378 lines
49 KiB
XML
378 lines
49 KiB
XML
<document id="C160E6416AE2D3D8B6C22696EA89D456" ID-DOI="10.11646/zootaxa.5026.4.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="5300792" ID-ZooBank="E712BC8E-3E75-422C-8587-69ED17B6CDD5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630041845836" checkinUser="plazi" docAuthor="Fomichev, Alexander A." docDate="2021" docId="D220E4309B18A400BFED8FACFA1BFE7D" docLanguage="en" docName="zootaxa.5026.4.7.pdf" docOrigin="Zootaxa 5026 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Acantholycosa levinae Marusik, Azarkina & Koponen 2004" docType="treatment" docVersion="4" lastPageNumber="578" masterDocId="2E199C489B12A40BBF7A8D7DFF9FFFAC" masterDocTitle="New data on the wolf spiders from theAcantholycosa-complex (Araneae: Lycosidae) from the South Siberia" masterLastPageNumber="585" masterPageNumber="567" pageNumber="577" updateTime="1699139093203" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="8575B447B1469C8A382EE694195A575C">New data on the wolf spiders from theAcantholycosa-complex (Araneae: Lycosidae) from the South Siberia</mods:title>
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<mods:namePart id="BF2A0CBFE725D9081E1C01B5CD0AA5D3">Fomichev, Alexander A.</mods:namePart>
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<mods:affiliation id="4EF0DD6BFDA72A25ADBF8EF48FB2B330">Altai State University, Lenina Pr., 61, Barnaul, RF- 656049, Russia. & Tomsk State University, Lenina Pr., 36, Tomsk, RF- 634050, Russia.</mods:affiliation>
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<mods:title id="3FACFFC1F08D471A03E89BB023A50302">Zootaxa</mods:title>
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<mods:date id="CEFE94F1293EF4CCC13176C0CFE6CD68">2021</mods:date>
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<mods:number id="83072BEE720369C22E6EF7DE1DE18E35">2021-08-27</mods:number>
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<taxonomicName id="9D892EA59B18A401BFED8FACFCE0FD40" ID-CoL="64GDM" authority="Marusik, Azarkina & Koponen, 2004" authorityName="Marusik, Azarkina & Koponen" authorityYear="2004" box="[151,895,721,748]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="levinae">
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<emphasis id="68FD89349B18A401BFED8FACFE39FD40" bold="true" box="[151,422,721,748]" italics="true" pageId="10" pageNumber="577">Acantholycosa levinae</emphasis>
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<bibRefCitation id="3E1828D79B18A401BED48FACFCE0FD40" author="Marusik, Y. M. & Azarkina, G. N. & Koponen, S." box="[430,895,721,748]" pageId="10" pageNumber="577" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. & Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">Marusik, Azarkina & Koponen, 2004</bibRefCitation>
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</emphasis>
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<subSubSection id="129306AD9B18A401BFED8F8BFD03FCBD" box="[151,668,758,785]" pageId="10" pageNumber="577" type="description">
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<paragraph id="5A3655269B18A401BFED8F8BFD03FCBD" blockId="10.[151,895,721,785]" box="[151,668,758,785]" pageId="10" pageNumber="577">
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<figureCitation id="C2B249A39B18A401BFED8F8BFF64FCBD" box="[151,251,758,785]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1832,1856]" captionTargetBox="[157,1436,189,1799]" captionTargetId="figure-33@2.[151,1436,181,1809]" captionTargetPageId="2" captionText="FIGURES 1–9. General appearance of Acantholycosa sayanensis (1–2), A. kulikovi sp. n. (3), A. levinae (4–5), A. sergeevi sp. n. (6), Mongolicosa lizae sp. n. (8–9) and leg I of A. sergeevi sp. n. (7). 1, 4, 6, 8—male, dorsal; 2, 3, 5, 9—female, dorsal; 7—leg I of male, prolateral. Scale bars=2 mm." figureDoi="http://doi.org/10.5281/zenodo.5300794" httpUri="https://zenodo.org/record/5300794/files/figure.png" pageId="10" pageNumber="577">Figs 4–5</figureCitation>
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,
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, 28–31, 36–37, 66–68, 75–76
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<emphasis id="68FD89349B18A401BFED8E46FF63FCFE" box="[151,252,826,850]" italics="true" pageId="10" pageNumber="577">A. levinae</emphasis>
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, f. 61–65 (
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<emphasis id="68FD89349B18A401BFED8E24FF63FCDD" box="[151,252,857,881]" italics="true" pageId="10" pageNumber="577">A. levinae</emphasis>
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—
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<bibRefCitation id="3E1828D79B18A401BE668E24FDC8FCDD" author="Marusik, Y. M. & Omelko, M. M." box="[284,599,857,881]" pageId="10" pageNumber="577" pagination="597 - 600" refId="ref10588" refString="Marusik, Y. M. & Omelko, M. M. (2017) A new species of Acantholycosa baltoroi - group (Araneae: Lycosidae, Pardosinae) from the Russian Far East. Zootaxa, 4232 (4), 597 - 600. https: // doi. org / 10.11646 / zootaxa. 4232.4.12" type="journal article" year="2017">Marusik & Omelko, 2017: 597</bibRefCitation>
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, f. 11 (
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).
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<emphasis id="68FD89349B18A401BFED8EE2FE1CFC15" bold="true" box="[151,387,927,953]" pageId="10" pageNumber="577">Material examined.</emphasis>
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, scree and lichen pillows,
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,
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;
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(
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, watershed of
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and
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,
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<geoCoordinate id="3FBD33E19B18A401BEF88976FD8CFB8A" box="[386,531,1035,1062]" degrees="85" direction="east" minutes="51.096" orientation="longitude" pageId="10" pageNumber="577" precision="1" value="85.8516">85°51.096’E</geoCoordinate>
|
||
, scree,
|
||
<quantity id="9D71F8C39B18A401BD1C8976FD23FB8A" box="[614,700,1035,1062]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="10" pageNumber="577" unit="m" value="2400.0">
|
||
<elevation id="D1A4B2159B18A401BD1C8976FD23FB8A" box="[614,700,1035,1062]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="10" pageNumber="577" unit="m" value="2400.0">2400 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="2E3773E69B18A401BDB28976FCD6FB89" box="[712,841,1035,1061]" pageId="10" pageNumber="577" value="2019-07-26">
|
||
<collectingDate id="3E738A0E9B18A401BDB28976FCD6FB89" box="[712,841,1035,1061]" pageId="10" pageNumber="577" value="2019-07-26">26.07.2019</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="F77C30F09B18A401BC288976FB9EFB89" box="[850,1025,1035,1061]" pageId="10" pageNumber="577">A.A. Fomichev</collectorName>
|
||
,
|
||
<collectorName id="F77C30F09B18A401BB718971FB61FB89" box="[1035,1278,1035,1062]" pageId="10" pageNumber="577">Yu.V. Dyachkov, E.A</collectorName>
|
||
.
|
||
<collectorName id="F77C30F09B18A401BA738971FAF0FB8A" box="[1289,1391,1036,1062]" pageId="10" pageNumber="577">Nepaeva</collectorName>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="EAE15F7B9B18A401BA038976FEB1FBE5" collectionCode="ZMMU" country="Russia" pageId="10" pageNumber="577" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Altay">
|
||
<specimenCount id="4C8F9EAF9B18A401BA038976FA02FB89" box="[1401,1437,1035,1061]" pageId="10" pageNumber="577" type="male">3♂</specimenCount>
|
||
<specimenCount id="4C8F9EAF9B18A401BFED8952FF25FBE5" box="[151,186,1071,1097]" pageId="10" pageNumber="577" type="female">1♀</specimenCount>
|
||
(
|
||
<collectionCode id="3C98CDE39B18A401BFB38952FEB9FBE5" box="[201,294,1071,1097]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="10" pageNumber="577" type="Museum">ZMMU</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="EAE15F7B9B18A401BE438952FE57FBC1" collectingDate="2018-07-21" collectorName="A. E. Naidenov & K. E. Naidenova" country="Russia" elevation="1942" latitude="49.916668" location="Poperechnoe Lake" longLatPrecision="1100" longitude="85.88333" municipality="Katunskyi Mt. Range" pageId="10" pageNumber="577" specimenCount="1" stateProvince="Altay">
|
||
<collectingMunicipality id="BA52CF5C9B18A401BE438952FDB4FBE6" box="[313,555,1071,1098]" pageId="10" pageNumber="577">Katunskyi Mt. Range</collectingMunicipality>
|
||
,
|
||
<location id="5F5603FD9B18A401BD4D8952FC96FBE5" LSID="urn:lsid:plazi:treatment:D220E4309B18A400BFED8FACFA1BFE7D:5F5603FD9B18A401BD4D8952FC96FBE5" box="[567,777,1071,1097]" country="Russia" latitude="49.916668" longLatPrecision="1100" longitude="85.88333" municipality="Katunskyi Mt. Range" name="Poperechnoe Lake" pageId="10" pageNumber="577" stateProvince="Altay">Poperechnoe Lake</location>
|
||
,
|
||
<geoCoordinate id="3FBD33E19B18A401BC6E8952FCE8FBE6" box="[788,887,1071,1098]" degrees="49" direction="north" minutes="55" orientation="latitude" pageId="10" pageNumber="577" precision="925" value="49.916668">49°55’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="3FBD33E19B18A401BCF98952FC7CFBE6" box="[899,995,1071,1098]" degrees="85" direction="east" minutes="53" orientation="longitude" pageId="10" pageNumber="577" precision="925" value="85.88333">85°53’E</geoCoordinate>
|
||
, scree,
|
||
<quantity id="9D71F8C39B18A401BB4C8952FB4CFBE6" box="[1078,1235,1071,1098]" metricMagnitude="3" metricUnit="m" metricValue="1.9425" metricValueMax="2.0" metricValueMin="1.885" pageId="10" pageNumber="577" unit="m" value="1942.5" valueMax="2000.0" valueMin="1885.0">
|
||
<elevation id="D1A4B2159B18A401BB4C8952FB4CFBE6" box="[1078,1235,1071,1098]" metricMagnitude="3" metricUnit="m" metricValue="1.9425" metricValueMax="2.0" metricValueMin="1.885" pageId="10" pageNumber="577" unit="m" value="1942.5" valueMax="2000.0" valueMin="1885.0">1885–2000 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="2E3773E69B18A401BBA58952FAC0FBE5" box="[1247,1375,1071,1097]" pageId="10" pageNumber="577" value="2018-07-21">
|
||
<collectingDate id="3E738A0E9B18A401BBA58952FAC0FBE5" box="[1247,1375,1071,1097]" pageId="10" pageNumber="577" value="2018-07-21">21.07.2018</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="F77C30F09B18A401BA138952FE9AFBC1" pageId="10" pageNumber="577">A.E. Naidenov</collectorName>
|
||
,
|
||
<collectorName id="F77C30F09B18A401BE6A8929FE57FBC1" box="[272,456,1107,1134]" pageId="10" pageNumber="577">K.E. Naidenova</collectorName>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="129306AD9B18A401BFBD890AFADEF9C9" pageId="10" pageNumber="577" type="diagnosis">
|
||
<paragraph id="5A3655269B18A401BFBD890AFADEF9C9" blockId="10.[151,1437,927,2034]" pageId="10" pageNumber="577">
|
||
<emphasis id="68FD89349B18A401BFBD890AFEDDFB3D" bold="true" box="[199,322,1143,1169]" pageId="10" pageNumber="577">Diagnosis.</emphasis>
|
||
<taxonomicName id="9D892EA59B18A401BE32890AFDDBFB3D" authorityName="Marusik, Azarkina & Koponen" authorityYear="2004" box="[328,580,1143,1169]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="levinae">
|
||
<emphasis id="68FD89349B18A401BE32890AFDDBFB3D" box="[328,580,1143,1169]" italics="true" pageId="10" pageNumber="577">Acantholycosa levinae</emphasis>
|
||
</taxonomicName>
|
||
resembles
|
||
<taxonomicName id="9D892EA59B18A401BDBB890AFCA7FB3D" baseAuthorityName="Caporiacco" baseAuthorityYear="1935" box="[705,824,1143,1169]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="baltoroi">
|
||
<emphasis id="68FD89349B18A401BDBB890AFCA7FB3D" box="[705,824,1143,1169]" italics="true" pageId="10" pageNumber="577">A. baltoroi</emphasis>
|
||
</taxonomicName>
|
||
. Both species have protruding palea (
|
||
<emphasis id="68FD89349B18A401BBA7890AFB69FB3D" box="[1245,1270,1143,1169]" italics="true" pageId="10" pageNumber="577">Pl</emphasis>
|
||
) with triangleshaped projection (
|
||
<emphasis id="68FD89349B18A401BE1789E1FE1AFB19" box="[365,389,1180,1205]" italics="true" pageId="10" pageNumber="577">Ts</emphasis>
|
||
), robust terminal apophysis (
|
||
<emphasis id="68FD89349B18A401BDB689E1FD78FB19" box="[716,743,1180,1205]" italics="true" pageId="10" pageNumber="577">Ta</emphasis>
|
||
) and relatively straight embolus (
|
||
<emphasis id="68FD89349B18A401BB2589E1FB1BFB19" box="[1119,1156,1180,1205]" italics="true" pageId="10" pageNumber="577">Em</emphasis>
|
||
). The males of these two species differ by the tip of the embolus (bilobated in
|
||
<taxonomicName id="9D892EA59B18A401BD8989C2FCFBFB75" authorityName="Marusik, Azarkina & Koponen" authorityYear="2004" box="[755,868,1215,1241]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="levinae">
|
||
<emphasis id="68FD89349B18A401BD8989C2FCFBFB75" box="[755,868,1215,1241]" italics="true" pageId="10" pageNumber="577">A. levinae</emphasis>
|
||
</taxonomicName>
|
||
, and single-blade in
|
||
<taxonomicName id="9D892EA59B18A401BB2889C2FB53FB75" baseAuthorityName="Caporiacco" baseAuthorityYear="1935" box="[1106,1228,1215,1241]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="baltoroi">
|
||
<emphasis id="68FD89349B18A401BB2889C2FB53FB75" box="[1106,1228,1215,1241]" italics="true" pageId="10" pageNumber="577">A. baltoroi</emphasis>
|
||
</taxonomicName>
|
||
) and by the shape of the terminal apophysis (curved and bifurcated in
|
||
<taxonomicName id="9D892EA59B18A401BDA6899EFCD2FB51" authorityName="Marusik, Azarkina & Koponen" authorityYear="2004" box="[732,845,1251,1277]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="levinae">
|
||
<emphasis id="68FD89349B18A401BDA6899EFCD2FB51" box="[732,845,1251,1277]" italics="true" pageId="10" pageNumber="577">A. levinae</emphasis>
|
||
</taxonomicName>
|
||
, and straight and blunt in
|
||
<taxonomicName id="9D892EA59B18A401BB09899EFB74FB51" baseAuthorityName="Caporiacco" baseAuthorityYear="1935" box="[1139,1259,1251,1277]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="baltoroi">
|
||
<emphasis id="68FD89349B18A401BB09899EFB74FB51" box="[1139,1259,1251,1277]" italics="true" pageId="10" pageNumber="577">A. baltoroi</emphasis>
|
||
</taxonomicName>
|
||
, cf. Fig. 29 and fig.
|
||
<quantity id="9D71F8C39B18A401BFBF887AFF61FA8D" box="[197,254,1287,1313]" metricMagnitude="0" metricUnit="m" metricValue="1.524" pageId="10" pageNumber="577" unit="in" value="60.0">60 in</quantity>
|
||
<bibRefCitation id="3E1828D79B18A401BE7C887AFE77FA8D" author="Marusik, Y. M. & Azarkina, G. N. & Koponen, S." box="[262,488,1287,1313]" pageId="10" pageNumber="577" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. & Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">
|
||
Marusik
|
||
<emphasis id="68FD89349B18A401BE118875FE3BFA8D" box="[363,420,1287,1313]" italics="true" pageId="10" pageNumber="577">et al.</emphasis>
|
||
2004
|
||
</bibRefCitation>
|
||
). The female of
|
||
<taxonomicName id="9D892EA59B18A401BDDA887AFC8EFA8D" authorityName="Marusik, Azarkina & Koponen" authorityYear="2004" box="[672,785,1287,1313]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="levinae">
|
||
<emphasis id="68FD89349B18A401BDDA887AFC8EFA8D" box="[672,785,1287,1313]" italics="true" pageId="10" pageNumber="577">A. levinae</emphasis>
|
||
</taxonomicName>
|
||
differs from those of
|
||
<taxonomicName id="9D892EA59B18A401BB72887AFB1FFA8D" baseAuthorityName="Caporiacco" baseAuthorityYear="1935" box="[1032,1152,1287,1313]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="baltoroi">
|
||
<emphasis id="68FD89349B18A401BB72887AFB1FFA8D" box="[1032,1152,1287,1313]" italics="true" pageId="10" pageNumber="577">A. baltoroi</emphasis>
|
||
</taxonomicName>
|
||
by the shape of epigynal fovea (
|
||
<emphasis id="68FD89349B18A401BF9D8851FE99FAE9" box="[231,262,1324,1349]" italics="true" pageId="10" pageNumber="577">Fo</emphasis>
|
||
), which is rectangular in the former species (vs. heart-shaped) and by rectangular septal base (
|
||
<emphasis id="68FD89349B18A401BA3F8856FAFEFAE9" box="[1349,1377,1323,1349]" italics="true" pageId="10" pageNumber="577">Sb</emphasis>
|
||
) (vs. anchor-shaped, cf.
|
||
<figureCitation id="C2B249A39B18A401BE0A8832FE5DFAC5" box="[368,450,1359,1385]" captionStart="FIGURES 63–71" captionStartId="11.[151,264,1861,1885]" captionTargetBox="[151,1436,489,1838]" captionTargetId="figure-147@11.[151,1436,489,1838]" captionTargetPageId="11" captionText="FIGURES 63–71. Epigyne and female internal genitalia of Acantholycosa kulikovi sp. n. (63–65), A. levinae (66–68) and Mongolicosa lizae sp. n. (69–71). 63, 66, 69—intact, ventral; 64, 67, 70—macerated, ventral; 65, 68, 71—macerated, dorsal. Scale bars=0.2 mm. Abbreviations: Al apical lips of the epigyne, Ap apical pocket, Cd copulatory ducts, Fo epigynal fovea, Li lip of the epigyne, Re receptacle, Sa septal base arm, Sb septal base, Ss septal stem." figureDoi="http://doi.org/10.5281/zenodo.5300804" httpUri="https://zenodo.org/record/5300804/files/figure.png" pageId="10" pageNumber="577">Fig. 66</figureCitation>
|
||
and fig. 2D in
|
||
<bibRefCitation id="3E1828D79B18A401BD0C8832FC92FAC6" author="Buchar, J." box="[630,781,1359,1386]" pageId="10" pageNumber="577" pagination="201 - 227" refId="ref9724" refString="Buchar, J. (1976) Uber einige Lycosiden (Araneae) aus Nepal. Ergebnisse des Forschungsunternehmens Nepal Himalaya, 5, 201 - 227." type="journal article" year="1976">Buchar 1976</bibRefCitation>
|
||
). The epigyne of
|
||
<taxonomicName id="9D892EA59B18A401BCA38832FBD4FAC5" authorityName="Marusik, Azarkina & Koponen" authorityYear="2004" box="[985,1099,1359,1385]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="levinae">
|
||
<emphasis id="68FD89349B18A401BCA38832FBD4FAC5" box="[985,1099,1359,1385]" italics="true" pageId="10" pageNumber="577">A. levinae</emphasis>
|
||
</taxonomicName>
|
||
is similar to those of species from the
|
||
<taxonomicName id="9D892EA59B18A401BF85880EFEC0FA21" authorityName="Marusik, Azarkina & Koponen" authorityYear="2004" box="[255,351,1395,1421]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="plumalis">
|
||
<emphasis id="68FD89349B18A401BF85880EFEC0FA21" box="[255,351,1395,1421]" italics="true" pageId="10" pageNumber="577">plumalis</emphasis>
|
||
</taxonomicName>
|
||
-group:
|
||
<taxonomicName id="9D892EA59B18A401BEC3880EFC78FA21" authority="Marusik, Azarkina & Koponen, 2004" authorityName="Marusik, Azarkina & Koponen" authorityYear="2004" box="[441,999,1395,1422]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="logunovi">
|
||
<emphasis id="68FD89349B18A401BEC3880EFDA5FA21" box="[441,570,1395,1421]" italics="true" pageId="10" pageNumber="577">A. logunovi</emphasis>
|
||
<bibRefCitation id="3E1828D79B18A401BD38880EFC78FA21" author="Marusik, Y. M. & Azarkina, G. N. & Koponen, S." box="[578,999,1395,1422]" pageId="10" pageNumber="577" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. & Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">Marusik, Azarkina & Koponen, 2004</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="9D892EA59B18A401BC88880EFEDAFA1D" authority="Marusik, Azarkina & Koponen, 2004" authorityName="Marusik, Azarkina & Koponen" authorityYear="2004" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="paraplumalis">
|
||
<emphasis id="68FD89349B18A401BC88880EFB38FA21" box="[1010,1191,1395,1421]" italics="true" pageId="10" pageNumber="577">A. paraplumalis</emphasis>
|
||
<bibRefCitation id="3E1828D79B18A401BBD4880EFEDAFA1D" author="Marusik, Y. M. & Azarkina, G. N. & Koponen, S." pageId="10" pageNumber="577" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. & Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">Marusik, Azarkina & Koponen, 2004</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="9D892EA59B18A401BE0788EAFC32FA1D" authority="Marusik, Azarkina & Koponen, 2004" authorityName="Marusik, Azarkina & Koponen" authorityYear="2004" box="[381,941,1431,1458]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="plumalis">
|
||
<emphasis id="68FD89349B18A401BE0788EAFE62FA1D" box="[381,509,1431,1457]" italics="true" pageId="10" pageNumber="577">A. plumalis</emphasis>
|
||
<bibRefCitation id="3E1828D79B18A401BD7F88EAFC32FA1D" author="Marusik, Y. M. & Azarkina, G. N. & Koponen, S." box="[517,941,1431,1458]" pageId="10" pageNumber="577" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. & Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">Marusik, Azarkina & Koponen, 2004</bibRefCitation>
|
||
</taxonomicName>
|
||
. The similarity includes a rectangular shape of the fovea as well as septal base and apical pocket (
|
||
<emphasis id="68FD89349B18A401BD8C88C6FC8AFA79" box="[758,789,1467,1493]" italics="true" pageId="10" pageNumber="577">Ap</emphasis>
|
||
) consisting of one hood. The female of
|
||
<taxonomicName id="9D892EA59B18A401BBA188C6FAD3FA79" authorityName="Marusik, Azarkina & Koponen" authorityYear="2004" box="[1243,1356,1467,1493]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="levinae">
|
||
<emphasis id="68FD89349B18A401BBA188C6FAD3FA79" box="[1243,1356,1467,1493]" italics="true" pageId="10" pageNumber="577">A. levinae</emphasis>
|
||
</taxonomicName>
|
||
differs from all aforementioned species by a reduced septal stem. In addition, it can be separated from
|
||
<taxonomicName id="9D892EA59B18A401BBB188A2FAD2FA55" authorityName="Marusik, Azarkina & Koponen" authorityYear="2004" box="[1227,1357,1503,1529]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="logunovi">
|
||
<emphasis id="68FD89349B18A401BBB188A2FAD2FA55" box="[1227,1357,1503,1529]" italics="true" pageId="10" pageNumber="577">A. logunovi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="9D892EA59B18A401BAFE88A2FEB3F9B1" authorityName="Marusik, Azarkina & Koponen" authorityYear="2004" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="paraplumalis">
|
||
<emphasis id="68FD89349B18A401BAFE88A2FEB3F9B1" italics="true" pageId="10" pageNumber="577">A. paraplumalis</emphasis>
|
||
</taxonomicName>
|
||
by having the apical pocket as wide as the septal base (vs. twice narrower). Finally, the female of
|
||
<taxonomicName id="9D892EA59B18A401BAFE8B7EFF77F9ED" authorityName="Marusik, Azarkina & Koponen" authorityYear="2004" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="levinae">
|
||
<emphasis id="68FD89349B18A401BAFE8B7EFF77F9ED" italics="true" pageId="10" pageNumber="577">A. levinae</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName id="9D892EA59B18A401BD5A8B5AFD3EF9ED" authorityName="Marusik, Azarkina & Koponen" authorityYear="2004" box="[544,673,1575,1601]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="plumalis">
|
||
<emphasis id="68FD89349B18A401BD5A8B5AFD3EF9ED" box="[544,673,1575,1601]" italics="true" pageId="10" pageNumber="577">A. plumalis</emphasis>
|
||
</taxonomicName>
|
||
by having receptacles (
|
||
<emphasis id="68FD89349B18A401BCC88B55FC4FF9ED" box="[946,976,1576,1601]" italics="true" pageId="10" pageNumber="577">Re</emphasis>
|
||
) almost touching each other (vs. widely spaced, cf.
|
||
<figureCitation id="C2B249A39B18A401BE6F8B36FE11F9C9" box="[277,398,1611,1637]" captionStart="FIGURES 63–71" captionStartId="11.[151,264,1861,1885]" captionTargetBox="[151,1436,489,1838]" captionTargetId="figure-147@11.[151,1436,489,1838]" captionTargetPageId="11" captionText="FIGURES 63–71. Epigyne and female internal genitalia of Acantholycosa kulikovi sp. n. (63–65), A. levinae (66–68) and Mongolicosa lizae sp. n. (69–71). 63, 66, 69—intact, ventral; 64, 67, 70—macerated, ventral; 65, 68, 71—macerated, dorsal. Scale bars=0.2 mm. Abbreviations: Al apical lips of the epigyne, Ap apical pocket, Cd copulatory ducts, Fo epigynal fovea, Li lip of the epigyne, Re receptacle, Sa septal base arm, Sb septal base, Ss septal stem." figureDoi="http://doi.org/10.5281/zenodo.5300804" httpUri="https://zenodo.org/record/5300804/files/figure.png" pageId="10" pageNumber="577">Fig. 66, 68</figureCitation>
|
||
and fig.
|
||
<quantity id="9D71F8C39B18A401BE888B36FD82F9C9" box="[498,541,1611,1637]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="10" pageNumber="577" unit="in" value="3.0">3 in</quantity>
|
||
<bibRefCitation id="3E1828D79B18A401BD5E8B36FCC9F9C9" author="Fomichev, A. A. & Marusik, Y. M." box="[548,854,1611,1637]" pageId="10" pageNumber="577" pagination="65 - 68" refId="ref9802" refString="Fomichev, A. A. & Marusik, Y. M. (2011) First description of the female of Acantholycosa logunovi (Araneae: Lycosidae). Zootaxa, 2813 (1), 65 - 68. https: // doi. org / 10.11646 / zootaxa. 2813.1.3" type="journal article" year="2011">Fomichev & Marusik 2011</bibRefCitation>
|
||
, and figs 47, 50–
|
||
<quantity id="9D71F8C39B18A401BB6C8B36FBD1F9C9" box="[1046,1102,1611,1638]" metricMagnitude="0" metricUnit="m" metricValue="1.3208" pageId="10" pageNumber="577" unit="in" value="52.0">52 in</quantity>
|
||
<bibRefCitation id="3E1828D79B18A401BB2F8B36FAAAF9C9" author="Marusik, Y. M. & Azarkina, G. N. & Koponen, S." box="[1109,1333,1611,1637]" pageId="10" pageNumber="577" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. & Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">
|
||
Marusik
|
||
<emphasis id="68FD89349B18A401BBC38B31FB6DF9C9" box="[1209,1266,1611,1637]" italics="true" pageId="10" pageNumber="577">et al.</emphasis>
|
||
2004
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="129306AD9B18A400BFBD8B12FDB8FF55" lastPageId="11" lastPageNumber="578" pageId="10" pageNumber="577" type="description">
|
||
<paragraph id="5A3655269B18A401BFBD8B12FABBF892" blockId="10.[151,1437,927,2034]" pageId="10" pageNumber="577">
|
||
<emphasis id="68FD89349B18A401BFBD8B12FEC6F925" bold="true" box="[199,345,1647,1673]" pageId="10" pageNumber="577">Description.</emphasis>
|
||
Female.Total length 8.6. Carapace: 4.15 long, 3.3 wide. General appearance as in
|
||
<figureCitation id="C2B249A39B18A401BBAE8B12FA8BF926" box="[1236,1300,1647,1674]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1832,1856]" captionTargetBox="[157,1436,189,1799]" captionTargetId="figure-33@2.[151,1436,181,1809]" captionTargetPageId="2" captionText="FIGURES 1–9. General appearance of Acantholycosa sayanensis (1–2), A. kulikovi sp. n. (3), A. levinae (4–5), A. sergeevi sp. n. (6), Mongolicosa lizae sp. n. (8–9) and leg I of A. sergeevi sp. n. (7). 1, 4, 6, 8—male, dorsal; 2, 3, 5, 9—female, dorsal; 7—leg I of male, prolateral. Scale bars=2 mm." figureDoi="http://doi.org/10.5281/zenodo.5300794" httpUri="https://zenodo.org/record/5300794/files/figure.png" pageId="10" pageNumber="577">Fig. 5</figureCitation>
|
||
.Coloration. Carapace, sternum and labium dark brown. Eye field black. Chelicerae, endites and palps yellow-brown. Coxae yellow-gray. Legs yellow-brown, femora with transverse dark gray stripes. Abdomen dark gray dorsally, with dark brown cardiac mark. Venter of abdomen yellow-gray. Spinnerets dark gray. Spination of leg I: Fe d1-1-1 p0-0-2 r0- 1-1; Ti p1-0-0 r1-0-0 v1-2-2-2-2-2; Mt p0-1-0 r0-1-0 v2-2-0. Leg measurements: I 3.9, 1.8, 3.65, 3.45, 1.7 (14.5); II 3.9, 1.75, 3.4, 3.55, 1.75 (14.35); III 3.8, 1.6, 3.15, 4.3, 1.85 (14.7); IV 4.85, 1.8, 4.3, 6.85, 2.55 (20.35).
|
||
</paragraph>
|
||
<paragraph id="5A3655269B18A401BFBD8A3AFC47F805" blockId="10.[151,1437,927,2034]" pageId="10" pageNumber="577">
|
||
Epigyne and internal genitalia as shown in
|
||
<figureCitation id="C2B249A39B18A401BDCB8A3AFCACF8CD" box="[689,819,1863,1889]" captionStart="FIGURES 63–71" captionStartId="11.[151,264,1861,1885]" captionTargetBox="[151,1436,489,1838]" captionTargetId="figure-147@11.[151,1436,489,1838]" captionTargetPageId="11" captionText="FIGURES 63–71. Epigyne and female internal genitalia of Acantholycosa kulikovi sp. n. (63–65), A. levinae (66–68) and Mongolicosa lizae sp. n. (69–71). 63, 66, 69—intact, ventral; 64, 67, 70—macerated, ventral; 65, 68, 71—macerated, dorsal. Scale bars=0.2 mm. Abbreviations: Al apical lips of the epigyne, Ap apical pocket, Cd copulatory ducts, Fo epigynal fovea, Li lip of the epigyne, Re receptacle, Sa septal base arm, Sb septal base, Ss septal stem." figureDoi="http://doi.org/10.5281/zenodo.5300804" httpUri="https://zenodo.org/record/5300804/files/figure.png" pageId="10" pageNumber="577">Figs 66–68</figureCitation>
|
||
. Fovea as wide as long. Septal base smoothly declines anteriorly. Lips of the epigyne (
|
||
<emphasis id="68FD89349B18A401BE808A11FD8EF829" box="[506,529,1900,1925]" italics="true" pageId="10" pageNumber="577">Li</emphasis>
|
||
) separated by one of their wide. Receptacles clavate, not clearly delineated from copulatory ducts (
|
||
<emphasis id="68FD89349B18A401BE1B8AF2FE1DF805" box="[353,386,1935,1961]" italics="true" pageId="10" pageNumber="577">Cd</emphasis>
|
||
). Copulatory ducts 1.5 times longer than receptacles.
|
||
</paragraph>
|
||
<paragraph id="5A3655269B18A400BFBD8ACEFDB8FF55" blockId="10.[151,1437,927,2034]" lastBlockId="11.[151,1437,151,465]" lastPageId="11" lastPageNumber="578" pageId="10" pageNumber="577">
|
||
Male. See
|
||
<bibRefCitation id="3E1828D79B18A401BE448ACEFDB4F861" author="Marusik, Y. M. & Azarkina, G. N. & Koponen, S." box="[318,555,1971,1997]" pageId="10" pageNumber="577" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. & Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">
|
||
Marusik
|
||
<emphasis id="68FD89349B18A401BED88AC9FE45F861" box="[418,474,1971,1997]" italics="true" pageId="10" pageNumber="577">et al.</emphasis>
|
||
(2004)
|
||
</bibRefCitation>
|
||
for description. General appearance as in
|
||
<figureCitation id="C2B249A39B18A401BB788ACEFBD9F861" box="[1026,1094,1971,1997]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1832,1856]" captionTargetBox="[157,1436,189,1799]" captionTargetId="figure-33@2.[151,1436,181,1809]" captionTargetPageId="2" captionText="FIGURES 1–9. General appearance of Acantholycosa sayanensis (1–2), A. kulikovi sp. n. (3), A. levinae (4–5), A. sergeevi sp. n. (6), Mongolicosa lizae sp. n. (8–9) and leg I of A. sergeevi sp. n. (7). 1, 4, 6, 8—male, dorsal; 2, 3, 5, 9—female, dorsal; 7—leg I of male, prolateral. Scale bars=2 mm." figureDoi="http://doi.org/10.5281/zenodo.5300794" httpUri="https://zenodo.org/record/5300794/files/figure.png" pageId="10" pageNumber="577">Fig. 4</figureCitation>
|
||
. Palp as in
|
||
<figureCitation id="C2B249A39B18A401BBBE8ACEFADBF861" box="[1220,1348,1971,1997]" captionStart="FIGURES 10–19" captionStartId="3.[151,264,1849,1873]" captionTargetBox="[155,1425,188,1810]" captionTargetId="figure-15@3.[151,1436,181,1825]" captionTargetPageId="3" captionText="FIGURES 10–19. Male palp (10–13) and bulb (14–19) of Acantholycosa sergeevi sp. n. (10–11), A. sayanensis (14–15), A. sidorovi sp. n. (16–17), A. levinae (18-19) and Mongolicosa lizae sp. n. (12–13). 10, 12—retrolateral; 11, 13, 14, 16, 18— ventral; 15, 17, 19—anterior. Scale bars=0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5300796" httpUri="https://zenodo.org/record/5300796/files/figure.png" pageId="10" pageNumber="577">Figs 18–19</figureCitation>
|
||
, 28–31, 36–37. Tegular apophysis (
|
||
<emphasis id="68FD89349B18A401BEBB8AA5FE40F85D" box="[449,479,2008,2033]" italics="true" pageId="10" pageNumber="577">Tg</emphasis>
|
||
) almost oval, with reduced apical and basal arms. Palea with abrupt retrolateral margin. Triangle shaped projection almost straight, with a hooked apical part. Terminal apophysis massive, subdivided into two lobes. Embolus with a spine (
|
||
<emphasis id="68FD89349B19A400BD2B8DC6FDF2FF79" box="[593,621,187,213]" italics="true" pageId="11" pageNumber="578">Sp</emphasis>
|
||
) in the basal part. Distal part of the embolus subdivided into triangular secondary part and embolus proper.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="129306AD9B19A400BFBD8C7FFF5BFEEE" pageId="11" pageNumber="578" type="distribution">
|
||
<paragraph id="5A3655269B19A400BFBD8C7FFF5BFEEE" blockId="11.[151,1437,151,465]" pageId="11" pageNumber="578">
|
||
<emphasis id="68FD89349B19A400BFBD8C7FFEFEFEB0" bold="true" box="[199,353,258,284]" pageId="11" pageNumber="578">Distribution.</emphasis>
|
||
Known from several close localities in the Katunskyi Mt. Range in
|
||
<collectingRegion id="984D9BC49B19A400BB158C7EFB38FEB1" box="[1135,1191,259,285]" country="Russia" name="Altay" pageId="11" pageNumber="578">Altai</collectingRegion>
|
||
Mountains (
|
||
<figureCitation id="C2B249A39B19A400BA428C7EFF28FEEE" captionStart="FIGURES 72–77" captionStartId="13.[151,264,1676,1700]" captionTargetBox="[151,1436,309,1652]" captionTargetId="figure-76@13.[151,1436,309,1652]" captionTargetPageId="13" captionText="FIGURES 72–77. Male of Acantholycosa sayanensis from Borus Mt. Range (72), female of Acantholycosa sp. (A. azyuzini or A. levinae) with egg sac, from Katunskyi Mt. Range (73), habitat of A. sayanensis and A. sergeevi sp. n. in Borus Mt. Range (74), collecting localities of studied species from the Acantholycosa-complex (75–76) and map showing the main mountain systems of South Siberia and adjacent regions (77). Diamond—A. sayanensis, square—A. sidorovi sp. n., triangle—A. kulikovi sp. n., star—A. levinae, circle—A. sergeevi sp. n., inverted triangle—Mongolicosa lizae sp. n. The frame of Fig. 76 refers to the content of Fig. 75." figureDoi="http://doi.org/10.5281/zenodo.5300806" httpUri="https://zenodo.org/record/5300806/files/figure.png" pageId="11" pageNumber="578">Figs 75– 76</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="129306AD9B19A400BFBD8C37FA1BFE7D" pageId="11" pageNumber="578" type="discussion">
|
||
<paragraph id="5A3655269B19A400BFBD8C37FA1BFE7D" blockId="11.[151,1437,151,465]" pageId="11" pageNumber="578">
|
||
<emphasis id="68FD89349B19A400BFBD8C37FED0FEC8" bold="true" box="[199,335,330,356]" pageId="11" pageNumber="578">Comments.</emphasis>
|
||
There are some doubts if the females match with males due to the morphology of the epigyne. Since
|
||
<taxonomicName id="9D892EA59B19A400BFED8C12FE97FE25" authorityName="Marusik, Azarkina & Koponen" authorityYear="2004" box="[151,264,367,393]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="578" phylum="Arthropoda" rank="species" species="levinae">
|
||
<emphasis id="68FD89349B19A400BFED8C12FE97FE25" box="[151,264,367,393]" italics="true" pageId="11" pageNumber="578">A. levinae</emphasis>
|
||
</taxonomicName>
|
||
belongs to the
|
||
<taxonomicName id="9D892EA59B19A400BEC28C12FD8EFE25" baseAuthorityName="Caporiacco" baseAuthorityYear="1935" box="[440,529,367,393]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="578" phylum="Arthropoda" rank="species" species="baltoroi">
|
||
<emphasis id="68FD89349B19A400BEC28C12FD8EFE25" box="[440,529,367,393]" italics="true" pageId="11" pageNumber="578">baltoroi</emphasis>
|
||
</taxonomicName>
|
||
-group, this species epigyne would be expected to have an anchor-shaped septal base. Instead, the epigyne of the females described herein are similar to those of species from the
|
||
<taxonomicName id="9D892EA59B19A400BB908CEEFAD5FE01" authorityName="Marusik, Azarkina & Koponen" authorityYear="2004" box="[1258,1354,403,429]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="578" phylum="Arthropoda" rank="species" species="plumalis">
|
||
<emphasis id="68FD89349B19A400BB908CEEFAD5FE01" box="[1258,1354,403,429]" italics="true" pageId="11" pageNumber="578">plumalis</emphasis>
|
||
</taxonomicName>
|
||
-group. Nevertheless, males and females were collected together in two different places, thus I treat them as conspecific.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |