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<document id="4F01A2AF3BEC9235E8AC192AC821B5F6" ID-DOI="10.5281/zenodo.13344847" ID-ISSN="2533-4069" ID-Zenodo-Dep="13344847" IM.bibliography_approvedBy="carolina" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722606393481" checkinUser="felipe" docAuthor="ZÁGORŠEK, KAMIL" docDate="2010" docId="316187B25325FFB80678F99D2940FA03" docLanguage="en" docName="ActaMusNatPraSerB.66.3-4.139-255.pdf" docOrigin="Acta Musei Nationalis Pragae Series B 66 (3 - 4)" docStyle="DocumentStyle:90C434725D1F5A377F8E8E9FAA5647D0.1:ActaMusNatPraSerB.2007-2013.journal_article.4cover" docStyleId="90C434725D1F5A377F8E8E9FAA5647D0" docStyleName="ActaMusNatPraSerB.2007-2013.journal_article.4cover" docStyleVersion="1" docTitle="Umbonula granulata ZÁGORŠEK 2010, sp. n." docType="treatment" docVersion="4" lastPageNumber="255" masterDocId="CD58FFCA5328FFB50763FFEB2D41FF81" masterDocTitle="BRYOZOA FROM THE LANGHIAN (MIOCENE) OF THE CZECH REPUBLIC PART II: SYSTEMATIC DESCRIPTION OF THE SUBORDER ASCOPHORA LEVINSEN, 1909 AND PALEOECOLOGICAL RECONSTRUCTION OF THE STUDIED PALEOENVIRONMENT" masterLastPageNumber="255" masterPageNumber="139" pageNumber="139" updateTime="1724090549386" updateUser="ExternalLinkService">
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<mods:title id="A617A923FE900A14D2EE19C85A8EB1DA">BRYOZOA FROM THE LANGHIAN (MIOCENE) OF THE CZECH REPUBLIC PART II: SYSTEMATIC DESCRIPTION OF THE SUBORDER ASCOPHORA LEVINSEN, 1909 AND PALEOECOLOGICAL RECONSTRUCTION OF THE STUDIED PALEOENVIRONMENT</mods:title>
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<mods:name id="2970D4D3F22513FE4E9F7A95DC12FFDF" type="personal">
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<mods:namePart id="67DAD0CD856DB4A587C2D7506799828B">ZÁGORŠEK, KAMIL</mods:namePart>
<mods:affiliation id="8D45F8871D1DA0F2FB74DDFF5E9558F9">National Museum, Prague, Václavské nám. 68,115 79 Praha 1, Czech Republic</mods:affiliation>
<mods:nameIdentifier id="84517E812166B2778FF601F2F1E51678" type="email">kamil_zagorsek@nm.cz</mods:nameIdentifier>
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<mods:typeOfResource id="8F1A7A35F6A46AF0C3D472D6D3BCD4E8">text</mods:typeOfResource>
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<mods:title id="8E1252367D5E2639214A6C70E69FED12">Acta Musei Nationalis Pragae Series B</mods:title>
</mods:titleInfo>
<mods:part id="DFF27C9CA5836B2198AD61B40E13FC90">
<mods:date id="83C27755B47CCD5FFE6CB12892876560">2010</mods:date>
<mods:detail id="B4B29EE9D5BD4869E54BDEB3DA89F192" type="volume">
<mods:number id="5C01C02E8ACF73D25B8AB75B894361E8">66</mods:number>
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<mods:number id="C7E31BF96A4B6D87B10D82ABFAC1A0BA">3 - 4</mods:number>
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<mods:identifier id="59D6941EB62D9C5907A05F00F2580E79" type="DOI">10.5281/zenodo.13344847</mods:identifier>
<mods:identifier id="49831CA9BEE3C4B45EB700924034C9B1" type="ISSN">2533-4069</mods:identifier>
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<treatment id="316187B25325FFB80678F99D2940FA03" ID-DOI="http://doi.org/10.5281/zenodo.13227656" ID-Zenodo-Dep="13227656" LSID="urn:lsid:plazi:treatment:316187B25325FFB80678F99D2940FA03" httpUri="http://treatment.plazi.org/id/316187B25325FFB80678F99D2940FA03" pageId="13">
<subSubSection id="F1D2652F5325FFB80678F99D2CBBF934" pageId="13" type="nomenclature">
<paragraph id="B97736A45325FFB80678F99D2F06F90C" blockId="13.[283,583,1654,1677]" box="[283,583,1654,1677]" pageId="13">
<heading id="E23F81C85325FFB80678F99D2F06F90C" box="[283,583,1654,1677]" centered="true" fontSize="8" level="2" pageId="13" reason="2">
<emphasis id="8BBCEAB65325FFB80678F99D2F06F90C" bold="true" box="[283,583,1654,1677]" pageId="13">
<taxonomicName id="7EC84D275325FFB80678F99D2F45F90C" ID-CoL="86V5D" authority="ZÁGORŠEK, 2010" authorityName="ZÁGORŠEK" authorityYear="2010" box="[283,516,1654,1677]" class="Gymnolaemata" family="Umbonulidae" genus="Umbonula" kingdom="Animalia" order="Cheilostomatida" pageId="13" phylum="Bryozoa" rank="species" species="granulata" status="sp. nov.">
<emphasis id="8BBCEAB65325FFB80678F99D2F45F90C" bold="true" box="[283,516,1654,1677]" italics="true" pageId="13">Umbonula granulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="908F57CD5325FFB80568F99D2F06F90C" box="[523,583,1654,1677]" pageId="13" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="B97736A45325FFB8060AF94B2CBBF934" blockId="13.[361,506,1696,1717]" box="[361,506,1696,1717]" pageId="13">Pl. 95, Fig. 1-6</paragraph>
</subSubSection>
<subSubSection id="F1D2652F5325FFB807F9F9262940FA03" pageId="13" type="description">
<paragraph id="B97736A45325FFB807F9F9262F83F8E5" blockId="13.[116,752,1741,2099]" pageId="13">
D i a g n o s i s: Colony encrusting. Autozooecia rhomboidal with granular frontal wall. Marginal areolar pores small, about 10-15 on the border of each autozooecium. Aperture with a large lyrula. Avicularia of
<specimenCount id="AFCEFD2D5325FFB8051CF8C62FAAF8C5" box="[639,747,1837,1860]" count="2" pageId="13" type="generic" typeStatus="types">two types</specimenCount>
, small suboral and giant spatulate. No ovicell observed.
</paragraph>
<paragraph id="B97736A45325FFB807F9F89E2FA1F84D" blockId="13.[116,752,1741,2099]" pageId="13">
H o l o t y p e: The specimen illustrated in Pl. 95, Fig. 1 and 4-6, from the section Podbřežice, is deposited in the National Museum
<collectingRegion id="7B0CF8465325FFB8065FF85E2CC7F84D" box="[316,390,1973,1996]" country="Czech Republic" name="Praha" pageId="13">Prague</collectingRegion>
under number PM2 P 01903.
</paragraph>
<paragraph id="B97736A45325FFB807F9F8372FAEF7B2" blockId="13.[116,752,1741,2099]" pageId="13">
<materialsCitation id="09A03CF95325FFB807F9F8372F44F7B2" collectionCode="PM" location="National Museum" municipality="Podbrezice" pageId="13" specimenCode="PM2" specimenCount="4" stateProvince="Praha">
P a r a t y p e s: Altogether
<specimenCount id="AFCEFD2D5325FFB806B5F8372F1AF872" box="[470,603,2012,2035]" count="4" pageId="13" type="generic">4 specimens</specimenCount>
were studied from the locality
<collectingMunicipality id="5913ACDE5325FFB80621F8172CFAF792" box="[322,443,2044,2067]" pageId="13">Podbřežice</collectingMunicipality>
, deposited in the
<location id="BC17607F5325FFB805F0F8172D90F7B2" LSID="urn:lsid:plazi:treatment:316187B25325FFB80678F99D2940FA03:BC17607F5325FFB805F0F8172D90F7B2" municipality="Podbrezice" name="National Museum" pageId="13" stateProvince="Praha">National Museum</location>
<collectingRegion id="7B0CF8465325FFB807BAF7F72C62F7B2" box="[217,291,2076,2099]" country="Czech Republic" name="Praha" pageId="13">Prague</collectingRegion>
under numbers
<specimenCode id="E96E9EDF5325FFB806B2F7F72F44F7B2" box="[465,517,2076,2099]" collectionCode="PM" country="USA" name="Pratt Museum" pageId="13">PM2</specimenCode>
</materialsCitation>
P 01904-P 01907.
</paragraph>
<paragraph id="B97736A45325FFB80717F78D2DDFF7FF" blockId="13.[116,158,2150,2174]" box="[116,158,2150,2174]" pageId="13">148</paragraph>
<paragraph id="B97736A45325FFB80403FF722EFFFF51" blockId="13.[827,1463,153,1410]" pageId="13">D e r i v a t i o n o m i n i s: Due to the granular frontal wall.</paragraph>
<paragraph id="B97736A45325FFB80403FF0B295FFE97" blockId="13.[827,1463,153,1410]" pageId="13">L o c u s t y p i c u s: section Podbřežice, sample Podbřežice village-1.</paragraph>
<paragraph id="B97736A45325FFB80403FECD28F6FEBC" blockId="13.[827,1463,153,1410]" box="[864,1463,294,317]" pageId="13">S t r a t u m t y p i c u m: Langhian Lower Badenian.</paragraph>
<paragraph id="B97736A45325FFB80403FEA62EF4FE05" blockId="13.[827,1463,153,1410]" pageId="13">M e a s u r e m e n t s: (in micro meters = µm; x = = average):</paragraph>
<paragraph id="B97736A45325FFB80403FE672850FE22" blockId="13.[827,1463,153,1410]" box="[864,1297,396,419]" pageId="13">length of autozooecia: 499-654; x = 614</paragraph>
<paragraph id="B97736A45325FFB80403FE47284AFE42" blockId="13.[827,1463,153,1410]" box="[864,1291,428,451]" pageId="13">width of autozooecia: 333-492; x = 396</paragraph>
<paragraph id="B97736A45325FFB80403FE202836FE63" blockId="13.[827,1463,153,1410]" box="[864,1399,459,482]" pageId="13">length of autozooecial aperture: 152-231; x = 180</paragraph>
<paragraph id="B97736A45325FFB80403FE002830FD83" blockId="13.[827,1463,153,1410]" box="[864,1393,491,514]" pageId="13">width of autozooecial aperture: 157-193; x = 176</paragraph>
<paragraph id="B97736A45325FFB80403FDE1280AFDA0" blockId="13.[827,1463,153,1410]" box="[864,1355,522,545]" pageId="13">length of suboral avicularium: 65-112; x = 85</paragraph>
<paragraph id="B97736A45325FFB80403FDC12878FDC0" blockId="13.[827,1463,153,1410]" box="[864,1337,554,577]" pageId="13">width of suboral avicularium: 57-85; x = 72</paragraph>
<paragraph id="B97736A45325FFB80403FDA22836FDE1" blockId="13.[827,1463,153,1410]" box="[864,1399,585,608]" pageId="13">length of spatulate avicularium: 275-299; x = 287</paragraph>
<paragraph id="B97736A45325FFB80403FD822830FD01" blockId="13.[827,1463,153,1410]" box="[864,1393,617,640]" pageId="13">width of spatulate avicularium: 164-197; x = 181</paragraph>
<paragraph id="B97736A45325FFB80403FD632844FD1E" blockId="13.[827,1463,153,1410]" box="[864,1285,648,671]" pageId="13">diameter of areolar pore: 20-68; x = 48</paragraph>
<paragraph id="B97736A45325FFB80403FD44295BFB80" blockId="13.[827,1463,153,1410]" pageId="13">D e s c r i p t i o n: Colony encrusting. Autozooecia rhomboidal with flat, granular frontal walls. Granules small, regularly arranged in quincuncial pattern. Marginal areolar pores small, there are about 10-15 pores on the bor- der of each autozooecium. Aperture large, oval with a large lyrula. Adventitious avicularia suboral or spatulate. Suboral avicularia with a pivotal bar (Pl. 95, Fig. 6); they are situat- ed on the middle of the frontal wall, proximally from the aperture, at the top of the umbo. The spatulate avicularia are giant, with an oval rostrum and a pivotal bar (Pl. 95, Fig. 3). No ovicell observed.</paragraph>
<paragraph id="B97736A45325FFB80403FBFA2940FA03" blockId="13.[827,1463,153,1410]" pageId="13">
C o m p a r i s o n: This new species is very similar to
<taxonomicName id="7EC84D275325FFB80458FBDB29F9FBC6" authority="Manzoni, 1877" authorityName="Manzoni" authorityYear="1877" box="[827,1208,1072,1095]" class="Gymnolaemata" family="Flustridae" genus="Eschara" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Cheilostomatida" pageId="13" phylum="Bryozoa" rank="species" species="chiragra">
<emphasis id="8BBCEAB65325FFB80458FBDB2EBEFBC6" box="[827,1023,1072,1095]" italics="true" pageId="13">Eschara chiragra</emphasis>
<bibRefCitation id="DD594B555325FFB8036DFBDB29F9FBC6" author="Manzoni, A." box="[1038,1208,1072,1095]" pageId="13" pagination="49 - 77" refId="ref41646" refString="Manzoni, A. (1877): I Briozoi fossili del Miocene d'Austria ed Ungheria, 2. - Denkschr. k. Akad. Wiss., math-naturwiss. Kl., 37 (2): 49 - 77." type="journal article" year="1877">
Manzoni,
<quantity id="7E309B415325FFB803E0FBDB29F9FBC6" box="[1155,1208,1072,1095]" metricMagnitude="1" metricUnit="m" metricValue="4.76758" pageId="13" unit="in" value="1877.0">1877</quantity>
</bibRefCitation>
</taxonomicName>
in having a granular frontal wall, but it differs in having giant avicularia and encrusting colonies (
<taxonomicName id="7EC84D275325FFB8034EFB8428F4FB07" authority="MANZONI, 1877" authorityName="Manzoni" authorityYear="1877" box="[1069,1461,1135,1158]" class="Gymnolaemata" family="Flustridae" genus="Eschara" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Cheilostomatida" pageId="13" phylum="Bryozoa" rank="species" species="chiragra">
<emphasis id="8BBCEAB65325FFB8034EFB8429B2FB07" box="[1069,1267,1135,1158]" italics="true" pageId="13">Eschara chiragra</emphasis>
<bibRefCitation id="DD594B555325FFB80267FB8428F4FB07" author="Manzoni, A." box="[1284,1461,1135,1158]" pageId="13" pagination="49 - 77" refId="ref41646" refString="Manzoni, A. (1877): I Briozoi fossili del Miocene d'Austria ed Ungheria, 2. - Denkschr. k. Akad. Wiss., math-naturwiss. Kl., 37 (2): 49 - 77." type="journal article" year="1877">
<smallCapsWord id="BF91A0785325FFB80267FB842828FB07" baselines="1152,1153" box="[1284,1385,1135,1158]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Manzoni" pageId="13">MANZONI</smallCapsWord>
, 1877
</bibRefCitation>
</taxonomicName>
forms bilaminar colonies).
<taxonomicName id="7EC84D275325FFB80339FB642E38FB44" authority="(PROCHAZKA, 1893)" authorityName="Pl." baseAuthorityName="PROCHAZKA" baseAuthorityYear="1893" class="Gymnolaemata" family="Umbonulidae" genus="Umbonula" kingdom="Animalia" order="Cheilostomatida" pageId="13" phylum="Bryozoa" rank="species" species="spinosa">
<emphasis id="8BBCEAB65325FFB80339FB642860FB27" box="[1114,1313,1167,1190]" italics="true" pageId="13">Umbonula spinosa</emphasis>
(
<bibRefCitation id="DD594B555325FFB80253FB642E33FB44" author="Prochazka, V. J." pageId="13" pagination="1 - 84" refId="ref42258" refString="Prochazka, V. J. (1893): Miocen kralicky u Nameste na Morave [Miocene around Kralice by Namest na Morave]. - Vestnik Kralovske Ceske Spolecnosti Nauk, trida mathematicko-prirodovedna, 6: 1 - 84. (in Czech)." type="journal article" year="1893">
<smallCapsWord id="BF91A0785325FFB80253FB6428EEFB24" baselines="1184,1184" box="[1328,1455,1167,1190]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Prochazka" pageId="13">PROCHÁZKA</smallCapsWord>
, 1893
</bibRefCitation>
)
</taxonomicName>
resembles this new species in having the “spines” similar to granules around the autozooecia. The granules are however much smaller than the tubercles on
<taxonomicName id="7EC84D275325FFB80226FB062932FAA5" authority="(PROCHAZKA, 1893)" authorityName="Pl." baseAuthorityName="PROCHAZKA" baseAuthorityYear="1893" class="Gymnolaemata" family="Umbonulidae" genus="Umbonula" kingdom="Animalia" order="Cheilostomatida" pageId="13" phylum="Bryozoa" rank="species" species="spinosa">
<emphasis id="8BBCEAB65325FFB80226FB062ECDFAA5" italics="true" pageId="13">Umbonula spinosa</emphasis>
(
<bibRefCitation id="DD594B555325FFB804C3FAE6292AFAA5" author="Prochazka, V. J." box="[928,1131,1293,1316]" pageId="13" pagination="1 - 84" refId="ref42258" refString="Prochazka, V. J. (1893): Miocen kralicky u Nameste na Morave [Miocene around Kralice by Namest na Morave]. - Vestnik Kralovske Ceske Spolecnosti Nauk, trida mathematicko-prirodovedna, 6: 1 - 84. (in Czech)." type="journal article" year="1893">
<smallCapsWord id="BF91A0785325FFB804C3FAE6295EFAA2" baselines="1310,1310" box="[928,1055,1293,1316]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Prochazka" pageId="13">PROCHÁZKA</smallCapsWord>
, 1893
</bibRefCitation>
)
</taxonomicName>
. There are no ridges on the frontal wall as in
<taxonomicName id="7EC84D275325FFB80491FAC728C8FAC2" authority="(REUSS, 1847)" authorityName="Pl." baseAuthorityName="REUSS" baseAuthorityYear="1847" box="[1010,1417,1324,1347]" class="Gymnolaemata" family="Umbonulidae" genus="Umbonula" kingdom="Animalia" order="Cheilostomatida" pageId="13" phylum="Bryozoa" rank="species" species="macrocheila">
<emphasis id="8BBCEAB65325FFB80491FAC729ACFAC2" box="[1010,1261,1324,1347]" italics="true" pageId="13">Umbonula macrocheila</emphasis>
(
<bibRefCitation id="DD594B555325FFB80399FAC728C3FAC2" author="Reuss, A. E." box="[1274,1410,1324,1347]" pageId="13" pagination="1 - 109" refId="ref42305" refString="Reuss, A. E. (1847): Die fossilen Polyparien des Wiener Tertiarbeckens. Ein monographischer Versuch. - Naturwiss. Abhadlung., 2 (1): 1 - 109." type="journal article" year="1847">
<smallCapsWord id="BF91A0785325FFB80399FAC7287FFAC3" baselines="1341,1341" box="[1274,1342,1324,1347]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Reuss" pageId="13">REUSS</smallCapsWord>
, 1847
</bibRefCitation>
)
</taxonomicName>
and the autozooecia in
<taxonomicName id="7EC84D275325FFB80360FAA029A2FAE3" authority="ZÁGORŠEK, 2010" authorityName="ZÁGORŠEK" authorityYear="2010" box="[1027,1251,1355,1378]" class="Gymnolaemata" family="Umbonulidae" genus="Umbonula" kingdom="Animalia" order="Cheilostomatida" pageId="13" phylum="Bryozoa" rank="species" species="granulata" status="sp. nov.">
<emphasis id="8BBCEAB65325FFB80360FAA029A2FAE3" box="[1027,1251,1355,1378]" italics="true" pageId="13">Umbonula granulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="908F57CD5325FFB8038AFAA02863FAE3" box="[1257,1314,1355,1378]" pageId="13" rank="species">sp. n.</taxonomicNameLabel>
are of a more rhomboidal shape.
</paragraph>
</subSubSection>
</treatment>
</document>