884,1906]" captionTargetBox="[151,1436,445,1863]" captionTargetId="figure@34.[151,1436,445,1863]" captionTargetPageId="34" captionText="FIGURE 12. Borojevia pirella sp.nov., a, habitus in situ of holotype RMNH Por. 11622, from Rodrigues, the easternmost of the Mascarene Islands (photo N.J. de Voogd), a1, preserved holotype (scale bar = 1 cm), b, habitus in situ of paratype RMNH Por. 11622b (photo N.J. de Voogd), b1, preserved paratype." httpUri="https://zenodo.org/record/1271289/files/figure.png" pageId="33" pageNumber="34">Figs 12a–b</figureCitation>
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||
) consisting of a mass of moderately tightly anastomosed tubuli separated by rounded or polyangular spaces. Color white, semitransparent. Size approximately
|
||
<quantity id="773DDAE5DC53A510FAFF8B48FA92FCC3" box="[1295,1372,834,859]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="33" pageNumber="34" unit="cm" value="1.5">1.5 cm</quantity>
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||
high,
|
||
<quantity id="773DDAE5DC53A510FF678B6DFF00FCE7" box="[151,206,871,895]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="33" pageNumber="34" unit="cm" value="1.0">1 cm</quantity>
|
||
in diameter. Tubuli approximately
|
||
<quantity id="773DDAE5DC53A510FDAB8B6DFD7FFCE7" box="[603,689,871,895]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="33" pageNumber="34" unit="mm" value="0.2">0.2 mm</quantity>
|
||
in diameter. The narrow end is provided with an oscule of about
|
||
<quantity id="773DDAE5DC53A510FA7E8B6DFF0DFC3B" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="33" pageNumber="34" unit="mm" value="2.0">2 mm</quantity>
|
||
diameter, leading into a short water-collecting tube and a pseudoatrium. Preserved material consists of two specimens (
|
||
<figureCitation id="28FE6B85DC53A510FED28BA4FE5FFC5E" box="[290,401,942,967]" captionStart="FIGURE 12" captionStartId="34.[151,249,1884,1906]" captionTargetBox="[151,1436,445,1863]" captionTargetId="figure@34.[151,1436,445,1863]" captionTargetPageId="34" captionText="FIGURE 12. Borojevia pirella sp.nov., a, habitus in situ of holotype RMNH Por. 11622, from Rodrigues, the easternmost of the Mascarene Islands (photo N.J. de Voogd), a1, preserved holotype (scale bar = 1 cm), b, habitus in situ of paratype RMNH Por. 11622b (photo N.J. de Voogd), b1, preserved paratype." httpUri="https://zenodo.org/record/1271289/files/figure.png" pageId="33" pageNumber="34">Figs 12a</figureCitation>
|
||
<figureCitation id="28FE6B85DC53A510FE618BA5FE55FC5E" box="[401,411,943,967]" captionStart="FIGURE 1" captionStartId="5.[151,249,1813,1835]" captionTargetBox="[151,1436,268,1791]" captionTargetId="figure@5.[151,1436,265,1791]" captionTargetPageId="5" captionText="FIGURE 1. Marine Ecoregions in the Western Indian Ocean with localities (numbered blue squares) from where the studied calcareous sponges were obtained. 1, Israelian Red Sea, Gulf of Aqaba, 2, Saudi Arabian Red Sea, off Jeddah, 3, Eritrean Red Sea, Dahlak Archipelago, 4, Oman, Gulf of Arabia, 5, Maldives, 6, India, Lakshadweep, 7, Seychelles, Mahé region, 8, Seychelles, Amirantes, 9, Mayotte, 10, Mozambique Channel, 11, Mauritius, 12, Rodrigues, 13, South Africa, Kwazulu Natal, 14. South Africa, Port Elizabeth, 15. Nosy Be, Madagascar." httpUri="https://zenodo.org/record/1271247/files/figure.png" pageId="33" pageNumber="34">1</figureCitation>
|
||
,
|
||
<typeStatus id="6F7EC9A2DC53A510FE5F8BA4FDDAFC5E" box="[431,532,942,967]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
|
||
, and 12b1,
|
||
<typeStatus id="6F7EC9A2DC53A510FD5D8BA5FCDCFC5E" box="[685,786,943,967]" pageId="33" pageNumber="34" type="paratype">paratype</typeStatus>
|
||
), which collapsed to form flattened objects with soft consistency.
|
||
</paragraph>
|
||
<paragraph id="B07A7700DC53A510FF378BFCFDEBFB96" blockId="33.[151,1437,654,2011]" box="[199,549,1014,1039]" pageId="33" pageNumber="34">
|
||
<emphasis id="82B1AB12DC53A510FF378BFCFE65FB96" bold="true" box="[199,427,1014,1039]" pageId="33" pageNumber="34">Aquiferous system.</emphasis>
|
||
Asconoid.
|
||
</paragraph>
|
||
<paragraph id="B07A7700DC53A510FF378C10FC41FBE2" blockId="33.[151,1437,654,2011]" pageId="33" pageNumber="34">
|
||
<emphasis id="82B1AB12DC53A510FF378C10FEFBFBAA" bold="true" box="[199,309,1050,1075]" pageId="33" pageNumber="34">Skeleton.</emphasis>
|
||
The pseudoatrium has no special skeleton, it is merely a space between the anastomosed tubuli (
|
||
<figureCitation id="28FE6B85DC53A510FA9F8C10FF0CFBCE" captionStart="FIGURE 13" captionStartId="35.[151,249,1605,1627]" captionTargetBox="[177,1421,199,1578]" captionTargetId="figure@35.[177,1421,199,1578]" captionTargetPageId="35" captionText="FIGURE 13. Borojevia pirella sp.nov., holotype RMNH Por. 11622, from Rodrigues, a, light microscopic cross section of oscular region showing pseudoatrium and surrounding tubuli, b–e, SEM images of the spicules, b, regular triactine, c, rare sagittal irregular triactine, d, tetractines, e, details of apical actines of the tetractines showing characteristic verticil of three conical spines." httpUri="https://zenodo.org/record/1271295/files/figure.png" pageId="33" pageNumber="34">Fig. 13a</figureCitation>
|
||
). The wall of these is thin, having one or two spicules comprising a mixture of triactines and tetractines, the latter with the apical actines protruding into the lumen of the tubuli.
|
||
</paragraph>
|
||
<paragraph id="B07A7700DC53A510FF378C8CFE2EFB5B" blockId="33.[151,1437,654,2011]" pageId="33" pageNumber="34">
|
||
<emphasis id="82B1AB12DC53A510FF378C8CFEFEFB06" bold="true" box="[199,304,1158,1183]" pageId="33" pageNumber="34">Spicules.</emphasis>
|
||
(
|
||
<figureCitation id="28FE6B85DC53A510FEB08C8CFE0EFB06" box="[320,448,1158,1183]" captionStart="FIGURE 13" captionStartId="35.[151,249,1605,1627]" captionTargetBox="[177,1421,199,1578]" captionTargetId="figure@35.[177,1421,199,1578]" captionTargetPageId="35" captionText="FIGURE 13. Borojevia pirella sp.nov., holotype RMNH Por. 11622, from Rodrigues, a, light microscopic cross section of oscular region showing pseudoatrium and surrounding tubuli, b–e, SEM images of the spicules, b, regular triactine, c, rare sagittal irregular triactine, d, tetractines, e, details of apical actines of the tetractines showing characteristic verticil of three conical spines." httpUri="https://zenodo.org/record/1271295/files/figure.png" pageId="33" pageNumber="34">Figs 13b–e</figureCitation>
|
||
) Triactines and tetractines of approximately the same size, no differentiated larger and smaller triactines are present.
|
||
</paragraph>
|
||
<paragraph id="B07A7700DC53A510FF378CC4FCB5FAB6" blockId="33.[151,1437,654,2011]" pageId="33" pageNumber="34">
|
||
Triactines (
|
||
<figureCitation id="28FE6B85DC53A510FEB88CC4FE0BFB7E" box="[328,453,1230,1255]" captionStart="FIGURE 13" captionStartId="35.[151,249,1605,1627]" captionTargetBox="[177,1421,199,1578]" captionTargetId="figure@35.[177,1421,199,1578]" captionTargetPageId="35" captionText="FIGURE 13. Borojevia pirella sp.nov., holotype RMNH Por. 11622, from Rodrigues, a, light microscopic cross section of oscular region showing pseudoatrium and surrounding tubuli, b–e, SEM images of the spicules, b, regular triactine, c, rare sagittal irregular triactine, d, tetractines, e, details of apical actines of the tetractines showing characteristic verticil of three conical spines." httpUri="https://zenodo.org/record/1271295/files/figure.png" pageId="33" pageNumber="34">Fig. 13b–c</figureCitation>
|
||
) equiangular and equiradiate (
|
||
<figureCitation id="28FE6B85DC53A510FCD78CC4FC45FB7E" box="[807,907,1230,1255]" captionStart="FIGURE 13" captionStartId="35.[151,249,1605,1627]" captionTargetBox="[177,1421,199,1578]" captionTargetId="figure@35.[177,1421,199,1578]" captionTargetPageId="35" captionText="FIGURE 13. Borojevia pirella sp.nov., holotype RMNH Por. 11622, from Rodrigues, a, light microscopic cross section of oscular region showing pseudoatrium and surrounding tubuli, b–e, SEM images of the spicules, b, regular triactine, c, rare sagittal irregular triactine, d, tetractines, e, details of apical actines of the tetractines showing characteristic verticil of three conical spines." httpUri="https://zenodo.org/record/1271295/files/figure.png" pageId="33" pageNumber="34">Fig. 13b</figureCitation>
|
||
), with a tendency to become slightly sagittal, with a longer unpaired actine; occasionally triactines are irregular with one of the actines wavy and distinctly sagittal (
|
||
<figureCitation id="28FE6B85DC53A510FF078D1CFE99FAB6" box="[247,343,1302,1327]" captionStart="FIGURE 13" captionStartId="35.[151,249,1605,1627]" captionTargetBox="[177,1421,199,1578]" captionTargetId="figure@35.[177,1421,199,1578]" captionTargetPageId="35" captionText="FIGURE 13. Borojevia pirella sp.nov., holotype RMNH Por. 11622, from Rodrigues, a, light microscopic cross section of oscular region showing pseudoatrium and surrounding tubuli, b–e, SEM images of the spicules, b, regular triactine, c, rare sagittal irregular triactine, d, tetractines, e, details of apical actines of the tetractines showing characteristic verticil of three conical spines." httpUri="https://zenodo.org/record/1271295/files/figure.png" pageId="33" pageNumber="34">Fig. 13c</figureCitation>
|
||
). Actine dimensions 51–
|
||
<emphasis id="82B1AB12DC53A510FD848D1DFD6BFAB6" box="[628,677,1303,1327]" italics="true" pageId="33" pageNumber="34">81.3</emphasis>
|
||
–
|
||
<date id="C47B51C0DC53A510FD448D1DFD39FAB6" box="[692,759,1302,1327]" pageId="33" pageNumber="34" value="1993-10-05">93 x 5</date>
|
||
–
|
||
<emphasis id="82B1AB12DC53A510FCF78D1CFCE4FAB6" box="[775,810,1302,1327]" italics="true" pageId="33" pageNumber="34">6.7</emphasis>
|
||
–8 µm.
|
||
</paragraph>
|
||
<paragraph id="B07A7700DC53A510FF378D30FA43FA02" blockId="33.[151,1437,654,2011]" pageId="33" pageNumber="34">
|
||
Tetractines (
|
||
<figureCitation id="28FE6B85DC53A510FEA08D30FE02FACA" box="[336,460,1338,1363]" captionStart="FIGURE 13" captionStartId="35.[151,249,1605,1627]" captionTargetBox="[177,1421,199,1578]" captionTargetId="figure@35.[177,1421,199,1578]" captionTargetPageId="35" captionText="FIGURE 13. Borojevia pirella sp.nov., holotype RMNH Por. 11622, from Rodrigues, a, light microscopic cross section of oscular region showing pseudoatrium and surrounding tubuli, b–e, SEM images of the spicules, b, regular triactine, c, rare sagittal irregular triactine, d, tetractines, e, details of apical actines of the tetractines showing characteristic verticil of three conical spines." httpUri="https://zenodo.org/record/1271295/files/figure.png" pageId="33" pageNumber="34">Fig. 13d–e</figureCitation>
|
||
), similar in size and shape to the triactines (
|
||
<figureCitation id="28FE6B85DC53A510FC418D30FBD8FACA" box="[945,1046,1338,1363]" captionStart="FIGURE 13" captionStartId="35.[151,249,1605,1627]" captionTargetBox="[177,1421,199,1578]" captionTargetId="figure@35.[177,1421,199,1578]" captionTargetPageId="35" captionText="FIGURE 13. Borojevia pirella sp.nov., holotype RMNH Por. 11622, from Rodrigues, a, light microscopic cross section of oscular region showing pseudoatrium and surrounding tubuli, b–e, SEM images of the spicules, b, regular triactine, c, rare sagittal irregular triactine, d, tetractines, e, details of apical actines of the tetractines showing characteristic verticil of three conical spines." httpUri="https://zenodo.org/record/1271295/files/figure.png" pageId="33" pageNumber="34">Figs 13d</figureCitation>
|
||
), with apical actines provided with three prominent conical spines in one verticil the position of which is matching the basal triradiate system (
|
||
<figureCitation id="28FE6B85DC53A510FA9A8D54FF0CFA02" captionStart="FIGURE 13" captionStartId="35.[151,249,1605,1627]" captionTargetBox="[177,1421,199,1578]" captionTargetId="figure@35.[177,1421,199,1578]" captionTargetPageId="35" captionText="FIGURE 13. Borojevia pirella sp.nov., holotype RMNH Por. 11622, from Rodrigues, a, light microscopic cross section of oscular region showing pseudoatrium and surrounding tubuli, b–e, SEM images of the spicules, b, regular triactine, c, rare sagittal irregular triactine, d, tetractines, e, details of apical actines of the tetractines showing characteristic verticil of three conical spines." httpUri="https://zenodo.org/record/1271295/files/figure.png" pageId="33" pageNumber="34">Figs 13e</figureCitation>
|
||
). Actines of the basal triadiate system 72–
|
||
<emphasis id="82B1AB12DC53A510FD6C8D89FD03FA02" box="[668,717,1411,1435]" italics="true" pageId="33" pageNumber="34">85.7</emphasis>
|
||
–
|
||
<date id="C47B51C0DC53A510FD2C8D88FCC2FA03" box="[732,780,1410,1435]" pageId="33" pageNumber="34" value="1996-10">96 x</date>
|
||
5.5–
|
||
<emphasis id="82B1AB12DC53A510FCB48D89FCA9FA02" box="[836,871,1411,1435]" italics="true" pageId="33" pageNumber="34">7.1</emphasis>
|
||
–8 µm, apical actines 31–
|
||
<emphasis id="82B1AB12DC53A510FB778D89FB76FA02" box="[1159,1208,1411,1435]" italics="true" pageId="33" pageNumber="34">58.4</emphasis>
|
||
–
|
||
<date id="C47B51C0DC53A510FB368D88FAC4FA02" box="[1222,1290,1410,1435]" pageId="33" pageNumber="34" value="1976-10-05">76 x 5</date>
|
||
–
|
||
<emphasis id="82B1AB12DC53A510FAEA8D89FAF3FA02" box="[1306,1341,1411,1435]" italics="true" pageId="33" pageNumber="34">5.9</emphasis>
|
||
–7 µm.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F8DF248BDC53A510FF378DACFB56FA26" box="[199,1176,1446,1471]" pageId="33" pageNumber="34" type="distribution">
|
||
<paragraph id="B07A7700DC53A510FF378DACFB56FA26" blockId="33.[151,1437,654,2011]" box="[199,1176,1446,1471]" pageId="33" pageNumber="34">
|
||
<emphasis id="82B1AB12DC53A510FF378DACFE3CFA26" bold="true" box="[199,498,1446,1471]" pageId="33" pageNumber="34">Distribution and ecology.</emphasis>
|
||
<collectingRegion id="7201B9E2DC53A510FE098DACFDA1FA26" box="[505,623,1446,1471]" country="Mauritius" name="Rodrigues" pageId="33" pageNumber="34">Rodrigues</collectingRegion>
|
||
, Mascarene Islands, on coral reefs at
|
||
<quantity id="773DDAE5DC53A510FBE38DADFB82FA27" box="[1043,1100,1447,1471]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="33" pageNumber="34" unit="m" value="12.0">12 m</quantity>
|
||
depth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F8DF248BDC53A510FF378DC0FA8CFA7A" box="[199,1346,1482,1507]" pageId="33" pageNumber="34" type="etymology">
|
||
<paragraph id="B07A7700DC53A510FF378DC0FA8CFA7A" blockId="33.[151,1437,654,2011]" box="[199,1346,1482,1507]" pageId="33" pageNumber="34">
|
||
<emphasis id="82B1AB12DC53A510FF378DC0FE80FA7A" bold="true" box="[199,334,1482,1507]" pageId="33" pageNumber="34">Etymology.</emphasis>
|
||
Pirella (L.), a feminine noun, meaning a small pear, referring to the habitus of the sponge.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F8DF248BDC53A512FF378DE4FDA0F86A" lastPageId="35" lastPageNumber="36" pageId="33" pageNumber="34" type="discussion">
|
||
<paragraph id="B07A7700DC53A510FF378DE4FE9EF82E" blockId="33.[151,1437,654,2011]" pageId="33" pageNumber="34">
|
||
<emphasis id="82B1AB12DC53A510FF378DE4FEF5F99E" bold="true" box="[199,315,1518,1543]" pageId="33" pageNumber="34">Remarks.</emphasis>
|
||
Despite the presence of a pseudoatrium and spined apical actines of the tetractines, molecular sequence data indicate clearly (
|
||
<figureCitation id="28FE6B85DC53A510FE078E18FD83F9B2" box="[503,589,1554,1579]" captionStart="FIGURE 2" captionStartId="8.[151,250,1826,1848]" captionTargetBox="[180,1383,196,1782]" captionTargetId="figure@8.[172,1416,195,1804]" captionTargetPageId="8" captionText="FIGURE 2. Maximum-likelihood tree based on analysis of sequences of the partial 28S gene (C2–D2 region, comprising 442 sites) of Western Indian Ocean Calcinea newly generated by the Naturalis Biodiversity Center and sequences additionally obtained from GenBank, the Sponge Barcode Project and from Oliver Voigt (München) from the Red Sea, Indonesia, Australia, and a few sequences from other parts of the world, to demonstrate the affiliation of the compared species. Please note that Arturia sequences do not appear in the same clade, indicating Arturia is non-monophyletic. Numbers at the root of clades are bootstrap values of 1000 replicates. Since the high number of sequences were difficult to read in detail, we subdivided the present tree in three detailed sections 2A, 2B, and 2C." httpUri="https://zenodo.org/record/1271249/files/figure.png" pageId="33" pageNumber="34">Fig. 2B</figureCitation>
|
||
) that this species is closely related to
|
||
<taxonomicName id="77C50C83DC53A510FC078E1EFACDF9B2" baseAuthorityName="Haeckel" baseAuthorityYear="1872" box="[1015,1283,1554,1579]" class="Calcarea" family="Clathrinidae" genus="Borojevia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clathrinida" pageId="33" pageNumber="34" phylum="Porifera" rank="species" species="cerebrum">
|
||
<emphasis id="82B1AB12DC53A510FC078E1EFBAAF9B2" box="[1015,1124,1556,1579]" italics="true" pageId="33" pageNumber="34">Borojevia</emphasis>
|
||
aff.
|
||
<emphasis id="82B1AB12DC53A510FB698E18FACDF9B2" box="[1177,1283,1554,1579]" italics="true" pageId="33" pageNumber="34">cerebrum</emphasis>
|
||
</taxonomicName>
|
||
and ‘
|
||
<taxonomicName id="77C50C83DC53A510FAB38E18FA52F9B2" box="[1347,1436,1554,1579]" class="Calcarea" family="Clathrinidae" genus="Ascaltis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clathrinida" pageId="33" pageNumber="34" phylum="Porifera" rank="genus">
|
||
<emphasis id="82B1AB12DC53A510FAB38E18FA52F9B2" box="[1347,1436,1554,1579]" italics="true" pageId="33" pageNumber="34">Ascaltis</emphasis>
|
||
</taxonomicName>
|
||
OV-2012’, both reassigned to
|
||
<taxonomicName id="77C50C83DC53A510FE168E3DFD9DF9D7" authorityName="Van & De" authorityYear="2018" box="[486,595,1591,1614]" class="Calcarea" family="Clathrinidae" genus="Borojevia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clathrinida" pageId="33" pageNumber="34" phylum="Porifera" rank="genus">
|
||
<emphasis id="82B1AB12DC53A510FE168E3DFD9DF9D7" box="[486,595,1591,1614]" italics="true" pageId="33" pageNumber="34">Borojevia</emphasis>
|
||
</taxonomicName>
|
||
(cf. Klautau
|
||
<emphasis id="82B1AB12DC53A510FD158E3DFCD3F9D6" box="[741,797,1590,1615]" italics="true" pageId="33" pageNumber="34">et al.</emphasis>
|
||
2016), and not to
|
||
<taxonomicName id="77C50C83DC53A510FC178E3CFB7EF9D6" baseAuthorityName="Schmidt" baseAuthorityYear="1862" box="[999,1200,1590,1615]" class="Calcarea" family="Clathrinidae" genus="Ascaltis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clathrinida" pageId="33" pageNumber="34" phylum="Porifera" rank="species" species="reticulum">
|
||
<emphasis id="82B1AB12DC53A510FC178E3CFB7EF9D6" box="[999,1200,1590,1615]" italics="true" pageId="33" pageNumber="34">Ascaltis reticulum</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="D4540AF1DC53A510FB4E8E3CFA52F9D6" box="[1214,1436,1590,1615]" pageId="33" pageNumber="34" refId="ref70121" refString="Voigt, O., Wulfing, E. & Worheide, G. (2012) Molecular phylogenetic evaluation of classification and scenarios of character evolution in calcareous sponges (Porifera, Class Calcarea). PLOS One, 7 (3), e 33417. https: // doi. org / 10.1371 / journal. pone. 0033417" type="journal article">
|
||
Voigt
|
||
<emphasis id="82B1AB12DC53A510FAF18E3DFAF6F9D6" box="[1281,1336,1590,1615]" italics="true" pageId="33" pageNumber="34">et al.</emphasis>
|
||
’s (2012)
|
||
</bibRefCitation>
|
||
Australian species ‘
|
||
<taxonomicName id="77C50C83DC53A510FE868E50FE01F9EA" box="[374,463,1626,1651]" class="Calcarea" family="Clathrinidae" genus="Ascaltis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clathrinida" pageId="33" pageNumber="34" phylum="Porifera" rank="genus">
|
||
<emphasis id="82B1AB12DC53A510FE868E50FE01F9EA" box="[374,463,1626,1651]" italics="true" pageId="33" pageNumber="34">Ascaltis</emphasis>
|
||
</taxonomicName>
|
||
OV 2012’ has a clear pseudoatrium illustrated in their Supporting
|
||
<figureCitation id="28FE6B85DC53A510FB378E50FAD7F9EA" box="[1223,1305,1626,1651]" captionStart="FIGURE 1" captionStartId="5.[151,249,1813,1835]" captionTargetBox="[151,1436,268,1791]" captionTargetId="figure@5.[151,1436,265,1791]" captionTargetPageId="5" captionText="FIGURE 1. Marine Ecoregions in the Western Indian Ocean with localities (numbered blue squares) from where the studied calcareous sponges were obtained. 1, Israelian Red Sea, Gulf of Aqaba, 2, Saudi Arabian Red Sea, off Jeddah, 3, Eritrean Red Sea, Dahlak Archipelago, 4, Oman, Gulf of Arabia, 5, Maldives, 6, India, Lakshadweep, 7, Seychelles, Mahé region, 8, Seychelles, Amirantes, 9, Mayotte, 10, Mozambique Channel, 11, Mauritius, 12, Rodrigues, 13, South Africa, Kwazulu Natal, 14. South Africa, Port Elizabeth, 15. Nosy Be, Madagascar." httpUri="https://zenodo.org/record/1271247/files/figure.png" pageId="33" pageNumber="34">Fig. S1</figureCitation>
|
||
inset J, but its sequence is grouped unequivocally among the
|
||
<taxonomicName id="77C50C83DC53A510FD308E75FCE3F90F" authorityName="Van & De" authorityYear="2018" box="[704,813,1663,1686]" class="Calcarea" family="Clathrinidae" genus="Borojevia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clathrinida" pageId="33" pageNumber="34" phylum="Porifera" rank="genus">
|
||
<emphasis id="82B1AB12DC53A510FD308E75FCE3F90F" box="[704,813,1663,1686]" italics="true" pageId="33" pageNumber="34">Borojevia</emphasis>
|
||
</taxonomicName>
|
||
’s in our
|
||
<figureCitation id="28FE6B85DC53A510FC7D8E74FC2EF90E" box="[909,992,1662,1687]" captionStart="FIGURE 2" captionStartId="8.[151,250,1826,1848]" captionTargetBox="[180,1383,196,1782]" captionTargetId="figure@8.[172,1416,195,1804]" captionTargetPageId="8" captionText="FIGURE 2. Maximum-likelihood tree based on analysis of sequences of the partial 28S gene (C2–D2 region, comprising 442 sites) of Western Indian Ocean Calcinea newly generated by the Naturalis Biodiversity Center and sequences additionally obtained from GenBank, the Sponge Barcode Project and from Oliver Voigt (München) from the Red Sea, Indonesia, Australia, and a few sequences from other parts of the world, to demonstrate the affiliation of the compared species. Please note that Arturia sequences do not appear in the same clade, indicating Arturia is non-monophyletic. Numbers at the root of clades are bootstrap values of 1000 replicates. Since the high number of sequences were difficult to read in detail, we subdivided the present tree in three detailed sections 2A, 2B, and 2C." httpUri="https://zenodo.org/record/1271249/files/figure.png" pageId="33" pageNumber="34">Fig. 2B</figureCitation>
|
||
phylogeny. This justifies assignment of the present species to the genus
|
||
<taxonomicName id="77C50C83DC53A510FDFE8EAEFDB5F922" authorityName="Van & De" authorityYear="2018" box="[526,635,1700,1723]" class="Calcarea" family="Clathrinidae" genus="Borojevia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clathrinida" pageId="33" pageNumber="34" phylum="Porifera" rank="genus">
|
||
<emphasis id="82B1AB12DC53A510FDFE8EAEFDB5F922" box="[526,635,1700,1723]" italics="true" pageId="33" pageNumber="34">Borojevia</emphasis>
|
||
</taxonomicName>
|
||
. The triactine complement of
|
||
<taxonomicName id="77C50C83DC53A510FC2C8EAEFB89F922" authority="Van & De, 2018" authorityName="Van & De" authorityYear="2018" box="[988,1095,1698,1723]" class="Calcarea" family="Clathrinidae" genus="Borojevia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clathrinida" pageId="33" pageNumber="34" phylum="Porifera" rank="species" species="pirella" status="sp. nov.">
|
||
<emphasis id="82B1AB12DC53A510FC2C8EAEFB89F922" box="[988,1095,1698,1723]" italics="true" pageId="33" pageNumber="34">B. pirella</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="82B1AB12DC53A510FBA18EA8FB6DF922" bold="true" box="[1105,1187,1698,1723]" pageId="33" pageNumber="34">
|
||
<taxonomicNameLabel id="99821669DC53A510FBA18EA8FB6DF922" box="[1105,1187,1698,1723]" pageId="33" pageNumber="34" rank="species">sp.nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
does not contain any clear tripods, as is one of the additional features described for the genus
|
||
<taxonomicName id="77C50C83DC53A510FBD58ECDFB5CF947" authorityName="Van & De" authorityYear="2018" box="[1061,1170,1735,1758]" class="Calcarea" family="Clathrinidae" genus="Borojevia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clathrinida" pageId="33" pageNumber="34" phylum="Porifera" rank="genus">
|
||
<emphasis id="82B1AB12DC53A510FBD58ECDFB5CF947" box="[1061,1170,1735,1758]" italics="true" pageId="33" pageNumber="34">Borojevia</emphasis>
|
||
</taxonomicName>
|
||
. The presence of the pseudoatrium is shared with the above described
|
||
<taxonomicName id="77C50C83DC53A510FD378EE6FC8AF89A" authority="Van & De, 2018" authorityName="Van & De" authorityYear="2018" box="[711,836,1770,1795]" class="Calcarea" family="Clathrinidae" genus="Borojevia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clathrinida" pageId="33" pageNumber="34" phylum="Porifera" rank="species" species="tubulata" status="sp. nov.">
|
||
<emphasis id="82B1AB12DC53A510FD378EE6FC8AF89A" box="[711,836,1770,1795]" italics="true" pageId="33" pageNumber="34">B. tubulata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="99821669DC53A510FCBC8EE0FC50F89B" box="[844,926,1770,1795]" pageId="33" pageNumber="34" rank="species">
|
||
<emphasis id="82B1AB12DC53A510FCBC8EE0FC56F89A" bold="true" box="[844,920,1770,1795]" pageId="33" pageNumber="34">sp.nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, but apparently is absent in
|
||
<taxonomicName id="77C50C83DC53A510FB118EE6FA8FF89A" authority="Van & De, 2018" authorityName="Van & De" authorityYear="2018" box="[1249,1345,1772,1795]" class="Calcarea" family="Clathrinidae" genus="Borojevia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clathrinida" pageId="33" pageNumber="34" phylum="Porifera" rank="species" species="voigti" status="sp. nov.">
|
||
<emphasis id="82B1AB12DC53A510FB118EE6FA8FF89A" box="[1249,1345,1772,1795]" italics="true" pageId="33" pageNumber="34">B. voigti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="82B1AB12DC53A510FAB98EE0FA52F89A" bold="true" box="[1353,1436,1770,1795]" pageId="33" pageNumber="34">
|
||
<taxonomicNameLabel id="99821669DC53A510FAB98EE0FA52F89A" box="[1353,1436,1770,1795]" pageId="33" pageNumber="34" rank="species">sp.nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and other
|
||
<taxonomicName id="77C50C83DC53A510FEFD8F05FEB4F8BF" authorityName="Van & De" authorityYear="2018" box="[269,378,1807,1830]" class="Calcarea" family="Clathrinidae" genus="Borojevia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clathrinida" pageId="33" pageNumber="34" phylum="Porifera" rank="genus">
|
||
<emphasis id="82B1AB12DC53A510FEFD8F05FEB4F8BF" box="[269,378,1807,1830]" italics="true" pageId="33" pageNumber="34">Borojevia</emphasis>
|
||
</taxonomicName>
|
||
spp. The relationship with
|
||
<taxonomicName id="77C50C83DC53A510FD4C8F04FCDBF8BE" box="[700,789,1806,1831]" class="Calcarea" family="Clathrinidae" genus="Ascaltis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clathrinida" pageId="33" pageNumber="34" phylum="Porifera" rank="genus">
|
||
<emphasis id="82B1AB12DC53A510FD4C8F04FCDBF8BE" box="[700,789,1806,1831]" italics="true" pageId="33" pageNumber="34">Ascaltis</emphasis>
|
||
</taxonomicName>
|
||
remains unresolved. The three prominent spines on the apical actines of the tetractines are a special feature of the new species. It is shared with the recently described Brazilian
|
||
<taxonomicName id="77C50C83DC53A510FEF68F5DFD2FF8F6" authority="Azevedo et al., 2017" authorityName="Azevedo et al." authorityYear="2017" box="[262,737,1878,1903]" class="Calcarea" family="Clathrinidae" genus="Borojevia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clathrinida" pageId="33" pageNumber="34" phylum="Porifera" rank="species" species="trispinata">
|
||
<emphasis id="82B1AB12DC53A510FEF68F5DFE27F8F7" box="[262,489,1879,1902]" italics="true" pageId="33" pageNumber="34">Borojevia trispinata</emphasis>
|
||
<bibRefCitation id="D4540AF1DC53A510FE018F5CFD2FF8F6" box="[497,737,1878,1903]" pageId="33" pageNumber="34" refId="ref63142" refString="Azevedo, F., Padua, A., Moraes, F., Rossi, A., Muricy, G. & Klautau, M. (2017) Taxonomy and phylogeny of calcareous sponges (Porifera: Calcarea: Calcinea) from Brazilian mid-shelf and oceanic islands. Zootaxa, 4311 (3), 301 - 344. https: // doi. org / 10.11646 / zootaxa. 4311.3.1" type="journal article">
|
||
Azevedo
|
||
<emphasis id="82B1AB12DC53A510FDAD8F5DFD59F8F6" box="[605,663,1878,1903]" italics="true" pageId="33" pageNumber="34">et al.</emphasis>
|
||
, 2017
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
. This is generally similar in spicule sizes, but on average has slightly smaller actines. The habitus differs in lacking a central pseudoatrium and being more irregular in shape, not pear-shaped.
|
||
</paragraph>
|
||
<paragraph id="B07A7700DC53A513FF378FC9FDBAFE11" blockId="33.[151,1437,654,2011]" lastBlockId="34.[151,1437,151,392]" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
|
||
We obtained sequences for
|
||
<taxonomicName id="77C50C83DC53A510FDF18FCEFD1AF842" authority="Van & De, 2018" authorityName="Van & De" authorityYear="2018" box="[513,724,1986,2011]" class="Calcarea" family="Clathrinidae" genus="Borojevia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clathrinida" pageId="33" pageNumber="34" phylum="Porifera" rank="species" species="tubulata" status="sp. nov.">
|
||
<emphasis id="82B1AB12DC53A510FDF18FCEFD1AF842" box="[513,724,1986,2011]" italics="true" pageId="33" pageNumber="34">Borojevia tubulata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="82B1AB12DC53A510FD2D8FC8FCFEF842" bold="true" box="[733,816,1986,2011]" pageId="33" pageNumber="34">
|
||
<taxonomicNameLabel id="99821669DC53A510FD2D8FC8FCFEF842" box="[733,816,1986,2011]" pageId="33" pageNumber="34" rank="species">sp.nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(both
|
||
<typeStatus id="6F7EC9A2DC53A510FC8D8FC8FC11F842" box="[893,991,1986,2011]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
|
||
and non-type
|
||
<collectingCountry id="C8D23790DC53A510FB758FC8FB33F842" box="[1157,1277,1986,2011]" name="Seychelles" pageId="33" pageNumber="34">Seychelles</collectingCountry>
|
||
specimen, cf. above) and
|
||
<taxonomicName id="77C50C83DC50A513FEEA8893FE18FF29" authority="Van & De, 2018" authorityName="Van & De" authorityYear="2018" box="[282,470,151,176]" class="Calcarea" family="Clathrinidae" genus="Borojevia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clathrinida" pageId="34" pageNumber="35" phylum="Porifera" rank="species" species="pirella" status="sp. nov.">
|
||
<emphasis id="82B1AB12DC50A513FEEA8893FE18FF29" box="[282,470,151,176]" italics="true" pageId="34" pageNumber="35">Borojevia pirella</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="82B1AB12DC50A513FE2F889DFDFCFF29" bold="true" box="[479,562,151,176]" pageId="34" pageNumber="35">
|
||
<taxonomicNameLabel id="99821669DC50A513FE2F889DFDFCFF29" box="[479,562,151,176]" pageId="34" pageNumber="35" rank="species">sp.nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(both
|
||
<typeStatus id="6F7EC9A2DC50A513FD8B889DFD13FF29" box="[635,733,151,176]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="6F7EC9A2DC50A513FCE48892FCB4FF29" box="[788,890,152,176]" pageId="34" pageNumber="35" type="paratype">paratype</typeStatus>
|
||
). We compared these with two sequences of
|
||
<collectingRegion id="7201B9E2DC50A513FA9F889DFF0EFF4D" country="Eritrea" name="Semenawi Keyih Bahri" pageId="34" pageNumber="35">Red Sea</collectingRegion>
|
||
specimens of
|
||
<taxonomicName id="77C50C83DC50A513FEAF88B7FD85FF4D" box="[351,587,188,213]" class="Calcarea" family="Clathrinidae" genus="Borojevia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clathrinida" pageId="34" pageNumber="35" phylum="Porifera" rank="species" species="aspina">
|
||
<emphasis id="82B1AB12DC50A513FEAF88B7FE02FF4D" box="[351,460,189,212]" italics="true" pageId="34" pageNumber="35">Borojevia</emphasis>
|
||
aff.
|
||
<emphasis id="82B1AB12DC50A513FDF088B7FD85FF4D" box="[512,587,189,212]" italics="true" pageId="34" pageNumber="35">aspina</emphasis>
|
||
</taxonomicName>
|
||
provided by Oliver Voigt in a separate restricted phylogenetic analysis. The aligned and trimmed sequences (length 426 sites) revealed that both identical sequences of
|
||
<taxonomicName id="77C50C83DC50A513FB2188EBFAF0FF61" authority="Van & De, 2018" authorityName="Van & De" authorityYear="2018" box="[1233,1342,223,248]" class="Calcarea" family="Clathrinidae" genus="Borojevia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clathrinida" pageId="34" pageNumber="35" phylum="Porifera" rank="species" species="pirella" status="sp. nov.">
|
||
<emphasis id="82B1AB12DC50A513FB2188EBFAF0FF61" box="[1233,1342,223,248]" italics="true" pageId="34" pageNumber="35">B. pirella</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="82B1AB12DC50A513FAB988D5FA52FF61" bold="true" box="[1353,1436,223,248]" pageId="34" pageNumber="35">
|
||
<taxonomicNameLabel id="99821669DC50A513FAB988D5FA52FF61" box="[1353,1436,223,248]" pageId="34" pageNumber="35" rank="species">sp.nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
showed 4–5 sites difference with
|
||
<taxonomicName id="77C50C83DC50A513FDEB890FFC00FE85" authority="sensu Voigt et al. 2017" authorityName="Voigt et al." authorityYear="2017" box="[539,974,260,285]" class="Calcarea" family="Clathrinidae" genus="Borojevia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clathrinida" pageId="34" pageNumber="35" phylum="Porifera" rank="species" species="aspina">
|
||
<emphasis id="82B1AB12DC50A513FDEB890FFDFAFE85" box="[539,564,261,284]" italics="true" pageId="34" pageNumber="35">B.</emphasis>
|
||
aff.
|
||
<emphasis id="82B1AB12DC50A513FD9C890FFD79FE85" box="[620,695,261,284]" italics="true" pageId="34" pageNumber="35">aspina</emphasis>
|
||
sensu
|
||
<bibRefCitation id="D4540AF1DC50A513FCF9890EFC00FE85" box="[777,974,260,285]" pageId="34" pageNumber="35" refId="ref70183" refString="Voigt, O., Erpenbeck, D., Gonzalez-Pech, R. A., Al-Aidaroos, A. M., Berumen, M. L. & Worheide, G. (2017) Calcinea of the Red Sea: providing a DNA barcode inventory with description of four new species. Marine Biodiversity, 47 (4), 1009 - 1034. https: // doi. org / 10.1007 / s 12526 - 017 - 0671 - x" type="journal article">
|
||
Voigt
|
||
<emphasis id="82B1AB12DC50A513FCBE890FFC44FE84" box="[846,906,260,285]" italics="true" pageId="34" pageNumber="35">et al.</emphasis>
|
||
2017
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
, 12–22 sites difference with
|
||
<taxonomicName id="77C50C83DC50A513FAED890FFA55FE84" authority="Van & De, 2018" authorityName="Van & De" authorityYear="2018" box="[1309,1435,260,285]" class="Calcarea" family="Clathrinidae" genus="Borojevia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clathrinida" pageId="34" pageNumber="35" phylum="Porifera" rank="species" species="tubulata" status="sp. nov.">
|
||
<emphasis id="82B1AB12DC50A513FAED890FFA55FE84" box="[1309,1435,260,285]" italics="true" pageId="34" pageNumber="35">B. tubulata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="82B1AB12DC50A513FF67892DFF27FED9" bold="true" box="[151,233,295,320]" pageId="34" pageNumber="35">
|
||
<taxonomicNameLabel id="99821669DC50A513FF67892DFF27FED9" box="[151,233,295,320]" pageId="34" pageNumber="35" rank="species">sp.nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(the higher difference being with the
|
||
<collectingCountry id="C8D23790DC50A513FD67892DFCC1FED9" box="[663,783,295,320]" name="Seychelles" pageId="34" pageNumber="35">Seychelles</collectingCountry>
|
||
specimen, as the
|
||
<typeStatus id="6F7EC9A2DC50A513FC29892DFBF5FED9" box="[985,1083,295,320]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
|
||
and Voigt’s
|
||
<taxonomicName id="77C50C83DC50A513FB3A8923FAF9FED9" authorityName="Van & De" authorityYear="2018" box="[1226,1335,297,320]" class="Calcarea" family="Clathrinidae" genus="Borojevia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clathrinida" pageId="34" pageNumber="35" phylum="Porifera" rank="genus">
|
||
<emphasis id="82B1AB12DC50A513FB3A8923FAF9FED9" box="[1226,1335,297,320]" italics="true" pageId="34" pageNumber="35">Borojevia</emphasis>
|
||
</taxonomicName>
|
||
sp. have identical sequences). It may be concluded that the new species and
|
||
<taxonomicName id="77C50C83DC50A513FC668947FB4AFEFD" box="[918,1156,332,357]" class="Calcarea" family="Clathrinidae" genus="Borojevia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clathrinida" pageId="34" pageNumber="35" phylum="Porifera" rank="species" species="aspina">
|
||
<emphasis id="82B1AB12DC50A513FC668947FBCDFEFD" box="[918,1027,333,356]" italics="true" pageId="34" pageNumber="35">Borojevia</emphasis>
|
||
aff.
|
||
<emphasis id="82B1AB12DC50A513FBC98947FB4AFEFD" box="[1081,1156,333,356]" italics="true" pageId="34" pageNumber="35">aspina</emphasis>
|
||
</taxonomicName>
|
||
are more closely related than each of them with
|
||
<taxonomicName id="77C50C83DC50A513FE6E897BFDD4FE11" authority="Van & De, 2018" authorityName="Van & De" authorityYear="2018" box="[414,538,367,392]" class="Calcarea" family="Clathrinidae" genus="Borojevia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clathrinida" pageId="34" pageNumber="35" phylum="Porifera" rank="species" species="tubulata" status="sp. nov.">
|
||
<emphasis id="82B1AB12DC50A513FE6E897BFDD4FE11" box="[414,538,367,392]" italics="true" pageId="34" pageNumber="35">B. tubulata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="82B1AB12DC50A513FDD28965FDBAFE11" bold="true" box="[546,628,367,392]" pageId="34" pageNumber="35">
|
||
<taxonomicNameLabel id="99821669DC50A513FDD28965FDBAFE11" box="[546,628,367,392]" pageId="34" pageNumber="35" rank="species">sp.nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<caption id="E4BA2788DC50A513FF678F56FD3EF836" httpUri="https://zenodo.org/record/1271289/files/figure.png" pageId="34" pageNumber="35" startId="34.[151,249,1884,1906]" targetBox="[151,1436,445,1863]" targetPageId="34">
|
||
<paragraph id="B07A7700DC50A513FF678F56FD3EF836" blockId="34.[151,1436,1884,1967]" pageId="34" pageNumber="35">
|
||
<emphasis id="82B1AB12DC50A513FF678F56FEEEF8E8" bold="true" box="[151,288,1884,1906]" pageId="34" pageNumber="35">FIGURE 12.</emphasis>
|
||
<taxonomicName id="77C50C83DC50A513FED78F57FE1AF8EB" authority="Van & De, 2018" authorityName="Van & De" authorityYear="2018" box="[295,468,1884,1906]" class="Calcarea" family="Clathrinidae" genus="Borojevia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clathrinida" pageId="34" pageNumber="35" phylum="Porifera" rank="species" species="pirella" status="sp. nov.">
|
||
<emphasis id="82B1AB12DC50A513FED78F57FE1AF8EB" box="[295,468,1884,1906]" italics="true" pageId="34" pageNumber="35">Borojevia pirella</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="82B1AB12DC50A513FE2C8F56FDE3F8E8" bold="true" box="[476,557,1884,1905]" pageId="34" pageNumber="35">
|
||
<taxonomicNameLabel id="99821669DC50A513FE2C8F56FDE6F8E8" box="[476,552,1884,1905]" pageId="34" pageNumber="35" rank="species">sp.nov.</taxonomicNameLabel>
|
||
,
|
||
</emphasis>
|
||
a, habitus in situ of holotype RMNH Por. 11622, from Rodrigues, the easternmost of the Mascarene Islands (photo N.J. de Voogd), a1, preserved holotype (scale bar = 1 cm), b, habitus in situ of paratype RMNH Por. 11622b (photo N.J. de Voogd), b1, preserved paratype.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="E4BA2788DC51A512FF678E4FFEE3F92E" httpUri="https://zenodo.org/record/1271295/files/figure.png" pageId="35" pageNumber="36" startId="35.[151,249,1605,1627]" targetBox="[177,1421,199,1578]" targetPageId="35">
|
||
<paragraph id="B07A7700DC51A512FF678E4FFEE3F92E" blockId="35.[151,1436,1605,1719]" pageId="35" pageNumber="36">
|
||
<emphasis id="82B1AB12DC51A512FF678E4FFEECF9C3" bold="true" box="[151,290,1605,1627]" pageId="35" pageNumber="36">FIGURE 13.</emphasis>
|
||
<taxonomicName id="77C50C83DC51A512FEDA8E4CFE17F9C2" authority="Van & De, 2018" authorityName="Van & De" authorityYear="2018" box="[298,473,1605,1627]" class="Calcarea" family="Clathrinidae" genus="Borojevia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clathrinida" pageId="35" pageNumber="36" phylum="Porifera" rank="species" species="pirella" status="sp. nov.">
|
||
<emphasis id="82B1AB12DC51A512FEDA8E4CFE17F9C2" box="[298,473,1605,1627]" italics="true" pageId="35" pageNumber="36">Borojevia pirella</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="82B1AB12DC51A512FE128E4FFDFCF9C3" bold="true" box="[482,562,1605,1626]" pageId="35" pageNumber="36">
|
||
<taxonomicNameLabel id="99821669DC51A512FE128E4FFDE0F9C3" box="[482,558,1605,1626]" pageId="35" pageNumber="36" rank="species">sp.nov.</taxonomicNameLabel>
|
||
,
|
||
</emphasis>
|
||
holotype RMNH Por. 11622, from Rodrigues, a, light microscopic cross section of oscular region showing pseudoatrium and surrounding tubuli, b–e, SEM images of the spicules, b, regular triactine, c, rare sagittal irregular triactine, d, tetractines, e, details of apical actines of the tetractines showing characteristic verticil of three conical spines.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="B07A7700DC51A512FF378ED4FDA0F86A" blockId="35.[151,1436,1758,2035]" pageId="35" pageNumber="36">
|
||
Molecular comparison of
|
||
<taxonomicName id="77C50C83DC51A512FE1D8ED5FD99F96E" authority="Van & De, 2018" authorityName="Van & De" authorityYear="2018" box="[493,599,1758,1783]" class="Calcarea" family="Clathrinidae" genus="Borojevia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clathrinida" pageId="35" pageNumber="36" phylum="Porifera" rank="species" species="pirella" status="sp. nov.">
|
||
<emphasis id="82B1AB12DC51A512FE1D8ED5FD99F96E" box="[493,599,1758,1783]" italics="true" pageId="35" pageNumber="36">B. pirella</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="82B1AB12DC51A512FDAF8ED4FD7CF96E" bold="true" box="[607,690,1758,1783]" pageId="35" pageNumber="36">
|
||
<taxonomicNameLabel id="99821669DC51A512FDAF8ED4FD7CF96E" box="[607,690,1758,1783]" pageId="35" pageNumber="36" rank="species">sp.nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
with the similar Brazilian species
|
||
<taxonomicName id="77C50C83DC51A512FBCE8ED5FB05F96F" box="[1086,1227,1759,1782]" class="Calcarea" family="Clathrinidae" genus="Borojevia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clathrinida" pageId="35" pageNumber="36" phylum="Porifera" rank="species" species="trispinata">
|
||
<emphasis id="82B1AB12DC51A512FBCE8ED5FB05F96F" box="[1086,1227,1759,1782]" italics="true" pageId="35" pageNumber="36">B. trispinata</emphasis>
|
||
</taxonomicName>
|
||
is complicated by differences in the target genes (our 28S vs
|
||
<bibRefCitation id="D4540AF1DC51A512FD8F8F08FCA4F882" box="[639,874,1794,1819]" pageId="35" pageNumber="36" refId="ref63142" refString="Azevedo, F., Padua, A., Moraes, F., Rossi, A., Muricy, G. & Klautau, M. (2017) Taxonomy and phylogeny of calcareous sponges (Porifera: Calcarea: Calcinea) from Brazilian mid-shelf and oceanic islands. Zootaxa, 4311 (3), 301 - 344. https: // doi. org / 10.11646 / zootaxa. 4311.3.1" type="journal article">
|
||
Azevedo
|
||
<emphasis id="82B1AB12DC51A512FD1B8F0EFCEBF882" box="[747,805,1794,1819]" italics="true" pageId="35" pageNumber="36">et al.</emphasis>
|
||
2017
|
||
</bibRefCitation>
|
||
’s ITS). In a special effort, we obtained additional ITS data for
|
||
<taxonomicName id="77C50C83DC51A512FEC28F2DFE6EF8A6" authority="Van & De, 2018" authorityName="Van & De" authorityYear="2018" box="[306,416,1830,1855]" class="Calcarea" family="Clathrinidae" genus="Borojevia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clathrinida" pageId="35" pageNumber="36" phylum="Porifera" rank="species" species="pirella" status="sp. nov.">
|
||
<emphasis id="82B1AB12DC51A512FEC28F2DFE6EF8A6" box="[306,416,1830,1855]" italics="true" pageId="35" pageNumber="36">B. pirella</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="82B1AB12DC51A512FE5B8F2CFE33F8A6" bold="true" box="[427,509,1830,1855]" pageId="35" pageNumber="36">
|
||
<taxonomicNameLabel id="99821669DC51A512FE5B8F2CFE33F8A6" box="[427,509,1830,1855]" pageId="35" pageNumber="36" rank="species">sp.nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and aligned two ITS sequences (
|
||
<typeStatus id="6F7EC9A2DC51A512FC7A8F2CFC20F8A6" box="[906,1006,1830,1855]" pageId="35" pageNumber="36" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="6F7EC9A2DC51A512FBDD8F2DFB5EF8A6" box="[1069,1168,1831,1855]" pageId="35" pageNumber="36" type="paratype">paratype</typeStatus>
|
||
, which appeared to be identical, not submitted to GenBank) with two sequences of
|
||
<taxonomicName id="77C50C83DC51A512FC548F46FBF5F8FA" box="[932,1083,1868,1891]" class="Calcarea" family="Clathrinidae" genus="Borojevia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clathrinida" pageId="35" pageNumber="36" phylum="Porifera" rank="species" species="trispinata">
|
||
<emphasis id="82B1AB12DC51A512FC548F46FBF5F8FA" box="[932,1083,1868,1891]" italics="true" pageId="35" pageNumber="36">B. trispinata</emphasis>
|
||
</taxonomicName>
|
||
downloaded from GenBank (UFRJPOR6419 and 6487, showing several site differences). A trimmed 792 sites alignment of the four sequences shows considerable differences (25 non–conserved sites differed between the two species, not counting a gap of 27 sites). Nevertheless, the morphological resemblance of the two species from the
|
||
<collectingRegion id="7201B9E2DC51A512FBCF8FBDFB54F857" box="[1087,1178,1975,1998]" country="Sri Lanka" name="Western" pageId="35" pageNumber="36">Western</collectingRegion>
|
||
Indian Ocean and the
|
||
<collectingRegion id="7201B9E2DC51A512FF678FD1FF3CF86B" box="[151,242,2011,2034]" country="Sri Lanka" name="Western" pageId="35" pageNumber="36">Western</collectingRegion>
|
||
Atlantic Ocean region is striking.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |