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<document id="AF6B8B6EC64D1208179ADAFAF1A81A53" ID-CLB-Dataset="23965" ID-DOI="10.1080/00222933.2019.1657194" ID-GBIF-Dataset="2d874abb-081f-40a2-a358-3686c451b9b2" ID-ISSN="1464-5262" ID-Zenodo-Dep="3670139" ID-ZooBank="FDB9EDDA-63A8-43C9-903C-48BF15C45F03" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1581941320443" checkinUser="valdenar" docAuthor="Cruz-López, Jesús A. &amp; Francke, Oscar F." docDate="2019" docId="03CDC35CBC1DFFFCFE15FBD7FBC9FC15" docLanguage="en" docName="JNATHIST.53.2324.14511464.pdf.imf" docOrigin="Journal of Natural History 53 (23)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Huasteca kardia Cruz-López &amp; Francke 2019, sp. nov." docType="treatment" docVersion="10" lastPageNumber="1459" masterDocId="FFF4BB24BC1EFFF5FFEFFFEFFFAAFFDB" masterDocTitle="New species of the cave-dwelling genus Huasteca (Opiliones: Stygnopsidae: Karosinae) ı from Northern Oaxacaı Mexicoı with a SEM survey of the sexually dimorphic areas on legs and structures related to chelicerae in the genus" masterLastPageNumber="1464" masterPageNumber="1451" pageNumber="1453" updateTime="1698783545986" updateUser="ExternalLinkService">
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<mods:title id="24C11CF68599F777823D93E8A5DB3C3C">New species of the cave-dwelling genus Huasteca (Opiliones: Stygnopsidae: Karosinae) ı from Northern Oaxacaı Mexicoı with a SEM survey of the sexually dimorphic areas on legs and structures related to chelicerae in the genus</mods:title>
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<mods:namePart id="81E4D0D2AFFA3160C22CD6D5D88A2359">Cruz-López, Jesús A.</mods:namePart>
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<mods:namePart id="86A9C255803C69919C8E979CC6D8AE5E">Francke, Oscar F.</mods:namePart>
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<mods:number id="A383EFCBB2E0F389D92B2D2A66EFDDB5">2019-09-05</mods:number>
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<treatment id="03CDC35CBC1DFFFCFE15FBD7FBC9FC15" ID-DOI="http://doi.org/10.5281/zenodo.3679548" ID-GBIF-Taxon="162163861" ID-Zenodo-Dep="3679548" LSID="urn:lsid:plazi:treatment:03CDC35CBC1DFFFCFE15FBD7FBC9FC15" httpUri="http://treatment.plazi.org/id/03CDC35CBC1DFFFCFE15FBD7FBC9FC15" lastPageId="9" lastPageNumber="1459" pageId="3" pageNumber="1453">
<subSubSection id="C37E21C1BC1DFFF6FE15FBD7FC80FB89" box="[506,810,1080,1106]" pageId="3" pageNumber="1453" type="nomenclature">
<paragraph id="8BDB724ABC1DFFF6FE15FBD7FC80FB89" blockId="3.[506,810,1080,1106]" box="[506,810,1080,1106]" pageId="3" pageNumber="1453">
<heading id="D093C526BC1DFFF6FE15FBD7FC80FB89" box="[506,810,1080,1106]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="1453" reason="2">
<taxonomicName id="4C6409C9BC1DFFF6FE15FBD7FD69FB89" ID-CoL="7WNHG" authority="Cruz-López &amp; Francke, 2019" authorityName="Cruz-López &amp; Francke" authorityYear="2019" box="[506,707,1080,1106]" class="Arachnida" family="Stygnopsidae" genus="Huasteca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="1453" phylum="Arthropoda" rank="species" species="kardia" status="sp. nov.">
<emphasis id="B910AE58BC1DFFF6FE15FBD7FD69FB89" bold="true" box="[506,707,1080,1106]" italics="true" pageId="3" pageNumber="1453">Huasteca kardia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2231323BC1DFFF6FD22FBD6FC80FB89" box="[717,810,1081,1106]" pageId="3" pageNumber="1453" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37E21C1BC1DFFF6FF4FFB82FC84FB3F" pageId="3" pageNumber="1453" type="reference_group">
<paragraph id="8BDB724ABC1DFFF6FF4FFB82FD3EFB72" blockId="3.[160,1156,1133,1193]" pageId="3" pageNumber="1453">
<treatmentCitationGroup id="AB745564BC1DFFF6FF4FFB82FD3EFB72" pageId="3" pageNumber="1453">
<taxonomicName id="4C6409C9BC1DFFF6FF4FFB82FEECFB5D" authority="Goodnight and Goodnight 1971" authorityName="Goodnight and Goodnight" authorityYear="1971" box="[160,326,1133,1158]" class="Arachnida" family="Stygnopsidae" genus="Karos" higherTaxonomySource="CoL" kingdom="Plantae" order="Opiliones" pageId="3" pageNumber="1453" phylum="Arthropoda" rank="species" species="gratiosus">
<emphasis id="B910AE58BC1DFFF6FF4FFB82FEECFB5D" box="[160,326,1133,1158]" italics="true" pageId="3" pageNumber="1453">Karos gratiosus</emphasis>
</taxonomicName>
(in part):
<treatmentCitation id="0AC5545BBC1DFFF6FE5EFB81FCE9FB5D" author="Goodnight CJi Goodnight ML" box="[433,835,1134,1158]" page="35" pageId="3" pageNumber="1453" year="1971">
<bibRefCitation id="EFF50FBBBC1DFFF6FE5EFB81FCE9FB5D" author="Goodnight CJi Goodnight ML" box="[433,835,1134,1158]" pageId="3" pageNumber="1453" pagination="33 - 45" refId="ref4670" refString="Goodnight CJi Goodnight ML. 1971. Opilionids (Phalangida) of the family Phalangodidae from Mexican caves. Bull Assoc Mex Stud. 4: 33 - 45." type="journal article" year="1971">Goodnight and Goodnight 1971: 35</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AC5545BBC1DFFF6FCA2FB81FBBFFB5D" author="Reddell JR" box="[845,1045,1134,1158]" page="161" pageId="3" pageNumber="1453" year="1981">
<bibRefCitation id="EFF50FBBBC1DFFF6FCA2FB81FBBFFB5D" author="Reddell JR" box="[845,1045,1134,1158]" pageId="3" pageNumber="1453" pagination="257" refId="ref4992" refString="Reddell JR. 1981. A review of the cavernicole fauna of Mexicoi Guatemala and Belize. Mus Espeleol Monogr. 26: 257." type="journal article" year="1981">Reddell 1981: 161</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AC5545BBC1DFFF6FBF0FB81FE72FB72" author="Kury ABi Cokendolpher JC" page="155" pageId="3" pageNumber="1453" year="2000">
<bibRefCitation id="EFF50FBBBC1DFFF6FBF0FB81FE72FB72" author="Kury ABi Cokendolpher JC" pageId="3" pageNumber="1453" pagination="137 - 157" refId="ref4754" refString="Kury ABi Cokendolpher JC. 2000. Opiliones. In: Llorente-Bousquets Ji Gonzalez-Soriano JEi Papavero Ni editors. Biodiversidadi taxonomia y biogeografia de artropodos de Mexico: hacia una sintesis de su conocimiento. Vol. II. Mexico: Facultad de Cienciasi UNAMi CONABIO and Bayer; p. 137 - 157." type="book chapter" year="2000">Kury and Cokendolpher 2000: 155</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AC5545BBC1DFFF6FE0BFB7EFD3AFB72" author="Kury AB" box="[484,656,1169,1193]" page="238" pageId="3" pageNumber="1453" year="2003">
<bibRefCitation id="EFF50FBBBC1DFFF6FE0BFB7EFD3AFB72" author="Kury AB" box="[484,656,1169,1193]" pageId="3" pageNumber="1453" pagination="337" refId="ref4726" refString="Kury AB. 2003. Annotated catalogue of the Laniatores of the New World. Rev Iber Aracnol. Vol. Special Monogr. 1: 337." type="journal article" year="2003">Kury 2003: 238</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<subSection id="E2EB6BA1BC1DFFF6FE1AFB24FC84FB3F" box="[501,814,1226,1252]" pageId="3" pageNumber="1453" type="multiple">
<paragraph id="8BDB724ABC1DFFF6FE1AFB24FC84FB3F" blockId="3.[501,814,1226,1252]" box="[501,814,1226,1252]" pageId="3" pageNumber="1453">
<heading id="D093C526BC1DFFF6FE1AFB24FC84FB3F" box="[501,814,1226,1252]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="1453" reason="5">
(
<figureCitation id="135F6ECFBC1DFFF6FE12FB24FDF0FB3F" box="[509,602,1227,1252]" captionStart="Figure 1" captionStartId="4.[160,225,1624,1647]" captionTargetBox="[163,1153,142,1592]" captionTargetId="figure@4.[163,1153,142,1592]" captionTargetPageId="4" captionText="Figure 1. Habitus of Huasteca species. (a) male of Huasteca gratiosa. (b) male of Huasteca rugosa. (c) female of Huasteca silhavyi. (d) female of Huasteca kardia sp. nov. Scale bars = 0.5 cm." figureDoi="http://doi.org/10.5281/zenodo.3670141" httpUri="https://zenodo.org/record/3670141/files/figure.png" pageId="3" pageNumber="1453">Figure 1</figureCitation>
(d)ı 2
<emphasis id="B910AE58BC1DFFF6FD73FB25FD00FB38" box="[668,682,1226,1251]" italics="true" pageId="3" pageNumber="1453"></emphasis>
5ı 6(dıe)ı 7)
</heading>
</paragraph>
</subSection>
</subSubSection>
<subSubSection id="C37E21C1BC1DFFF0FF4FFACAFD2CFE42" lastPageId="5" lastPageNumber="1455" pageId="3" pageNumber="1453" type="materials_examined">
<paragraph id="8BDB724ABC1DFFF6FF4FFACAFEEFFAE5" blockId="3.[160,1156,1317,1550]" box="[160,325,1317,1342]" pageId="3" pageNumber="1453">
<emphasis id="B910AE58BC1DFFF6FF4FFACAFEEFFAE5" bold="true" box="[160,325,1317,1342]" italics="true" pageId="3" pageNumber="1453">
<typeStatus id="54DFCCE8BC1DFFF6FF4FFACAFF72FAE5" box="[160,216,1317,1342]" pageId="3" pageNumber="1453">Type</typeStatus>
material
</emphasis>
</paragraph>
<paragraph id="8BDB724ABC1DFFF6FF4FFAA6FC49F9D5" blockId="3.[160,1156,1317,1550]" pageId="3" pageNumber="1453">
<typeStatus id="54DFCCE8BC1DFFF6FF4FFAA6FEAFFABA" box="[160,261,1353,1377]" pageId="3" pageNumber="1453" type="holotype">Holotype</typeStatus>
male (CNAN-T01302)ı Mexicoı Oaxacaı Huautla de Jiménezı Puente de Fierro caveı 18.15307Nı 96.85320Wı leg. O. Franckeı J. Mendozaı G. Contrerası R. Monjarazı
<date id="FFDA548ABC1DFFF6FF4FFA61FE97FA7D" box="[160,317,1422,1446]" pageId="3" pageNumber="1453" value="2015-04-13">13 April 2015</date>
.
<materialsCitation id="3B0C7817BC1DFFF6FEA4FA61FD37FA30" ID-GBIF-Occurrence="2571153597" collectingDate="2014-04-12" collectionCode="CNAN- T01303, CNAN- T01304" collectorName="Paratypes &amp; O. Franckei J. Cruzi" location="Huautla de Jimenezi Li Nita" municipality="Ni" pageId="3" pageNumber="1453" specimenCount="1" typeStatus="holotype">
<collectorName id="2691179CBC1DFFF6FEA4FA61FE13FA7D" box="[331,441,1422,1446]" pageId="3" pageNumber="1453">Paratypes</collectorName>
: female (
<collectionCode id="ED75EA8FBC1DFFF6FDCEFA61FD61FA7D" box="[545,715,1422,1446]" pageId="3" pageNumber="1453">CNAN- T01303</collectionCode>
) same data as
<typeStatus id="54DFCCE8BC1DFFF6FC91FA61FC49FA7D" box="[894,995,1422,1446]" pageId="3" pageNumber="1453" type="holotype">holotype</typeStatus>
; male (
<collectionCode id="ED75EA8FBC1DFFF6FBD8FA61FF5FFA12" pageId="3" pageNumber="1453">CNAN- T01304</collectionCode>
)ı
<location id="8EBB2491BC1DFFF6FEE4FA5EFDFFFA12" LSID="urn:lsid:plazi:treatment:03CDC35CBC1DFFFCFE15FBD7FBC9FC15:8EBB2491BC1DFFF6FEE4FA5EFDFFFA12" box="[267,597,1457,1481]" municipality="Ni" name="Huautla de Jimenezi Li Nita" pageId="3" pageNumber="1453">Huautla de Jiménezı Li Nita</location>
caveı
<geoCoordinate id="EE50148DBC1DFFF6FD47FA5EFC8BFA12" box="[680,801,1457,1481]" pageId="3" pageNumber="1453">18.14770Nı</geoCoordinate>
<geoCoordinate id="EE50148DBC1DFFF6FCAFFA5EFC15FA12" box="[832,959,1457,1481]" pageId="3" pageNumber="1453">96.79844Wı</geoCoordinate>
leg.
<collectorName id="2691179CBC1DFFF6FBECFA5EFF46FA30" pageId="3" pageNumber="1453">O. Franckeı J. Cruzı</collectorName>
J. Mendozaı G. Contrerası
<date id="FFDA548ABC1DFFF6FDE5FA3CFD37FA30" box="[522,669,1491,1515]" pageId="3" pageNumber="1453" value="2014-04-12">
<collectingDate id="EF9EAD62BC1DFFF6FDE5FA3CFD37FA30" box="[522,669,1491,1515]" pageId="3" pageNumber="1453" value="2014-04-12">12 April 2014</collectingDate>
</date>
</materialsCitation>
; two females (CNAN-T01305)ı same locationı leg. J. Cruzı J. Mendozaı G. Contrerası D. Barralesı R. Monjarazı
<date id="FFDA548ABC1DFFF6FCA5FA19FC74F9D5" box="[842,990,1526,1550]" pageId="3" pageNumber="1453" value="2016-04-11">11 April 2016</date>
.
</paragraph>
<paragraph id="8BDB724ABC1DFFF6FF4FF9D1FE67F98C" blockId="3.[160,1156,1598,1727]" box="[160,461,1598,1623]" pageId="3" pageNumber="1453">
<emphasis id="B910AE58BC1DFFF6FF4FF9D1FE67F98C" bold="true" box="[160,461,1598,1623]" italics="true" pageId="3" pageNumber="1453">Other material examined</emphasis>
</paragraph>
<paragraph id="8BDB724ABC1DFFF0FF4FF98DFD2CFE42" blockId="3.[160,1156,1598,1727]" lastBlockId="5.[160,1156,142,409]" lastPageId="5" lastPageNumber="1455" pageId="3" pageNumber="1453">
Immature of
<taxonomicName id="4C6409C9BC1DFFF6FEDCF98EFE3CF9A1" authority="Cruz-López &amp; Francke, 2019" authorityName="Cruz-López &amp; Francke" authorityYear="2019" box="[307,406,1633,1658]" class="Arachnida" family="Stygnopsidae" genus="Huasteca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="1453" phylum="Arthropoda" rank="species" species="kardia" status="sp. nov.">
<emphasis id="B910AE58BC1DFFF6FEDCF98EFE3CF9A1" box="[307,406,1633,1658]" italics="true" pageId="3" pageNumber="1453">H. kardia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2231323BC1DFFF6FE72F98DFE52F9A1" box="[413,504,1634,1658]" pageId="3" pageNumber="1453" rank="species">sp. nov.</taxonomicNameLabel>
ı designated as
<typeStatus id="54DFCCE8BC1DFFF6FD4AF98DFCA3F9A1" box="[677,777,1634,1658]" pageId="3" pageNumber="1453" type="paratype">paratype</typeStatus>
of
<taxonomicName id="4C6409C9BC1DFFF6FCC0F98EFC07F9A1" authority="Goodnight and Goodnight" authorityName="Goodnight and Goodnight" authorityYear="1971" box="[815,941,1633,1658]" class="Arachnida" family="Stygnopsidae" genus="Karos" higherTaxonomySource="CoL" kingdom="Plantae" order="Opiliones" pageId="3" pageNumber="1453" phylum="Arthropoda" rank="species" species="gratiosus">
<emphasis id="B910AE58BC1DFFF6FCC0F98EFC07F9A1" box="[815,941,1633,1658]" italics="true" pageId="3" pageNumber="1453">K. gratiosus</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFF50FBBBC1DFFF6FC37F98DFEF0F946" author="Goodnight CJi Goodnight ML" pageId="3" pageNumber="1453" pagination="33 - 45" refId="ref4670" refString="Goodnight CJi Goodnight ML. 1971. Opilionids (Phalangida) of the family Phalangodidae from Mexican caves. Bull Assoc Mex Stud. 4: 33 - 45." type="journal article" year="1971">Goodnight and Goodnight 1971</bibRefCitation>
(AMNH)ı Millipede Caveı
<quantity id="4C9CDFAFBC1DFFF6FD6CF96AFD14F946" box="[643,702,1669,1693]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="1453" unit="km" value="8.0">8 km</quantity>
W of Huautlaı Oaxacaı Juneı
<date id="FFDA548ABC1DFFF6FBFAF96AFBF9F946" box="[1045,1107,1669,1693]" pageId="3" pageNumber="1453" value="1965-01">1965ı</date>
leg. W. Russell. Male of
<taxonomicName id="4C6409C9BC1DFFF6FE7FF949FDA6F964" baseAuthorityName="Goodnight and Goodnighti" baseAuthorityYear="1971" box="[400,524,1702,1727]" class="Arachnida" family="Stygnopsidae" genus="Huasteca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="1453" phylum="Arthropoda" rank="species" species="gratiosa">
<emphasis id="B910AE58BC1DFFF6FE7FF949FDA6F964" box="[400,524,1702,1727]" italics="true" pageId="3" pageNumber="1453">H. gratiosa</emphasis>
</taxonomicName>
(CNAN-SEM-Op0042)ı El Salitre caveı Xilitlaı San Luís Potosíı 21.38733Nı 98.98370Wı leg. O. Franckeı J. Cruzı G. Contrerası R. Monjarazı D. Barralesı
<date id="FFDA548ABC1BFFF0FEC7FF5EFE7FFF12" box="[296,469,177,201]" pageId="5" pageNumber="1455" value="2011-03-25">25 March 2011</date>
. Male of
<taxonomicName id="4C6409C9BC1BFFF0FDADFF5FFD06FF12" baseAuthorityName="Goodnight and Goodnighti" baseAuthorityYear="1971" box="[578,684,176,201]" class="Arachnida" family="Stygnopsidae" genus="Huasteca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="1455" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="B910AE58BC1BFFF0FDADFF5FFD06FF12" box="[578,684,176,201]" italics="true" pageId="5" pageNumber="1455">H. rugosa</emphasis>
</taxonomicName>
(CNAN-SEM-Op0207)ı Ojo de Agua caveı Tlilapanı Veracruzı 18.8065Nı 97.10148Wı leg. O. Franckeı J. Cruzı G. Contrerası
<date id="FFDA548ABC1BFFF0FF4FFF19FE2DFED5" box="[160,391,246,270]" pageId="5" pageNumber="1455" value="2016-09-19">19 September 2016</date>
. Female of
<taxonomicName id="4C6409C9BC1BFFF0FDFAFF1AFD2DFED5" authorityName="Cruz-Lopez and Franckei" authorityYear="2015" box="[533,647,245,270]" class="Arachnida" family="Stygnopsidae" genus="Huasteca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="1455" phylum="Arthropoda" rank="species" species="silhavyi">
<emphasis id="B910AE58BC1BFFF0FDFAFF1AFD2DFED5" box="[533,647,245,270]" italics="true" pageId="5" pageNumber="1455">H. silhavyi</emphasis>
</taxonomicName>
(CNAN-SEM-Op0220)ı Ojo de Agua Grande caveı Amatlán de los Reyesı Veracruzı 18.92651Nı 96.87593Wı leg. P. Sprouseı
<date id="FFDA548ABC1BFFF0FF4FFED3FEF9FE8F" box="[160,339,316,340]" pageId="5" pageNumber="1455" value="2007-03-13">13 March 2007</date>
. Female of
<taxonomicName id="4C6409C9BC1BFFF0FE0BFED4FDE6FE8F" authority="Cruz-López &amp; Francke, 2019" authorityName="Cruz-López &amp; Francke" authorityYear="2019" box="[484,588,315,340]" class="Arachnida" family="Stygnopsidae" genus="Huasteca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="1455" phylum="Arthropoda" rank="species" species="kardia" status="sp. nov.">
<emphasis id="B910AE58BC1BFFF0FE0BFED4FDE6FE8F" box="[484,588,315,340]" italics="true" pageId="5" pageNumber="1455">H. kardia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2231323BC1BFFF0FDB6FED3FD1FFE8F" box="[601,693,316,340]" pageId="5" pageNumber="1455" rank="species">sp. nov.</taxonomicNameLabel>
(CNAN-SEM-Op0280)ı Puente de Fierro caveı Huautla de Jiménezı Oaxacaı 18.15307Nı 96.85320Wı leg. R. Paredesı A. Valdezı C. Santibáñezı J. Cruzı
<date id="FFDA548ABC1BFFF0FE4FFE6EFD28FE42" box="[416,642,385,409]" pageId="5" pageNumber="1455" value="2009-09-20">20 September 2009</date>
.
</paragraph>
</subSubSection>
<caption id="DF1B22C2BC1AFFF1FF4FF9B7FBA7F957" ID-DOI="http://doi.org/10.5281/zenodo.3670141" ID-Zenodo-Dep="3670141" httpUri="https://zenodo.org/record/3670141/files/figure.png" pageId="4" pageNumber="1454" startId="4.[160,225,1624,1647]" targetBox="[163,1153,142,1592]" targetPageId="4">
<paragraph id="8BDB724ABC1AFFF1FF4FF9B7FBA7F957" blockId="4.[160,1155,1623,1676]" pageId="4" pageNumber="1454">
<emphasis id="B910AE58BC1AFFF1FF4FF9B7FF51F9B4" bold="true" box="[160,251,1624,1647]" pageId="4" pageNumber="1454">Figure 1.</emphasis>
Habitus of
<taxonomicName id="4C6409C9BC1AFFF1FE81F9B8FE6CF9B5" authorityName="Cruz-Lopez and Franckei" authorityYear="2015" box="[366,454,1623,1646]" class="Arachnida" family="Stygnopsidae" genus="Huasteca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="1454" phylum="Arthropoda" rank="genus">
<emphasis id="B910AE58BC1AFFF1FE81F9B8FE6CF9B5" box="[366,454,1623,1646]" italics="true" pageId="4" pageNumber="1454">Huasteca</emphasis>
</taxonomicName>
species. (a) male of
<taxonomicName id="4C6409C9BC1AFFF1FD7DF9B8FC95F9B5" baseAuthorityName="Goodnight and Goodnighti" baseAuthorityYear="1971" box="[658,831,1623,1646]" class="Arachnida" family="Stygnopsidae" genus="Huasteca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="1454" phylum="Arthropoda" rank="species" species="gratiosa">
<emphasis id="B910AE58BC1AFFF1FD7DF9B8FC95F9B5" box="[658,831,1623,1646]" italics="true" pageId="4" pageNumber="1454">Huasteca gratiosa</emphasis>
</taxonomicName>
. (b) male of
<taxonomicName id="4C6409C9BC1AFFF1FC50F9B8FBF5F9B5" baseAuthorityName="Goodnight and Goodnighti" baseAuthorityYear="1971" box="[959,1119,1623,1646]" class="Arachnida" family="Stygnopsidae" genus="Huasteca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="1454" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="B910AE58BC1AFFF1FC50F9B8FBF5F9B5" box="[959,1119,1623,1646]" italics="true" pageId="4" pageNumber="1454">Huasteca rugosa</emphasis>
</taxonomicName>
. (c) female of
<taxonomicName id="4C6409C9BC1AFFF1FEE8F99AFE04F957" authorityName="Cruz-Lopez and Franckei" authorityYear="2015" box="[263,430,1653,1676]" class="Arachnida" family="Stygnopsidae" genus="Huasteca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="1454" phylum="Arthropoda" rank="species" species="silhavyi">
<emphasis id="B910AE58BC1AFFF1FEE8F99AFE04F957" box="[263,430,1653,1676]" italics="true" pageId="4" pageNumber="1454">Huasteca silhavyi</emphasis>
</taxonomicName>
. (d) female of
<taxonomicName id="4C6409C9BC1AFFF1FDAAF99AFD4BF957" authority="Cruz-López &amp; Francke, 2019" authorityName="Cruz-López &amp; Francke" authorityYear="2019" box="[581,737,1653,1676]" class="Arachnida" family="Stygnopsidae" genus="Huasteca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="1454" phylum="Arthropoda" rank="species" species="kardia" status="sp. nov.">
<emphasis id="B910AE58BC1AFFF1FDAAF99AFD4BF957" box="[581,737,1653,1676]" italics="true" pageId="4" pageNumber="1454">Huasteca kardia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2231323BC1AFFF1FD05F99AFC93F957" box="[746,825,1653,1676]" pageId="4" pageNumber="1454" rank="species">sp. nov.</taxonomicNameLabel>
Scale bars = 0.5 cm.
</paragraph>
</caption>
<subSubSection id="C37E21C1BC1BFFF0FF4FFE3AFDBDFDEF" pageId="5" pageNumber="1455" type="etymology">
<paragraph id="8BDB724ABC1BFFF0FF4FFE3AFEB7FE35" blockId="5.[160,1155,469,564]" box="[160,285,469,494]" pageId="5" pageNumber="1455">
<emphasis id="B910AE58BC1BFFF0FF4FFE3AFEB7FE35" bold="true" box="[160,285,469,494]" italics="true" pageId="5" pageNumber="1455">Etymology</emphasis>
</paragraph>
<paragraph id="8BDB724ABC1BFFF0FF4FFE16FDBDFDEF" blockId="5.[160,1155,469,564]" pageId="5" pageNumber="1455">
Noun in apposition from the Greek Καρδία (
<taxonomicName id="4C6409C9BC1BFFF0FD61FE17FD79FDCA" authorityName="Cruz-López &amp; Francke" authorityYear="2019" box="[654,723,504,529]" class="Arachnida" family="Stygnopsidae" genus="Huasteca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="1455" phylum="Arthropoda" rank="species" species="kardia">
<emphasis id="B910AE58BC1BFFF0FD61FE17FD79FDCA" box="[654,723,504,529]" italics="true" pageId="5" pageNumber="1455">kardia</emphasis>
</taxonomicName>
)
<emphasis id="B910AE58BC1BFFF0FD0EFE16FD4DFDCA" box="[737,743,505,529]" italics="true" pageId="5" pageNumber="1455"></emphasis>
heart
<emphasis id="B910AE58BC1BFFF0FCCEFE16FC8DFDCA" box="[801,807,505,529]" italics="true" pageId="5" pageNumber="1455"></emphasis>
ı in allusion of the shape of the lateral channels on mesotergum.
</paragraph>
</subSubSection>
<subSubSection id="C37E21C1BC1BFFF0FF4FFD9FFC17FC82" pageId="5" pageNumber="1455" type="diagnosis">
<paragraph id="8BDB724ABC1BFFF0FF4FFD9FFEBEFD52" blockId="5.[160,1156,624,857]" box="[160,276,624,649]" pageId="5" pageNumber="1455">
<emphasis id="B910AE58BC1BFFF0FF4FFD9FFEBEFD52" bold="true" box="[160,276,624,649]" italics="true" pageId="5" pageNumber="1455">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BDB724ABC1BFFF0FF4FFD7CFC17FC82" blockId="5.[160,1156,624,857]" pageId="5" pageNumber="1455">
<taxonomicName id="4C6409C9BC1BFFF0FF4FFD7CFEFAFD77" authority="Cruz-López &amp; Francke, 2019" authorityName="Cruz-López &amp; Francke" authorityYear="2019" box="[160,336,659,684]" class="Arachnida" family="Stygnopsidae" genus="Huasteca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="1455" phylum="Arthropoda" rank="species" species="kardia" status="sp. nov.">
<emphasis id="B910AE58BC1BFFF0FF4FFD7CFEFAFD77" box="[160,336,659,684]" italics="true" pageId="5" pageNumber="1455">Huasteca kardia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2231323BC1BFFF0FEB9FD7BFE07FD77" box="[342,429,660,684]" pageId="5" pageNumber="1455" rank="species">sp. nov.</taxonomicNameLabel>
is easily recognisable from the remaining members of the genus by the extreme modification of the lateral channels of scutumı running along the central portion of mesotergumı over areas II and III forming a pair of heart-shaped ditches (
<figureCitation id="135F6ECFBC1BFFF0FF48FD13FEA9FCCF" box="[167,259,764,788]" captionStart="Figure 1" captionStartId="4.[160,225,1624,1647]" captionTargetBox="[163,1153,142,1592]" captionTargetId="figure@4.[163,1153,142,1592]" captionTargetPageId="4" captionText="Figure 1. Habitus of Huasteca species. (a) male of Huasteca gratiosa. (b) male of Huasteca rugosa. (c) female of Huasteca silhavyi. (d) female of Huasteca kardia sp. nov. Scale bars = 0.5 cm." figureDoi="http://doi.org/10.5281/zenodo.3670141" httpUri="https://zenodo.org/record/3670141/files/figure.png" pageId="5" pageNumber="1455">Figure 1</figureCitation>
(d)ı 6(dıe)). In additionı the dorsal ornamentation of scutum with one pair of paramedian humps on mesotergal areas I and II eachı and the extremely swollen patella II in males are exclusive characters of this species (
<figureCitation id="135F6ECFBC1BFFF0FD0AFCAEFC95FC82" box="[741,831,833,857]" captionStart="Figure 1" captionStartId="4.[160,225,1624,1647]" captionTargetBox="[163,1153,142,1592]" captionTargetId="figure@4.[163,1153,142,1592]" captionTargetPageId="4" captionText="Figure 1. Habitus of Huasteca species. (a) male of Huasteca gratiosa. (b) male of Huasteca rugosa. (c) female of Huasteca silhavyi. (d) female of Huasteca kardia sp. nov. Scale bars = 0.5 cm." figureDoi="http://doi.org/10.5281/zenodo.3670141" httpUri="https://zenodo.org/record/3670141/files/figure.png" pageId="5" pageNumber="1455">Figure 1</figureCitation>
(d)ı 4(gıh)).
</paragraph>
</subSubSection>
<subSubSection id="C37E21C1BC1BFFF3FF4FFC7AFEECF983" lastPageId="6" lastPageNumber="1456" pageId="5" pageNumber="1455" type="description">
<paragraph id="8BDB724ABC1BFFF0FF4FFC7AFE8CFC75" blockId="5.[160,1156,917,1670]" box="[160,294,917,942]" pageId="5" pageNumber="1455">
<emphasis id="B910AE58BC1BFFF0FF4FFC7AFE8CFC75" bold="true" box="[160,294,917,942]" italics="true" pageId="5" pageNumber="1455">Description</emphasis>
</paragraph>
<paragraph id="8BDB724ABC1BFFF0FF4FFC57FDC7F95D" blockId="5.[160,1156,917,1670]" pageId="5" pageNumber="1455">
<emphasis id="B910AE58BC1BFFF0FF4FFC57FE91FC0A" box="[160,315,952,977]" italics="true" pageId="5" pageNumber="1455">
Male
<typeStatus id="54DFCCE8BC1BFFF0FF32FC57FE91FC0A" box="[221,315,952,977]" pageId="5" pageNumber="1455" type="holotype">holotype</typeStatus>
</emphasis>
. Scutum zeta type (ζ) with the lateral projections on opisthosomal tergite 1 very elongateı with one pair of paramedian humps on mesotergal areas I and II eachı these humps are connected by the lateral channels which run over the mesotergumı forming two heart-shaped ditches on each side of scutum. Posterior corners of scutum and free tergites with lighter and rounded projections. Posterior margin of scutum and free tergites with a transverse row of small tubercles (
<figureCitation id="135F6ECFBC1BFFF0FCF4FB89FCDDFBA5" box="[795,887,1126,1150]" captionStart="Figure 1" captionStartId="4.[160,225,1624,1647]" captionTargetBox="[163,1153,142,1592]" captionTargetId="figure@4.[163,1153,142,1592]" captionTargetPageId="4" captionText="Figure 1. Habitus of Huasteca species. (a) male of Huasteca gratiosa. (b) male of Huasteca rugosa. (c) female of Huasteca silhavyi. (d) female of Huasteca kardia sp. nov. Scale bars = 0.5 cm." figureDoi="http://doi.org/10.5281/zenodo.3670141" httpUri="https://zenodo.org/record/3670141/files/figure.png" pageId="5" pageNumber="1455">Figure 1</figureCitation>
(d)ı 2(a)ı 6(eıd)). Venter covered by small tuberclesı lateral margins of stigmatic area poorly markedı genital operculum sub-triangular (
<figureCitation id="135F6ECFBC1BFFF0FE38FB43FD93FB1F" box="[471,569,1196,1220]" captionStart="Figure 2" captionStartId="6.[160,225,1380,1403]" captionTargetBox="[163,1153,142,1349]" captionTargetId="figure@6.[163,1153,142,1349]" captionTargetPageId="6" captionText="Figure 2. Habitus and venter of Huasteca kardia sp. nov. (a) habitus holotype. (b) habitus female paratype. (c) venter holotype. (d) venter female paratype. Dotted line indicates swollen portions of leg II. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3670143" httpUri="https://zenodo.org/record/3670143/files/figure.png" pageId="5" pageNumber="1455">Figure 2</figureCitation>
(c)). Chelicerae smallı with the bulla rounded and smoothı cheliceral fingers thinı fixed finger with three contiguous teethı cheliceral comb with 18 lamellae (
<figureCitation id="135F6ECFBC1BFFF0FE88FB1EFE69FAD2" box="[359,451,1265,1289]" captionStart="Figure 3" captionStartId="7.[160,225,1458,1481]" captionTargetBox="[163,1153,142,1426]" captionTargetId="figure@7.[163,1153,142,1426]" captionTargetPageId="7" captionText="Figure 3. Chelicera and pedipalp of Huasteca kardia sp. nov. (a) chelicera frontal, male. (b) chelicera mesal, male. (c) cheliceral comb, male. (d) chelicera ectal, male. (e) pedipalp mesal. Scale bars = a and b: 150 µm, c: 50 µm, d and e: 250 µm." figureDoi="http://doi.org/10.5281/zenodo.3670145" httpUri="https://zenodo.org/record/3670145/files/figure.png" pageId="5" pageNumber="1455">Figure 3</figureCitation>
(a-d)). Pedipalp strongly armedı femur with a ventral row of six setiferous tubercles plus a mesoapical oneı patella with two mesal setiferous tuberclesı tibia with IiIi (1 = 3&gt; 2 = 4) and II (1 = 2) tubercles on mesal and ectal sidesı respectivelyı and tarsus with III (1&gt; 2&gt; 3) on both sides (
<figureCitation id="135F6ECFBC1BFFF0FD41FAB6FCA6FAAA" box="[686,780,1369,1393]" captionStart="Figure 3" captionStartId="7.[160,225,1458,1481]" captionTargetBox="[163,1153,142,1426]" captionTargetId="figure@7.[163,1153,142,1426]" captionTargetPageId="7" captionText="Figure 3. Chelicera and pedipalp of Huasteca kardia sp. nov. (a) chelicera frontal, male. (b) chelicera mesal, male. (c) cheliceral comb, male. (d) chelicera ectal, male. (e) pedipalp mesal. Scale bars = a and b: 150 µm, c: 50 µm, d and e: 250 µm." figureDoi="http://doi.org/10.5281/zenodo.3670145" httpUri="https://zenodo.org/record/3670145/files/figure.png" pageId="5" pageNumber="1455">Figure 3</figureCitation>
(e)). Legs are slender covered by small tubercles and setae; the distalmost half of femur II slightly swollen and lighter on prolateral sideı patella II swollen and lighter prolaterallyı tibia II without modifications. Tarsal count: 4(2):8(3):6:6ı basitarsomeres I of legs III and IV approximately one-third of the length of their respective tarsus (
<figureCitation id="135F6ECFBC1BFFF0FDB9FA0BFD1FFA27" box="[598,693,1508,1532]" captionStart="Figure 2" captionStartId="6.[160,225,1380,1403]" captionTargetBox="[163,1153,142,1349]" captionTargetId="figure@6.[163,1153,142,1349]" captionTargetPageId="6" captionText="Figure 2. Habitus and venter of Huasteca kardia sp. nov. (a) habitus holotype. (b) habitus female paratype. (c) venter holotype. (d) venter female paratype. Dotted line indicates swollen portions of leg II. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3670143" httpUri="https://zenodo.org/record/3670143/files/figure.png" pageId="5" pageNumber="1455">Figure 2</figureCitation>
(a)ı 4(gıh)). Penis with the flimsy lamina convex apicallyı two pairs of macrosetae C and A forming a lateral rowı two pairs of macrosetae D on the base of follisı small macrosetae E1 and E2 at the same level of C2 and A1ı respectivelyı additional macrosetae E slightly below A2; lateral projection of follis lobed and apically blunt (
<figureCitation id="135F6ECFBC1BFFF0FDECF981FDCAF95D" box="[515,608,1646,1670]" captionStart="Figure 5" captionStartId="9.[160,225,769,792]" captionTargetBox="[163,1153,142,738]" captionTargetId="figure@9.[163,1153,142,738]" captionTargetPageId="9" captionText="Figure 5. Male genitalia of Huasteca kardia sp. nov. (a) dorsal. (b) lateral. (c) ventral. Macrosetae A-E indicated in the figures, adE = additional macrosetae E. Scale bars = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3670149" httpUri="https://zenodo.org/record/3670149/files/figure.png" pageId="5" pageNumber="1455">Figure 5</figureCitation>
).
</paragraph>
<caption id="DF1B22C2BC18FFF3FF4FFA8BFE1CFA6D" ID-DOI="http://doi.org/10.5281/zenodo.3670143" ID-Zenodo-Dep="3670143" httpUri="https://zenodo.org/record/3670143/files/figure.png" pageId="6" pageNumber="1456" startId="6.[160,225,1380,1403]" targetBox="[163,1153,142,1349]" targetPageId="6">
<paragraph id="8BDB724ABC18FFF3FF4FFA8BFE1CFA6D" blockId="6.[160,1156,1380,1462]" pageId="6" pageNumber="1456">
<emphasis id="B910AE58BC18FFF3FF4FFA8BFF57FAA0" bold="true" box="[160,253,1380,1403]" pageId="6" pageNumber="1456">Figure 2.</emphasis>
Habitus and venter of
<taxonomicName id="4C6409C9BC18FFF3FE00FA8BFD26FAA0" authority="Cruz-López &amp; Francke, 2019" authorityName="Cruz-López &amp; Francke" authorityYear="2019" box="[495,652,1380,1403]" class="Arachnida" family="Stygnopsidae" genus="Huasteca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="1456" phylum="Arthropoda" rank="species" species="kardia" status="sp. nov.">
<emphasis id="B910AE58BC18FFF3FE00FA8BFD26FAA0" box="[495,652,1380,1403]" italics="true" pageId="6" pageNumber="1456">Huasteca kardia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2231323BC18FFF3FD79FA8BFD4CFAA0" box="[662,742,1380,1403]" pageId="6" pageNumber="1456" rank="species">sp. nov.</taxonomicNameLabel>
(a) habitus holotype. (b) habitus female paratype. (c) venter holotype. (d) venter female paratype. Dotted line indicates swollen portions of leg II. Scale bars = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BDB724ABC18FFF3FF4FFA15FF5CF9C8" blockId="6.[160,1156,1530,1624]" box="[160,246,1530,1555]" pageId="6" pageNumber="1456">
<emphasis id="B910AE58BC18FFF3FF4FFA15FF5CF9C8" bold="true" box="[160,246,1530,1555]" italics="true" pageId="6" pageNumber="1456">Female</emphasis>
</paragraph>
<paragraph id="8BDB724ABC18FFF3FF4FF9F2FEECF983" blockId="6.[160,1156,1530,1624]" pageId="6" pageNumber="1456">
Similar to the maleı but without the modifications on legs II and stigmatic area broader (
<figureCitation id="135F6ECFBC18FFF3FF48F9AFFEAFF983" box="[167,261,1600,1624]" captionStart="Figure 2" captionStartId="6.[160,225,1380,1403]" captionTargetBox="[163,1153,142,1349]" captionTargetId="figure@6.[163,1153,142,1349]" captionTargetPageId="6" captionText="Figure 2. Habitus and venter of Huasteca kardia sp. nov. (a) habitus holotype. (b) habitus female paratype. (c) venter holotype. (d) venter female paratype. Dotted line indicates swollen portions of leg II. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3670143" httpUri="https://zenodo.org/record/3670143/files/figure.png" pageId="6" pageNumber="1456">Figure 2</figureCitation>
(bıd)).
</paragraph>
</subSubSection>
<caption id="DF1B22C2BC19FFF2FF4FFA5DFDC1F9DF" ID-DOI="http://doi.org/10.5281/zenodo.3670145" ID-Zenodo-Dep="3670145" httpUri="https://zenodo.org/record/3670145/files/figure.png" pageId="7" pageNumber="1457" startId="7.[160,225,1458,1481]" targetBox="[163,1153,142,1426]" targetPageId="7">
<paragraph id="8BDB724ABC19FFF2FF4FFA5DFDC1F9DF" blockId="7.[160,1155,1457,1540]" pageId="7" pageNumber="1457">
<emphasis id="B910AE58BC19FFF2FF4FFA5DFF51FA12" bold="true" box="[160,251,1458,1481]" pageId="7" pageNumber="1457">Figure 3.</emphasis>
Chelicera and pedipalp of
<taxonomicName id="4C6409C9BC19FFF2FDE5FA5EFD0FFA13" authority="Cruz-López &amp; Francke, 2019" authorityName="Cruz-López &amp; Francke" authorityYear="2019" box="[522,677,1457,1480]" class="Arachnida" family="Stygnopsidae" genus="Huasteca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="1457" phylum="Arthropoda" rank="species" species="kardia" status="sp. nov.">
<emphasis id="B910AE58BC19FFF2FDE5FA5EFD0FFA13" box="[522,677,1457,1480]" italics="true" pageId="7" pageNumber="1457">Huasteca kardia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2231323BC19FFF2FD43FA5DFD50FA12" box="[684,762,1458,1481]" pageId="7" pageNumber="1457" rank="species">sp. nov.</taxonomicNameLabel>
(a) chelicera frontal, male. (b) chelicera mesal, male. (c) cheliceral comb, male. (d) chelicera ectal, male. (e) pedipalp mesal. Scale bars = a and b: 150 µm, c: 50 µm, d and e: 250 µm.
</paragraph>
</caption>
<subSubSection id="C37E21C1BC19FFFDFF4FF9A6FE50F965" lastPageId="8" lastPageNumber="1458" pageId="7" pageNumber="1457" type="biology_ecology">
<paragraph id="8BDB724ABC19FFF2FF4FF9A6FEFEF9B9" blockId="7.[160,1155,1609,1703]" box="[160,340,1609,1634]" pageId="7" pageNumber="1457">
<emphasis id="B910AE58BC19FFF2FF4FF9A6FEFEF9B9" bold="true" box="[160,340,1609,1634]" italics="true" pageId="7" pageNumber="1457">Natural history</emphasis>
</paragraph>
<paragraph id="8BDB724ABC19FFFDFF4FF983FE50F965" blockId="7.[160,1155,1609,1703]" lastBlockId="8.[160,1155,1668,1727]" lastPageId="8" lastPageNumber="1458" pageId="7" pageNumber="1457">
All specimens were found walking alone on the floor and walls in the caves. In the same cavesı specimens of undescribed harvestmenı species of
<taxonomicName id="4C6409C9BC19FFF2FD51F961FC81F97C" authorityName="Sorensen" authorityYear="1902" box="[702,811,1678,1703]" class="Arachnida" family="Stygnopsidae" genus="Stygnopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="1457" phylum="Arthropoda" rank="genus">
<emphasis id="B910AE58BC19FFF2FD51F961FC81F97C" box="[702,811,1678,1703]" italics="true" pageId="7" pageNumber="1457">Stygnopsis</emphasis>
</taxonomicName>
sp. (
<taxonomicName id="4C6409C9BC19FFF2FCB2F960FC5EF97C" baseAuthorityName="Cruz-Lopez" baseAuthorityYear="2018" box="[861,1012,1679,1703]" class="Arachnida" family="Stygnopsidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="1457" phylum="Arthropoda" rank="family">Stygnopsidae</taxonomicName>
)ı
<taxonomicName id="4C6409C9BC19FFF2FBEBF961FB29F97C" box="[1028,1155,1678,1703]" class="Arachnida" family="Stygnommatidae" genus="Stygnomma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="1457" phylum="Arthropoda" rank="genus">
<emphasis id="B910AE58BC19FFF2FBEBF961FB29F97C" box="[1028,1155,1678,1703]" italics="true" pageId="7" pageNumber="1457">Stygnomma</emphasis>
</taxonomicName>
sp. (
<taxonomicName id="4C6409C9BC16FFFDFF22F96BFE23F947" authorityName="Roewer" authorityYear="1923" box="[205,393,1668,1692]" class="Arachnida" family="Stygnommatidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="1458" phylum="Arthropoda" rank="family">Stygnommatidae</taxonomicName>
) and a species of
<taxonomicName id="4C6409C9BC16FFFDFDB8F96BFC33F947" box="[599,921,1668,1692]" class="Arachnida" family="Sclerosomatidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="1458" phylum="Arthropoda" rank="subFamily" subFamily="Gagrellinae">Gagrellinae (Sclerosomatidae)</taxonomicName>
were found living in sympatry with
<taxonomicName id="4C6409C9BC16FFFDFEAEF949FE35F964" authority="Cruz-López &amp; Francke, 2019" authorityName="Cruz-López &amp; Francke" authorityYear="2019" box="[321,415,1702,1727]" class="Arachnida" family="Stygnopsidae" genus="Huasteca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="1458" phylum="Arthropoda" rank="species" species="kardia" status="sp. nov.">
<emphasis id="B910AE58BC16FFFDFEAEF949FE35F964" box="[321,415,1702,1727]" italics="true" pageId="8" pageNumber="1458">H. kardia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2231323BC16FFFDFE49F949FE50F965" box="[422,506,1702,1726]" pageId="8" pageNumber="1458" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<caption id="DF1B22C2BC16FFFDFF4FFA52FC18F992" ID-DOI="http://doi.org/10.5281/zenodo.3670147" ID-Zenodo-Dep="3670147" httpUri="https://zenodo.org/record/3670147/files/figure.png" pageId="8" pageNumber="1458" startId="8.[160,225,1469,1492]" targetBox="[163,1153,142,1437]" targetPageId="8">
<paragraph id="8BDB724ABC16FFFDFF4FFA52FC18F992" blockId="8.[160,1156,1468,1609]" pageId="8" pageNumber="1458">
<emphasis id="B910AE58BC16FFFDFF4FFA52FF50FA0F" bold="true" box="[160,250,1469,1492]" pageId="8" pageNumber="1458">Figure 4.</emphasis>
Leg II of males of
<taxonomicName id="4C6409C9BC16FFFDFE5FFA53FDA2FA08" authorityName="Cruz-Lopez and Franckei" authorityYear="2015" box="[432,520,1468,1491]" class="Arachnida" family="Stygnopsidae" genus="Huasteca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="1458" phylum="Arthropoda" rank="genus">
<emphasis id="B910AE58BC16FFFDFE5FFA53FDA2FA08" box="[432,520,1468,1491]" italics="true" pageId="8" pageNumber="1458">Huasteca</emphasis>
</taxonomicName>
. (a) apical femur, patella and basal tibia of
<taxonomicName id="4C6409C9BC16FFFDFC5EFA53FBF7FA08" baseAuthorityName="Goodnight and Goodnighti" baseAuthorityYear="1971" box="[945,1117,1468,1491]" class="Arachnida" family="Stygnopsidae" genus="Huasteca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="1458" phylum="Arthropoda" rank="species" species="gratiosa">
<emphasis id="B910AE58BC16FFFDFC5EFA53FBF7FA08" box="[945,1117,1468,1491]" italics="true" pageId="8" pageNumber="1458">Huasteca gratiosa</emphasis>
</taxonomicName>
. (b) basal tibia of
<taxonomicName id="4C6409C9BC16FFFDFEC7FA35FE2FFA2A" baseAuthorityName="Goodnight and Goodnighti" baseAuthorityYear="1971" box="[296,389,1498,1521]" class="Arachnida" family="Stygnopsidae" genus="Huasteca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="1458" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="B910AE58BC16FFFDFEC7FA35FE2FFA2A" box="[296,389,1498,1521]" italics="true" pageId="8" pageNumber="1458">H. rugosa</emphasis>
</taxonomicName>
. (c) apical femur, patella and basal tibia of
<taxonomicName id="4C6409C9BC16FFFDFCD7FA35FC72FA2A" baseAuthorityName="Goodnight and Goodnighti" baseAuthorityYear="1971" box="[824,984,1498,1521]" class="Arachnida" family="Stygnopsidae" genus="Huasteca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="1458" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="B910AE58BC16FFFDFCD7FA35FC72FA2A" box="[824,984,1498,1521]" italics="true" pageId="8" pageNumber="1458">Huasteca rugosa</emphasis>
</taxonomicName>
. (d) apical femur of
<taxonomicName id="4C6409C9BC16FFFDFF54FA18FEBDF9D5" baseAuthorityName="Goodnight and Goodnighti" baseAuthorityYear="1971" box="[187,279,1527,1550]" class="Arachnida" family="Stygnopsidae" genus="Huasteca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="1458" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="B910AE58BC16FFFDFF54FA18FEBDF9D5" box="[187,279,1527,1550]" italics="true" pageId="8" pageNumber="1458">H. rugosa</emphasis>
</taxonomicName>
. (e) apical femur, patella and basal tibia of
<taxonomicName id="4C6409C9BC16FFFDFD27FA18FCC4F9D5" authorityName="Cruz-Lopez and Franckei" authorityYear="2015" box="[712,878,1527,1550]" class="Arachnida" family="Stygnopsidae" genus="Huasteca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="1458" phylum="Arthropoda" rank="species" species="silhavyi">
<emphasis id="B910AE58BC16FFFDFD27FA18FCC4F9D5" box="[712,878,1527,1550]" italics="true" pageId="8" pageNumber="1458">Huasteca silhavyi</emphasis>
</taxonomicName>
. (f) patella of
<taxonomicName id="4C6409C9BC16FFFDFC15FA18FBF6F9D5" authorityName="Cruz-Lopez and Franckei" authorityYear="2015" box="[1018,1116,1527,1550]" class="Arachnida" family="Stygnopsidae" genus="Huasteca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="1458" phylum="Arthropoda" rank="species" species="silhavyi">
<emphasis id="B910AE58BC16FFFDFC15FA18FBF6F9D5" box="[1018,1116,1527,1550]" italics="true" pageId="8" pageNumber="1458">H. silhavyi</emphasis>
</taxonomicName>
. (g) apical femur, patella and basal tibia of
<taxonomicName id="4C6409C9BC16FFFDFDCBF9FBFD14F9F0" authority="Cruz-López &amp; Francke, 2019" authorityName="Cruz-López &amp; Francke" authorityYear="2019" box="[548,702,1556,1579]" class="Arachnida" family="Stygnopsidae" genus="Huasteca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="1458" phylum="Arthropoda" rank="species" species="kardia" status="sp. nov.">
<emphasis id="B910AE58BC16FFFDFDCBF9FBFD14F9F0" box="[548,702,1556,1579]" italics="true" pageId="8" pageNumber="1458">Huasteca kardia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2231323BC16FFFDFD29F9FAFCB9F9F7" box="[710,787,1557,1580]" pageId="8" pageNumber="1458" rank="species">sp. nov.</taxonomicNameLabel>
(h) patella of
<taxonomicName id="4C6409C9BC16FFFDFC70F9FBFC5CF9F0" authority="Cruz-López &amp; Francke, 2019" authorityName="Cruz-López &amp; Francke" authorityYear="2019" box="[927,1014,1556,1579]" class="Arachnida" family="Stygnopsidae" genus="Huasteca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="1458" phylum="Arthropoda" rank="species" species="kardia" status="sp. nov.">
<emphasis id="B910AE58BC16FFFDFC70F9FBFC5CF9F0" box="[927,1014,1556,1579]" italics="true" pageId="8" pageNumber="1458">H. kardia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2231323BC16FFFDFC12F9FAFBE1F9F7" box="[1021,1099,1557,1580]" pageId="8" pageNumber="1458" rank="species">sp. nov.</taxonomicNameLabel>
Scale bars = a: 200 µm, c, e and g: 250 µm, b and d: 50 µm, h: 150 µm, f: 100 µm.
</paragraph>
</caption>
<caption id="DF1B22C2BC17FFFCFF4FFCEEFC67FCED" ID-DOI="http://doi.org/10.5281/zenodo.3670149" ID-Zenodo-Dep="3670149" httpUri="https://zenodo.org/record/3670149/files/figure.png" pageId="9" pageNumber="1459" startId="9.[160,225,769,792]" targetBox="[163,1153,142,738]" targetPageId="9">
<paragraph id="8BDB724ABC17FFFCFF4FFCEEFC67FCED" blockId="9.[160,1155,768,822]" pageId="9" pageNumber="1459">
<emphasis id="B910AE58BC17FFFCFF4FFCEEFF57FCC3" bold="true" box="[160,253,769,792]" pageId="9" pageNumber="1459">Figure 5.</emphasis>
Male genitalia of
<taxonomicName id="4C6409C9BC17FFFCFE52FCEFFDF1FCCC" authority="Cruz-López &amp; Francke, 2019" authorityName="Cruz-López &amp; Francke" authorityYear="2019" box="[445,603,768,791]" class="Arachnida" family="Stygnopsidae" genus="Huasteca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="1459" phylum="Arthropoda" rank="species" species="kardia" status="sp. nov.">
<emphasis id="B910AE58BC17FFFCFE52FCEFFDF1FCCC" box="[445,603,768,791]" italics="true" pageId="9" pageNumber="1459">Huasteca kardia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2231323BC17FFFCFD8AFCEEFD1CFCC3" box="[613,694,769,792]" pageId="9" pageNumber="1459" rank="species">sp. nov.</taxonomicNameLabel>
(a) dorsal. (b) lateral. (c) ventral. Macrosetae A-E indicated in the figures, adE = additional macrosetae E. Scale bars = 50 µm.
</paragraph>
</caption>
<subSubSection id="C37E21C1BC17FFFCFF4FFC9FFBC9FC15" pageId="9" pageNumber="1459" type="distribution">
<paragraph id="8BDB724ABC17FFFCFF4FFC9FFE86FC52" blockId="9.[160,1155,880,974]" box="[160,300,880,905]" pageId="9" pageNumber="1459">
<emphasis id="B910AE58BC17FFFCFF4FFC9FFE86FC52" bold="true" box="[160,300,880,905]" italics="true" pageId="9" pageNumber="1459">Distribution</emphasis>
</paragraph>
<paragraph id="8BDB724ABC17FFFCFF4FFC7CFBC9FC15" blockId="9.[160,1155,880,974]" pageId="9" pageNumber="1459">
This species is known from three different caves of the Huautla cave systemı although it is probably found in other caves of this extremely large and deep system (
<figureCitation id="135F6ECFBC17FFFCFC16FC59FBFDFC15" box="[1017,1111,950,974]" captionStart="Figure 7" captionStartId="11.[160,225,792,815]" captionTargetBox="[163,1153,142,760]" captionTargetId="figure@11.[163,1153,142,760]" captionTargetPageId="11" captionText="Figure 7. Distribution map of the species of Huasteca. Circles = Huasteca gratiosa, star = Huasteca rugosa, square = Huasteca silhavyi and triangles = Huasteca kardia sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3670153" httpUri="https://zenodo.org/record/3670153/files/figure.png" pageId="9" pageNumber="1459">Figure 7</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>