treatments-xml/data/95/7C/D8/957CD8C73D01CBDD714A864CA518BEE0.xml

<document ID-DOI="http://dx.doi.org/10.3897/mycokeys.29.22572" ID-GBIF-Dataset="04e3717e-c951-401c-8840-198a6f83de66" ID-PMC="PMC5804118" ID-Pensoft-Pub="1314-4049-29-63" ID-PubMed="29559826" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1314-4049-29-63" ModsDocOrigin="MycoKeys 29" ModsDocTitle="Phylogenetic affinities of the sequestrate genus Rhodactina (Boletaceae), with a new species, R.rostratispora from Thailand" checkinTime="1516918265336" checkinUser="pensoft" docAuthor="Vadthanarat, Santhiti, Raspe, Olivier &amp; Lumyong, Saisamorn" docDate="2018" docId="957CD8C73D01CBDD714A864CA518BEE0" docLanguage="en" docName="MycoKeys 29: 63-80" docOrigin="MycoKeys 29" docSource="http://dx.doi.org/10.3897/mycokeys.29.22572" docTitle="Rhodactina rostratispora Vadthanarat, Raspe &amp; Lumyong, sp. nov." docType="treatment" docVersion="4" lastPageNumber="69" masterDocId="FFA4FFA63F71BC6E7271FFC2FFC1FFBF" masterDocTitle="Phylogenetic affinities of the sequestrate genus Rhodactina (Boletaceae), with a new species, R. rostratispora from Thailand" masterLastPageNumber="80" masterPageNumber="63" pageNumber="67" updateTime="1668135894767" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Phylogenetic affinities of the sequestrate genus Rhodactina (Boletaceae), with a new species, R. rostratispora from Thailand</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vadthanarat, Santhiti</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Raspe, Olivier</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lumyong, Saisamorn</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>MycoKeys</mods:title>
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<mods:part>
<mods:date>2018</mods:date>
<mods:detail type="volume">
<mods:number>29</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>63</mods:start>
<mods:end>80</mods:end>
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<mods:url>http://dx.doi.org/10.3897/mycokeys.29.22572</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/mycokeys.29.22572</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-4049-29-63</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="140417908" LSID="urn:lsid:plazi:treatment:957CD8C73D01CBDD714A864CA518BEE0" httpUri="http://treatment.plazi.org/id/957CD8C73D01CBDD714A864CA518BEE0" lastPageId="6" lastPageNumber="69" pageId="4" pageNumber="67">
<subSubSection pageId="4" pageNumber="67" type="nomenclature">
<paragraph pageId="4" pageNumber="67">
<taxonomicName LSID="822126" authority="Vadthanarat, Raspe &amp; Lumyong" class="Agaricomycetes" family="Boletaceae" genus="Rhodactina" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhodactina rostratispora" order="Boletales" pageId="4" pageNumber="67" phylum="Basidiomycota" rank="species" species="rostratispora">
<pageBreakToken pageId="4" pageNumber="67" start="start">Rhodactina</pageBreakToken>
rostratispora Vadthanarat, 
<normalizedToken originalValue="Raspé">Raspe</normalizedToken>
&amp; Lumyong
</taxonomicName>
<taxonomicNameLabel pageId="4" pageNumber="67">sp. nov.</taxonomicNameLabel>
Figs 2, 3, 4
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="67" type="materials_examined">
<paragraph pageId="4" pageNumber="67">Type.</paragraph>
<paragraph pageId="4" pageNumber="67">
THAILAND, Ubon Ratchathani Province, Trakan Phuet Phon District, Don Khok Tam Lae community forest, 
<geoCoordinate direction="north" orientation="latitude" precision="15" value="15.59611">15°35'46&quot;N</geoCoordinate>
, 
<geoCoordinate direction="east" orientation="longitude" precision="15" value="105.11056">105°06'38&quot;E</geoCoordinate>
, elev. 150 m., 28 July 2015, S. Vadthanarat 170, (holotype: CMUB!; isotype: BR!).
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="67" type="etymology">
<paragraph pageId="4" pageNumber="67">Etymology.</paragraph>
<paragraph pageId="4" pageNumber="67">
From Latin 
<normalizedToken originalValue="“rostrati–”">&quot;rostrati-&quot;</normalizedToken>
meaning having beaked prow or a solid projection and 
<normalizedToken originalValue="“spora”">&quot;spora&quot;</normalizedToken>
meaning spores, referring to the basidiospores having a markedly prominent and large hilar appendage.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="67" type="description">
<paragraph pageId="4" pageNumber="67">Description.</paragraph>
<paragraph pageId="4" pageNumber="67">
Basidiomata small to medium-sized 0.8-2.5(4.5) cm diam., subglobose to ovoid with a rudimentary elongated basal attachment, with greyish white to pale brown rhizoids at the base and going up along the surface of basidiomata to about half of the height. Peridium surface (outer peridium) fibrillose to arachnoid, off-white to pinkish white (7A2-3 to 9A2), dull, moist, cracked in places. Peridium very thin, 0.1-0.2(0.4) mm thick. Hymenophore cartilaginous, completely enclosed, whitish orange to reddish orange (7A3-4 to 8A5-6) at first becoming orangey red to red (9 
<normalizedToken originalValue="D–E">D-E</normalizedToken>
8 to 10 
<normalizedToken originalValue="D–E">D-E</normalizedToken>
8) with age, then dark red when very old, irregular; Stipe-columella absent. Taste fungoid. Odour absent when young, very strongly fruity alcoholic when old.
</paragraph>
<paragraph pageId="4" pageNumber="67">Macrochemical reactions: hymenophore turned dark purplish (15F8) to greyish violet (19D3) with 5% KOH, slightly greyish violet (19D3) with NH4OH.</paragraph>
<paragraph pageId="4" pageNumber="67">
Basidiospores [404/8/8] (11.5 
<normalizedToken originalValue="–)12–13.6–15(–">-)12-13.6-15(-</normalizedToken>
16) 
<normalizedToken originalValue="×">x</normalizedToken>
(10 
<normalizedToken originalValue="–)10.5–11.7–13(–">-)10.5-11.7-13(-</normalizedToken>
14), Q = (1 
<normalizedToken originalValue="–)1.04–1.16–1.3(–">-)1.04-1.16-1.3(-</normalizedToken>
1.4), from the holotype, (12 
<normalizedToken originalValue="–)12–13.5–15.2(–">-)12-13.5-15.2(-</normalizedToken>
16) 
<normalizedToken originalValue="×">x</normalizedToken>
(10 
<normalizedToken originalValue="–)10–11.6–13.2(–">-)10-11.6-13.2(-</normalizedToken>
14) 
<normalizedToken originalValue="µm">µm</normalizedToken>
, Q = (1 
<normalizedToken originalValue="–)1–1.02–1.33(–">-)1-1.02-1.33(-</normalizedToken>
1.4), N = 50, ellipsoid to broadly ellipsoid with longitudinal ridges, stellate in polar-view, thick-walled (1-1.5 
<normalizedToken originalValue="µm">µm</normalizedToken>
thick), yellowish to orangey hyaline to reddish yellow at first, reddish to brownish yellow with age in water, slightly purplish and slightly more reddish to brownish in 5% KOH, slightly purplish hyaline in NH4OH, slightly dextrinoid to dextrinoid in 
<normalizedToken originalValue="Melzer’s">Melzer's</normalizedToken>
reagent; ornamentation (7)8-9 solid ridges regularly and longitudinally arranged under light microscope, sometimes anastomosing under SEM, 2-3 
<normalizedToken originalValue="µm">µm</normalizedToken>
tall and 2-2.5 
<normalizedToken originalValue="µm">µm</normalizedToken>
wide at the base; hilar appendage prominent, 2.5-5 
<normalizedToken originalValue="µm">µm</normalizedToken>
long with a terminal hilum. Basidia 4-spored, (26 
<normalizedToken originalValue="–)26.1–32.3–36(–">-)26.1-32.3-36(-</normalizedToken>
36) 
<normalizedToken originalValue="×">x</normalizedToken>
(8 
<normalizedToken originalValue="–)8–9.5–11(–">-)8-9.5-11(-</normalizedToken>
11) 
<normalizedToken originalValue="µm">µm</normalizedToken>
(n = 20; from holotype only), clavate to cylindrical, hyaline in water, 5% KOH and NH4OH, yellowish hyaline in 
<normalizedToken originalValue="Melzer’s">Melzer's</normalizedToken>
reagent; sterigmata broken by spore release, stout, 3-4 
<normalizedToken originalValue="µm">µm</normalizedToken>
long. Cystidia none observed. Hymenophoral trama 60-130 
<normalizedToken originalValue="µm">µm</normalizedToken>
thick, irregular, with a narrow, central layer of subparallel to loosely interwoven, 3-7(8) 
<normalizedToken originalValue="µm">µm</normalizedToken>
wide, thin-walled hyphae, slightly gelatinised, hyaline in water, 5% KOH and NH4OH. Peridiopellis a tomentum 45-120 
<normalizedToken originalValue="µm">µm</normalizedToken>
thick, poorly differentiated, composed of thin-walled, 3-10 
<normalizedToken originalValue="µm">µm</normalizedToken>
wide hyphae, anastomosing at places and covered with yellowish brown incrustations on the surface at places, otherwise hyaline in water, 5% KOH and NH4OH, inamyloid. Clamp connections not seen in any of the tissues.
</paragraph>
<caption pageId="4" pageNumber="67">
<paragraph pageId="4" pageNumber="67">
Figure 2. Basidiomata of 
<taxonomicName class="Agaricomycetes" family="Boletaceae" genus="Rhodactina" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhodactina rostratispora" order="Boletales" pageId="4" pageNumber="67" phylum="Basidiomycota" rank="species" species="rostratispora">Rhodactina rostratispora</taxonomicName>
A S. Vadthanarat 170 (holotype) B S. Vadthanarat 206 C S. Vadthanarat 208 D O. 
<normalizedToken originalValue="Raspé">Raspe</normalizedToken>
1055 E S. Vadthanarat 406, showing one basidioma (white arrow) that had a strong fruity alcoholic smell F Hymenophore turned dark purple to greyish violet with 5% KOH (white arrow). Scale bars: 
<normalizedToken originalValue="A–E">A-E</normalizedToken>
= 1 cm; F =0.5 cm.
</paragraph>
</caption>
<caption pageId="4" pageNumber="67">
<paragraph pageId="4" pageNumber="67">
Figure 3. Microscopic features of 
<taxonomicName class="Agaricomycetes" family="Boletaceae" genus="Rhodactina" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhodactina rostratispora" order="Boletales" pageId="4" pageNumber="67" phylum="Basidiomycota" rank="species" species="rostratispora">Rhodactina rostratispora</taxonomicName>
A Basidiospores in side view, polar view and optical section B Basidia C Hymenium showing basidia and basidioles D Peridiopellis covered with some encrustations. All drawings were made from the type. Scale bars: A = 10 
<normalizedToken originalValue="µm">µm</normalizedToken>
; 
<normalizedToken originalValue="B–C">B-C</normalizedToken>
= 20 
<normalizedToken originalValue="µm">µm</normalizedToken>
; D = 50 
<normalizedToken originalValue="µm">µm</normalizedToken>
.
</paragraph>
</caption>
<caption pageId="4" pageNumber="67">
<paragraph pageId="4" pageNumber="67">
Figure 4. Scanning electron micrographs of basidiospores 
<normalizedToken originalValue="A–B">A-B</normalizedToken>
<taxonomicName class="Agaricomycetes" family="Boletaceae" genus="Rhodactina" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhodactina himalayensis" order="Boletales" pageId="4" pageNumber="67" phylum="Basidiomycota" rank="species" species="himalayensis">Rhodactina himalayensis</taxonomicName>
(CMU25117) showing the basidiospores with 6-7 longitudinal ridges 
<normalizedToken originalValue="C–D">C-D</normalizedToken>
<taxonomicName class="Agaricomycetes" family="Boletaceae" genus="Rhodactina" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhodactina incarnata" order="Boletales" pageId="4" pageNumber="67" phylum="Basidiomycota" rank="species" species="incarnata">Rhodactina incarnata</taxonomicName>
(CMU25116, holotype) showing the basidiospores with 8-9 longitudinal ridges 
<normalizedToken originalValue="E–F">E-F</normalizedToken>
<taxonomicName class="Agaricomycetes" family="Boletaceae" genus="Rhodactina" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhodactina rostratispora" order="Boletales" pageId="4" pageNumber="67" phylum="Basidiomycota" rank="species" species="rostratispora">Rhodactina rostratispora</taxonomicName>
(O. 
<normalizedToken originalValue="Raspé">Raspe</normalizedToken>
1055) showing the basidiospores with 8-9 longitudinal ridges, the wide and prominent hilar appendage (ha), a terminal hilum (th) and anastomosing ridges in some spores (as).
</paragraph>
</caption>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="68" pageId="4" pageNumber="67" type="habit and habitat">
<paragraph pageId="4" pageNumber="67">Habit and habitat.</paragraph>
<paragraph lastPageId="5" lastPageNumber="68" pageId="4" pageNumber="67">
Subepigeal, solitary to gregarious (4-7 basidiomata), or fasciculate by 2-5 basidiomata, on sandy soil in dipterocarp forest dominated by 
<taxonomicName class="Magnoliopsida" family="Dipterocarpaceae" genus="Dipterocarpus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dipterocarpus tuberculatus" order="Malvales" pageId="5" pageNumber="68" phylum="Tracheophyta" rank="species" species="tuberculatus">
<pageBreakToken pageId="5" pageNumber="68" start="start">Dipterocarpus</pageBreakToken>
tuberculatus
</taxonomicName>
, 
<taxonomicName lsidName="D. intricatus" pageId="5" pageNumber="68" rank="species" species="intricatus">D. intricatus</taxonomicName>
, 
<taxonomicName lsidName="D. obtusifolius" pageId="5" pageNumber="68" rank="species" species="obtusifolius">D. obtusifolius</taxonomicName>
, 
<taxonomicName class="Magnoliopsida" family="Dipterocarpaceae" genus="Shorea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Shorea obtusa" order="Malvales" pageId="5" pageNumber="68" phylum="Tracheophyta" rank="species" species="obtusa">Shorea obtusa</taxonomicName>
, 
<taxonomicName lsidName="S. siamensis" pageId="5" pageNumber="68" rank="species" species="siamensis">S. siamensis</taxonomicName>
and 
<taxonomicName class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eucalyptus" order="Myrtales" pageId="5" pageNumber="68" phylum="Tracheophyta" rank="genus">Eucalyptus</taxonomicName>
sp.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="68" type="distribution">
<paragraph pageId="5" pageNumber="68">Known distribution.</paragraph>
<paragraph pageId="5" pageNumber="68">Currently found only from Ubon Ratchathani province, northeastern Thailand.</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="69" pageId="5" pageNumber="68" type="materials_examined">
<paragraph pageId="5" pageNumber="68">Additional specimens examined.</paragraph>
<paragraph pageId="5" pageNumber="68">
<taxonomicName class="Agaricomycetes" family="Boletaceae" genus="Rhodactina" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhodactina rostratispora" order="Boletales" pageId="5" pageNumber="68" phylum="Basidiomycota" rank="species" species="rostratispora">Rhodactina rostratispora</taxonomicName>
.-THAILAND, Ubon Ratchathani Province, Trakan Phuet Phon District, Don Khok Tam Lae community forest, 
<geoCoordinate direction="north" orientation="latitude" precision="1" value="15.5945">
15°35'40.2&quot; 
<normalizedToken originalValue="N–">N-</normalizedToken>
</geoCoordinate>
<geoCoordinate direction="east" orientation="longitude" precision="1" value="105.1105">105°06'37.8&quot;E</geoCoordinate>
, elev. 150 m., 28 July 2015, S. Vadthanarat 169, (CMUB, BR); ibid. 
<geoCoordinate direction="north" orientation="latitude" precision="1" value="15.594861">
15°35'41.5&quot; 
<normalizedToken originalValue="N–">N-</normalizedToken>
</geoCoordinate>
<geoCoordinate direction="east" orientation="longitude" precision="1" value="105.10983">105°06'35.4&quot;E</geoCoordinate>
, elev. 150 m., 28 July 2015, O. 
<normalizedToken originalValue="Raspé">Raspe</normalizedToken>
1055, (CMUB, BR); ibid. 
<geoCoordinate direction="north" orientation="latitude" precision="1" value="15.596749">15°35'48.3&quot;N</geoCoordinate>
- 
<geoCoordinate direction="east" orientation="longitude" precision="1" value="105.10997">105°06'35.9&quot;E</geoCoordinate>
, elev. 150 m., 6 August 2015, S. Vadthanarat 206, (CMUB, BR); ibid. 
<geoCoordinate direction="north" orientation="latitude" precision="1" value="15.597889">
15°35'52.4&quot; 
<normalizedToken originalValue="N–">N-</normalizedToken>
</geoCoordinate>
<geoCoordinate direction="east" orientation="longitude" precision="1" value="105.11144">105°06'41.2&quot;E</geoCoordinate>
, elev. 150 m., 6 August 2015, S. Vadthanarat 208, (CMUB, BR); ibid. 
<geoCoordinate direction="north" orientation="latitude" precision="1" value="15.598916">
15°35'56.1&quot; 
<normalizedToken originalValue="N–">N-</normalizedToken>
</geoCoordinate>
<geoCoordinate direction="east" orientation="longitude" precision="1" value="105.1108">105°06'38.9&quot;E</geoCoordinate>
, elev. 150 m., 6 August 2015, S. Vadthanarat 212, (CMUB, BR); ibid. 
<geoCoordinate direction="north" orientation="latitude" precision="1" value="15.600722">
15°36'2.6&quot; 
<normalizedToken originalValue="N–">N-</normalizedToken>
</geoCoordinate>
<geoCoordinate direction="east" orientation="longitude" precision="1" value="105.11019">105°06'36.7&quot;E</geoCoordinate>
, elev. 150 m., 14 May 2017, S. Vadthanarat 376, (CMUB, BR); Ban Huay Fai community forest, 
<geoCoordinate direction="north" orientation="latitude" precision="1" value="15.545195">
15°32'42.7&quot; 
<normalizedToken originalValue="N–">N-</normalizedToken>
</geoCoordinate>
<geoCoordinate direction="east" orientation="longitude" precision="1" value="105.17119">105°10'16.3&quot;E</geoCoordinate>
, elev. 160 m., 15 July 2017, S. Vadthanarat 406, (CMUB, BR).
</paragraph>
<paragraph pageId="5" pageNumber="68">
<taxonomicName lsidName="R. himalayensis" pageId="5" pageNumber="68" rank="species" species="himalayensis">R. himalayensis</taxonomicName>
. - THAILAND, Chiang Mai Province, Doi Suthep-Pui National Park, forest behind Channel 9 TV station, 4 August 2000, Saisamorn Lumyong, Pipob Lumyong, Rarunee Sanmee and B. Dell 2254 (CMU25117).
</paragraph>
<paragraph pageId="6" pageNumber="69">
<taxonomicName lsidName="R. incarnata" pageId="6" pageNumber="69" rank="species" species="incarnata">
<pageBreakToken pageId="6" pageNumber="69" start="start">R</pageBreakToken>
. incarnata
</taxonomicName>
. - THAILAND, Chiang Mai Province, Sanpatong District, Mae Wang, Conservation forest, Sanpatong-Ban Guard Rd., 24 July 2002, Saisamorn Lumyong, Pipob Lumyong, Rarunee Sanmee and Zhu L. Yang 45209 (CMU25116; holotype).
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="69" type="remarks">
<paragraph pageId="6" pageNumber="69">Remarks.</paragraph>
<paragraph pageId="6" pageNumber="69">
<taxonomicName class="Agaricomycetes" family="Boletaceae" genus="Rhodactina" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhodactina rostratispora" order="Boletales" pageId="6" pageNumber="69" phylum="Basidiomycota" rank="species" species="rostratispora">Rhodactina rostratispora</taxonomicName>
is characterised by its basidiospores having a markedly prominent hilar appendage (2.5-5 
<normalizedToken originalValue="µm">µm</normalizedToken>
long, 3.5-5 
<normalizedToken originalValue="µm">µm</normalizedToken>
wide), with a terminal hilum; ornamentation consisting of (7)8-9 longitudinal ridges, and (11.5 
<normalizedToken originalValue="–)12–13.6–15(–">-)12-13.6-15(-</normalizedToken>
16) 
<normalizedToken originalValue="×">x</normalizedToken>
(10 
<normalizedToken originalValue="–)10.5–11.7–13(–">-)10.5-11.7-13(-</normalizedToken>
14) 
<normalizedToken originalValue="µm">µm</normalizedToken>
. 
<taxonomicName lsidName="R. himalayensis" pageId="6" pageNumber="69" rank="species" species="himalayensis">R. himalayensis</taxonomicName>
has larger basidiospores (15-20 
<normalizedToken originalValue="×">x</normalizedToken>
12.5-18 
<normalizedToken originalValue="µm">µm</normalizedToken>
) without prominent hilar appendage, with fewer [(5)6-7(8)], broader ridges, while 
<taxonomicName lsidName="R. incarnata" pageId="6" pageNumber="69" rank="species" species="incarnata">R. incarnata</taxonomicName>
has a similar spore size (10-13 
<normalizedToken originalValue="×">x</normalizedToken>
10-12 
<normalizedToken originalValue="µm">µm</normalizedToken>
) and the same number of spore ridges [(7)8-9(10)] as the new species, but it does not have the prominent hilar appendage.
</paragraph>
<paragraph pageId="6" pageNumber="69">
In one 
<taxonomicName lsidName="R. rostratispora" pageId="6" pageNumber="69" rank="species" species="rostratispora">R. rostratispora</taxonomicName>
specimen (S. Vadthanarat 208), abnormal spores were observed. Those spores were elongated, 21-24 
<normalizedToken originalValue="×">x</normalizedToken>
4-8 
<normalizedToken originalValue="µm">µm</normalizedToken>
, thick-walled, narrowly fusiform to bacilliform, with or without longitudinal ridges, more or less constricted in the middle. They were usually found attached to apparently normal basidia with four sterigmata.
</paragraph>
</subSubSection>
</treatment>
</document>