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<document id="69A2CA2F98E9FC6CEF8909298691FA39" ID-CLB-Dataset="293300" ID-DOI="10.11646/zootaxa.5433.2.5" ID-GBIF-Dataset="d933d945-36e9-453c-979e-8e408d0e0330" ID-ISSN="1175-5326" ID-Zenodo-Dep="10954120" ID-ZooBank="AE99AF6B-787F-4E48-8DD3-F8B0ADD3A887" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1712751459482" checkinUser="plazi" docAuthor="Lee, Chi-Woo &amp; Min, Gi-Sik" docDate="2024" docId="AA1187DAFFD4F810FF1019ABFB56FC0C" docLanguage="en" docName="zootaxa.5433.2.5.pdf" docOrigin="Zootaxa 5433 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5433.2.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudocrangonyx danyangensis Lee &amp; Min 2024, sp. nov." docType="treatment" docUuid="514F443E-A246-4FFF-91B0-0540090CC498" docUuidSource="ZooBank" docVersion="2" lastPageNumber="264" masterDocId="5628FFA2FFDDF81FFF87193CFF92FF8D" masterDocTitle="Two new species of Pseudocrangonyx (Amphipoda: Pseudocrangonyctidae) from the hyporheic zones in South Korea" masterLastPageNumber="265" masterPageNumber="249" pageNumber="258" updateTime="1712752432382" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="F8FCA0BBDF218CA66CC8A2C1CA81B92E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="0AA93DA2D345583691C99A61F351D0C7">Two new species of Pseudocrangonyx (Amphipoda: Pseudocrangonyctidae) from the hyporheic zones in South Korea</mods:title>
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<mods:name id="0CF6362D44F11240C85937C07196B729" type="personal">
<mods:role id="DDA5A033A1C912A5ED6D86C2F577C70A">
<mods:roleTerm id="BC00F4FF4B7FD6D39816C1BFC1ACBEE9">Author</mods:roleTerm>
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<mods:namePart id="C5A77AF7249199ED144BAA754BB91A7F">Lee, Chi-Woo</mods:namePart>
<mods:affiliation id="877067FA15F7D101153DE04AD25E8E5D">Nakdonggang National Institute of Biological Resources, Sangju 37242, Korea &amp; Department of Biological Sciences, Inha University, Incheon 22212, Korea</mods:affiliation>
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<mods:roleTerm id="304690692C2C9880907D675791617633">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D52D0F287AF3820B2375AC172A3D5B75">Min, Gi-Sik</mods:namePart>
<mods:affiliation id="2F68F99914D95B771BD1CF56D6C48470">Department of Biological Sciences, Inha University, Incheon 22212, Korea</mods:affiliation>
</mods:name>
<mods:typeOfResource id="A3C47011590E5C5FC5C1E73E86F5A7BC">text</mods:typeOfResource>
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<mods:title id="4BDCFE412D01BCDA6E172540149F531B">Zootaxa</mods:title>
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<mods:part id="26C22EE88F727D8E533B8AE63DD1A061">
<mods:date id="8C86B60539E19E4F0CF1FB5DBEBADE00">2024</mods:date>
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<mods:number id="AB43B505D5C0A8A0B3955EFB39D98A03">2024-04-05</mods:number>
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<mods:number id="87EAFEC5446B910851D71771907CD012">5433</mods:number>
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<mods:number id="D8511C342BDB028CEEC3751B8DDB1A9C">2</mods:number>
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<mods:url id="DDB1F489BA105104C6C1F1ED8FA95E51">http://dx.doi.org/10.11646/zootaxa.5433.2.5</mods:url>
</mods:location>
<mods:classification id="C3A5FC7164BA2096A711D294738A417A">journal article</mods:classification>
<mods:identifier id="274C73E8ED679AB99DBB690EFAB52A93" type="CLB-Dataset">293300</mods:identifier>
<mods:identifier id="0FBDBE359E21A534C82E4F334E89B00F" type="DOI">10.11646/zootaxa.5433.2.5</mods:identifier>
<mods:identifier id="EE7CB5EA839C3E89E614515F3BEF1537" type="GBIF-Dataset">d933d945-36e9-453c-979e-8e408d0e0330</mods:identifier>
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<mods:identifier id="DF23743B437C95616E733E1F1FBF5573" type="Zenodo-Dep">10954120</mods:identifier>
<mods:identifier id="042087BBD37ED3AC5171DCF61B542A73" type="ZooBank">AE99AF6B-787F-4E48-8DD3-F8B0ADD3A887</mods:identifier>
</mods:mods>
<treatment id="AA1187DAFFD4F810FF1019ABFB56FC0C" ID-DOI="http://doi.org/10.5281/zenodo.10954109" ID-Zenodo-Dep="10954109" LSID="urn:lsid:zoobank.org:act:514F443E-A246-4FFF-91B0-0540090CC498" httpUri="http://treatment.plazi.org/id/AA1187DAFFD4F810FF1019ABFB56FC0C" lastPageId="15" lastPageNumber="264" pageId="9" pageNumber="258">
<subSubSection id="6AA26547FFD4F816FF1019ABFD0CFF3C" box="[151,670,151,178]" pageId="9" pageNumber="258" type="nomenclature">
<paragraph id="220736CCFFD4F816FF1019ABFD0CFF3C" blockId="9.[151,946,151,285]" box="[151,670,151,178]" pageId="9" pageNumber="258">
<heading id="794F81A0FFD4F816FF1019ABFD0CFF3C" bold="true" box="[151,670,151,178]" fontSize="11" level="1" pageId="9" pageNumber="258" reason="1">
<emphasis id="10CCEADEFFD4F816FF1019ABFD0CFF3C" bold="true" box="[151,670,151,178]" pageId="9" pageNumber="258">
2.
<taxonomicName id="E5B84D4FFFD4F816FF3219ABFDA8FF3F" authority="Lee &amp; Min, 2024" authorityName="Lee &amp; Min" authorityYear="2024" box="[181,570,151,178]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="258" phylum="Arthropoda" rank="species" species="danyangensis" status="sp. nov.">
<emphasis id="10CCEADEFFD4F816FF3219ABFDA8FF3F" bold="true" box="[181,570,151,178]" italics="true" pageId="9" pageNumber="258">Pseudocrangonyx danyangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0BFF57A5FFD4F816FDC619ABFD0CFF3C" box="[577,670,151,177]" pageId="9" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="6AA26547FFD4F816FF101987FEDDFE90" pageId="9" pageNumber="258" type="description">
<paragraph id="220736CCFFD4F816FF101987FC20FF5B" blockId="9.[151,946,151,285]" box="[151,946,187,214]" pageId="9" pageNumber="258">
<uri id="56293ACEFFD4F816FF101987FC20FF5B" box="[151,946,187,214]" pageId="9" pageNumber="258">
urn:lsid:zoobank.org:act:
<uuid id="561E0C19FFD4F816FE221987FC20FF5B" box="[421,946,187,214]" pageId="9" pageNumber="258">514F443E-A246-4FFF-91B0-0540090CC498</uuid>
</uri>
</paragraph>
<paragraph id="220736CCFFD4F816FF1019DCFD77FF74" blockId="9.[151,946,151,285]" box="[151,741,223,250]" pageId="9" pageNumber="258">[New Korean name: dan-yang-dong-gul-yeop-sae-u]</paragraph>
<paragraph id="220736CCFFD4F816FF10183FFEDDFE90" blockId="9.[151,946,151,285]" box="[151,335,259,285]" pageId="9" pageNumber="258">
(
<figureCitation id="BA832A49FFD4F816FF18183FFE90FE90" box="[159,258,259,285]" captionStart="FIGURE 2" captionStartId="3.[151,250,1908,1932]" captionTargetBox="[151,1436,1354,1884]" captionTargetId="figure-343@3.[151,1436,1354,1884]" captionTargetPageId="3" captionText="FIGURE 2. Habitus of two new species. A, Pseudocrangonyx seomjinensis sp. nov.; B, Pseudocrangonyx danyangensis sp. nov.." figureDoi="http://doi.org/10.5281/zenodo.10954124" httpUri="https://zenodo.org/record/10954124/files/figure.png" pageId="9" pageNumber="258">Figs. 2B</figureCitation>
,
<figureCitation id="BA832A49FFD4F816FE8A183FFED5FE90" box="[269,327,259,285]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStart-2="FIGURE 9" captionStart-3="FIGURE 10" captionStartId-0="10.[151,250,1909,1933]" captionStartId-1="11.[151,250,1909,1933]" captionStartId-2="13.[151,250,1909,1933]" captionStartId-3="14.[151,250,1943,1967]" captionTargetBox-0="[189,1399,186,1884]" captionTargetBox-1="[187,1399,186,1884]" captionTargetBox-2="[222,1366,187,1884]" captionTargetBox-3="[159,1428,181,1919]" captionTargetId-0="figure-33@10.[189,1399,181,1884]" captionTargetId-1="figure-17@11.[187,1399,181,1884]" captionTargetId-2="figure-17@13.[222,1366,181,1884]" captionTargetId-3="figure-33@14.[159,1428,181,1919]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="13" captionTargetPageId-3="14" captionText-0="FIGURE 7. Pseudocrangonyx danyangensis sp. nov., holotype female, NNIBRIV69645. A–G, I, J; paratype male, NNIBRIV69646, H. A, antenna 1, medial view; B, accessory flagellum of antenna 1, medial view; C, antenna 2, medial view; D, upper lip, posterior view; E, left mandible, medial view; F, incisor, lacinia mobilis of right mandible, medial view; G, lower lip, dorsal view; H, maxilla 1, dorsal view; I, maxilla 2, dorsal view; J, maxilliped, ventral view." captionText-1="FIGURE 8. Pseudocrangonyx danyangensis sp. nov., holotype female, NNIBRIV69645. A, gnathopod 1, lateral view; B, gnathopod 2, lateral view; C, pereopod 3, lateral view; D, dactylus of pereopod 3, lateral view; E, pereopod 4, lateral view; F, dactylus of pereopod 4, lateral view; G, pereopod 5, lateral view; H, dactylus of pereopod 5, lateral view; I, pereopod 6, lateral view; J, dactylus of pereopod 6, lateral view; K, pereopod 7, lateral view; L, dactylus of pereopod 7, lateral view." captionText-2="FIGURE 9. Pseudocrangonyx danyangensis sp. nov., holotype female, NNIBRIV69645. A–C, Pleopod 1–3, lateral view; D, uropod 1, ventral view; E, uropod 2, dorsal view; F, uropod 3, dorsal view; G, telson, ventral view." captionText-3="FIGURE 10. Pseudocrangonyx danyangensis sp. nov., paratype male, NNIBRIV69646.A, antenna 1, medial view; B, accessory flagellum of antenna 1, medial view; C, antenna 2, lateral view; D, gnathopod 1, lateral view; E, gnathopod 2, lateral view; F, uropod 1, ventral view; G, uropod 2, ventral view; H, uropod 3, dorsal view; I, telson, ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.10954134" figureDoi-1="http://doi.org/10.5281/zenodo.10954136" figureDoi-2="http://doi.org/10.5281/zenodo.10954140" figureDoi-3="http://doi.org/10.5281/zenodo.10954142" httpUri-0="https://zenodo.org/record/10954134/files/figure.png" httpUri-1="https://zenodo.org/record/10954136/files/figure.png" httpUri-2="https://zenodo.org/record/10954140/files/figure.png" httpUri-3="https://zenodo.org/record/10954142/files/figure.png" pageId="9" pageNumber="258">7–10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="6AA26547FFD4F816FF101877FED5FE5C" pageId="9" pageNumber="258" type="materials_examined">
<paragraph id="220736CCFFD4F816FF101877FAEBFE07" blockId="9.[151,1437,331,2014]" pageId="9" pageNumber="258">
<emphasis id="10CCEADEFFD4F816FF101877FE13FEE8" bold="true" box="[151,385,331,357]" pageId="9" pageNumber="258">Material examined.</emphasis>
<materialsCitation id="92D03C91FFD4F816FE0E1877FAEBFE07" collectingDate="2020-06-19" collectorName="C. - W. Lee." country="South Korea" county="Danyang-gun" latitude="37.084385" location="Namhan River" longLatPrecision="1" longitude="128.48405" municipality="Yeongchun-myeon" pageId="9" pageNumber="258" specimenCount="1" stateProvince="Chungbuk" typeStatus="holotype">
<typeStatus id="FD03886EFFD4F816FE0E1877FE64FEE8" box="[393,502,331,357]" pageId="9" pageNumber="258" type="holotype">Holotype</typeStatus>
: NNIBRIV69645, female, BL
<quantity id="E5409B29FFD4F816FCC81877FC3AFEEB" box="[847,936,331,358]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="9" pageNumber="258" unit="mm" value="5.2">5.2 mm</quantity>
, collected from
<location id="27676017FFD4F816FBD81877FA91FEE8" LSID="urn:lsid:plazi:treatment:AA1187DAFFD4F810FF1019ABFB56FC0C:27676017FFD4F816FBD81877FA91FEE8" box="[1119,1283,331,357]" country="South Korea" county="Danyang-gun" latitude="37.084385" longLatPrecision="1" longitude="128.48405" municipality="Yeongchun-myeon" name="Namhan River" pageId="9" pageNumber="258" stateProvince="Chungbuk">Namhan River</location>
(
<geoCoordinate id="478C500BFFD4F816FA941871FA0AFEE5" box="[1299,1432,333,360]" degrees="37" direction="north" minutes="5.063" orientation="latitude" pageId="9" pageNumber="258" precision="1" value="37.084385">37°5.063′N</geoCoordinate>
,
<geoCoordinate id="478C500BFFD4F816FF10184DFEA6FE01" box="[151,308,369,396]" degrees="128" direction="east" minutes="29.043" orientation="longitude" pageId="9" pageNumber="258" precision="1" value="128.48405">128°29.043′E</geoCoordinate>
),
<collectingMunicipality id="C263ACB6FFD4F816FEC01853FD8CFE04" box="[327,542,367,393]" pageId="9" pageNumber="258">Yeongchun-myeon</collectingMunicipality>
,
<collectingCounty id="CB664E40FFD4F816FDAE184CFD56FE07" box="[553,708,368,394]" pageId="9" pageNumber="258">Danyang-gun</collectingCounty>
,
<collectingRegion id="E07CF82EFFD4F816FD481853FC2FFE04" box="[719,957,367,393]" country="South Korea" name="Chungbuk" pageId="9" pageNumber="258">Chungcheongbuk-do</collectingRegion>
,
<collectingCountry id="5AAF765CFFD4F816FC4F184CFB9DFE07" box="[968,1039,368,394]" name="South Korea" pageId="9" pageNumber="258">Korea</collectingCountry>
, on
<date id="5606100CFFD4F816FBBA1853FB5BFE04" box="[1085,1225,367,394]" pageId="9" pageNumber="258" value="2020-06-19">
<collectingDate id="4642E9E4FFD4F816FBBA1853FB5BFE04" box="[1085,1225,367,394]" pageId="9" pageNumber="258" value="2020-06-19">19 Jun 2020</collectingDate>
</date>
, by
<collectorName id="8F4D531AFFD4F816FB701853FAEBFE07" box="[1271,1401,367,394]" pageId="9" pageNumber="258">C. -W. Lee.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="220736CCFFD4F816FF4018A8FED5FE5C" blockId="9.[151,1437,331,2014]" pageId="9" pageNumber="258">
<typeStatus id="FD03886EFFD4F816FF4018A8FEAAFE23" box="[199,312,404,430]" pageId="9" pageNumber="258" type="paratype">Paratypes</typeStatus>
: NNIBRIV69646,
<specimenCount id="34BEFD45FFD4F816FD8C18AFFDCBFE20" box="[523,601,403,429]" count="1" pageId="9" pageNumber="258" type="male">1 male</specimenCount>
, BL 4.0 mm, NNIBRIV69644,
<specimenCount id="34BEFD45FFD4F816FC3C18AFFB8DFE20" box="[955,1055,403,429]" count="1" pageId="9" pageNumber="258" type="female">1 female</specimenCount>
, BL
<quantity id="E5409B29FFD4F816FBD418AFFB3EFE23" box="[1107,1196,403,430]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="9" pageNumber="258" unit="mm" value="4.9">4.9 mm</quantity>
;
<materialsCitation id="92D03C91FFD4F816FB3F18AFFED1FE5C" collectingDate="2020-06-19" collectorName="C. - W. Lee." country="South Korea" county="Danyang-gun" latitude="37.084385" location="Namhan River" longLatPrecision="1" longitude="128.48405" municipality="Yeongchun-myeon" pageId="9" pageNumber="258" specimenCount="1" stateProvince="Chungbuk" typeStatus="paratype">collection data same as for holotype</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="6AA26547FFD4F816FF4018E7FD54FC9B" pageId="9" pageNumber="258" type="diagnosis">
<paragraph id="220736CCFFD4F816FF4018E7FD54FC9B" blockId="9.[151,1437,331,2014]" pageId="9" pageNumber="258">
<emphasis id="10CCEADEFFD4F816FF4018E7FED0FE78" bold="true" box="[199,322,475,501]" pageId="9" pageNumber="258">Diagnosis.</emphasis>
Antennal sinus with rounded angle; eyes absent; pereonites 1–7 with short dorsal setae; dorsal margin of urosomite 1–3 with setae; sternal gill absent; antenna 1 being 0.38 times as long as body length; male antenna 2 with calceoli; maxilla 1 outer plate with 7 serrate teeth; maxilla 2 inner plate with oblique inner row of 2 setae; palmar margins of propodi of gnathopods 1 and 2 with 5–6 and 7 robust setae, respectively; pleopods inner margin of inner rami with bifid setae; uropod 1 inner ramus 0.74 times as long as peduncle, inner margin with 2 robust setae, outer margin bare, basal part with 2 slender setae, outer ramus 0.67 times as long as inner, inner margin and marginal with robust seta, respectively; uropod 2 peduncle outer and inner margin with 6 and 2 robust setae, respectively; uropod 3 terminal article longer than adjacent robust setae; telson each lobe with 2 lateral penicillate setae, 1 apical robust and 1 apical penicillate seta.
</paragraph>
</subSubSection>
<subSubSection id="6AA26547FFD4F813FF401A23FBF9FAF4" lastPageId="12" lastPageNumber="261" pageId="9" pageNumber="258" type="description">
<paragraph id="220736CCFFD4F816FF401A23FBB9FC60" blockId="9.[151,1437,331,2014]" pageId="9" pageNumber="258">
<emphasis id="10CCEADEFFD4F816FF401A23FECBFCB4" bold="true" box="[199,345,799,825]" pageId="9" pageNumber="258">Description.</emphasis>
Female
<typeStatus id="FD03886EFFD4F816FE381A23FDB3FCB4" box="[447,545,799,825]" pageId="9" pageNumber="258" type="holotype">holotype</typeStatus>
(NNIBRIV69645): Head (
<figureCitation id="BA832A49FFD4F816FCDF1A23FC23FCB4" box="[856,945,799,825]" captionStart="FIGURE 2" captionStartId="3.[151,250,1908,1932]" captionTargetBox="[151,1436,1354,1884]" captionTargetId="figure-343@3.[151,1436,1354,1884]" captionTargetPageId="3" captionText="FIGURE 2. Habitus of two new species. A, Pseudocrangonyx seomjinensis sp. nov.; B, Pseudocrangonyx danyangensis sp. nov.." figureDoi="http://doi.org/10.5281/zenodo.10954124" httpUri="https://zenodo.org/record/10954124/files/figure.png" pageId="9" pageNumber="258">Fig. 2B</figureCitation>
) with short dorsal setae; rostrum reduced; lateral cephalic lobe rounded; antennal sinus shallow with rounded angle; eyes absent. Dorsal margin of pleonites 1–7 and urosomites 1–2 with setae (
<figureCitation id="BA832A49FFD4F816FDBA1A5BFD0AFC0C" box="[573,664,871,897]" captionStart="FIGURE 2" captionStartId="3.[151,250,1908,1932]" captionTargetBox="[151,1436,1354,1884]" captionTargetId="figure-343@3.[151,1436,1354,1884]" captionTargetPageId="3" captionText="FIGURE 2. Habitus of two new species. A, Pseudocrangonyx seomjinensis sp. nov.; B, Pseudocrangonyx danyangensis sp. nov.." figureDoi="http://doi.org/10.5281/zenodo.10954124" httpUri="https://zenodo.org/record/10954124/files/figure.png" pageId="9" pageNumber="258">Fig. 2B</figureCitation>
). posteroventral corner of urosomite 1 with seta (
<figureCitation id="BA832A49FFD4F816FB5E1A5BFAA6FC0C" box="[1241,1332,871,897]" captionStart="FIGURE 2" captionStartId="3.[151,250,1908,1932]" captionTargetBox="[151,1436,1354,1884]" captionTargetId="figure-343@3.[151,1436,1354,1884]" captionTargetPageId="3" captionText="FIGURE 2. Habitus of two new species. A, Pseudocrangonyx seomjinensis sp. nov.; B, Pseudocrangonyx danyangensis sp. nov.." figureDoi="http://doi.org/10.5281/zenodo.10954124" httpUri="https://zenodo.org/record/10954124/files/figure.png" pageId="9" pageNumber="258">Fig. 2B</figureCitation>
). Ventral margin of epimeral plate 1 without seta, posteroventral corner rounded with seta (
<figureCitation id="BA832A49FFD4F816FBC71AB7FB08FC28" box="[1088,1178,907,933]" captionStart="FIGURE 2" captionStartId="3.[151,250,1908,1932]" captionTargetBox="[151,1436,1354,1884]" captionTargetId="figure-343@3.[151,1436,1354,1884]" captionTargetPageId="3" captionText="FIGURE 2. Habitus of two new species. A, Pseudocrangonyx seomjinensis sp. nov.; B, Pseudocrangonyx danyangensis sp. nov.." figureDoi="http://doi.org/10.5281/zenodo.10954124" httpUri="https://zenodo.org/record/10954124/files/figure.png" pageId="9" pageNumber="258">Fig. 2B</figureCitation>
); ventral and posterior margins of plate 2 each with seta, posteroventral corner rounded with seta (
<figureCitation id="BA832A49FFD4F816FC621A93FBAFFC44" box="[997,1085,943,969]" captionStart="FIGURE 2" captionStartId="3.[151,250,1908,1932]" captionTargetBox="[151,1436,1354,1884]" captionTargetId="figure-343@3.[151,1436,1354,1884]" captionTargetPageId="3" captionText="FIGURE 2. Habitus of two new species. A, Pseudocrangonyx seomjinensis sp. nov.; B, Pseudocrangonyx danyangensis sp. nov.." figureDoi="http://doi.org/10.5281/zenodo.10954124" httpUri="https://zenodo.org/record/10954124/files/figure.png" pageId="9" pageNumber="258">Fig. 2B</figureCitation>
); ventral and posterior margins of plate 3 each with 2 setae, posteroventral corner rounded without seta (
<figureCitation id="BA832A49FFD4F816FC411AEFFB8CFC60" box="[966,1054,979,1005]" captionStart="FIGURE 2" captionStartId="3.[151,250,1908,1932]" captionTargetBox="[151,1436,1354,1884]" captionTargetId="figure-343@3.[151,1436,1354,1884]" captionTargetPageId="3" captionText="FIGURE 2. Habitus of two new species. A, Pseudocrangonyx seomjinensis sp. nov.; B, Pseudocrangonyx danyangensis sp. nov.." figureDoi="http://doi.org/10.5281/zenodo.10954124" httpUri="https://zenodo.org/record/10954124/files/figure.png" pageId="9" pageNumber="258">Fig. 2B</figureCitation>
).
</paragraph>
<paragraph id="220736CCFFD4F816FF401ACBFBE5FBF0" blockId="9.[151,1437,331,2014]" pageId="9" pageNumber="258">
Antenna 1 (
<figureCitation id="BA832A49FFD4F816FECB1ACBFE37FB9C" box="[332,421,1015,1041]" captionStart="FIGURE 7" captionStartId="10.[151,250,1909,1933]" captionTargetBox="[189,1399,186,1884]" captionTargetId="figure-33@10.[189,1399,181,1884]" captionTargetPageId="10" captionText="FIGURE 7. Pseudocrangonyx danyangensis sp. nov., holotype female, NNIBRIV69645. A–G, I, J; paratype male, NNIBRIV69646, H. A, antenna 1, medial view; B, accessory flagellum of antenna 1, medial view; C, antenna 2, medial view; D, upper lip, posterior view; E, left mandible, medial view; F, incisor, lacinia mobilis of right mandible, medial view; G, lower lip, dorsal view; H, maxilla 1, dorsal view; I, maxilla 2, dorsal view; J, maxilliped, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10954134" httpUri="https://zenodo.org/record/10954134/files/figure.png" pageId="9" pageNumber="258">Fig. 7A</figureCitation>
) 0.38 times as long as body length, peduncular articles
<quantity id="E5409B29FFD4F816FB991ACBFBECFB9C" box="[1054,1150,1015,1042]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" metricValueMax="7.62" metricValueMin="2.54" pageId="9" pageNumber="258" unit="in" value="2.0" valueMax="3.0" valueMin="1.0">1 to 3 in</quantity>
length ratio of 1.0: 0.6: 0.34; accessory flagellum (
<figureCitation id="BA832A49FFD4F816FE461D27FD84FBB8" box="[449,534,1051,1077]" captionStart="FIGURE 7" captionStartId="10.[151,250,1909,1933]" captionTargetBox="[189,1399,186,1884]" captionTargetId="figure-33@10.[189,1399,181,1884]" captionTargetPageId="10" captionText="FIGURE 7. Pseudocrangonyx danyangensis sp. nov., holotype female, NNIBRIV69645. A–G, I, J; paratype male, NNIBRIV69646, H. A, antenna 1, medial view; B, accessory flagellum of antenna 1, medial view; C, antenna 2, medial view; D, upper lip, posterior view; E, left mandible, medial view; F, incisor, lacinia mobilis of right mandible, medial view; G, lower lip, dorsal view; H, maxilla 1, dorsal view; I, maxilla 2, dorsal view; J, maxilliped, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10954134" httpUri="https://zenodo.org/record/10954134/files/figure.png" pageId="9" pageNumber="258">Fig. 7B</figureCitation>
) 2-articulate, terminal article with 3 setae and 1 aesthetasc; primary flagellum 11- articulate, 1 aesthetasc on some articles. Antenna 2 (
<figureCitation id="BA832A49FFD4F816FD651D03FCAAFBD4" box="[738,824,1087,1113]" captionStart="FIGURE 7" captionStartId="10.[151,250,1909,1933]" captionTargetBox="[189,1399,186,1884]" captionTargetId="figure-33@10.[189,1399,181,1884]" captionTargetPageId="10" captionText="FIGURE 7. Pseudocrangonyx danyangensis sp. nov., holotype female, NNIBRIV69645. A–G, I, J; paratype male, NNIBRIV69646, H. A, antenna 1, medial view; B, accessory flagellum of antenna 1, medial view; C, antenna 2, medial view; D, upper lip, posterior view; E, left mandible, medial view; F, incisor, lacinia mobilis of right mandible, medial view; G, lower lip, dorsal view; H, maxilla 1, dorsal view; I, maxilla 2, dorsal view; J, maxilliped, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10954134" httpUri="https://zenodo.org/record/10954134/files/figure.png" pageId="9" pageNumber="258">Fig. 7C</figureCitation>
) 0.65 times as long as antenna 1; flagellum 0.69 times as long as peduncular articles 4 and 5 combined, consisting of 5 articles; calceoli absent.
</paragraph>
<paragraph id="220736CCFFD4F816FF401DB4FC4AFA4C" blockId="9.[151,1437,331,2014]" pageId="9" pageNumber="258">
Upper lip (
<figureCitation id="BA832A49FFD4F816FEC71DBBFE0AFB2C" box="[320,408,1159,1185]" captionStart="FIGURE 7" captionStartId="10.[151,250,1909,1933]" captionTargetBox="[189,1399,186,1884]" captionTargetId="figure-33@10.[189,1399,181,1884]" captionTargetPageId="10" captionText="FIGURE 7. Pseudocrangonyx danyangensis sp. nov., holotype female, NNIBRIV69645. A–G, I, J; paratype male, NNIBRIV69646, H. A, antenna 1, medial view; B, accessory flagellum of antenna 1, medial view; C, antenna 2, medial view; D, upper lip, posterior view; E, left mandible, medial view; F, incisor, lacinia mobilis of right mandible, medial view; G, lower lip, dorsal view; H, maxilla 1, dorsal view; I, maxilla 2, dorsal view; J, maxilliped, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10954134" httpUri="https://zenodo.org/record/10954134/files/figure.png" pageId="9" pageNumber="258">Fig. 7D</figureCitation>
) with rounded anterior margin, bearing fine setae. Mandibles (
<figureCitation id="BA832A49FFD4F816FBD41DBBFB57FB2C" box="[1107,1221,1159,1186]" captionStart="FIGURE 7" captionStartId="10.[151,250,1909,1933]" captionTargetBox="[189,1399,186,1884]" captionTargetId="figure-33@10.[189,1399,181,1884]" captionTargetPageId="10" captionText="FIGURE 7. Pseudocrangonyx danyangensis sp. nov., holotype female, NNIBRIV69645. A–G, I, J; paratype male, NNIBRIV69646, H. A, antenna 1, medial view; B, accessory flagellum of antenna 1, medial view; C, antenna 2, medial view; D, upper lip, posterior view; E, left mandible, medial view; F, incisor, lacinia mobilis of right mandible, medial view; G, lower lip, dorsal view; H, maxilla 1, dorsal view; I, maxilla 2, dorsal view; J, maxilliped, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10954134" httpUri="https://zenodo.org/record/10954134/files/figure.png" pageId="9" pageNumber="258">Fig. 7E, F</figureCitation>
) with left and right incisors with 6- and 4- dentate, respectively; left lacinia mobilis 5- dentate, right lacinia bifid, bearing many teeth; molar process triturative; palp 3-articulate, article 3 with 1 A-, 5 D-, and 4 E-setae. Lower lip (
<figureCitation id="BA832A49FFD4F816FB3B1DF3FA86FB64" box="[1212,1300,1231,1257]" captionStart="FIGURE 7" captionStartId="10.[151,250,1909,1933]" captionTargetBox="[189,1399,186,1884]" captionTargetId="figure-33@10.[189,1399,181,1884]" captionTargetPageId="10" captionText="FIGURE 7. Pseudocrangonyx danyangensis sp. nov., holotype female, NNIBRIV69645. A–G, I, J; paratype male, NNIBRIV69646, H. A, antenna 1, medial view; B, accessory flagellum of antenna 1, medial view; C, antenna 2, medial view; D, upper lip, posterior view; E, left mandible, medial view; F, incisor, lacinia mobilis of right mandible, medial view; G, lower lip, dorsal view; H, maxilla 1, dorsal view; I, maxilla 2, dorsal view; J, maxilliped, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10954134" httpUri="https://zenodo.org/record/10954134/files/figure.png" pageId="9" pageNumber="258">Fig. 7G</figureCitation>
) with broad outer lobes with fine setae, mandibular process of outer lobe rounded apically; inner lobes indistinct. Maxilla 1 (Male
<typeStatus id="FD03886EFFD4F816FF641C24FED4FABF" box="[227,326,1304,1330]" pageId="9" pageNumber="258" type="paratype">paratype</typeStatus>
,
<figureCitation id="BA832A49FFD4F816FED41C2BFE3EFABC" box="[339,428,1303,1329]" captionStart="FIGURE 7" captionStartId="10.[151,250,1909,1933]" captionTargetBox="[189,1399,186,1884]" captionTargetId="figure-33@10.[189,1399,181,1884]" captionTargetPageId="10" captionText="FIGURE 7. Pseudocrangonyx danyangensis sp. nov., holotype female, NNIBRIV69645. A–G, I, J; paratype male, NNIBRIV69646, H. A, antenna 1, medial view; B, accessory flagellum of antenna 1, medial view; C, antenna 2, medial view; D, upper lip, posterior view; E, left mandible, medial view; F, incisor, lacinia mobilis of right mandible, medial view; G, lower lip, dorsal view; H, maxilla 1, dorsal view; I, maxilla 2, dorsal view; J, maxilliped, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10954134" httpUri="https://zenodo.org/record/10954134/files/figure.png" pageId="9" pageNumber="258">Fig. 7H</figureCitation>
) with inner and outer plates, and palp; inner plate subovate with 3 plumose setae; outer plate subrectangular with 7 serrate teeth apically; palp 2-articulate, longer than outer plate, article 2 with 3 apical robust setae. Maxilla 2 (
<figureCitation id="BA832A49FFD4F816FE251C63FE7CFAF4" box="[418,494,1375,1401]" captionStart="FIGURE 7" captionStartId="10.[151,250,1909,1933]" captionTargetBox="[189,1399,186,1884]" captionTargetId="figure-33@10.[189,1399,181,1884]" captionTargetPageId="10" captionText="FIGURE 7. Pseudocrangonyx danyangensis sp. nov., holotype female, NNIBRIV69645. A–G, I, J; paratype male, NNIBRIV69646, H. A, antenna 1, medial view; B, accessory flagellum of antenna 1, medial view; C, antenna 2, medial view; D, upper lip, posterior view; E, left mandible, medial view; F, incisor, lacinia mobilis of right mandible, medial view; G, lower lip, dorsal view; H, maxilla 1, dorsal view; I, maxilla 2, dorsal view; J, maxilliped, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10954134" httpUri="https://zenodo.org/record/10954134/files/figure.png" pageId="9" pageNumber="258">Fig. 7I</figureCitation>
) with oblique inner row of 2 setae on inner plate. Maxilliped (
<figureCitation id="BA832A49FFD4F816FB271C63FB7CFAF4" box="[1184,1262,1375,1401]" captionStart="FIGURE 7" captionStartId="10.[151,250,1909,1933]" captionTargetBox="[189,1399,186,1884]" captionTargetId="figure-33@10.[189,1399,181,1884]" captionTargetPageId="10" captionText="FIGURE 7. Pseudocrangonyx danyangensis sp. nov., holotype female, NNIBRIV69645. A–G, I, J; paratype male, NNIBRIV69646, H. A, antenna 1, medial view; B, accessory flagellum of antenna 1, medial view; C, antenna 2, medial view; D, upper lip, posterior view; E, left mandible, medial view; F, incisor, lacinia mobilis of right mandible, medial view; G, lower lip, dorsal view; H, maxilla 1, dorsal view; I, maxilla 2, dorsal view; J, maxilliped, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10954134" httpUri="https://zenodo.org/record/10954134/files/figure.png" pageId="9" pageNumber="258">Fig. 7J</figureCitation>
) with inner and outer plates, and palp; inner plate with apical robust seta; outer plate with plumose seta and some medial setae; palp 4-articulate, medial margin of article 2 lined with setae, article 4 with nail.
</paragraph>
<paragraph id="220736CCFFD4F816FF401CF7FCA4F930" blockId="9.[151,1437,331,2014]" pageId="9" pageNumber="258">
Gnathopod 1 (
<figureCitation id="BA832A49FFD4F816FEE01CF7FE2CFA68" box="[359,446,1483,1509]" captionStart="FIGURE 8" captionStartId="11.[151,250,1909,1933]" captionTargetBox="[187,1399,186,1884]" captionTargetId="figure-17@11.[187,1399,181,1884]" captionTargetPageId="11" captionText="FIGURE 8. Pseudocrangonyx danyangensis sp. nov., holotype female, NNIBRIV69645. A, gnathopod 1, lateral view; B, gnathopod 2, lateral view; C, pereopod 3, lateral view; D, dactylus of pereopod 3, lateral view; E, pereopod 4, lateral view; F, dactylus of pereopod 4, lateral view; G, pereopod 5, lateral view; H, dactylus of pereopod 5, lateral view; I, pereopod 6, lateral view; J, dactylus of pereopod 6, lateral view; K, pereopod 7, lateral view; L, dactylus of pereopod 7, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10954136" httpUri="https://zenodo.org/record/10954136/files/figure.png" pageId="9" pageNumber="258">Fig. 8A</figureCitation>
) with subquadrate coxa, bearing setae on anterodistal corner and margin, width 1.8 times as long as depth; posterior margin of basis with 3 setae; posterodistal corner of carpus with slender setae, some weakly pectinate; propodus stout, subtriangular, palmar margin with 5 robust setae, some distally notched; posterior margin of dactylus dentate. Gnathopod 2 (
<figureCitation id="BA832A49FFD4F816FDE81F0BFD57F9DC" box="[623,709,1591,1617]" captionStart="FIGURE 8" captionStartId="11.[151,250,1909,1933]" captionTargetBox="[187,1399,186,1884]" captionTargetId="figure-17@11.[187,1399,181,1884]" captionTargetPageId="11" captionText="FIGURE 8. Pseudocrangonyx danyangensis sp. nov., holotype female, NNIBRIV69645. A, gnathopod 1, lateral view; B, gnathopod 2, lateral view; C, pereopod 3, lateral view; D, dactylus of pereopod 3, lateral view; E, pereopod 4, lateral view; F, dactylus of pereopod 4, lateral view; G, pereopod 5, lateral view; H, dactylus of pereopod 5, lateral view; I, pereopod 6, lateral view; J, dactylus of pereopod 6, lateral view; K, pereopod 7, lateral view; L, dactylus of pereopod 7, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10954136" httpUri="https://zenodo.org/record/10954136/files/figure.png" pageId="9" pageNumber="258">Fig. 8B</figureCitation>
) with subquadrate coxa, bearing setae on anterodistal corner and posteroventral margin, width 1.6 times as long as depth; posterior margin of basis with 4 setae; posterodistal corner of carpus with slender setae, some weakly pectinate; propodus stout, subchelate, palmar margin with 7 robust setae, some distally notched; posterior margin of dactylus dentate.
</paragraph>
<paragraph id="220736CCFFD4F816FF401FFBFB6BF850" blockId="9.[151,1437,331,2014]" pageId="9" pageNumber="258">
Pereopod 3 (
<figureCitation id="BA832A49FFD4F816FED41FFBFE35F96C" box="[339,423,1735,1761]" captionStart="FIGURE 8" captionStartId="11.[151,250,1909,1933]" captionTargetBox="[187,1399,186,1884]" captionTargetId="figure-17@11.[187,1399,181,1884]" captionTargetPageId="11" captionText="FIGURE 8. Pseudocrangonyx danyangensis sp. nov., holotype female, NNIBRIV69645. A, gnathopod 1, lateral view; B, gnathopod 2, lateral view; C, pereopod 3, lateral view; D, dactylus of pereopod 3, lateral view; E, pereopod 4, lateral view; F, dactylus of pereopod 4, lateral view; G, pereopod 5, lateral view; H, dactylus of pereopod 5, lateral view; I, pereopod 6, lateral view; J, dactylus of pereopod 6, lateral view; K, pereopod 7, lateral view; L, dactylus of pereopod 7, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10954136" httpUri="https://zenodo.org/record/10954136/files/figure.png" pageId="9" pageNumber="258">Fig. 8C</figureCitation>
) with subquadrate coxa bearing setae on anterior to posteroventral margins, width 1.6 times as long as depth; anterior and posterior margins of basis with 2 and 4 setae, respectively; merus, carpus, and propodus in length ratio of 1.0: 0.7: 0.8; posterior margin and submargin of dactylus each with seta (
<figureCitation id="BA832A49FFD4F816FBFC1E33FB40F8A4" box="[1147,1234,1807,1833]" captionStart="FIGURE 8" captionStartId="11.[151,250,1909,1933]" captionTargetBox="[187,1399,186,1884]" captionTargetId="figure-17@11.[187,1399,181,1884]" captionTargetPageId="11" captionText="FIGURE 8. Pseudocrangonyx danyangensis sp. nov., holotype female, NNIBRIV69645. A, gnathopod 1, lateral view; B, gnathopod 2, lateral view; C, pereopod 3, lateral view; D, dactylus of pereopod 3, lateral view; E, pereopod 4, lateral view; F, dactylus of pereopod 4, lateral view; G, pereopod 5, lateral view; H, dactylus of pereopod 5, lateral view; I, pereopod 6, lateral view; J, dactylus of pereopod 6, lateral view; K, pereopod 7, lateral view; L, dactylus of pereopod 7, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10954136" httpUri="https://zenodo.org/record/10954136/files/figure.png" pageId="9" pageNumber="258">Fig. 8D</figureCitation>
). Pereopod 4 (
<figureCitation id="BA832A49FFD4F816FAE81E33FF25F8C0" captionStart="FIGURE 8" captionStartId="11.[151,250,1909,1933]" captionTargetBox="[187,1399,186,1884]" captionTargetId="figure-17@11.[187,1399,181,1884]" captionTargetPageId="11" captionText="FIGURE 8. Pseudocrangonyx danyangensis sp. nov., holotype female, NNIBRIV69645. A, gnathopod 1, lateral view; B, gnathopod 2, lateral view; C, pereopod 3, lateral view; D, dactylus of pereopod 3, lateral view; E, pereopod 4, lateral view; F, dactylus of pereopod 4, lateral view; G, pereopod 5, lateral view; H, dactylus of pereopod 5, lateral view; I, pereopod 6, lateral view; J, dactylus of pereopod 6, lateral view; K, pereopod 7, lateral view; L, dactylus of pereopod 7, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10954136" httpUri="https://zenodo.org/record/10954136/files/figure.png" pageId="9" pageNumber="258">Fig. 8E</figureCitation>
) with subquadrate coxa bearing setae on anterior to ventral corner, width 1.86 times as long as depth; anterior and posterior margins of basis each with 3 and 4 setae; merus, carpus, and propodus in length ratio of 1.0: 0.8: 0.8; posterior margin and submargin of dactylus each with seta (
<figureCitation id="BA832A49FFD4F816FCAE1E47FCEFF818" box="[809,893,1915,1941]" captionStart="FIGURE 8" captionStartId="11.[151,250,1909,1933]" captionTargetBox="[187,1399,186,1884]" captionTargetId="figure-17@11.[187,1399,181,1884]" captionTargetPageId="11" captionText="FIGURE 8. Pseudocrangonyx danyangensis sp. nov., holotype female, NNIBRIV69645. A, gnathopod 1, lateral view; B, gnathopod 2, lateral view; C, pereopod 3, lateral view; D, dactylus of pereopod 3, lateral view; E, pereopod 4, lateral view; F, dactylus of pereopod 4, lateral view; G, pereopod 5, lateral view; H, dactylus of pereopod 5, lateral view; I, pereopod 6, lateral view; J, dactylus of pereopod 6, lateral view; K, pereopod 7, lateral view; L, dactylus of pereopod 7, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10954136" httpUri="https://zenodo.org/record/10954136/files/figure.png" pageId="9" pageNumber="258">Fig. 8F</figureCitation>
). Pereopod 5 (
<figureCitation id="BA832A49FFD4F816FB9C1E47FBE3F818" box="[1051,1137,1915,1941]" captionStart="FIGURE 8" captionStartId="11.[151,250,1909,1933]" captionTargetBox="[187,1399,186,1884]" captionTargetId="figure-17@11.[187,1399,181,1884]" captionTargetPageId="11" captionText="FIGURE 8. Pseudocrangonyx danyangensis sp. nov., holotype female, NNIBRIV69645. A, gnathopod 1, lateral view; B, gnathopod 2, lateral view; C, pereopod 3, lateral view; D, dactylus of pereopod 3, lateral view; E, pereopod 4, lateral view; F, dactylus of pereopod 4, lateral view; G, pereopod 5, lateral view; H, dactylus of pereopod 5, lateral view; I, pereopod 6, lateral view; J, dactylus of pereopod 6, lateral view; K, pereopod 7, lateral view; L, dactylus of pereopod 7, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10954136" httpUri="https://zenodo.org/record/10954136/files/figure.png" pageId="9" pageNumber="258">Fig. 8G</figureCitation>
) with weakly bilobed coxa bearing setae on anterior to posterior lobes; anterior and posterior margins of basis with 3 setae, respectively; merus, carpus, and propodus in length ratio of 1.0: 0.7: 0.8; anterior margin of dactylus with seta (
<figureCitation id="BA832A49FFD4F816FB141EFFFB7EF850" box="[1171,1260,1987,2013]" captionStart="FIGURE 8" captionStartId="11.[151,250,1909,1933]" captionTargetBox="[187,1399,186,1884]" captionTargetId="figure-17@11.[187,1399,181,1884]" captionTargetPageId="11" captionText="FIGURE 8. Pseudocrangonyx danyangensis sp. nov., holotype female, NNIBRIV69645. A, gnathopod 1, lateral view; B, gnathopod 2, lateral view; C, pereopod 3, lateral view; D, dactylus of pereopod 3, lateral view; E, pereopod 4, lateral view; F, dactylus of pereopod 4, lateral view; G, pereopod 5, lateral view; H, dactylus of pereopod 5, lateral view; I, pereopod 6, lateral view; J, dactylus of pereopod 6, lateral view; K, pereopod 7, lateral view; L, dactylus of pereopod 7, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10954136" httpUri="https://zenodo.org/record/10954136/files/figure.png" pageId="9" pageNumber="258">Fig. 8H</figureCitation>
).
</paragraph>
<caption id="76C76644FFD7F815FF101E49FBF4F874" ID-DOI="http://doi.org/10.5281/zenodo.10954134" ID-Zenodo-Dep="10954134" httpUri="https://zenodo.org/record/10954134/files/figure.png" pageId="10" pageNumber="259" startId="10.[151,250,1909,1933]" targetBox="[189,1399,186,1884]" targetPageId="10" targetType="figure">
<paragraph id="220736CCFFD7F815FF101E49FBF4F874" blockId="10.[151,1437,1909,2041]" pageId="10" pageNumber="259">
<emphasis id="10CCEADEFFD7F815FF101E49FE8EF800" bold="true" box="[151,284,1909,1933]" pageId="10" pageNumber="259">FIGURE 7.</emphasis>
<taxonomicName id="E5B84D4FFFD7F815FEAC1E49FDE8F800" authority="Lee &amp; Min, 2024" authorityName="Lee &amp; Min" authorityYear="2024" box="[299,634,1909,1933]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="259" phylum="Arthropoda" rank="species" species="danyangensis" status="sp. nov.">
<emphasis id="10CCEADEFFD7F815FEAC1E49FDE8F800" box="[299,634,1909,1933]" italics="true" pageId="10" pageNumber="259">Pseudocrangonyx danyangensis</emphasis>
</taxonomicName>
<emphasis id="10CCEADEFFD7F815FD0E1E49FD71F800" bold="true" box="[649,739,1909,1933]" pageId="10" pageNumber="259">
<taxonomicNameLabel id="0BFF57A5FFD7F815FD0E1E49FD71F800" box="[649,739,1909,1933]" pageId="10" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype female, NNIBRIV69645. A–G, I, J; paratype male, NNIBRIV69646, H. A, antenna 1, medial view; B, accessory flagellum of antenna 1, medial view; C, antenna 2, medial view; D, upper lip, posterior view; E, left mandible, medial view; F, incisor, lacinia mobilis of right mandible, medial view; G, lower lip, dorsal view; H, maxilla 1, dorsal view; I, maxilla 2, dorsal view; J, maxilliped, ventral view.
</paragraph>
</caption>
<caption id="76C76644FFD6F814FF101E49FA81F874" ID-DOI="http://doi.org/10.5281/zenodo.10954136" ID-Zenodo-Dep="10954136" httpUri="https://zenodo.org/record/10954136/files/figure.png" pageId="11" pageNumber="260" startId="11.[151,250,1909,1933]" targetBox="[187,1399,186,1884]" targetPageId="11" targetType="figure">
<paragraph id="220736CCFFD6F814FF101E49FA81F874" blockId="11.[151,1437,1909,2041]" pageId="11" pageNumber="260">
<emphasis id="10CCEADEFFD6F814FF101E49FE84F800" bold="true" box="[151,278,1909,1933]" pageId="11" pageNumber="260">FIGURE 8.</emphasis>
<taxonomicName id="E5B84D4FFFD6F814FEA71E49FDFBF800" authority="Lee &amp; Min, 2024" authorityName="Lee &amp; Min" authorityYear="2024" box="[288,617,1909,1933]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" kingdom="Animalia" order="Amphipoda" pageId="11" pageNumber="260" phylum="Arthropoda" rank="species" species="danyangensis" status="sp. nov.">
<emphasis id="10CCEADEFFD6F814FEA71E49FDFBF800" box="[288,617,1909,1933]" italics="true" pageId="11" pageNumber="260">Pseudocrangonyx danyangensis</emphasis>
</taxonomicName>
<emphasis id="10CCEADEFFD6F814FDF41E49FD55F800" bold="true" box="[627,711,1909,1933]" pageId="11" pageNumber="260">
<taxonomicNameLabel id="0BFF57A5FFD6F814FDF41E49FD55F800" box="[627,711,1909,1933]" pageId="11" pageNumber="260" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype female, NNIBRIV69645. A, gnathopod 1, lateral view; B, gnathopod 2, lateral view; C, pereopod 3, lateral view; D, dactylus of pereopod 3, lateral view; E, pereopod 4, lateral view; F, dactylus of pereopod 4, lateral view; G, pereopod 5, lateral view; H, dactylus of pereopod 5, lateral view; I, pereopod 6, lateral view; J, dactylus of pereopod 6, lateral view; K, pereopod 7, lateral view; L, dactylus of pereopod 7, lateral view.
</paragraph>
</caption>
<paragraph id="220736CCFFD1F813FF1019ABFC91FECC" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="261">
Pereopod 6 (
<figureCitation id="BA832A49FFD1F813FEAE19ABFEE4FF3C" box="[297,374,151,177]" captionStart="FIGURE 8" captionStartId="11.[151,250,1909,1933]" captionTargetBox="[187,1399,186,1884]" captionTargetId="figure-17@11.[187,1399,181,1884]" captionTargetPageId="11" captionText="FIGURE 8. Pseudocrangonyx danyangensis sp. nov., holotype female, NNIBRIV69645. A, gnathopod 1, lateral view; B, gnathopod 2, lateral view; C, pereopod 3, lateral view; D, dactylus of pereopod 3, lateral view; E, pereopod 4, lateral view; F, dactylus of pereopod 4, lateral view; G, pereopod 5, lateral view; H, dactylus of pereopod 5, lateral view; I, pereopod 6, lateral view; J, dactylus of pereopod 6, lateral view; K, pereopod 7, lateral view; L, dactylus of pereopod 7, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10954136" httpUri="https://zenodo.org/record/10954136/files/figure.png" pageId="12" pageNumber="261">Fig. 8I</figureCitation>
) with weakly bilobed coxa bearing setae on posterior lobe; anterior and posterior margins of basis with 4 and 2 setae, respectively; merus, carpus, and propodus in length ratio of 1.0: 0.7: 0.9; anterior margin of dactylus with seta (
<figureCitation id="BA832A49FFD1F813FE0019E3FE47FF74" box="[391,469,223,249]" captionStart="FIGURE 8" captionStartId="11.[151,250,1909,1933]" captionTargetBox="[187,1399,186,1884]" captionTargetId="figure-17@11.[187,1399,181,1884]" captionTargetPageId="11" captionText="FIGURE 8. Pseudocrangonyx danyangensis sp. nov., holotype female, NNIBRIV69645. A, gnathopod 1, lateral view; B, gnathopod 2, lateral view; C, pereopod 3, lateral view; D, dactylus of pereopod 3, lateral view; E, pereopod 4, lateral view; F, dactylus of pereopod 4, lateral view; G, pereopod 5, lateral view; H, dactylus of pereopod 5, lateral view; I, pereopod 6, lateral view; J, dactylus of pereopod 6, lateral view; K, pereopod 7, lateral view; L, dactylus of pereopod 7, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10954136" httpUri="https://zenodo.org/record/10954136/files/figure.png" pageId="12" pageNumber="261">Fig. 8J</figureCitation>
). Pereopod 7 (
<figureCitation id="BA832A49FFD1F813FDF319E3FD5BFF74" box="[628,713,223,249]" captionStart="FIGURE 8" captionStartId="11.[151,250,1909,1933]" captionTargetBox="[187,1399,186,1884]" captionTargetId="figure-17@11.[187,1399,181,1884]" captionTargetPageId="11" captionText="FIGURE 8. Pseudocrangonyx danyangensis sp. nov., holotype female, NNIBRIV69645. A, gnathopod 1, lateral view; B, gnathopod 2, lateral view; C, pereopod 3, lateral view; D, dactylus of pereopod 3, lateral view; E, pereopod 4, lateral view; F, dactylus of pereopod 4, lateral view; G, pereopod 5, lateral view; H, dactylus of pereopod 5, lateral view; I, pereopod 6, lateral view; J, dactylus of pereopod 6, lateral view; K, pereopod 7, lateral view; L, dactylus of pereopod 7, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10954136" httpUri="https://zenodo.org/record/10954136/files/figure.png" pageId="12" pageNumber="261">Fig. 8K</figureCitation>
) with subtriangular coxa, bearing seta on posteroproximal corner; anterior and posterior margins of basis with 3 setae, respectively; merus, carpus, and propodus in length ratio of 1.0: 0.9: 1.1; anterior margin of dactylus with seta (
<figureCitation id="BA832A49FFD1F813FD26181BFD64FECC" box="[673,758,295,321]" captionStart="FIGURE 8" captionStartId="11.[151,250,1909,1933]" captionTargetBox="[187,1399,186,1884]" captionTargetId="figure-17@11.[187,1399,181,1884]" captionTargetPageId="11" captionText="FIGURE 8. Pseudocrangonyx danyangensis sp. nov., holotype female, NNIBRIV69645. A, gnathopod 1, lateral view; B, gnathopod 2, lateral view; C, pereopod 3, lateral view; D, dactylus of pereopod 3, lateral view; E, pereopod 4, lateral view; F, dactylus of pereopod 4, lateral view; G, pereopod 5, lateral view; H, dactylus of pereopod 5, lateral view; I, pereopod 6, lateral view; J, dactylus of pereopod 6, lateral view; K, pereopod 7, lateral view; L, dactylus of pereopod 7, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10954136" httpUri="https://zenodo.org/record/10954136/files/figure.png" pageId="12" pageNumber="261">Fig. 8L</figureCitation>
).
</paragraph>
<paragraph id="220736CCFFD1F813FF401877FC1CFE04" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="261">
Coxal gills (
<figureCitation id="BA832A49FFD1F813FECA1877FD84FEE8" box="[333,534,331,357]" captionStart="FIGURE 8" captionStartId="11.[151,250,1909,1933]" captionTargetBox="[187,1399,186,1884]" captionTargetId="figure-17@11.[187,1399,181,1884]" captionTargetPageId="11" captionText="FIGURE 8. Pseudocrangonyx danyangensis sp. nov., holotype female, NNIBRIV69645. A, gnathopod 1, lateral view; B, gnathopod 2, lateral view; C, pereopod 3, lateral view; D, dactylus of pereopod 3, lateral view; E, pereopod 4, lateral view; F, dactylus of pereopod 4, lateral view; G, pereopod 5, lateral view; H, dactylus of pereopod 5, lateral view; I, pereopod 6, lateral view; J, dactylus of pereopod 6, lateral view; K, pereopod 7, lateral view; L, dactylus of pereopod 7, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10954136" httpUri="https://zenodo.org/record/10954136/files/figure.png" pageId="12" pageNumber="261">Fig. 8B, C, E, G, I</figureCitation>
) on gnathopod 2 and pereopods 3–6; sternal gills absent. Brood plates (
<figureCitation id="BA832A49FFD1F813FAA31877FF3EFE04" captionStart="FIGURE 8" captionStartId="11.[151,250,1909,1933]" captionTargetBox="[187,1399,186,1884]" captionTargetId="figure-17@11.[187,1399,181,1884]" captionTargetPageId="11" captionText="FIGURE 8. Pseudocrangonyx danyangensis sp. nov., holotype female, NNIBRIV69645. A, gnathopod 1, lateral view; B, gnathopod 2, lateral view; C, pereopod 3, lateral view; D, dactylus of pereopod 3, lateral view; E, pereopod 4, lateral view; F, dactylus of pereopod 4, lateral view; G, pereopod 5, lateral view; H, dactylus of pereopod 5, lateral view; I, pereopod 6, lateral view; J, dactylus of pereopod 6, lateral view; K, pereopod 7, lateral view; L, dactylus of pereopod 7, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10954136" httpUri="https://zenodo.org/record/10954136/files/figure.png" pageId="12" pageNumber="261">Fig. 8C, E, G</figureCitation>
) slender, with numerous setae, on gnathopod 2 and pereopods 3–5.
</paragraph>
<paragraph id="220736CCFFD1F813FF4018AFFF7BFD94" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="261">
Peduncles of pleopod 1 and 2 (
<figureCitation id="BA832A49FFD1F813FDA318AFFD0FFE20" box="[548,669,403,429]" captionStart="FIGURE 9" captionStartId="13.[151,250,1909,1933]" captionTargetBox="[222,1366,187,1884]" captionTargetId="figure-17@13.[222,1366,181,1884]" captionTargetPageId="13" captionText="FIGURE 9. Pseudocrangonyx danyangensis sp. nov., holotype female, NNIBRIV69645. A–C, Pleopod 1–3, lateral view; D, uropod 1, ventral view; E, uropod 2, dorsal view; F, uropod 3, dorsal view; G, telson, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10954140" httpUri="https://zenodo.org/record/10954140/files/figure.png" pageId="12" pageNumber="261">Fig. 9A, B</figureCitation>
) anterodistal corner with seta; peduncle of pleopod 3 (
<figureCitation id="BA832A49FFD1F813FA8218AFFAC8FE20" box="[1285,1370,403,429]" captionStart="FIGURE 9" captionStartId="13.[151,250,1909,1933]" captionTargetBox="[222,1366,187,1884]" captionTargetId="figure-17@13.[222,1366,181,1884]" captionTargetPageId="13" captionText="FIGURE 9. Pseudocrangonyx danyangensis sp. nov., holotype female, NNIBRIV69645. A–C, Pleopod 1–3, lateral view; D, uropod 1, ventral view; E, uropod 2, dorsal view; F, uropod 3, dorsal view; G, telson, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10954140" httpUri="https://zenodo.org/record/10954140/files/figure.png" pageId="12" pageNumber="261">Fig. 9C</figureCitation>
) with marginal setae. Pleopods 1–3 with paired retinacula, and with bifid seta (clothes-pin setae) on inner basal margin of inner ramus; pleopods 1–3 inner ramus 4-, 3-, and 3-articulate; pleopods 1–3 outer ramus 5-, 4-, and 3-articulate (
<figureCitation id="BA832A49FFD1F813FAF718E7FF4FFD94" captionStart="FIGURE 9" captionStartId="13.[151,250,1909,1933]" captionTargetBox="[222,1366,187,1884]" captionTargetId="figure-17@13.[222,1366,181,1884]" captionTargetPageId="13" captionText="FIGURE 9. Pseudocrangonyx danyangensis sp. nov., holotype female, NNIBRIV69645. A–C, Pleopod 1–3, lateral view; D, uropod 1, ventral view; E, uropod 2, dorsal view; F, uropod 3, dorsal view; G, telson, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10954140" httpUri="https://zenodo.org/record/10954140/files/figure.png" pageId="12" pageNumber="261">Fig. 9A–C</figureCitation>
).
</paragraph>
<paragraph id="220736CCFFD1F813FF401B1FFB5FFC98" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="261">
Uropod 1 (
<figureCitation id="BA832A49FFD1F813FEBA1B1FFE00FDB0" box="[317,402,547,573]" captionStart="FIGURE 9" captionStartId="13.[151,250,1909,1933]" captionTargetBox="[222,1366,187,1884]" captionTargetId="figure-17@13.[222,1366,181,1884]" captionTargetPageId="13" captionText="FIGURE 9. Pseudocrangonyx danyangensis sp. nov., holotype female, NNIBRIV69645. A–C, Pleopod 1–3, lateral view; D, uropod 1, ventral view; E, uropod 2, dorsal view; F, uropod 3, dorsal view; G, telson, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10954140" httpUri="https://zenodo.org/record/10954140/files/figure.png" pageId="12" pageNumber="261">Fig. 9D</figureCitation>
) with 2 basofacial setae on peduncle; inner ramus 0.74 times as long as peduncle, inner margin with 2 robust setae, outer margin bare, basal part with 2 slender setae; outer ramus 0.67 times as long as inner, inner margin and marginal with robust seta, respectively. Uropod 2 (
<figureCitation id="BA832A49FFD1F813FCC21B57FC05FD08" box="[837,919,619,645]" captionStart="FIGURE 9" captionStartId="13.[151,250,1909,1933]" captionTargetBox="[222,1366,187,1884]" captionTargetId="figure-17@13.[222,1366,181,1884]" captionTargetPageId="13" captionText="FIGURE 9. Pseudocrangonyx danyangensis sp. nov., holotype female, NNIBRIV69645. A–C, Pleopod 1–3, lateral view; D, uropod 1, ventral view; E, uropod 2, dorsal view; F, uropod 3, dorsal view; G, telson, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10954140" httpUri="https://zenodo.org/record/10954140/files/figure.png" pageId="12" pageNumber="261">Fig. 9E</figureCitation>
) with inner and outer rami; peduncle outer and inner margin with 6 and 2 robust setae, respectively; inner ramus 0.76 times as long as peduncle, inner margin with 2 robust setae; outer ramus 0.73 times as long as inner ramus, marginally bare. Uropod 3 (
<figureCitation id="BA832A49FFD1F813FBEF1B8FFB28FD40" box="[1128,1210,691,717]" captionStart="FIGURE 9" captionStartId="13.[151,250,1909,1933]" captionTargetBox="[222,1366,187,1884]" captionTargetId="figure-17@13.[222,1366,181,1884]" captionTargetPageId="13" captionText="FIGURE 9. Pseudocrangonyx danyangensis sp. nov., holotype female, NNIBRIV69645. A–C, Pleopod 1–3, lateral view; D, uropod 1, ventral view; E, uropod 2, dorsal view; F, uropod 3, dorsal view; G, telson, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10954140" httpUri="https://zenodo.org/record/10954140/files/figure.png" pageId="12" pageNumber="261">Fig. 9F</figureCitation>
) with peduncle 0.32 times as long as outer ramus, with 2 robust and 1 slender setae; inner ramus absent; outer ramus 2-articulate, proximal article with robust setae, terminal article 0.38 times as long as proximal article, with 3 distal setae.
</paragraph>
<paragraph id="220736CCFFD1F813FF401A23FD29FCD0" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="261">
Telson (
<figureCitation id="BA832A49FFD1F813FEA61A23FEEAFCB4" box="[289,376,799,825]" captionStart="FIGURE 9" captionStartId="13.[151,250,1909,1933]" captionTargetBox="[222,1366,187,1884]" captionTargetId="figure-17@13.[222,1366,181,1884]" captionTargetPageId="13" captionText="FIGURE 9. Pseudocrangonyx danyangensis sp. nov., holotype female, NNIBRIV69645. A–C, Pleopod 1–3, lateral view; D, uropod 1, ventral view; E, uropod 2, dorsal view; F, uropod 3, dorsal view; G, telson, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10954140" httpUri="https://zenodo.org/record/10954140/files/figure.png" pageId="12" pageNumber="261">Fig. 9G</figureCitation>
) length 1.53 times as long as wide, almost not cleft, each telson lobe with 2 lateral penicillate setae, 1 apical robust and 1 apical penicillate seta.
</paragraph>
<paragraph id="220736CCFFD1F813FF401A5BFEA0FC60" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="261">
Male
<typeStatus id="FD03886EFFD1F813FE891A5BFEE2FC0C" box="[270,368,871,897]" pageId="12" pageNumber="261" type="paratype">paratype</typeStatus>
(NNIBRIV69646): Antenna 1 (
<figureCitation id="BA832A49FFD1F813FD681A5BFC13FC0C" box="[751,897,871,897]" captionStart="FIGURE 10" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[159,1428,181,1919]" captionTargetId="figure-33@14.[159,1428,181,1919]" captionTargetPageId="14" captionText="FIGURE 10. Pseudocrangonyx danyangensis sp. nov., paratype male, NNIBRIV69646.A, antenna 1, medial view; B, accessory flagellum of antenna 1, medial view; C, antenna 2, lateral view; D, gnathopod 1, lateral view; E, gnathopod 2, lateral view; F, uropod 1, ventral view; G, uropod 2, ventral view; H, uropod 3, dorsal view; I, telson, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10954142" httpUri="https://zenodo.org/record/10954142/files/figure.png" pageId="12" pageNumber="261">Fig. 10A, B</figureCitation>
) 0.34 times as long as body length, primary flagellum 10-articulate, 1 aesthetasc on some articles. Antenna 2 (
<figureCitation id="BA832A49FFD1F813FC251AB7FB99FC28" box="[930,1035,907,933]" captionStart="FIGURE 10" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[159,1428,181,1919]" captionTargetId="figure-33@14.[159,1428,181,1919]" captionTargetPageId="14" captionText="FIGURE 10. Pseudocrangonyx danyangensis sp. nov., paratype male, NNIBRIV69646.A, antenna 1, medial view; B, accessory flagellum of antenna 1, medial view; C, antenna 2, lateral view; D, gnathopod 1, lateral view; E, gnathopod 2, lateral view; F, uropod 1, ventral view; G, uropod 2, ventral view; H, uropod 3, dorsal view; I, telson, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10954142" httpUri="https://zenodo.org/record/10954142/files/figure.png" pageId="12" pageNumber="261">Fig. 10C</figureCitation>
) 0.73 times as long as antenna 1; flagellum 0.66 times as long as peduncular articles 4 and 5 combined, consisting of 4 articles, first 2 of which with calceoli.
</paragraph>
<paragraph id="220736CCFFD1F813FF401ACBFD32FBD4" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="261">
Gnathopod 1 (
<figureCitation id="BA832A49FFD1F813FEEE1ACBFE5DFB9C" box="[361,463,1015,1041]" captionStart="FIGURE 10" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[159,1428,181,1919]" captionTargetId="figure-33@14.[159,1428,181,1919]" captionTargetPageId="14" captionText="FIGURE 10. Pseudocrangonyx danyangensis sp. nov., paratype male, NNIBRIV69646.A, antenna 1, medial view; B, accessory flagellum of antenna 1, medial view; C, antenna 2, lateral view; D, gnathopod 1, lateral view; E, gnathopod 2, lateral view; F, uropod 1, ventral view; G, uropod 2, ventral view; H, uropod 3, dorsal view; I, telson, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10954142" httpUri="https://zenodo.org/record/10954142/files/figure.png" pageId="12" pageNumber="261">Fig. 10D</figureCitation>
) with coxa width 1.94 times as long as depth; palmar margin with 6 robust setae in 2 rows, some distally notched. Gnathopod 2 (
<figureCitation id="BA832A49FFD1F813FDF31D27FD47FBB8" box="[628,725,1051,1077]" captionStart="FIGURE 10" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[159,1428,181,1919]" captionTargetId="figure-33@14.[159,1428,181,1919]" captionTargetPageId="14" captionText="FIGURE 10. Pseudocrangonyx danyangensis sp. nov., paratype male, NNIBRIV69646.A, antenna 1, medial view; B, accessory flagellum of antenna 1, medial view; C, antenna 2, lateral view; D, gnathopod 1, lateral view; E, gnathopod 2, lateral view; F, uropod 1, ventral view; G, uropod 2, ventral view; H, uropod 3, dorsal view; I, telson, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10954142" httpUri="https://zenodo.org/record/10954142/files/figure.png" pageId="12" pageNumber="261">Fig. 10E</figureCitation>
) with coxa width 1.6 times as long as depth; palmar margin with 7 robust setae in 2 rows, some distally notched.
</paragraph>
<paragraph id="220736CCFFD1F813FF401D5FFCCAFAD8" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="261">
Uropod 1 (
<figureCitation id="BA832A49FFD1F813FEC71D5FFE32FBF0" box="[320,416,1123,1149]" captionStart="FIGURE 10" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[159,1428,181,1919]" captionTargetId="figure-33@14.[159,1428,181,1919]" captionTargetPageId="14" captionText="FIGURE 10. Pseudocrangonyx danyangensis sp. nov., paratype male, NNIBRIV69646.A, antenna 1, medial view; B, accessory flagellum of antenna 1, medial view; C, antenna 2, lateral view; D, gnathopod 1, lateral view; E, gnathopod 2, lateral view; F, uropod 1, ventral view; G, uropod 2, ventral view; H, uropod 3, dorsal view; I, telson, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10954142" httpUri="https://zenodo.org/record/10954142/files/figure.png" pageId="12" pageNumber="261">Fig. 10F</figureCitation>
) with basofacial seta on peduncle; inner ramus 0.78 times as long as peduncle, outer margin with 2 robust setae, inner margin bare, basal part with slender seta; outer ramus 0.71 times as long as inner ramus, inner margin with robust seta. Uropod 2 (
<figureCitation id="BA832A49FFD1F813FDE61D97FD54FB48" box="[609,710,1195,1221]" captionStart="FIGURE 10" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[159,1428,181,1919]" captionTargetId="figure-33@14.[159,1428,181,1919]" captionTargetPageId="14" captionText="FIGURE 10. Pseudocrangonyx danyangensis sp. nov., paratype male, NNIBRIV69646.A, antenna 1, medial view; B, accessory flagellum of antenna 1, medial view; C, antenna 2, lateral view; D, gnathopod 1, lateral view; E, gnathopod 2, lateral view; F, uropod 1, ventral view; G, uropod 2, ventral view; H, uropod 3, dorsal view; I, telson, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10954142" httpUri="https://zenodo.org/record/10954142/files/figure.png" pageId="12" pageNumber="261">Fig. 10G</figureCitation>
) with inner and outer rami; peduncle outer and inner margin with 7 and 2 robust setae, respectively; inner ramus 0.75 times as long as peduncle, inner and outer margins with 1 and 2 robust setae, respectively; outer ramus 0.83 times as long as inner ramus. Uropod 3 (
<figureCitation id="BA832A49FFD1F813FBD61DCFFB24FA80" box="[1105,1206,1267,1293]" captionStart="FIGURE 10" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[159,1428,181,1919]" captionTargetId="figure-33@14.[159,1428,181,1919]" captionTargetPageId="14" captionText="FIGURE 10. Pseudocrangonyx danyangensis sp. nov., paratype male, NNIBRIV69646.A, antenna 1, medial view; B, accessory flagellum of antenna 1, medial view; C, antenna 2, lateral view; D, gnathopod 1, lateral view; E, gnathopod 2, lateral view; F, uropod 1, ventral view; G, uropod 2, ventral view; H, uropod 3, dorsal view; I, telson, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10954142" httpUri="https://zenodo.org/record/10954142/files/figure.png" pageId="12" pageNumber="261">Fig. 10H</figureCitation>
) with peduncle 0.32 times as long as outer ramus; inner ramus absent; outer ramus 2-articulate, proximal article with robust setae, terminal article 0.39 times as long as proximal article, with 3 distal setae.
</paragraph>
<paragraph id="220736CCFFD1F813FF401C63FBF9FAF4" blockId="12.[151,1437,151,2013]" box="[199,1131,1374,1401]" pageId="12" pageNumber="261">
Telson (
<figureCitation id="BA832A49FFD1F813FE981C63FEEAFAF4" box="[287,376,1375,1401]" captionStart="FIGURE 10" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[159,1428,181,1919]" captionTargetId="figure-33@14.[159,1428,181,1919]" captionTargetPageId="14" captionText="FIGURE 10. Pseudocrangonyx danyangensis sp. nov., paratype male, NNIBRIV69646.A, antenna 1, medial view; B, accessory flagellum of antenna 1, medial view; C, antenna 2, lateral view; D, gnathopod 1, lateral view; E, gnathopod 2, lateral view; F, uropod 1, ventral view; G, uropod 2, ventral view; H, uropod 3, dorsal view; I, telson, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10954142" httpUri="https://zenodo.org/record/10954142/files/figure.png" pageId="12" pageNumber="261">Fig. 10I</figureCitation>
) length 1.45 times as long as width, subrectangular, almost not cleft.
</paragraph>
</subSubSection>
<subSubSection id="6AA26547FFD1F813FF401CBEFD74FA10" box="[199,742,1410,1437]" pageId="12" pageNumber="261" type="distribution">
<paragraph id="220736CCFFD1F813FF401CBEFD74FA10" blockId="12.[151,1437,151,2013]" box="[199,742,1410,1437]" pageId="12" pageNumber="261">
<emphasis id="10CCEADEFFD1F813FF401CBEFECCFA11" bold="true" box="[199,350,1410,1436]" pageId="12" pageNumber="261">Distribution.</emphasis>
Known only from the
<typeStatus id="FD03886EFFD1F813FDDF1CBFFD15FA10" box="[600,647,1411,1437]" pageId="12" pageNumber="261">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="6AA26547FFD1F813FF401C9BFA15FA4C" box="[199,1415,1447,1473]" pageId="12" pageNumber="261" type="etymology">
<paragraph id="220736CCFFD1F813FF401C9BFA15FA4C" blockId="12.[151,1437,151,2013]" box="[199,1415,1447,1473]" pageId="12" pageNumber="261">
<emphasis id="10CCEADEFFD1F813FF401C9BFED9FA4C" bold="true" box="[199,331,1447,1473]" pageId="12" pageNumber="261">Etymology.</emphasis>
The specific name is an adjective derived from Danyang-gun, the
<typeStatus id="FD03886EFFD1F813FBA11C9BFBC7FA4C" box="[1062,1109,1447,1473]" pageId="12" pageNumber="261">type</typeStatus>
locality of the new species.
</paragraph>
</subSubSection>
<subSubSection id="6AA26547FFD1F810FF401CF7FB56FC0C" lastPageId="15" lastPageNumber="264" pageId="12" pageNumber="261" type="discussion">
<paragraph id="220736CCFFD1F813FF401CF7FB16F96C" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="261">
<emphasis id="10CCEADEFFD1F813FF401CF7FEABFA68" bold="true" box="[199,313,1483,1509]" pageId="12" pageNumber="261">Remarks.</emphasis>
<taxonomicName id="E5B84D4FFFD1F813FEB81CF7FD0CFA68" authority="Lee &amp; Min, 2024" authorityName="Lee &amp; Min" authorityYear="2024" box="[319,670,1483,1509]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="261" phylum="Arthropoda" rank="species" species="danyangensis" status="sp. nov.">
<emphasis id="10CCEADEFFD1F813FEB81CF7FD0CFA68" box="[319,670,1483,1509]" italics="true" pageId="12" pageNumber="261">Pseudocrangonyx danyangensis</emphasis>
</taxonomicName>
<emphasis id="10CCEADEFFD1F813FD221CF7FD6EFA68" bold="true" box="[677,764,1483,1509]" pageId="12" pageNumber="261">
<taxonomicNameLabel id="0BFF57A5FFD1F813FD221CF7FD6EFA68" box="[677,764,1483,1509]" pageId="12" pageNumber="261" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is morphologically most similar to
<taxonomicName id="E5B84D4FFFD1F813FBF81CF7FF5CF984" authority="Lee et al., 2018" authorityName="Lee" authorityYear="2018" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="261" phylum="Arthropoda" rank="species" species="daejeonensis">
<emphasis id="10CCEADEFFD1F813FBF81CF7FB02FA69" box="[1151,1168,1483,1508]" italics="true" pageId="12" pageNumber="261">P</emphasis>
.
<emphasis id="10CCEADEFFD1F813FB1B1CF7FAB8FA68" box="[1180,1322,1483,1509]" italics="true" pageId="12" pageNumber="261">daejeonensis</emphasis>
<bibRefCitation id="46294B3DFFD1F813FAB71CF7FF5CF984" author="Lee, C. - W. &amp; Tomikawa, K. &amp; Nakano, T. &amp; Min, G. - S." pageId="12" pageNumber="261" pagination="27 - 44" refId="ref7864" refString="Lee, C. - W., Tomikawa, K., Nakano, T. &amp; Min, G. - S. (2018) A new species of the genus Pseudocrangonyx (Crustacea, Amphipoda, Pseudocrangonyctidae) from Korea. ZooKeys, 735, 27 - 44. https: // doi. org / 10.3897 / zookeys. 735.21697" type="journal article" year="2018">
Lee
<emphasis id="10CCEADEFFD1F813FAD81CF7FA1CFA68" box="[1375,1422,1483,1509]" italics="true" pageId="12" pageNumber="261">et al</emphasis>
.,
<quantity id="E5409B29FFD1F813FF101CD3FF5CF984" box="[151,206,1519,1545]" metricMagnitude="1" metricUnit="m" metricValue="5.125719999999999" pageId="12" pageNumber="261" unit="in" value="2018.0">2018</quantity>
</bibRefCitation>
</taxonomicName>
in having 1) antenna 1 length shorter than body length half, 2) antenna 1 accessory flagellum first article exceed main flagellum first article, 3) male antenna 2 with calceoli, 4) uropod 2 outer ramus marginally bare, 5) uropod 3 terminal article longer than adjacent robust setae, and 6) telson lobe with 2 lateral penicillate setae. However, the new species is distinguished from
<taxonomicName id="E5B84D4FFFD1F813FE5E1F67FD13F9F8" box="[473,641,1627,1653]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="261" phylum="Arthropoda" rank="species" species="daejeonensis">
<emphasis id="10CCEADEFFD1F813FE5E1F67FD13F9F8" box="[473,641,1627,1653]" italics="true" pageId="12" pageNumber="261">P. daejeonensis</emphasis>
</taxonomicName>
by the following features (features of
<taxonomicName id="E5B84D4FFFD1F813FBA61F67FB5FF9F8" box="[1057,1229,1627,1653]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="261" phylum="Arthropoda" rank="species" species="daejeonensis">
<emphasis id="10CCEADEFFD1F813FBA61F67FBA0F9F9" box="[1057,1074,1627,1652]" italics="true" pageId="12" pageNumber="261">P</emphasis>
.
<emphasis id="10CCEADEFFD1F813FBB81F67FB5FF9F8" box="[1087,1229,1627,1653]" italics="true" pageId="12" pageNumber="261">daejeonensis</emphasis>
</taxonomicName>
in parentheses): 1) inner rami of pleopods with (without) bifid setae on inner margin, 2) female and male uropod 1 margin with (without) robust setae, 3) female uropod 2 peduncle shorter (longer) than inner ramus, 4) male uropod 2 peduncle outer margin with 7 (1 robust setae) robust setae, and 5) telson (cleft for 8.6% of length) almost not cleft.
</paragraph>
<paragraph id="220736CCFFD1F813FF401FD7FBC8F850" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="261">
The new species is also similar to
<taxonomicName id="E5B84D4FFFD1F813FDBA1FD7FD75F889" authority="Lee &amp; Min, 2024" authorityName="Lee &amp; Min" authorityYear="2024" box="[573,743,1771,1797]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="261" phylum="Arthropoda" rank="species" species="seomjinensis" status="sp. nov.">
<emphasis id="10CCEADEFFD1F813FDBA1FD7FDDCF889" box="[573,590,1771,1796]" italics="true" pageId="12" pageNumber="261">P</emphasis>
.
<emphasis id="10CCEADEFFD1F813FDDC1FD7FD75F889" box="[603,743,1771,1796]" italics="true" pageId="12" pageNumber="261">seomjinensis</emphasis>
</taxonomicName>
<emphasis id="10CCEADEFFD1F813FD6A1FD7FCD7F888" bold="true" box="[749,837,1771,1797]" pageId="12" pageNumber="261">
<taxonomicNameLabel id="0BFF57A5FFD1F813FD6A1FD7FCD7F888" box="[749,837,1771,1797]" pageId="12" pageNumber="261" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in having 1) eyes completely absent, 2) maxilla 2 with oblique inner row of 2 setae on inner plate, 3) uropod 1 with same number robust setae, and 4) uropod 2 outer ramus marginally bare. However,
<taxonomicName id="E5B84D4FFFD1F813FE4A1E0FFD16F8C0" authority="Lee &amp; Min, 2024" authorityName="Lee &amp; Min" authorityYear="2024" box="[461,644,1843,1869]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="261" phylum="Arthropoda" rank="species" species="danyangensis" status="sp. nov.">
<emphasis id="10CCEADEFFD1F813FE4A1E0FFE4CF8C1" box="[461,478,1843,1868]" italics="true" pageId="12" pageNumber="261">P</emphasis>
.
<emphasis id="10CCEADEFFD1F813FE681E0FFD16F8C0" box="[495,644,1843,1869]" italics="true" pageId="12" pageNumber="261">danyangensis</emphasis>
</taxonomicName>
<emphasis id="10CCEADEFFD1F813FD081E0FFD78F8C0" bold="true" box="[655,746,1843,1869]" pageId="12" pageNumber="261">
<taxonomicNameLabel id="0BFF57A5FFD1F813FD081E0FFD78F8C0" box="[655,746,1843,1869]" pageId="12" pageNumber="261" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is distinguished from
<taxonomicName id="E5B84D4FFFD1F813FC691E0FFB0EF8C1" authority="Lee &amp; Min, 2024" authorityName="Lee &amp; Min" authorityYear="2024" box="[1006,1180,1843,1869]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="261" phylum="Arthropoda" rank="species" species="seomjinensis" status="sp. nov.">
<emphasis id="10CCEADEFFD1F813FC691E0FFC6DF8C1" box="[1006,1023,1843,1868]" italics="true" pageId="12" pageNumber="261">P</emphasis>
.
<emphasis id="10CCEADEFFD1F813FB971E0FFB0EF8C1" box="[1040,1180,1843,1868]" italics="true" pageId="12" pageNumber="261">seomjinensis</emphasis>
</taxonomicName>
<emphasis id="10CCEADEFFD1F813FB211E0FFA93F8C0" bold="true" box="[1190,1281,1843,1869]" pageId="12" pageNumber="261">
<taxonomicNameLabel id="0BFF57A5FFD1F813FB211E0FFA93F8C0" box="[1190,1281,1843,1869]" pageId="12" pageNumber="261" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by following features (features of
<taxonomicName id="E5B84D4FFFD1F813FE071E6BFDBFF8FD" authority="Lee &amp; Min, 2024" authorityName="Lee &amp; Min" authorityYear="2024" box="[384,557,1879,1905]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="261" phylum="Arthropoda" rank="species" species="seomjinensis" status="sp. nov.">
<emphasis id="10CCEADEFFD1F813FE071E6BFE03F8FD" box="[384,401,1879,1904]" italics="true" pageId="12" pageNumber="261">P</emphasis>
.
<emphasis id="10CCEADEFFD1F813FE261E6BFDBFF8FD" box="[417,557,1879,1904]" italics="true" pageId="12" pageNumber="261">seomjinensis</emphasis>
</taxonomicName>
<emphasis id="10CCEADEFFD1F813FDB11E6BFD02F8FC" bold="true" box="[566,656,1879,1905]" pageId="12" pageNumber="261">
<taxonomicNameLabel id="0BFF57A5FFD1F813FDB11E6BFD02F8FC" box="[566,656,1879,1905]" pageId="12" pageNumber="261" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in parentheses): 1) antenna 1 accessory flagellum first article exceed (not exceed) main flagellum of first article, 2) inner rami of pleopods with (without) bifid setae on inner margin, 3) female uropod 2 peduncle outer margin with 6 robust setae (2 robust setae), 4) male uropod 2 outer ramus almost reach (shorter than) inner ramus end, and 5) male telson shape subrectangular (subtriangular).
</paragraph>
<caption id="76C76644FFD0F812FF101E49FBEDF83C" ID-DOI="http://doi.org/10.5281/zenodo.10954140" ID-Zenodo-Dep="10954140" httpUri="https://zenodo.org/record/10954140/files/figure.png" pageId="13" pageNumber="262" startId="13.[151,250,1909,1933]" targetBox="[222,1366,187,1884]" targetPageId="13" targetType="figure">
<paragraph id="220736CCFFD0F812FF101E49FBEDF83C" blockId="13.[151,1436,1909,1969]" pageId="13" pageNumber="262">
<emphasis id="10CCEADEFFD0F812FF101E49FE9CF800" bold="true" box="[151,270,1909,1933]" pageId="13" pageNumber="262">FIGURE 9</emphasis>
.
<taxonomicName id="E5B84D4FFFD0F812FE9B1E49FDF0F800" authority="Lee &amp; Min, 2024" authorityName="Lee &amp; Min" authorityYear="2024" box="[284,610,1909,1933]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" kingdom="Animalia" order="Amphipoda" pageId="13" pageNumber="262" phylum="Arthropoda" rank="species" species="danyangensis" status="sp. nov.">
<emphasis id="10CCEADEFFD0F812FE9B1E49FDF0F800" box="[284,610,1909,1933]" italics="true" pageId="13" pageNumber="262">Pseudocrangonyx danyangensis</emphasis>
</taxonomicName>
<emphasis id="10CCEADEFFD0F812FDED1E49FD29F800" bold="true" box="[618,699,1909,1933]" pageId="13" pageNumber="262">
<taxonomicNameLabel id="0BFF57A5FFD0F812FDED1E49FD29F800" box="[618,699,1909,1933]" pageId="13" pageNumber="262" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype female, NNIBRIV69645. A–C, Pleopod 1–3, lateral view; D, uropod 1, ventral view; E, uropod 2, dorsal view; F, uropod 3, dorsal view; G, telson, ventral view.
</paragraph>
</caption>
<caption id="76C76644FFD3F811FF101EABFB15F875" ID-DOI="http://doi.org/10.5281/zenodo.10954142" ID-Zenodo-Dep="10954142" httpUri="https://zenodo.org/record/10954142/files/figure.png" pageId="14" pageNumber="263" startId="14.[151,250,1943,1967]" targetBox="[159,1428,181,1919]" targetPageId="14" targetType="figure">
<paragraph id="220736CCFFD3F811FF101EABFB15F875" blockId="14.[151,1436,1943,2040]" pageId="14" pageNumber="263">
<emphasis id="10CCEADEFFD3F811FF101EABFE8CF822" bold="true" box="[151,286,1943,1967]" pageId="14" pageNumber="263">FIGURE 10.</emphasis>
<taxonomicName id="E5B84D4FFFD3F811FEA41EA4FDF5F83D" authority="Lee &amp; Min, 2024" authorityName="Lee &amp; Min" authorityYear="2024" box="[291,615,1944,1968]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" kingdom="Animalia" order="Amphipoda" pageId="14" pageNumber="263" phylum="Arthropoda" rank="species" species="danyangensis" status="sp. nov.">
<emphasis id="10CCEADEFFD3F811FEA41EA4FDF5F83D" box="[291,615,1944,1968]" italics="true" pageId="14" pageNumber="263">Pseudocrangonyx danyangensis</emphasis>
</taxonomicName>
<emphasis id="10CCEADEFFD3F811FDEC1EA4FD28F83D" bold="true" box="[619,698,1944,1968]" pageId="14" pageNumber="263">
<taxonomicNameLabel id="0BFF57A5FFD3F811FDEC1EA4FD28F83D" box="[619,698,1944,1968]" pageId="14" pageNumber="263" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype male, NNIBRIV69646. A, antenna 1, medial view; B, accessory flagellum of antenna 1, medial view; C, antenna 2, lateral view; D, gnathopod 1, lateral view; E, gnathopod 2, lateral view; F, uropod 1, ventral view; G, uropod 2, ventral view; H, uropod 3, dorsal view; I, telson, ventral view.
</paragraph>
</caption>
<paragraph id="220736CCFFD2F810FF4019AAFC54FECC" blockId="15.[151,1437,150,897]" pageId="15" pageNumber="264">
<emphasis id="10CCEADEFFD2F810FF4019AAFD47FF3D" bold="true" box="[199,725,150,177]" pageId="15" pageNumber="264">Phylogenetic position and genetic distances.</emphasis>
Phylogenetic relationships within Korean pseudocrangonyctid species were identified using 28S rRNA and COI gene sequences. The uncorrected COI
<emphasis id="10CCEADEFFD2F810FBEA1987FBE9FF59" box="[1133,1147,187,212]" italics="true" pageId="15" pageNumber="264">p</emphasis>
-distance between the two new species (
<taxonomicName id="E5B84D4FFFD2F810FEB619E3FE4FFF75" authority="Lee &amp; Min, 2024" authorityName="Lee &amp; Min" authorityYear="2024" box="[305,477,223,248]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" kingdom="Animalia" order="Amphipoda" pageId="15" pageNumber="264" phylum="Arthropoda" rank="species" species="seomjinensis" status="sp. nov.">
<emphasis id="10CCEADEFFD2F810FEB619E3FE4FFF75" box="[305,477,223,248]" italics="true" pageId="15" pageNumber="264">P. seomjinensis</emphasis>
</taxonomicName>
<emphasis id="10CCEADEFFD2F810FE6119E3FDD3FF74" bold="true" box="[486,577,223,249]" pageId="15" pageNumber="264">
<taxonomicNameLabel id="0BFF57A5FFD2F810FE6119E3FDD3FF74" box="[486,577,223,249]" pageId="15" pageNumber="264" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="E5B84D4FFFD2F810FDFD19E3FCBDFF74" authority="Lee &amp; Min, 2024" authorityName="Lee &amp; Min" authorityYear="2024" box="[634,815,223,249]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" kingdom="Animalia" order="Amphipoda" pageId="15" pageNumber="264" phylum="Arthropoda" rank="species" species="danyangensis" status="sp. nov.">
<emphasis id="10CCEADEFFD2F810FDFD19E3FCBDFF74" box="[634,815,223,249]" italics="true" pageId="15" pageNumber="264">P. danyangensis</emphasis>
</taxonomicName>
<emphasis id="10CCEADEFFD2F810FCBF19E3FC01FF74" bold="true" box="[824,915,223,249]" pageId="15" pageNumber="264">
<taxonomicNameLabel id="0BFF57A5FFD2F810FCBF19E3FC01FF74" box="[824,915,223,249]" pageId="15" pageNumber="264" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) was 15.2%, calculated based on 609 aligned positions from the dataset. This distance is the range of COI differentiation that can distinguish a distinct species in the genus
<taxonomicName id="E5B84D4FFFD2F810FE8E181BFE40FECC" authorityName="Akatsuka &amp; Komai" authorityYear="1922" box="[265,466,295,321]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" kingdom="Animalia" order="Amphipoda" pageId="15" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="10CCEADEFFD2F810FE8E181BFE40FECC" box="[265,466,295,321]" italics="true" pageId="15" pageNumber="264">Pseudocrangonyx</emphasis>
</taxonomicName>
, as reported previously (
<bibRefCitation id="46294B3DFFD2F810FD60181BFC28FECC" author="Zhao, S. &amp; Hou, Z." box="[743,954,295,321]" pageId="15" pageNumber="264" pagination="1 - 22" refId="ref8384" refString="Zhao, S. &amp; Hou, Z. (2017) A new subterranean species of Pseudocrangonyx from China with an identification key to all species of the genus (Crustacea, Amphipoda, Pseudocrangonyctidae). ZooKeys, 647, 1 - 22. https: // doi. org / 10.3897 / zookeys. 647.11192" type="journal article" year="2017">Zhao &amp; Hou 2017</bibRefCitation>
).
</paragraph>
<paragraph id="220736CCFFD2F810FF401877FCF5FD40" blockId="15.[151,1437,150,897]" pageId="15" pageNumber="264">
In phylogenetic analyses, the topologies of the BI and ML trees were almost identical.The molecular phylogenetic tree (
<figureCitation id="BA832A49FFD2F810FF571853FEB1FE04" box="[208,291,367,393]" captionStart="FIGURE 11" captionStartId="15.[151,250,1689,1713]" captionTargetBox="[197,1397,924,1661]" captionTargetId="figure-359@15.[189,1398,921,1666]" captionTargetPageId="15" captionText="FIGURE 11. Maximum likelihood and Bayesian inference analyses based on nuclear 28S rRNA and mitochondrial COI sequences. Numbers on nodes represent bootstrap values for maximum likelihood and Bayesian posterior probabilities." figureDoi="http://doi.org/10.5281/zenodo.10954144" httpUri="https://zenodo.org/record/10954144/files/figure.png" pageId="15" pageNumber="264">Fig. 11</figureCitation>
; ML tree not shown) showed that the Korean
<taxonomicName id="E5B84D4FFFD2F810FCA31853FC7DFE04" authorityName="Akatsuka &amp; Komai" authorityYear="1922" box="[804,1007,367,393]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" kingdom="Animalia" order="Amphipoda" pageId="15" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="10CCEADEFFD2F810FCA31853FC7DFE04" box="[804,1007,367,393]" italics="true" pageId="15" pageNumber="264">Pseudocrangonyx</emphasis>
</taxonomicName>
split into two clades: interstitial water (in the gray circle,
<taxonomicName id="E5B84D4FFFD2F810FEF618AFFD4DFE20" authority="Lee et al., 2018" authorityName="Lee" authorityYear="2018" box="[369,735,403,429]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" kingdom="Animalia" order="Amphipoda" pageId="15" pageNumber="264" phylum="Arthropoda" rank="species" species="daejeonensis">
<emphasis id="10CCEADEFFD2F810FEF618AFFDB2FE20" box="[369,544,403,429]" italics="true" pageId="15" pageNumber="264">P. daejeonensis</emphasis>
<bibRefCitation id="46294B3DFFD2F810FDAF18AFFD4DFE20" author="Lee, C. - W. &amp; Tomikawa, K. &amp; Nakano, T. &amp; Min, G. - S." box="[552,735,403,429]" pageId="15" pageNumber="264" pagination="27 - 44" refId="ref7864" refString="Lee, C. - W., Tomikawa, K., Nakano, T. &amp; Min, G. - S. (2018) A new species of the genus Pseudocrangonyx (Crustacea, Amphipoda, Pseudocrangonyctidae) from Korea. ZooKeys, 735, 27 - 44. https: // doi. org / 10.3897 / zookeys. 735.21697" type="journal article" year="2018">
Lee
<emphasis id="10CCEADEFFD2F810FDDC18AFFD1FFE20" box="[603,653,403,429]" italics="true" pageId="15" pageNumber="264">et al</emphasis>
., 2018
</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="E5B84D4FFFD2F810FD6B18AFFC0BFE21" authority="Lee &amp; Min, 2024" authorityName="Lee &amp; Min" authorityYear="2024" box="[748,921,403,428]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" kingdom="Animalia" order="Amphipoda" pageId="15" pageNumber="264" phylum="Arthropoda" rank="species" species="seomjinensis" status="sp. nov.">
<emphasis id="10CCEADEFFD2F810FD6B18AFFC0BFE21" box="[748,921,403,428]" italics="true" pageId="15" pageNumber="264">P. seomjinensis</emphasis>
</taxonomicName>
<emphasis id="10CCEADEFFD2F810FC2618AFFC6EFE20" bold="true" box="[929,1020,403,429]" pageId="15" pageNumber="264">
<taxonomicNameLabel id="0BFF57A5FFD2F810FC2618AFFC6EFE20" box="[929,1020,403,429]" pageId="15" pageNumber="264" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<taxonomicName id="E5B84D4FFFD2F810FB8B18AFFB51FE20" authority="Lee &amp; Min, 2024" authorityName="Lee &amp; Min" authorityYear="2024" box="[1036,1219,403,429]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" kingdom="Animalia" order="Amphipoda" pageId="15" pageNumber="264" phylum="Arthropoda" rank="species" species="danyangensis" status="sp. nov.">
<emphasis id="10CCEADEFFD2F810FB8B18AFFB51FE20" box="[1036,1219,403,429]" italics="true" pageId="15" pageNumber="264">P. danyangensis</emphasis>
</taxonomicName>
<emphasis id="10CCEADEFFD2F810FB4C18AFFAB4FE20" bold="true" box="[1227,1318,403,429]" pageId="15" pageNumber="264">
<taxonomicNameLabel id="0BFF57A5FFD2F810FB4C18AFFAB4FE20" box="[1227,1318,403,429]" pageId="15" pageNumber="264" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) and cave groundwater (in the gray square,
<taxonomicName id="E5B84D4FFFD2F810FD91188BFCD0FE5C" authority="Lee et al., 2020" authorityName="Lee" authorityYear="2020" box="[534,834,439,465]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" kingdom="Animalia" order="Amphipoda" pageId="15" pageNumber="264" phylum="Arthropoda" rank="species" species="joolaei">
<emphasis id="10CCEADEFFD2F810FD91188BFD10FE5C" box="[534,642,439,465]" italics="true" pageId="15" pageNumber="264">P. joolaei</emphasis>
Lee
<emphasis id="10CCEADEFFD2F810FD3A188BFD7DFE5C" box="[701,751,439,465]" italics="true" pageId="15" pageNumber="264">et al</emphasis>
., 2020
</taxonomicName>
;
<taxonomicName id="E5B84D4FFFD2F810FCC8188BFB1FFE5C" authority="Lee et al., 2020" authorityName="Lee" authorityYear="2020" box="[847,1165,439,465]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" kingdom="Animalia" order="Amphipoda" pageId="15" pageNumber="264" phylum="Arthropoda" rank="species" species="wonkimi">
<emphasis id="10CCEADEFFD2F810FCC8188BFC5EFE5C" box="[847,972,439,465]" italics="true" pageId="15" pageNumber="264">P. wonkimi</emphasis>
Lee
<emphasis id="10CCEADEFFD2F810FB80188BFBA8FE5C" box="[1031,1082,439,465]" italics="true" pageId="15" pageNumber="264">et al</emphasis>
., 2020
</taxonomicName>
;
<taxonomicName id="E5B84D4FFFD2F810FB1D188BFE82FE78" authority="Lee &amp; Min, 2021" authorityName="Lee &amp; Min" authorityYear="2021" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" kingdom="Animalia" order="Amphipoda" pageId="15" pageNumber="264" phylum="Arthropoda" rank="species" species="deureunensis">
<emphasis id="10CCEADEFFD2F810FB1D188BFAD9FE5C" box="[1178,1355,439,465]" italics="true" pageId="15" pageNumber="264">P. deureunensis</emphasis>
Lee &amp; Min, 2021
</taxonomicName>
;
<taxonomicName id="E5B84D4FFFD2F810FE9C18E7FD6CFE78" authority="Lee &amp; Min, 2021" authorityName="Lee &amp; Min" authorityYear="2021" box="[283,766,475,501]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" kingdom="Animalia" order="Amphipoda" pageId="15" pageNumber="264" phylum="Arthropoda" rank="species" species="kwangcheonseonensis">
<emphasis id="10CCEADEFFD2F810FE9C18E7FDA3FE78" box="[283,561,475,501]" italics="true" pageId="15" pageNumber="264">P. kwangcheonseonensis</emphasis>
Lee &amp; Min, 2021
</taxonomicName>
;
<taxonomicName id="E5B84D4FFFD2F810FC8E18E7FB08FE78" authority="Lee &amp; Min, 2021" authorityName="Lee &amp; Min" authorityYear="2021" box="[777,1178,475,501]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" kingdom="Animalia" order="Amphipoda" pageId="15" pageNumber="264" phylum="Arthropoda" rank="species" species="hwanseonensis">
<emphasis id="10CCEADEFFD2F810FC8E18E7FC5DFE78" box="[777,975,475,501]" italics="true" pageId="15" pageNumber="264">P. hwanseonensis</emphasis>
Lee &amp; Min, 2021
</taxonomicName>
) clades. Moreover, the obtained phylogenies revealed that the Korean interstitial water species are close congeners of the Japanese species (
<taxonomicName id="E5B84D4FFFD2F810FF261B1FFDD9FDB0" authority="Tomikawa &amp; Nakano, 2019" authorityName="Tomikawa &amp; Nakano" authorityYear="2019" box="[161,587,547,573]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" kingdom="Animalia" order="Amphipoda" pageId="15" pageNumber="264" phylum="Arthropoda" rank="species" species="uenoi">
<emphasis id="10CCEADEFFD2F810FF261B1FFF6CFDB1" box="[161,254,547,572]" italics="true" pageId="15" pageNumber="264">P. uenoi</emphasis>
Tomikawa &amp; Nakano, 2019
</taxonomicName>
and
<taxonomicName id="E5B84D4FFFD2F810FD0F1B1FFC6DFDB0" authority="Tomikawa et al., 2022" authorityName="Tomikawa" authorityYear="2022" box="[648,1023,547,573]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" kingdom="Animalia" order="Amphipoda" pageId="15" pageNumber="264" phylum="Arthropoda" rank="species" species="dunan">
<emphasis id="10CCEADEFFD2F810FD0F1B1FFD7CFDB0" box="[648,750,547,573]" italics="true" pageId="15" pageNumber="264">P. dunan</emphasis>
Tomikawa
<emphasis id="10CCEADEFFD2F810FCFF1B1FFC3FFDB0" box="[888,941,547,573]" italics="true" pageId="15" pageNumber="264">et al</emphasis>
., 2022
</taxonomicName>
) and are not Korean cave-dwelling species. Meanwhile, the Korean cave-dwelling species formed a clade with the Japanese
<taxonomicName id="E5B84D4FFFD2F810FBFB1B7BFEAAFD08" authority="Tomikawa &amp; Nakano, 2018" authorityName="Tomikawa &amp; Nakano" authorityYear="2018" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" kingdom="Animalia" order="Amphipoda" pageId="15" pageNumber="264" phylum="Arthropoda" rank="species" species="akatsukai">
<emphasis id="10CCEADEFFD2F810FBFB1B7BFA91FDEC" box="[1148,1283,583,609]" italics="true" pageId="15" pageNumber="264">P. akatsukai</emphasis>
Tomikawa &amp; Nakano, 2018
</taxonomicName>
, which are morphologically similar. These relationships suggest that evolutionary history may have changed depending on the habitat environment, interstitial water, or cave groundwater. The Korean and Japanese
<taxonomicName id="E5B84D4FFFD2F810FF101B8FFEF0FD40" authorityName="Akatsuka &amp; Komai" authorityYear="1922" box="[151,354,691,717]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" kingdom="Animalia" order="Amphipoda" pageId="15" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="10CCEADEFFD2F810FF101B8FFEF0FD40" box="[151,354,691,717]" italics="true" pageId="15" pageNumber="264">Pseudocrangonyx</emphasis>
</taxonomicName>
may have diverged from a common ancestor.
</paragraph>
<paragraph id="220736CCFFD2F810FF401BEBFB56FC0C" blockId="15.[151,1437,150,897]" pageId="15" pageNumber="264">
Finally, the existence of pseudocrangonyctids is related to biogeographic interest, and the origin and evolution of subterranean amphipod fauna in the Far East region (
<bibRefCitation id="46294B3DFFD2F810FC921BC7FBDCFC98" author="Sidorov, D. &amp; Holsinger, J. R." box="[789,1102,763,789]" pageId="15" pageNumber="264" pagination="417 - 430" refId="ref8220" refString="Sidorov, D. &amp; Holsinger, J. R. (2007) Procrangonyx stygoedincus, a new species of subterranean amphipod (pseudocrangonyctidae) from the far east of Russia, with remarks on biogeographic relationships. Crustaceana, 80, 417 - 430. https: // doi. org / 10.1163 / 156854007780440984" type="journal article" year="2007">Sidorov &amp; Holsinger 2007</bibRefCitation>
). Further taxon sampling and molecular data are required to elucidate the origin and dispersal routes of the genus
<taxonomicName id="E5B84D4FFFD2F810FBD41A23FF4CFCD0" authority="(Lee et al. 2018)" baseAuthorityName="Lee" baseAuthorityYear="2018" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" kingdom="Animalia" order="Amphipoda" pageId="15" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="10CCEADEFFD2F810FBD41A23FAB0FCB4" box="[1107,1314,799,825]" italics="true" pageId="15" pageNumber="264">Pseudocrangonyx</emphasis>
(
<bibRefCitation id="46294B3DFFD2F810FAB71A23FF44FCD0" author="Lee, C. - W. &amp; Tomikawa, K. &amp; Nakano, T. &amp; Min, G. - S." pageId="15" pageNumber="264" pagination="27 - 44" refId="ref7864" refString="Lee, C. - W., Tomikawa, K., Nakano, T. &amp; Min, G. - S. (2018) A new species of the genus Pseudocrangonyx (Crustacea, Amphipoda, Pseudocrangonyctidae) from Korea. ZooKeys, 735, 27 - 44. https: // doi. org / 10.3897 / zookeys. 735.21697" type="journal article" year="2018">
Lee
<emphasis id="10CCEADEFFD2F810FAE41A23FA07FCB4" box="[1379,1429,799,825]" italics="true" pageId="15" pageNumber="264">et al</emphasis>
. 2018
</bibRefCitation>
)
</taxonomicName>
. Further molecular phylogenetic analyses of
<taxonomicName id="E5B84D4FFFD2F810FD611A7FFC27FCD0" box="[742,949,835,861]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" kingdom="Animalia" order="Amphipoda" pageId="15" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="10CCEADEFFD2F810FD611A7FFC27FCD0" box="[742,949,835,861]" italics="true" pageId="15" pageNumber="264">Pseudocrangonyx</emphasis>
</taxonomicName>
will provide a better understanding of the diversity and evolutionary history of subterranean Crangonyctoidea species in Far
<collectingCountry id="5AAF765CFFD2F810FBD71A5BFB53FC0C" box="[1104,1217,871,897]" name="China" pageId="15" pageNumber="264">East Asia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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