837 lines
103 KiB
XML
837 lines
103 KiB
XML
<document id="CDED1AFF0129C1205A66856438454269" ID-CLB-Dataset="9460" ID-DOI="10.11646/zootaxa.4885.2.2" ID-GBIF-Dataset="8c20ec52-8244-420c-8053-bd68d8fd4bc0" ID-ISSN="1175-5326" ID-Zenodo-Dep="4296526" ID-ZooBank="BD22C7FA-9861-4C90-9E36-ECFFB6BE9129" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1606675969140" checkinUser="plazi" docAuthor="Sayyadzadeh, Golnaz & Esmaeili, Hamid Reza" docDate="2020" docId="C54A8791C8215830FF49FE5CFC81FA20" docLanguage="en" docName="zootaxa.4885.2.2.pdf" docOrigin="Zootaxa 4885 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oxynoemacheilus marunensis Sayyadzadeh & Esmaeili 2020, new species" docType="treatment" docVersion="9" lastPageNumber="200" masterDocId="3973FFE9C827583BFFDEFF89FFC4FFE3" masterDocTitle="Oxynoemacheilus marunensis, a new loach species from the Persian Gulf basin with remarks on O. frenatus (Teleostei: Nemacheilidae)" masterLastPageNumber="206" masterPageNumber="189" pageNumber="195" updateTime="1698846764549" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="D6E1B1EFBBFE5D06C4CFADB99C71F337">
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<mods:title id="4BEAE2628D8DE3EE7D79AD0160A62CA5">Oxynoemacheilus marunensis, a new loach species from the Persian Gulf basin with remarks on O. frenatus (Teleostei: Nemacheilidae)</mods:title>
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<mods:name id="215BBAD6E0C05E4EF78FC9B6136942A6" type="personal">
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<mods:roleTerm id="ACF8B8545E2CE72BEFFCC83ACB6DD14B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D6068733835AB981BD665BF98213EBFD">Sayyadzadeh, Golnaz</mods:namePart>
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<mods:affiliation id="7736B249F70F8FE4A2B7D09375B66720">Ichthyology and Molecular Systematics Research Laboratory, Department of Biology, School of Science, Shiraz University, Shiraz, Iran. g. sayyadzadeh 92 @ gmail. com; https: // orcid. org / 0000 - 0003 - 0758 - 6502</mods:affiliation>
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<mods:roleTerm id="54554101CFC47FD44767A6134A65DA5D">Author</mods:roleTerm>
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<mods:namePart id="0D08708A01A077D8192E6CAF1A41CD6B">Esmaeili, Hamid Reza</mods:namePart>
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<mods:affiliation id="F4492C3FB11ABA3DDDAC16E41CD47A91">Ichthyology and Molecular Systematics Research Laboratory, Department of Biology, School of Science, Shiraz University, Shiraz, Iran. g. sayyadzadeh 92 @ gmail. com; https: // orcid. org / 0000 - 0003 - 0758 - 6502 & Center for Hydrobiology and Aquatic Biotechnology, Shiraz University, Shiraz, Iran.</mods:affiliation>
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<mods:typeOfResource id="7E3B38A9AFBF856A8EF67DF1EA72AC45">text</mods:typeOfResource>
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<mods:title id="CEA2DB92CFA82DBF8C083DA4E75202EC">Zootaxa</mods:title>
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<mods:part id="C5EE0194FBBFFEB5802F1B5F62EE4768">
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<mods:date id="7EF53262F7FF0F179C0CCEB5C40670C8">2020</mods:date>
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<mods:number id="A1E72E941008168B88E7275171AF7C28">2020-11-25</mods:number>
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<mods:detail id="022A2238C8224499388F1135018DCDEC" type="volume">
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<mods:number id="41EA3BE033320AE73C3EABBF3E51DF95">4885</mods:number>
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<mods:detail id="17D8210B35D8E2248E36FBE103D8689C" type="issue">
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<mods:identifier id="2BD8B94AA75AD8F013706820C8C281C9" type="ZooBank">BD22C7FA-9861-4C90-9E36-ECFFB6BE9129</mods:identifier>
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<treatment id="C54A8791C8215830FF49FE5CFC81FA20" ID-DOI="http://doi.org/10.5281/zenodo.4324013" ID-GBIF-Taxon="170606583" ID-Zenodo-Dep="4324013" LSID="urn:lsid:plazi:treatment:C54A8791C8215830FF49FE5CFC81FA20" httpUri="http://treatment.plazi.org/id/C54A8791C8215830FF49FE5CFC81FA20" lastPageId="11" lastPageNumber="200" pageId="6" pageNumber="195">
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<subSubSection id="05F9650CC821583DFF49FE5CFD60FE0C" box="[151,676,469,496]" pageId="6" pageNumber="195" type="nomenclature">
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<paragraph id="4D5C3687C821583DFF49FE5CFD60FE0C" blockId="6.[151,676,469,531]" box="[151,676,469,496]" pageId="6" pageNumber="195">
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<heading id="161481EBC821583DFF49FE5CFD60FE0C" bold="true" box="[151,676,469,496]" fontSize="11" level="1" pageId="6" pageNumber="195" reason="1">
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<emphasis id="7F97EA95C821583DFF49FE5CFD60FE0C" bold="true" box="[151,676,469,496]" pageId="6" pageNumber="195">
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<taxonomicName id="8AE34D04C821583DFF49FE5CFDC1FE0C" authority="Sayyadzadeh & Esmaeili, 2020" authorityName="Sayyadzadeh & Esmaeili" authorityYear="2020" box="[151,517,469,496]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="195" phylum="Chordata" rank="species" species="marunensis" status="sp. nov.">
|
||
<emphasis id="7F97EA95C821583DFF49FE5CFDC1FE0C" bold="true" box="[151,517,469,496]" italics="true" pageId="6" pageNumber="195">Oxynoemacheilus marunensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="64A457EEC821583DFDCAFE5CFD60FE0C" box="[532,676,469,495]" pageId="6" pageNumber="195" rank="species">new species</taxonomicNameLabel>
|
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</emphasis>
|
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="05F9650CC821583DFF49FE70FED6FDF0" box="[151,274,505,531]" pageId="6" pageNumber="195" type="description">
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<paragraph id="4D5C3687C821583DFF49FE70FED6FDF0" blockId="6.[151,676,469,531]" box="[151,274,505,531]" pageId="6" pageNumber="195">
|
||
(
|
||
<figureCitation id="D5D82A02C821583DFF41FE70FECFFDF0" box="[159,267,505,531]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStartId-0="7.[151,250,1050,1074]" captionStartId-1="7.[151,250,1979,2003]" captionStartId-2="8.[151,250,1044,1068]" captionStartId-3="8.[151,250,1981,2005]" captionStartId-4="9.[151,250,2007,2031]" captionStartId-5="10.[151,250,1144,1168]" captionTargetBox-0="[211,1376,181,1026]" captionTargetBox-1="[215,1372,1099,1956]" captionTargetBox-2="[209,1378,181,1020]" captionTargetBox-3="[209,1378,1128,1957]" captionTargetBox-4="[403,1184,885,1984]" captionTargetBox-5="[170,1417,705,1120]" captionTargetId-0="figure-19@7.[211,1376,181,1026]" captionTargetId-1="figure-36@7.[215,1372,1099,1956]" captionTargetId-2="figure-25@8.[209,1378,181,1020]" captionTargetId-3="figure-55@8.[209,1378,1128,1957]" captionTargetId-4="figure-357@9.[403,1184,885,1984]" captionTargetId-5="figure-267@10.[170,1417,705,1120]" captionTargetPageId-0="7" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="10" captionText-0="FIGURE 2. Oxynoemacheilus marunensis, ZM-CBSU H2509, holotype, 43 mm SL; Iran: Marun River." captionText-1="FIGURE 3. Oxynoemacheilus marunensis, paratypes; a, ZM-CBSU H2510, 36 mm SL; b, ZM-CBSU H2518, 32 mm SL; c, ZM-CBSU H2519, 27 mm SL; Iran: Marun River." captionText-2="FIGURE 4. Oxynoemacheilus marunensis, paratypes; a, ZM-CBSU H2510, 36 mm SL; b, ZM-CBSU H2518, 32 mm SL; c, ZM-CBSU H2519, 27 mm SL; Iran: Marun River." captionText-3="FIGURE 5. Oxynoemacheilus marunensis, paratypes; a, ZM-CBSU H2510, 36 mm SL; b, ZM-CBSU H2518, 32 mm SL; c, ZM-CBSU H2519, 27 mm SL; Iran: Marun River." captionText-4="FIGURE 6. Oxynoemacheilus marunensis, ZM-CBSU H2509, holotype, 43 mm SL; ventral view of Head, Iran: Marun River." captionText-5="FIGURE 7. Oxynoemacheilus marunensis, Uncatalogued, about 35 mm SL; Iran: Marun River." figureDoi-0="http://doi.org/10.5281/zenodo.4296530" figureDoi-1="http://doi.org/10.5281/zenodo.4296532" figureDoi-2="http://doi.org/10.5281/zenodo.4296534" figureDoi-3="http://doi.org/10.5281/zenodo.4296536" figureDoi-4="http://doi.org/10.5281/zenodo.4296538" figureDoi-5="http://doi.org/10.5281/zenodo.4296540" httpUri-0="https://zenodo.org/record/4296530/files/figure.png" httpUri-1="https://zenodo.org/record/4296532/files/figure.png" httpUri-2="https://zenodo.org/record/4296534/files/figure.png" httpUri-3="https://zenodo.org/record/4296536/files/figure.png" httpUri-4="https://zenodo.org/record/4296538/files/figure.png" httpUri-5="https://zenodo.org/record/4296540/files/figure.png" pageId="6" pageNumber="195">Figs. 2–7</figureCitation>
|
||
)
|
||
</paragraph>
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||
</subSubSection>
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<subSubSection id="05F9650CC821583DFF49FDB5FC15FC94" pageId="6" pageNumber="195" type="materials_examined">
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<paragraph id="4D5C3687C821583DFF49FDB5FD3CFD99" blockId="6.[151,1437,572,2039]" pageId="6" pageNumber="195">
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<materialsCitation id="FD8B3CDAC821583DFF49FDB5FD3CFD99" ID-GBIF-Occurrence="2988202304" collectingDate="2019-12-11" collectorName="G. Sayyadzadeh & F. Zarei" country="Iran" county="Marun" latitude="30.65611" location="Kharestan" longLatPrecision="20" longitude="50.20861" municipality="Maroon River" pageId="6" pageNumber="195" specimenCount="1" stateProvince="Khuzestan" typeStatus="holotype">
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<emphasis id="7F97EA95C821583DFF49FDB5FEC9FDB5" bold="true" box="[151,269,572,598]" pageId="6" pageNumber="195">
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<typeStatus id="92588825C821583DFF49FDB5FECCFDB5" box="[151,264,572,598]" pageId="6" pageNumber="195" type="holotype">Holotype</typeStatus>
|
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:
|
||
</emphasis>
|
||
ZM-CBSU H2509, 43 mm SL;
|
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<collectingCountry id="35F47617C821583DFDA6FDB4FD6DFDB4" box="[632,681,573,599]" name="Iran" pageId="6" pageNumber="195">Iran</collectingCountry>
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||
:
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<collectingRegion id="8F27F865C821583DFD6AFDB5FCABFDB4" box="[692,879,572,599]" country="Iran" name="Khuzestan" pageId="6" pageNumber="195">Khuzestan prov.</collectingRegion>
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:
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<collectingCounty id="A43D4E0BC821583DFCA7FDB5FC07FDB5" box="[889,963,572,598]" pageId="6" pageNumber="195">Marun</collectingCounty>
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/
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<collectingMunicipality id="AD38ACFDC821583DFC12FDB5FBADFDB5" box="[972,1129,572,598]" pageId="6" pageNumber="195">Maroon River</collectingMunicipality>
|
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at
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||
<location id="483C605CC821583DFB52FDB5FAC4FDB5" LSID="urn:lsid:plazi:treatment:C54A8791C8215830FF49FE5CFC81FA20:483C605CC821583DFB52FDB5FAC4FDB5" box="[1164,1280,572,598]" country="Iran" county="Marun" latitude="30.65611" longLatPrecision="20" longitude="50.20861" municipality="Maroon River" name="Kharestan" pageId="6" pageNumber="195" stateProvince="Khuzestan">Kharestan</location>
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,
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<geoCoordinate id="28D75040C821583DFAD2FDB5FA58FDB4" box="[1292,1436,572,599]" degrees="30" direction="north" minutes="39" orientation="latitude" pageId="6" pageNumber="195" precision="15" seconds="22" value="30.65611">30°39’22” N</geoCoordinate>
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<geoCoordinate id="28D75040C821583DFF49FDE9FEE3FD98" box="[151,295,608,635]" degrees="50" direction="east" minutes="12" orientation="longitude" pageId="6" pageNumber="195" precision="15" seconds="31" value="50.20861">50°12’31” E</geoCoordinate>
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;
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<collectorName id="E0165351C821583DFEEDFDE9FE28FD99" box="[307,492,608,634]" pageId="6" pageNumber="195">G. Sayyadzadeh</collectorName>
|
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,
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||
<collectorName id="E0165351C821583DFE29FDE8FD94FD99" box="[503,592,608,635]" pageId="6" pageNumber="195">F. Zarei</collectorName>
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,
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<date id="395D1047C821583DFD85FDE9FD30FD99" box="[603,756,608,635]" pageId="6" pageNumber="195" value="2019-12-11">
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<collectingDate id="2919E9AFC821583DFD85FDE9FD30FD99" box="[603,756,608,635]" pageId="6" pageNumber="195" value="2019-12-11">11 Dec. 2019</collectingDate>
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</date>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="4D5C3687C821583DFF19FD0DFE01FD05" blockId="6.[151,1437,572,2039]" pageId="6" pageNumber="195">
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<materialsCitation id="FD8B3CDAC821583DFF19FD0DFE05FD05" ID-GBIF-Occurrence="2988202302" collectingDate="2013-12-23" collectionCode="R" collectorName="Esmaeili, G. Sayyadzadeh, M. Amini, M. Masoudi" country="Iran" latitude="30.659721" location="Kharestan" longLatPrecision="20" longitude="50.21472" municipality="Marun River" pageId="6" pageNumber="195" specimenCount="2" stateProvince="Khuzestan prov." typeStatus="paratype">
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<emphasis id="7F97EA95C821583DFF19FD0DFE8CFD7D" bold="true" box="[199,328,644,670]" pageId="6" pageNumber="195">
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<typeStatus id="92588825C821583DFF19FD0DFE87FD7D" box="[199,323,644,670]" pageId="6" pageNumber="195" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
ZM-CBSU H2510, 36,
|
||
<quantity id="8A1B9B62C821583DFD8FFD0DFD0DFD7C" box="[593,713,644,671]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" metricValueMax="3.7" metricValueMin="2.7" pageId="6" pageNumber="195" unit="mm" value="32.0" valueMax="37.0" valueMin="27.0">27–37 mm</quantity>
|
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SL; same as
|
||
<typeStatus id="92588825C821583DFC86FD0DFC7AFD7D" box="[856,958,644,670]" pageId="6" pageNumber="195" type="holotype">holotype</typeStatus>
|
||
. ZM-CBSU H1998, 9,
|
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<quantity id="8A1B9B62C821583DFB63FD0DFAF0FD7C" box="[1213,1332,644,671]" metricMagnitude="-2" metricUnit="m" metricValue="3.55" metricValueMax="4.1" metricValueMin="3.0" pageId="6" pageNumber="195" unit="mm" value="35.5" valueMax="41.0" valueMin="30.0">30–41 mm</quantity>
|
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SL;
|
||
<collectingCountry id="35F47617C821583DFAB9FD0CFA5CFD7C" box="[1383,1432,645,671]" name="Iran" pageId="6" pageNumber="195">Iran</collectingCountry>
|
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:
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<collectingRegion id="8F27F865C821583DFF49FD21FE95FD20" box="[151,337,680,707]" country="Iran" name="Khuzestan" pageId="6" pageNumber="195">Khuzestan prov.</collectingRegion>
|
||
:
|
||
<collectingMunicipality id="AD38ACFDC821583DFE85FD20FE2EFD21" box="[347,490,680,707]" pageId="6" pageNumber="195">Marun River</collectingMunicipality>
|
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at
|
||
<location id="483C605CC821583DFDD5FD21FDBBFD21" LSID="urn:lsid:plazi:treatment:C54A8791C8215830FF49FE5CFC81FA20:483C605CC821583DFDD5FD21FDBBFD21" box="[523,639,680,706]" country="Iran" latitude="30.659721" longLatPrecision="20" longitude="50.21472" municipality="Marun River" name="Kharestan" pageId="6" pageNumber="195" stateProvince="Khuzestan prov.">Kharestan</location>
|
||
,
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<geoCoordinate id="28D75040C821583DFD54FD21FCDEFD20" box="[650,794,680,707]" degrees="30" direction="north" minutes="39" orientation="latitude" pageId="6" pageNumber="195" precision="15" seconds="35" value="30.659721">30°39’35” N</geoCoordinate>
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<geoCoordinate id="28D75040C821583DFCFEFD21FC74FD20" box="[800,944,680,707]" degrees="50" direction="east" minutes="12" orientation="longitude" pageId="6" pageNumber="195" precision="15" seconds="53" value="50.21472">50°12’53” E</geoCoordinate>
|
||
; H.
|
||
<collectionCode id="2BF2AE42C821583DFC08FD20FC28FD20" box="[982,1004,681,707]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="195">R</collectionCode>
|
||
.
|
||
<collectorName id="E0165351C821583DFC28FD21FEE7FD05" pageId="6" pageNumber="195">Esmaeili, G. Sayyadzadeh, M. Amini, M. Masoudi</collectorName>
|
||
,
|
||
<date id="395D1047C821583DFEF0FD45FE05FD05" box="[302,449,716,743]" pageId="6" pageNumber="195" value="2013-12-23">
|
||
<collectingDate id="2919E9AFC821583DFEF0FD45FE05FD05" box="[302,449,716,743]" pageId="6" pageNumber="195" value="2013-12-23">23 Dec 2013</collectingDate>
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||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="4D5C3687C821583DFF19FD79FC15FC94" blockId="6.[151,1437,572,2039]" pageId="6" pageNumber="195">
|
||
<emphasis id="7F97EA95C821583DFF19FD79FD39FCE9" bold="true" box="[199,765,752,778]" pageId="6" pageNumber="195">Material used in the molecular genetic analysis:</emphasis>
|
||
<materialsCitation id="FD8B3CDAC821583DFCDDFD79FB59FCCC" ID-GBIF-Occurrence="2988202307" accessionNumber="MW136427" collectionCode="MW" country="Iran" latitude="30.670834" location="Emam Reza village" longLatPrecision="20" longitude="50.30361" municipality="Maroun River" pageId="6" pageNumber="195" specimenCount="1" stateProvince="Khuzestan">
|
||
ZM-CBSU M1735;
|
||
<collectingCountry id="35F47617C821583DFC3BFD78FBD2FCE8" box="[997,1046,753,779]" name="Iran" pageId="6" pageNumber="195">Iran</collectingCountry>
|
||
:
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<collectingRegion id="8F27F865C821583DFBFFFD79FB1FFCE8" box="[1057,1243,752,779]" country="Iran" name="Khuzestan" pageId="6" pageNumber="195">Khuzestan prov.</collectingRegion>
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||
:
|
||
<collectingMunicipality id="AD38ACFDC821583DFB3AFD78FA45FCE9" box="[1252,1409,752,779]" pageId="6" pageNumber="195">Maroun River</collectingMunicipality>
|
||
at
|
||
<location id="483C605CC821583DFF49FC9CFEB2FCCD" LSID="urn:lsid:plazi:treatment:C54A8791C8215830FF49FE5CFC81FA20:483C605CC821583DFF49FC9CFEB2FCCD" box="[151,374,788,815]" country="Iran" latitude="30.670834" longLatPrecision="20" longitude="50.30361" municipality="Maroun River" name="Emam Reza village" pageId="6" pageNumber="195" stateProvince="Khuzestan">Emam Reza village</location>
|
||
,
|
||
<geoCoordinate id="28D75040C821583DFE5CFC9DFDD7FCCC" box="[386,531,788,815]" degrees="30" direction="north" minutes="40" orientation="latitude" pageId="6" pageNumber="195" precision="15" seconds="15" value="30.670834">30°40‘15“ N</geoCoordinate>
|
||
<geoCoordinate id="28D75040C821583DFDC4FC9DFD6CFCCC" box="[538,680,788,815]" degrees="50" direction="east" minutes="18" orientation="longitude" pageId="6" pageNumber="195" precision="15" seconds="13" value="50.30361">50°18‘13“ E</geoCoordinate>
|
||
(GenBank accession numbers:
|
||
<accessionNumber id="52B0AB64C821583DFBD3FC9DFB51FCCC" box="[1037,1173,788,815]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MW136427" pageId="6" pageNumber="195">
|
||
<collectionCode id="2BF2AE42C821583DFBD3FC9DFB86FCCC" box="[1037,1090,788,815]" country="Netherlands" name="Museum Wasmann" pageId="6" pageNumber="195">MW</collectionCode>
|
||
136427
|
||
</accessionNumber>
|
||
)
|
||
</materialsCitation>
|
||
.—
|
||
<materialsCitation id="FD8B3CDAC821583DFB63FC9DFC09FC94" ID-GBIF-Occurrence="2988202305" accessionNumber="MW136428, MW136429, MW136430, MW136431" collectionCode="MW" country="Iran" latitude="30.659721" location="Kharestan" longLatPrecision="20" longitude="50.21472" municipality="Marun River" pageId="6" pageNumber="195" specimenCount="1" stateProvince="Khuzestan">
|
||
ZM-CBSU M1736, M1737, M2812, M2813;
|
||
<collectingCountry id="35F47617C821583DFE64FCB0FE2FFCB0" box="[442,491,825,851]" name="Iran" pageId="6" pageNumber="195">Iran</collectingCountry>
|
||
:
|
||
<collectingRegion id="8F27F865C821583DFE29FCB1FD77FCB0" box="[503,691,824,851]" country="Iran" name="Khuzestan" pageId="6" pageNumber="195">Khuzestan prov.</collectingRegion>
|
||
:
|
||
<collectingMunicipality id="AD38ACFDC821583DFD61FCB0FC94FCB1" box="[703,848,824,851]" pageId="6" pageNumber="195">Marun River</collectingMunicipality>
|
||
at
|
||
<location id="483C605CC821583DFCA8FCB1FC2EFCB1" LSID="urn:lsid:plazi:treatment:C54A8791C8215830FF49FE5CFC81FA20:483C605CC821583DFCA8FCB1FC2EFCB1" box="[886,1002,824,850]" country="Iran" latitude="30.659721" longLatPrecision="20" longitude="50.21472" municipality="Marun River" name="Kharestan" pageId="6" pageNumber="195" stateProvince="Khuzestan">Kharestan</location>
|
||
,
|
||
<geoCoordinate id="28D75040C821583DFC29FCB1FB4DFCB0" box="[1015,1161,824,851]" degrees="30" direction="north" minutes="39" orientation="latitude" pageId="6" pageNumber="195" precision="15" seconds="35" value="30.659721">30°39’35” N</geoCoordinate>
|
||
<geoCoordinate id="28D75040C821583DFB4CFCB1FAE4FCB0" box="[1170,1312,824,851]" degrees="50" direction="east" minutes="12" orientation="longitude" pageId="6" pageNumber="195" precision="15" seconds="53" value="50.21472">50°12’53” E</geoCoordinate>
|
||
(GenBank accession numbers:
|
||
<accessionNumber id="52B0AB64C821583DFEA7FCD5FDC6FC95" box="[377,514,860,886]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MW136428" pageId="6" pageNumber="195">
|
||
<collectionCode id="2BF2AE42C821583DFEA7FCD5FE6BFC95" box="[377,431,860,886]" country="Netherlands" name="Museum Wasmann" pageId="6" pageNumber="195">MW</collectionCode>
|
||
136428
|
||
</accessionNumber>
|
||
,
|
||
<accessionNumber id="52B0AB64C821583DFDD0FCD5FD5DFC95" box="[526,665,860,886]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MW136429" pageId="6" pageNumber="195">
|
||
<collectionCode id="2BF2AE42C821583DFDD0FCD5FD80FC95" box="[526,580,860,886]" country="Netherlands" name="Museum Wasmann" pageId="6" pageNumber="195">MW</collectionCode>
|
||
136429
|
||
</accessionNumber>
|
||
,
|
||
<accessionNumber id="52B0AB64C821583DFD7DFCD5FCEAFC95" box="[675,814,860,886]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MW136430" pageId="6" pageNumber="195">
|
||
<collectionCode id="2BF2AE42C821583DFD7DFCD5FD1CFC95" box="[675,728,860,886]" country="Netherlands" name="Museum Wasmann" pageId="6" pageNumber="195">MW</collectionCode>
|
||
136430
|
||
</accessionNumber>
|
||
,
|
||
<accessionNumber id="52B0AB64C821583DFCE7FCD5FC01FC94" box="[825,965,860,887]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MW136431" pageId="6" pageNumber="195">
|
||
<collectionCode id="2BF2AE42C821583DFCE7FCD5FCB4FC94" box="[825,880,860,887]" country="Netherlands" name="Museum Wasmann" pageId="6" pageNumber="195">MW</collectionCode>
|
||
136431
|
||
</accessionNumber>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="05F9650CC821583DFF19FC09FC69F831" pageId="6" pageNumber="195" type="diagnosis">
|
||
<paragraph id="4D5C3687C821583DFF19FC09FE7AFB75" blockId="6.[151,1437,572,2039]" pageId="6" pageNumber="195">
|
||
<emphasis id="7F97EA95C821583DFF19FC09FE80FC79" bold="true" box="[199,324,896,922]" pageId="6" pageNumber="195">Diagnosis:</emphasis>
|
||
<taxonomicName id="8AE34D04C821583DFE91FC09FD66FC79" authorityName="Sayyadzadeh & Esmaeili" authorityYear="2020" box="[335,674,896,922]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="195" phylum="Chordata" rank="species" species="marunensis">
|
||
<emphasis id="7F97EA95C821583DFE91FC09FD66FC79" box="[335,674,896,922]" italics="true" pageId="6" pageNumber="195">Oxynoemacheilus marunensis</emphasis>
|
||
</taxonomicName>
|
||
belongs to a group of species having two bold, black, round, or comma-shaped black spots on the caudal-fin base. Other species in this group in
|
||
<collectingCountry id="35F47617C821583DFBC4FC2CFB83FC5C" box="[1050,1095,933,959]" name="Iran" pageId="6" pageNumber="195">Iran</collectingCountry>
|
||
and the Persian Gulf basin are
|
||
<taxonomicName id="8AE34D04C821583DFF49FC41FEA6FC01" baseAuthorityName="Heckel" baseAuthorityYear="1847" box="[151,354,968,995]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="195" phylum="Chordata" rank="species" species="argyrogramma">
|
||
<emphasis id="7F97EA95C821583DFF49FC41FF6FFC01" box="[151,171,968,994]" italics="true" pageId="6" pageNumber="195">O</emphasis>
|
||
.
|
||
<emphasis id="7F97EA95C821583DFF65FC40FEA6FC01" box="[187,354,969,994]" italics="true" pageId="6" pageNumber="195">argyrogramma</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="8AE34D04C821583DFEADFC41FDDFFC01" baseAuthorityName="Bănărescu & Nalbant" baseAuthorityYear="1964" box="[371,539,968,995]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="195" phylum="Chordata" rank="species" species="euphraticus">
|
||
<emphasis id="7F97EA95C821583DFEADFC41FE43FC01" box="[371,391,968,994]" italics="true" pageId="6" pageNumber="195">O</emphasis>
|
||
.
|
||
<emphasis id="7F97EA95C821583DFE49FC41FDDFFC01" box="[407,539,968,994]" italics="true" pageId="6" pageNumber="195">euphraticus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="8AE34D04C821583DFDF5FC41FD57FC01" box="[555,659,968,995]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="195" phylum="Chordata" rank="species" species="hanae">
|
||
<emphasis id="7F97EA95C821583DFDF5FC41FDFBFC01" box="[555,575,968,994]" italics="true" pageId="6" pageNumber="195">O</emphasis>
|
||
.
|
||
<emphasis id="7F97EA95C821583DFD91FC41FD57FC01" box="[591,659,968,994]" italics="true" pageId="6" pageNumber="195">hanae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="8AE34D04C821583DFD7DFC41FC87FC01" authorityName=", ZM-CBSU H" authorityYear="1789" box="[675,835,968,995]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="195" phylum="Chordata" rank="species" species="karunensis">
|
||
<emphasis id="7F97EA95C821583DFD7DFC41FD73FC01" box="[675,695,968,994]" italics="true" pageId="6" pageNumber="195">O</emphasis>
|
||
.
|
||
<emphasis id="7F97EA95C821583DFD19FC41FC87FC01" box="[711,835,968,994]" italics="true" pageId="6" pageNumber="195">karunensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="7F97EA95C821583DFC9DFC41FC83FC01" box="[835,839,968,994]" italics="true" pageId="6" pageNumber="195">,</emphasis>
|
||
and
|
||
<taxonomicName id="8AE34D04C821583DFC5FFC41FBF8FC01" authorityName=": ZM-CBSU H" authorityYear="1802" box="[897,1084,968,995]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="195" phylum="Chordata" rank="species" species="kurdistanicus">
|
||
<emphasis id="7F97EA95C821583DFC5FFC41FC51FC01" box="[897,917,968,994]" italics="true" pageId="6" pageNumber="195">O</emphasis>
|
||
.
|
||
<emphasis id="7F97EA95C821583DFC78FC41FBF8FC01" box="[934,1084,968,994]" italics="true" pageId="6" pageNumber="195">kurdistanicus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="8AE34D04C821583DFB95FC41FA58FC01" authorityName="Sayyadzadeh & Esmaeili" authorityYear="2020" box="[1099,1436,968,994]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="195" phylum="Chordata" rank="species" species="marunensis">
|
||
<emphasis id="7F97EA95C821583DFB95FC41FA58FC01" box="[1099,1436,968,994]" italics="true" pageId="6" pageNumber="195">Oxynoemacheilus marunensis</emphasis>
|
||
</taxonomicName>
|
||
is most similar to
|
||
<taxonomicName id="8AE34D04C821583DFEB8FC65FDC7FBE5" authorityName=", ZM-CBSU H" authorityYear="1789" box="[358,515,1004,1031]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="195" phylum="Chordata" rank="species" species="karunensis">
|
||
<emphasis id="7F97EA95C821583DFEB8FC65FEBEFBE5" box="[358,378,1004,1030]" italics="true" pageId="6" pageNumber="195">O</emphasis>
|
||
.
|
||
<emphasis id="7F97EA95C821583DFE54FC65FDC7FBE5" box="[394,515,1004,1030]" italics="true" pageId="6" pageNumber="195">karunensis</emphasis>
|
||
</taxonomicName>
|
||
in morphological characters except the distance between pelvic and anal-fin origins, which is longer in
|
||
<taxonomicName id="8AE34D04C821583DFE76FB99FD88FBC9" authorityName="Sayyadzadeh & Esmaeili" authorityYear="2020" box="[424,588,1040,1066]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="195" phylum="Chordata" rank="species" species="marunensis">
|
||
<emphasis id="7F97EA95C821583DFE76FB99FD88FBC9" box="[424,588,1040,1066]" italics="true" pageId="6" pageNumber="195">O. marunensis</emphasis>
|
||
</taxonomicName>
|
||
[22.3–23.8 vs. 19.5–22.3 (% SL)]. Therefore, the distinguishing characters between
|
||
<taxonomicName id="8AE34D04C821583DFF22FBBDFE64FBAD" authorityName="Sayyadzadeh & Esmaeili" authorityYear="2020" box="[252,416,1076,1103]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="195" phylum="Chordata" rank="species" species="marunensis">
|
||
<emphasis id="7F97EA95C821583DFF22FBBDFED4FBAD" box="[252,272,1076,1102]" italics="true" pageId="6" pageNumber="195">O</emphasis>
|
||
.
|
||
<emphasis id="7F97EA95C821583DFEC1FBBCFE64FBAD" box="[287,416,1077,1102]" italics="true" pageId="6" pageNumber="195">marunensis</emphasis>
|
||
</taxonomicName>
|
||
and other species in this group are largely common with
|
||
<taxonomicName id="8AE34D04C821583DFBF5FBBDFB03FBAD" authorityName=", ZM-CBSU H" authorityYear="1789" box="[1067,1223,1076,1103]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="195" phylum="Chordata" rank="species" species="karunensis">
|
||
<emphasis id="7F97EA95C821583DFBF5FBBDFBFBFBAD" box="[1067,1087,1076,1102]" italics="true" pageId="6" pageNumber="195">O</emphasis>
|
||
.
|
||
<emphasis id="7F97EA95C821583DFB90FBBDFB03FBAD" box="[1102,1223,1076,1102]" italics="true" pageId="6" pageNumber="195">karunensis</emphasis>
|
||
</taxonomicName>
|
||
(see Freyhof 2016 and
|
||
<bibRefCitation id="29724B76C821583DFF18FBD1FE29FB91" author="Freyhof, J. & Abdullah, Y. S." box="[198,493,1112,1138]" pageId="6" pageNumber="195" pagination="73 - 87" refId="ref8649" refString="Freyhof, J. & Abdullah, Y. S. (2017) Two new species of Oxynoemacheilus from the Tigris drainage in Iraqi Kurdistan (Teleostei: Nemacheilidae). Zootaxa, 4238 (1), 73 - 87. https: // doi. org / 10.11646 / zootaxa. 4238.1.5" type="journal article" year="2017">Freyhof & Abdullah 2017</bibRefCitation>
|
||
).
|
||
<taxonomicName id="8AE34D04C821583DFE21FBD1FC89FB91" authorityName="Sayyadzadeh & Esmaeili" authorityYear="2020" box="[511,845,1112,1138]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="195" phylum="Chordata" rank="species" species="marunensis">
|
||
<emphasis id="7F97EA95C821583DFE21FBD1FC89FB91" box="[511,845,1112,1138]" italics="true" pageId="6" pageNumber="195">Oxynoemacheilus marunensis</emphasis>
|
||
</taxonomicName>
|
||
is also distinguished from
|
||
<taxonomicName id="8AE34D04C821583DFBA5FBD1FAD2FB91" authorityName=", ZM-CBSU H" authorityYear="1789" box="[1147,1302,1112,1139]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="195" phylum="Chordata" rank="species" species="karunensis">
|
||
<emphasis id="7F97EA95C821583DFBA5FBD1FB4BFB91" box="[1147,1167,1112,1138]" italics="true" pageId="6" pageNumber="195">O</emphasis>
|
||
.
|
||
<emphasis id="7F97EA95C821583DFB43FBD1FAD2FB91" box="[1181,1302,1112,1138]" italics="true" pageId="6" pageNumber="195">karunensis</emphasis>
|
||
</taxonomicName>
|
||
by 6% K2P COI sequence divergence.
|
||
</paragraph>
|
||
<paragraph id="4D5C3687C821583DFF19FB29FE72F9C0" blockId="6.[151,1437,572,2039]" pageId="6" pageNumber="195">
|
||
<taxonomicName id="8AE34D04C821583DFF19FB29FDD3FB59" authorityName="Sayyadzadeh & Esmaeili" authorityYear="2020" box="[199,535,1184,1210]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="195" phylum="Chordata" rank="species" species="marunensis">
|
||
<emphasis id="7F97EA95C821583DFF19FB29FDD3FB59" box="[199,535,1184,1210]" italics="true" pageId="6" pageNumber="195">Oxynoemacheilus marunensis</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="8AE34D04C821583DFCC8FB29FC0BFB59" authorityName=": ZM-CBSU H" authorityYear="1802" box="[790,975,1184,1211]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="195" phylum="Chordata" rank="species" species="kurdistanicus">
|
||
<emphasis id="7F97EA95C821583DFCC8FB29FCEEFB59" box="[790,810,1184,1210]" italics="true" pageId="6" pageNumber="195">O</emphasis>
|
||
.
|
||
<emphasis id="7F97EA95C821583DFCE7FB29FC0BFB59" box="[825,975,1184,1210]" italics="true" pageId="6" pageNumber="195">kurdistanicus</emphasis>
|
||
</taxonomicName>
|
||
, which is widespread in the Tigris River drainage, by having a more slender caudal peduncle [caudal peduncle depth 9.8–11.3 vs. 7–8.6 (% SL)], and from
|
||
<taxonomicName id="8AE34D04C821583DFF49FB61FEFBFAE1" baseAuthorityName="Bănărescu & Nalbant" baseAuthorityYear="1964" box="[151,319,1256,1283]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="195" phylum="Chordata" rank="species" species="euphraticus">
|
||
<emphasis id="7F97EA95C821583DFF49FB61FF6FFAE1" box="[151,171,1256,1282]" italics="true" pageId="6" pageNumber="195">O</emphasis>
|
||
.
|
||
<emphasis id="7F97EA95C821583DFF65FB61FEFBFAE1" box="[187,319,1256,1282]" italics="true" pageId="6" pageNumber="195">euphraticus</emphasis>
|
||
</taxonomicName>
|
||
by having longer distance between pelvic and anal-fin origins. [22.3–23.8 vs. 20.0–22.5 (% SL)] and shorter snout length [37.6–43.9 vs. 42.6–49.6 (% SL)]. It is also distinguished from those two species,
|
||
<taxonomicName id="8AE34D04C821583DFAE1FA85FF3FFAA9" authorityName=": ZM-CBSU H" authorityYear="1802" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="195" phylum="Chordata" rank="species" species="kurdistanicus">
|
||
<emphasis id="7F97EA95C821583DFAE1FA85FA97FAC5" box="[1343,1363,1292,1318]" italics="true" pageId="6" pageNumber="195">O</emphasis>
|
||
.
|
||
<emphasis id="7F97EA95C821583DFABFFA85FF3FFAA9" italics="true" pageId="6" pageNumber="195">kurdistanicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="8AE34D04C821583DFEEAFAB9FE1FFAA9" baseAuthorityName="Bănărescu & Nalbant" baseAuthorityYear="1964" box="[308,475,1328,1355]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="195" phylum="Chordata" rank="species" species="euphraticus">
|
||
<emphasis id="7F97EA95C821583DFEEAFAB9FE8CFAA9" box="[308,328,1328,1354]" italics="true" pageId="6" pageNumber="195">O</emphasis>
|
||
.
|
||
<emphasis id="7F97EA95C821583DFE89FAB9FE1FFAA9" box="[343,475,1328,1354]" italics="true" pageId="6" pageNumber="195">euphraticus</emphasis>
|
||
</taxonomicName>
|
||
, which are widespread in the Euphrates and Tigris River drainages, by having many minute dark-brown spots on the back, the flank above the lateral midline, and the caudal peduncle, with a mottled colour pattern in the interspaces of the saddles and large blotches on the back or these interspaces with vermiculated pattern (vs. without spots, vermiculation or mottling, in some individuals with small, vertically-elongated and transversely-positioned blotches), flank usually with a mid-lateral row of short, vertically-elongated blotches usually not confluent with the saddles on the back (vs. regularly or irregularly shaped bars on the flank behind the dorsal-fin base) (see Freyhof 2016).
|
||
</paragraph>
|
||
<paragraph id="4D5C3687C821583DFF19F9A5FDAAF950" blockId="6.[151,1437,572,2039]" pageId="6" pageNumber="195">
|
||
<taxonomicName id="8AE34D04C821583DFF19F9A5FDD2F9A5" authorityName="Sayyadzadeh & Esmaeili" authorityYear="2020" box="[199,534,1580,1606]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="195" phylum="Chordata" rank="species" species="marunensis">
|
||
<emphasis id="7F97EA95C821583DFF19F9A5FDD2F9A5" box="[199,534,1580,1606]" italics="true" pageId="6" pageNumber="195">Oxynoemacheilus marunensis</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="8AE34D04C821583DFCCEF9A5FC1DF9A5" baseAuthorityName="Heckel" baseAuthorityYear="1847" box="[784,985,1580,1607]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="195" phylum="Chordata" rank="species" species="argyrogramma">
|
||
<emphasis id="7F97EA95C821583DFCCEF9A5FCE0F9A5" box="[784,804,1580,1606]" italics="true" pageId="6" pageNumber="195">O</emphasis>
|
||
.
|
||
<emphasis id="7F97EA95C821583DFCECF9A4FC1DF9A5" box="[818,985,1581,1606]" italics="true" pageId="6" pageNumber="195">argyrogramma</emphasis>
|
||
</taxonomicName>
|
||
in the Euphrates and Qweik river drainages, by having a series of vertically elongated blotches or short bars along the lateral midline (vs. marbled or mottled colour pattern on the flank) and dark-brown blotches behind the dorsal-fin base being narrower than interspaces (vs. as wide or wider) (see Freyhof 2016).
|
||
</paragraph>
|
||
<paragraph id="4D5C3687C821583DFF19F934FAFDF869" blockId="6.[151,1437,572,2039]" pageId="6" pageNumber="195">
|
||
It is distinguished from
|
||
<taxonomicName id="8AE34D04C821583DFE05F935FD87F935" box="[475,579,1724,1751]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="195" phylum="Chordata" rank="species" species="hanae">
|
||
<emphasis id="7F97EA95C821583DFE05F935FE2BF935" box="[475,495,1724,1750]" italics="true" pageId="6" pageNumber="195">O</emphasis>
|
||
.
|
||
<emphasis id="7F97EA95C821583DFE21F935FD87F935" box="[511,579,1724,1750]" italics="true" pageId="6" pageNumber="195">hanae</emphasis>
|
||
</taxonomicName>
|
||
, the other species in this group from the Sirvan River drainage, a left side tributary of the Tigris River in
|
||
<collectingCountry id="35F47617C821583DFE2AF968FDE0F918" box="[500,548,1761,1787]" name="Iraq" pageId="6" pageNumber="195">Iraq</collectingCountry>
|
||
, by the two black blotches or spots at the caudal-fin base prominent in life and preserved fishes (vs. usually being invisible in life, indistinct and overlaid by a chevron-shaped, dark-brown or black bar), the lower flank with irregularly shaped and set brown blotches forming a continuous pattern with lateral and dorsal flank pattern (vs. flank below the lateral series of blotches without colour pattern except for a longitudinal series of isolated patches of blotches or a row of small dark-brown spots) (see
|
||
<bibRefCitation id="29724B76C821583DFBD8F8F9FAE9F869" author="Freyhof, J. & Abdullah, Y. S." box="[1030,1325,1904,1930]" pageId="6" pageNumber="195" pagination="73 - 87" refId="ref8649" refString="Freyhof, J. & Abdullah, Y. S. (2017) Two new species of Oxynoemacheilus from the Tigris drainage in Iraqi Kurdistan (Teleostei: Nemacheilidae). Zootaxa, 4238 (1), 73 - 87. https: // doi. org / 10.11646 / zootaxa. 4238.1.5" type="journal article" year="2017">Freyhof & Abdullah 2017</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="4D5C3687C821583DFF19F81DFC69F831" blockId="6.[151,1437,572,2039]" pageId="6" pageNumber="195">
|
||
Among the Iranian congeners studied for molecular characters,
|
||
<taxonomicName id="8AE34D04C821583DFC7CF81DFB8CF84D" authorityName="Sayyadzadeh & Esmaeili" authorityYear="2020" box="[930,1096,1940,1967]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="195" phylum="Chordata" rank="species" species="marunensis">
|
||
<emphasis id="7F97EA95C821583DFC7CF81DFC72F84D" box="[930,950,1940,1966]" italics="true" pageId="6" pageNumber="195">O</emphasis>
|
||
.
|
||
<emphasis id="7F97EA95C821583DFC19F81CFB8CF84D" box="[967,1096,1941,1966]" italics="true" pageId="6" pageNumber="195">marunensis</emphasis>
|
||
</taxonomicName>
|
||
is characterised by two fixed diagnostic nucleotide substitutions in the mtDNA COI barcode region.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="05F9650CC8215831FF19F855FD72FD82" lastPageId="10" lastPageNumber="199" pageId="6" pageNumber="195" type="description">
|
||
<paragraph id="4D5C3687C821583DFF19F855FB74F815" blockId="6.[151,1437,572,2039]" box="[199,1200,2012,2039]" pageId="6" pageNumber="195">
|
||
<emphasis id="7F97EA95C821583DFF19F855FE9FF815" bold="true" box="[199,347,2012,2038]" pageId="6" pageNumber="195">Description:</emphasis>
|
||
See
|
||
<figureCitation id="D5D82A02C821583DFE4FF855FDD2F815" box="[401,534,2012,2038]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStartId-0="7.[151,250,1050,1074]" captionStartId-1="7.[151,250,1979,2003]" captionStartId-2="8.[151,250,1044,1068]" captionStartId-3="8.[151,250,1981,2005]" captionStartId-4="9.[151,250,2007,2031]" captionStartId-5="10.[151,250,1144,1168]" captionTargetBox-0="[211,1376,181,1026]" captionTargetBox-1="[215,1372,1099,1956]" captionTargetBox-2="[209,1378,181,1020]" captionTargetBox-3="[209,1378,1128,1957]" captionTargetBox-4="[403,1184,885,1984]" captionTargetBox-5="[170,1417,705,1120]" captionTargetId-0="figure-19@7.[211,1376,181,1026]" captionTargetId-1="figure-36@7.[215,1372,1099,1956]" captionTargetId-2="figure-25@8.[209,1378,181,1020]" captionTargetId-3="figure-55@8.[209,1378,1128,1957]" captionTargetId-4="figure-357@9.[403,1184,885,1984]" captionTargetId-5="figure-267@10.[170,1417,705,1120]" captionTargetPageId-0="7" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="10" captionText-0="FIGURE 2. Oxynoemacheilus marunensis, ZM-CBSU H2509, holotype, 43 mm SL; Iran: Marun River." captionText-1="FIGURE 3. Oxynoemacheilus marunensis, paratypes; a, ZM-CBSU H2510, 36 mm SL; b, ZM-CBSU H2518, 32 mm SL; c, ZM-CBSU H2519, 27 mm SL; Iran: Marun River." captionText-2="FIGURE 4. Oxynoemacheilus marunensis, paratypes; a, ZM-CBSU H2510, 36 mm SL; b, ZM-CBSU H2518, 32 mm SL; c, ZM-CBSU H2519, 27 mm SL; Iran: Marun River." captionText-3="FIGURE 5. Oxynoemacheilus marunensis, paratypes; a, ZM-CBSU H2510, 36 mm SL; b, ZM-CBSU H2518, 32 mm SL; c, ZM-CBSU H2519, 27 mm SL; Iran: Marun River." captionText-4="FIGURE 6. Oxynoemacheilus marunensis, ZM-CBSU H2509, holotype, 43 mm SL; ventral view of Head, Iran: Marun River." captionText-5="FIGURE 7. Oxynoemacheilus marunensis, Uncatalogued, about 35 mm SL; Iran: Marun River." figureDoi-0="http://doi.org/10.5281/zenodo.4296530" figureDoi-1="http://doi.org/10.5281/zenodo.4296532" figureDoi-2="http://doi.org/10.5281/zenodo.4296534" figureDoi-3="http://doi.org/10.5281/zenodo.4296536" figureDoi-4="http://doi.org/10.5281/zenodo.4296538" figureDoi-5="http://doi.org/10.5281/zenodo.4296540" httpUri-0="https://zenodo.org/record/4296530/files/figure.png" httpUri-1="https://zenodo.org/record/4296532/files/figure.png" httpUri-2="https://zenodo.org/record/4296534/files/figure.png" httpUri-3="https://zenodo.org/record/4296536/files/figure.png" httpUri-4="https://zenodo.org/record/4296538/files/figure.png" httpUri-5="https://zenodo.org/record/4296540/files/figure.png" pageId="6" pageNumber="195">Figures 2–7</figureCitation>
|
||
for general appearance and
|
||
<tableCitation id="0061033CC821583DFC8CF855FC61F815" box="[850,933,2012,2038]" captionStart="TABLE 4" captionStartId="11.[151,239,151,175]" captionTargetPageId="11" captionText="TABLE 4. Morphometric data of Oxynoemacheilus marunensis ZM-CBSU H1998-H2006 and ZM-CBSU H2509-H2519, (n=20); O. karunensis, ZM-CBSU H1789, (n=13)." httpUri="http://table.plazi.org/id/199C660FC82C5830FF49FF1EFD76FF37" pageId="6" pageNumber="195" tableUuid="199C660FC82C5830FF49FF1EFD76FF37">Table 4</tableCitation>
|
||
for morphometric data.
|
||
</paragraph>
|
||
<caption id="199C660FC820583CFF49FB93FB7EFBD1" ID-DOI="http://doi.org/10.5281/zenodo.4296530" ID-Zenodo-Dep="4296530" box="[151,1210,1050,1075]" httpUri="https://zenodo.org/record/4296530/files/figure.png" pageId="7" pageNumber="196" startId="7.[151,250,1050,1074]" targetBox="[211,1376,181,1026]" targetPageId="7">
|
||
<paragraph id="4D5C3687C820583CFF49FB93FB7EFBD1" blockId="7.[151,1210,1050,1075]" box="[151,1210,1050,1075]" pageId="7" pageNumber="196">
|
||
<emphasis id="7F97EA95C820583CFF49FB93FED7FBD1" bold="true" box="[151,275,1050,1074]" pageId="7" pageNumber="196">FIGURE 2.</emphasis>
|
||
<taxonomicName id="8AE34D04C820583CFEC7FB93FCD0FBD1" authority=", ZM-CBSU H 2509" authorityName=", ZM-CBSU H" authorityYear="2509" box="[281,788,1050,1075]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="196" phylum="Chordata" rank="species" species="marunensis">
|
||
<emphasis id="7F97EA95C820583CFEC7FB93FD8DFBD1" box="[281,585,1050,1074]" italics="true" pageId="7" pageNumber="196">Oxynoemacheilus marunensis</emphasis>
|
||
, ZM-CBSU H2509
|
||
</taxonomicName>
|
||
, holotype, 43 mm SL; Iran: Marun River.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="199C660FC820583CFF49F832FD52F814" ID-DOI="http://doi.org/10.5281/zenodo.4296532" ID-Zenodo-Dep="4296532" httpUri="https://zenodo.org/record/4296532/files/figure.png" pageId="7" pageNumber="196" startId="7.[151,250,1979,2003]" targetBox="[215,1372,1099,1956]" targetPageId="7">
|
||
<paragraph id="4D5C3687C820583CFF49F832FD52F814" blockId="7.[151,1437,1979,2039]" pageId="7" pageNumber="196">
|
||
<emphasis id="7F97EA95C820583CFF49F832FED0F830" bold="true" box="[151,276,1979,2003]" pageId="7" pageNumber="196">FIGURE 3.</emphasis>
|
||
<taxonomicName id="8AE34D04C820583CFEC2F832FD88F830" authorityName=", ZM-CBSU H" authorityYear="2509" box="[284,588,1979,2003]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="196" phylum="Chordata" rank="species" species="marunensis">
|
||
<emphasis id="7F97EA95C820583CFEC2F832FD88F830" box="[284,588,1979,2003]" italics="true" pageId="7" pageNumber="196">Oxynoemacheilus marunensis</emphasis>
|
||
</taxonomicName>
|
||
, paratypes; a, ZM-CBSU H2510, 36 mm SL; b, ZM-CBSU H2518, 32 mm SL; c, ZM-CBSU H2519, 27 mm SL; Iran: Marun River.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="199C660FC82F5833FF49FB9DFD52FBB3" ID-DOI="http://doi.org/10.5281/zenodo.4296534" ID-Zenodo-Dep="4296534" httpUri="https://zenodo.org/record/4296534/files/figure.png" pageId="8" pageNumber="197" startId="8.[151,250,1044,1068]" targetBox="[209,1378,181,1020]" targetPageId="8">
|
||
<paragraph id="4D5C3687C82F5833FF49FB9DFD52FBB3" blockId="8.[151,1437,1044,1104]" pageId="8" pageNumber="197">
|
||
<emphasis id="7F97EA95C82F5833FF49FB9DFED0FBCF" bold="true" box="[151,276,1044,1068]" pageId="8" pageNumber="197">FIGURE 4.</emphasis>
|
||
<taxonomicName id="8AE34D04C82F5833FEC2FB9DFD88FBCF" authorityName=", ZM-CBSU H" authorityYear="2509" box="[284,588,1044,1068]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="197" phylum="Chordata" rank="species" species="marunensis">
|
||
<emphasis id="7F97EA95C82F5833FEC2FB9DFD88FBCF" box="[284,588,1044,1068]" italics="true" pageId="8" pageNumber="197">Oxynoemacheilus marunensis</emphasis>
|
||
</taxonomicName>
|
||
, paratypes; a, ZM-CBSU H2510, 36 mm SL; b, ZM-CBSU H2518, 32 mm SL; c, ZM-CBSU H2519, 27 mm SL; Iran: Marun River.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="199C660FC82F5833FF49F834FD52F819" ID-DOI="http://doi.org/10.5281/zenodo.4296536" ID-Zenodo-Dep="4296536" httpUri="https://zenodo.org/record/4296536/files/figure.png" pageId="8" pageNumber="197" startId="8.[151,250,1981,2005]" targetBox="[209,1378,1128,1957]" targetPageId="8">
|
||
<paragraph id="4D5C3687C82F5833FF49F834FD52F819" blockId="8.[151,1437,1981,2042]" pageId="8" pageNumber="197">
|
||
<emphasis id="7F97EA95C82F5833FF49F834FED0F835" bold="true" box="[151,276,1981,2006]" pageId="8" pageNumber="197">FIGURE 5.</emphasis>
|
||
<taxonomicName id="8AE34D04C82F5833FEC2F837FD88F836" authorityName=", ZM-CBSU H" authorityYear="2509" box="[284,588,1982,2006]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="197" phylum="Chordata" rank="species" species="marunensis">
|
||
<emphasis id="7F97EA95C82F5833FEC2F837FD88F836" box="[284,588,1982,2006]" italics="true" pageId="8" pageNumber="197">Oxynoemacheilus marunensis</emphasis>
|
||
</taxonomicName>
|
||
, paratypes; a, ZM-CBSU H2510, 36 mm SL; b, ZM-CBSU H2518, 32 mm SL; c, ZM-CBSU H2519, 27 mm SL; Iran: Marun River.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="4D5C3687C82E5832FF19FF1EFB9AFDFA" blockId="9.[151,1437,151,861]" pageId="9" pageNumber="198">
|
||
Medium-sized (
|
||
<quantity id="8A1B9B62C82E5832FEA6FF1EFE30FF52" box="[376,500,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="3.45" metricValueMax="4.2" metricValueMin="2.7" pageId="9" pageNumber="198" unit="mm" value="34.5" valueMax="42.0" valueMin="27.0">27–42 mm</quantity>
|
||
SL) and moderately-elongate (13.7–18.3 body depth % SL) species with a slightly pointed head. Head length 24.7–27.4% SL. Body deepest at dorsal-fin origin or about midline between nape and dorsal-fin origin, depth decreasing below dorsal-fin base and decreasing very slowly towards caudal-fin base. No hump at nape. Greatest body width at pectoral-fin base. Section of head roundish, flattened on the ventral surface. Caudal peduncle slender, compressed laterally, 1.4–1.9 (mean 1.7) times longer than deep. Small, usually triangular axillary lobe at base of pelvic fin, fully attached to the body. Pectoral fin equal or slightly longer than the head length and reaching approximately 89–111% of the distance from pectoral-fin origin to pelvic-fin origin. Pelvic-fin origin below first or second branched dorsal-fin ray, fin not reaching vertical of tip of last dorsal-fin ray, reaching to anus. Anal-fin origin below tip of last dorsal-fin ray. Anus about one eye diameter in front of anal-fin origin. Anal fin not reaching caudal-fin base. No dorsal or ventral adipose crest on caudal peduncle. Margin of the dorsal fin straight or slightly concave. Caudal fin deeply emarginated. Largest known specimen
|
||
<quantity id="8A1B9B62C82E5832FC3EFE76FBEBFDFA" box="[992,1071,511,537]" metricMagnitude="-2" metricUnit="m" metricValue="4.3" pageId="9" pageNumber="198" unit="mm" value="43.0">43 mm</quantity>
|
||
SL.
|
||
</paragraph>
|
||
<paragraph id="4D5C3687C82E5832FF19FDAAFDDFFCBE" blockId="9.[151,1437,151,861]" pageId="9" pageNumber="198">
|
||
Dorsal fin with 9½–10½ branched rays. Anal fin with 5½ branched rays. Caudal fin with 8+7–8+8 branched rays. Pectoral fin with 9–10 and pelvic fin with 6–7 branched rays. Body covered by embedded scales. Lateral line complete, reaching to caudal-fin base. Anterior nostril opening at end of low, pointed, and flap-like tube. Tube of anterior nostril slightly overlapping posterior nostril when folded back. Mouth small, arched (
|
||
<figureCitation id="D5D82A02C82E5832FB1BFD06FAC8FD49" box="[1221,1292,655,682]" captionStart="FIGURE 6" captionStartId="9.[151,250,2007,2031]" captionTargetBox="[403,1184,885,1984]" captionTargetId="figure-357@9.[403,1184,885,1984]" captionTargetPageId="9" captionText="FIGURE 6. Oxynoemacheilus marunensis, ZM-CBSU H2509, holotype, 43 mm SL; ventral view of Head, Iran: Marun River." figureDoi="http://doi.org/10.5281/zenodo.4296538" httpUri="https://zenodo.org/record/4296538/files/figure.png" pageId="9" pageNumber="198">Fig. 6</figureCitation>
|
||
). Lips thick, with poorly marked furrows. A deep median interruption in the lower lip. Median incision in upper lip very small and not visible in some individuals. Processus dentiformis narrow and pointed. No median notch in lower jaw. Inner rostral barbel reaching to base of maxillary barbel or slightly beyond; outer one reaching to vertical of anterior eye margin or slightly in front. Maxillary barbel reaching vertical through middle or posterior part of eye. Male with suborbital grove, absent in female.
|
||
</paragraph>
|
||
<caption id="199C660FC82E5832FF49F85EFA58F80C" ID-DOI="http://doi.org/10.5281/zenodo.4296538" ID-Zenodo-Dep="4296538" box="[151,1436,2007,2031]" httpUri="https://zenodo.org/record/4296538/files/figure.png" pageId="9" pageNumber="198" startId="9.[151,250,2007,2031]" targetBox="[403,1184,885,1984]" targetPageId="9">
|
||
<paragraph id="4D5C3687C82E5832FF49F85EFA58F80C" blockId="9.[151,1436,2007,2031]" box="[151,1436,2007,2031]" pageId="9" pageNumber="198">
|
||
<emphasis id="7F97EA95C82E5832FF49F85EFED7F80C" bold="true" box="[151,275,2007,2031]" pageId="9" pageNumber="198">FIGURE 6.</emphasis>
|
||
<taxonomicName id="8AE34D04C82E5832FEC7F85EFCD0F80C" authority=", ZM-CBSU H 2509" authorityName=", ZM-CBSU H" authorityYear="2509" box="[281,788,2007,2031]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="198" phylum="Chordata" rank="species" species="marunensis">
|
||
<emphasis id="7F97EA95C82E5832FEC7F85EFD8DF80C" box="[281,585,2007,2031]" italics="true" pageId="9" pageNumber="198">Oxynoemacheilus marunensis</emphasis>
|
||
, ZM-CBSU H2509
|
||
</taxonomicName>
|
||
, holotype, 43 mm SL; ventral view of Head, Iran: Marun River.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="4D5C3687C82D5831FF19FF1FFD72FD82" blockId="10.[151,1437,150,681]" pageId="10" pageNumber="199">
|
||
<emphasis id="7F97EA95C82D5831FF19FF1FFEA5FF53" bold="true" box="[199,353,150,176]" pageId="10" pageNumber="199">Colouration:</emphasis>
|
||
Head and body with yellowish background colour and brown pattern. Head brown on top to lower margin of eye or with spotted pattern, cheeks and ventral head surface without colour pattern. A dark-brown line between the anterior eye-margin and tip of the snout. Pre-dorsal with one to two, wide dark-brown blotches, equal or slightly wider than interspaces. Large, brown blotch at dorsal fin-origin. Two or three, wide dark-brown blotches on upper caudal peduncle, fused with blotches on flank forming saddles in few individuals. Flank below line between pectoral-fin base and anus without pattern. Flank with 7–10 dark-brown, irregularly shaped, vertically elongated blotches along lateral midline. Flank blotches narrower than interspaces, usually dissociated and often faded in front of dorsal-fin origin, more clearly set on caudal peduncle, rarely joined with saddles on the back or forming bars on the flank. Back, flank above the lateral midline and caudal peduncle with many, minute dark-brown spots, vermiculation, or even a mottled colour pattern at interspaces of saddles and large blotches. One distinct, small, roundish or comma-shaped, black spot at upper and lower posterior extremity of caudal peduncle. Dorsal fin with 2–3 and caudal fin with 3–5 brown bands of small, elongated blotches on fin-rays. Anal-, pelvic- and pectoral fins hyaline, with few dark-brown spots on rays.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="05F9650CC82D5831FF19FDE3FEA3FAC3" pageId="10" pageNumber="199" type="distribution">
|
||
<paragraph id="4D5C3687C82D5831FF19FDE3FC40FD4A" blockId="10.[151,1437,150,681]" pageId="10" pageNumber="199">
|
||
<emphasis id="7F97EA95C82D5831FF19FDE3FE3FFD67" bold="true" box="[199,507,618,645]" pageId="10" pageNumber="199">Distribution and Habitat:</emphasis>
|
||
<taxonomicName id="8AE34D04C82D5831FDDDFDE2FC97FD67" authorityName=", ZM-CBSU H" authorityYear="2509" box="[515,851,619,645]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="199" phylum="Chordata" rank="species" species="marunensis">
|
||
<emphasis id="7F97EA95C82D5831FDDDFDE2FC97FD67" box="[515,851,619,645]" italics="true" pageId="10" pageNumber="199">Oxynoemacheilus marunensis</emphasis>
|
||
</taxonomicName>
|
||
is known from the Marun, a tributary of the Jarrahi River, which flows just east of the Tigris River to the Persian Gulf.
|
||
</paragraph>
|
||
<caption id="199C660FC82D5831FF49FBF1FBA0FB73" ID-DOI="http://doi.org/10.5281/zenodo.4296540" ID-Zenodo-Dep="4296540" box="[151,1124,1144,1168]" httpUri="https://zenodo.org/record/4296540/files/figure.png" pageId="10" pageNumber="199" startId="10.[151,250,1144,1168]" targetBox="[170,1417,705,1120]" targetPageId="10">
|
||
<paragraph id="4D5C3687C82D5831FF49FBF1FBA0FB73" blockId="10.[151,1124,1144,1168]" box="[151,1124,1144,1168]" pageId="10" pageNumber="199">
|
||
<emphasis id="7F97EA95C82D5831FF49FBF1FED7FB73" bold="true" box="[151,275,1144,1168]" pageId="10" pageNumber="199">FIGURE 7.</emphasis>
|
||
<taxonomicName id="8AE34D04C82D5831FEC7FBF1FD20FB73" authority=", Uncatalogued" authorityName="Uncatalogued" box="[281,740,1144,1168]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="199" phylum="Chordata" rank="species" species="marunensis">
|
||
<emphasis id="7F97EA95C82D5831FEC7FBF1FD8DFB73" box="[281,585,1144,1168]" italics="true" pageId="10" pageNumber="199">Oxynoemacheilus marunensis</emphasis>
|
||
, Uncatalogued
|
||
</taxonomicName>
|
||
, about 35 mm SL; Iran: Marun River.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="4D5C3687C82D5831FF19FB37FEA3FAC3" blockId="10.[151,1437,1214,1997]" pageId="10" pageNumber="199">
|
||
At the sampling site, the river is about
|
||
<quantity id="8A1B9B62C82D5831FDA3FB37FD72FB3A" box="[637,694,1214,1241]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="199" unit="m" value="10.0">10 m</quantity>
|
||
wide, with a substrate consisting of coarse gravel and small boulders, with moderate flow and semi-transparent waters (
|
||
<figureCitation id="D5D82A02C82D5831FCC7FB6BFCA5FB1F" box="[793,865,1250,1276]" captionStart="FIGURE 8" captionStartId="12.[151,250,1159,1183]" captionTargetBox="[151,1436,181,1135]" captionTargetId="figure-25@12.[151,1436,181,1135]" captionTargetPageId="12" captionText="FIGURE 8. Habitat of Oxynoemacheilus marunensis, Iran: Marun River at Kharestan, a tributary of Jarrahi River drainage." figureDoi="http://doi.org/10.5281/zenodo.4296542" httpUri="https://zenodo.org/record/4296542/files/figure.png" pageId="10" pageNumber="199">Fig. 8</figureCitation>
|
||
). Drought, water use, and pollution are the main threats to this fish.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="05F9650CC82D5830FF19FAA3FC81FA20" lastPageId="11" lastPageNumber="200" pageId="10" pageNumber="199" type="etymology">
|
||
<paragraph id="4D5C3687C82D5831FF19FAA3FF0DFA8B" blockId="10.[151,1437,1214,1997]" pageId="10" pageNumber="199">
|
||
<emphasis id="7F97EA95C82D5831FF19FAA3FE96FAA7" bold="true" box="[199,338,1322,1348]" pageId="10" pageNumber="199">Etymology:</emphasis>
|
||
<taxonomicName id="8AE34D04C82D5831FE84FAA3FD6EFAA7" box="[346,682,1322,1348]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="199" phylum="Chordata" rank="species" species="marunensis">
|
||
<emphasis id="7F97EA95C82D5831FE84FAA3FD6EFAA7" box="[346,682,1322,1348]" italics="true" pageId="10" pageNumber="199">Oxynoemacheilus marunensis</emphasis>
|
||
</taxonomicName>
|
||
is named for the Marun River inhabited by this species. An adjective.
|
||
</paragraph>
|
||
<paragraph id="4D5C3687C82D5831FF19FAFBFB9CF94F" blockId="10.[151,1437,1214,1997]" pageId="10" pageNumber="199">
|
||
<emphasis id="7F97EA95C82D5831FF19FAFBFE1EFA6F" bold="true" box="[199,474,1394,1420]" pageId="10" pageNumber="199">Comparative remarks:</emphasis>
|
||
Freyhof (2016) considered the oxynoemacheilid population from the Marun River as
|
||
<taxonomicName id="8AE34D04C82D5831FF49FA1EFEF0FA53" authorityName=", ZM-CBSU H" authorityYear="1789" box="[151,308,1430,1457]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="199" phylum="Chordata" rank="species" species="karunensis">
|
||
<emphasis id="7F97EA95C82D5831FF49FA1EFF6FFA52" box="[151,171,1431,1457]" italics="true" pageId="10" pageNumber="199">O</emphasis>
|
||
.
|
||
<emphasis id="7F97EA95C82D5831FF65FA1FFEF0FA53" box="[187,308,1430,1456]" italics="true" pageId="10" pageNumber="199">karunensis</emphasis>
|
||
</taxonomicName>
|
||
along with some populations from the Karun River. He examined three specimens from the Marun River (SMF IR7, 3,
|
||
<quantity id="8A1B9B62C82D5831FEABFA33FE29FA36" box="[373,493,1466,1493]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="4.4" metricValueMin="3.6" pageId="10" pageNumber="199" unit="mm" value="40.0" valueMax="44.0" valueMin="36.0">36–44 mm</quantity>
|
||
SL) and provided the morphological characters. In a comprehensive molecular study based on the COI barcoding region including all the Iranian and other oxynoemacheilid sequences from the Middle East available in the GenBank (
|
||
<figureCitation id="D5D82A02C82D5831FE21F98BFD86F9FF" box="[511,578,1538,1564]" captionStart="FIGURE 1" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[359,1229,185,1917]" captionTargetId="figure-21@5.[353,1234,181,1920]" captionTargetPageId="5" captionText="FIGURE 1. Maximum Likelihood and Bayesian phylogeny reconstructed based on 617 bp of COI 5’ end. The values besides the branches before and after a slash are BI posterior and ML bootstrap probability values, respectively. Black bars right to the specimen labels indicates species delimitation results from PTP followed by the results of the mPTP approach as dashed bars." figureDoi="http://doi.org/10.5281/zenodo.4296528" httpUri="https://zenodo.org/record/4296528/files/figure.png" pageId="10" pageNumber="199">Fig. 1</figureCitation>
|
||
), we found two different clades for populations from the Marun and Karun Rivers with approximately 6% K2P COI sequence divergence whereas they are similar morphologically. In addition to molecular distinguishing characters,
|
||
<taxonomicName id="8AE34D04C82D5831FD5FF9C2FCE0F987" box="[641,804,1611,1637]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="199" phylum="Chordata" rank="species" species="marunensis">
|
||
<emphasis id="7F97EA95C82D5831FD5FF9C2FD51F986" box="[641,661,1611,1637]" italics="true" pageId="10" pageNumber="199">O</emphasis>
|
||
.
|
||
<emphasis id="7F97EA95C82D5831FD7DF9C2FCE0F987" box="[675,804,1611,1636]" italics="true" pageId="10" pageNumber="199">marunensis</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="8AE34D04C82D5831FBC0F9C2FB7DF987" authorityName=", ZM-CBSU H" authorityYear="1789" box="[1054,1209,1610,1637]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="199" phylum="Chordata" rank="species" species="karunensis">
|
||
<emphasis id="7F97EA95C82D5831FBC0F9C2FBF6F986" box="[1054,1074,1611,1637]" italics="true" pageId="10" pageNumber="199">O</emphasis>
|
||
.
|
||
<emphasis id="7F97EA95C82D5831FB9EF9C3FB7DF987" box="[1088,1209,1610,1636]" italics="true" pageId="10" pageNumber="199">karunensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="D5D82A02C82D5831FB16F9C3FACFF987" box="[1224,1291,1610,1636]" captionStart="FIGURE 9" captionStartId="12.[151,250,1663,1687]" captionTargetBox="[150,1437,1258,1638]" captionTargetId="figure-45@12.[150,1437,1207,1638]" captionTargetPageId="12" captionText="FIGURE 9. Oxynoemacheilus karunensis, uncatalogued, about 50 mm SL; Iran: Dehno spring." figureDoi="http://doi.org/10.5281/zenodo.4296544" httpUri="https://zenodo.org/record/4296544/files/figure.png" pageId="10" pageNumber="199">Fig. 9</figureCitation>
|
||
) by having a longer distance between pelvic and anal-fin origins. There are also some distingiushing (but overlapping) morphometric characters such as head length; pectoral-fin and pelvic-fin length (see
|
||
<tableCitation id="0061033CC82D5831FC2BF91BFB8FF94F" box="[1013,1099,1682,1708]" captionStart="TABLE 4" captionStartId="11.[151,239,151,175]" captionTargetPageId="11" captionText="TABLE 4. Morphometric data of Oxynoemacheilus marunensis ZM-CBSU H1998-H2006 and ZM-CBSU H2509-H2519, (n=20); O. karunensis, ZM-CBSU H1789, (n=13)." httpUri="http://table.plazi.org/id/199C660FC82C5830FF49FF1EFD76FF37" pageId="10" pageNumber="199" tableUuid="199C660FC82C5830FF49FF1EFD76FF37">Table 4</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="4D5C3687C82D5831FF19F93FFE60F82E" blockId="10.[151,1437,1214,1997]" pageId="10" pageNumber="199">
|
||
<taxonomicName id="8AE34D04C82D5831FF19F93FFDDDF933" box="[199,537,1718,1744]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="199" phylum="Chordata" rank="species" species="marunensis">
|
||
<emphasis id="7F97EA95C82D5831FF19F93FFDDDF933" box="[199,537,1718,1744]" italics="true" pageId="10" pageNumber="199">Oxynoemacheilus marunensis</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="8AE34D04C82D5831FCFFF93EFC71F933" authorityName=": ZM-CBSU H" authorityYear="1555" box="[801,949,1718,1745]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="199" phylum="Chordata" rank="species" species="bergianus">
|
||
<emphasis id="7F97EA95C82D5831FCFFF93EFCF1F932" box="[801,821,1719,1745]" italics="true" pageId="10" pageNumber="199">O</emphasis>
|
||
.
|
||
<emphasis id="7F97EA95C82D5831FC98F93FFC71F933" box="[838,949,1718,1744]" italics="true" pageId="10" pageNumber="199">bergianus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="8AE34D04C82D5831FC19F93EFB82F933" authorityName=": ZM-CBSU H" authorityYear="1749" box="[967,1094,1718,1745]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="199" phylum="Chordata" rank="species" species="brandtii">
|
||
<emphasis id="7F97EA95C82D5831FC19F93EFC1FF932" box="[967,987,1719,1745]" italics="true" pageId="10" pageNumber="199">O</emphasis>
|
||
.
|
||
<emphasis id="7F97EA95C82D5831FC32F93FFB82F933" box="[1004,1094,1718,1744]" italics="true" pageId="10" pageNumber="199">brandtii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="8AE34D04C82D5831FB86F93EFB3FF933" baseAuthorityName="Coad & Nalbant" baseAuthorityYear="2005" box="[1112,1275,1718,1745]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="199" phylum="Chordata" rank="species" species="longipinnis">
|
||
<emphasis id="7F97EA95C82D5831FB86F93EFBA8F932" box="[1112,1132,1719,1745]" italics="true" pageId="10" pageNumber="199">O</emphasis>
|
||
.
|
||
<emphasis id="7F97EA95C82D5831FBA3F93FFB3FF933" box="[1149,1275,1718,1744]" italics="true" pageId="10" pageNumber="199">longipinnis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="8AE34D04C82D5831FAD2F93EFA5CF933" authorityName=": ZM-CBSU H" authorityYear="1973" box="[1292,1432,1719,1745]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="199" phylum="Chordata" rank="species" species="parvinae">
|
||
<emphasis id="7F97EA95C82D5831FAD2F93EFAE4F932" box="[1292,1312,1719,1745]" italics="true" pageId="10" pageNumber="199">O</emphasis>
|
||
.
|
||
<emphasis id="7F97EA95C82D5831FAECF93EFA5CF933" box="[1330,1432,1719,1744]" italics="true" pageId="10" pageNumber="199">parvinae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="7F97EA95C82D5831FA46F93EFA58F933" box="[1432,1436,1719,1744]" italics="true" pageId="10" pageNumber="199">,</emphasis>
|
||
and
|
||
<taxonomicName id="8AE34D04C82D5831FF16F952FEEEF917" authorityName=": ZM-CBSU H" authorityYear="1852" box="[200,298,1755,1781]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="199" phylum="Chordata" rank="species" species="persa">
|
||
<emphasis id="7F97EA95C82D5831FF16F952FF18F916" box="[200,220,1755,1781]" italics="true" pageId="10" pageNumber="199">O</emphasis>
|
||
.
|
||
<emphasis id="7F97EA95C82D5831FF32F952FEEEF917" box="[236,298,1755,1780]" italics="true" pageId="10" pageNumber="199">persa</emphasis>
|
||
</taxonomicName>
|
||
, five species with forked or deeply emarginate caudal fins, by having two prominent black spots at the caudal-fin base (vs. spots absent). It is distinguished from
|
||
<taxonomicName id="8AE34D04C82D5831FC90F976FC24F8FB" authorityName=": ZM-CBSU H" authorityYear="1844" box="[846,992,1791,1817]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="199" phylum="Chordata" rank="species" species="tongiorgii">
|
||
<emphasis id="7F97EA95C82D5831FC90F976FCA6F8FA" box="[846,866,1791,1817]" italics="true" pageId="10" pageNumber="199">O</emphasis>
|
||
.
|
||
<emphasis id="7F97EA95C82D5831FCAFF976FC24F8FB" box="[881,992,1791,1816]" italics="true" pageId="10" pageNumber="199">tongiorgii</emphasis>
|
||
</taxonomicName>
|
||
from the Kor River drainage of
|
||
<collectingCountry id="35F47617C82D5831FA92F976FABDF8FA" box="[1356,1401,1791,1817]" name="Iran" pageId="10" pageNumber="199">Iran</collectingCountry>
|
||
by having two prominent black spots at the caudal-fin base (vs. spots absent), a deeply emarginate or forked caudal fin (vs. truncate), and a slenderer caudal peduncle (vs. deep).
|
||
<taxonomicName id="8AE34D04C82D5831FC8BF8CFFB61F883" box="[853,1189,1862,1888]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="199" phylum="Chordata" rank="species" species="marunensis">
|
||
<emphasis id="7F97EA95C82D5831FC8BF8CFFB61F883" box="[853,1189,1862,1888]" italics="true" pageId="10" pageNumber="199">Oxynoemacheilus marunensis</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="8AE34D04C82D5831FF49F8E2FE85F867" authorityName=": ZM-CBSU H" authorityYear="1814" box="[151,321,1898,1925]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="199" phylum="Chordata" rank="species" species="chomanicus">
|
||
<emphasis id="7F97EA95C82D5831FF49F8E2FF6FF866" box="[151,171,1899,1925]" italics="true" pageId="10" pageNumber="199">O</emphasis>
|
||
.
|
||
<emphasis id="7F97EA95C82D5831FF65F8E3FE85F867" box="[187,321,1898,1924]" italics="true" pageId="10" pageNumber="199">chomanicus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="8AE34D04C82D5831FE8EF8E2FE0AF867" baseAuthorityName="Heckel" baseAuthorityYear="1843" box="[336,462,1898,1925]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="199" phylum="Chordata" rank="species" species="frenatus">
|
||
<emphasis id="7F97EA95C82D5831FE8EF8E2FEA0F866" box="[336,356,1899,1925]" italics="true" pageId="10" pageNumber="199">O</emphasis>
|
||
.
|
||
<emphasis id="7F97EA95C82D5831FEADF8E3FE0AF867" box="[371,462,1898,1924]" italics="true" pageId="10" pageNumber="199">frenatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="8AE34D04C82D5831FE03F8E2FD85F867" authorityName=": ZM-CBSU H" authorityYear="1785" box="[477,577,1898,1925]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="199" phylum="Chordata" rank="species" species="kiabii">
|
||
<emphasis id="7F97EA95C82D5831FE03F8E2FE35F866" box="[477,497,1899,1925]" italics="true" pageId="10" pageNumber="199">O</emphasis>
|
||
.
|
||
<emphasis id="7F97EA95C82D5831FDDFF8E3FD85F867" box="[513,577,1898,1924]" italics="true" pageId="10" pageNumber="199">kiabii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="8AE34D04C82D5831FDA4F8E2FCDBF867" authorityName=": ZM-CBSU H" authorityYear="1829" box="[634,799,1899,1925]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="199" phylum="Chordata" rank="species" species="zagrosensis">
|
||
<emphasis id="7F97EA95C82D5831FDA4F8E2FD4AF866" box="[634,654,1899,1925]" italics="true" pageId="10" pageNumber="199">O</emphasis>
|
||
.
|
||
<emphasis id="7F97EA95C82D5831FD43F8E2FCDBF867" box="[669,799,1899,1924]" italics="true" pageId="10" pageNumber="199">zagrosensis</emphasis>
|
||
</taxonomicName>
|
||
, four additional species found in the Tigris River drainage, by having a suborbital groove in males (vs. absent) and a deeply emarginate or forked caudal fin (vs. slightly emarginate or truncate).
|
||
</paragraph>
|
||
<caption id="199C660FC82C5830FF49FF1EFD76FF37" ID-Table-UUID="199C660FC82C5830FF49FF1EFD76FF37" httpUri="http://table.plazi.org/id/199C660FC82C5830FF49FF1EFD76FF37" pageId="11" pageNumber="200" targetBox="[166,1417,227,1256]" targetIsTable="true" targetPageId="11">
|
||
<paragraph id="4D5C3687C82C5830FF49FF1EFD76FF37" blockId="11.[151,1437,151,212]" pageId="11" pageNumber="200">
|
||
<emphasis id="7F97EA95C82C5830FF49FF1EFECCFF4C" bold="true" box="[151,264,151,175]" pageId="11" pageNumber="200">TABLE 4.</emphasis>
|
||
Morphometric data of
|
||
<taxonomicName id="8AE34D04C82C5830FE26FF1EFC39FF4C" authority="ZM-CBSU H 1998" authorityName="ZM-CBSU H" authorityYear="1998" box="[504,1021,151,176]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="200" phylum="Chordata" rank="species" species="marunensis">
|
||
<emphasis id="7F97EA95C82C5830FE26FF1EFCF2FF4C" box="[504,822,151,175]" italics="true" pageId="11" pageNumber="200">Oxynoemacheilus marunensis</emphasis>
|
||
ZM-CBSU H1998
|
||
</taxonomicName>
|
||
-H2006 and ZM-CBSU H2509-H2519, (n=20);
|
||
<taxonomicName id="8AE34D04C82C5830FF30FF35FD9CFF37" authority=", ZM-CBSU H 1789" authorityName=", ZM-CBSU H" authorityYear="1789" box="[238,600,187,212]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="200" phylum="Chordata" rank="species" species="karunensis">
|
||
<emphasis id="7F97EA95C82C5830FF30FF35FEC5FF30" box="[238,257,188,211]" italics="true" pageId="11" pageNumber="200">O</emphasis>
|
||
.
|
||
<emphasis id="7F97EA95C82C5830FED1FF32FE47FF30" box="[271,387,187,211]" italics="true" pageId="11" pageNumber="200">karunensis</emphasis>
|
||
, ZM-CBSU H1789
|
||
</taxonomicName>
|
||
, (n=13).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="4D5C3687C82C5830FCE5FF6AFA4DFB0B" pageId="11" pageNumber="200">
|
||
<table id="3FE3C427C82CA7C4FF78FF6AFA4DFB0B" box="[166,1417,227,1256]" gridcols="7" gridrows="26" pageId="11" pageNumber="200">
|
||
<tr id="F3D334C5C82CA7C4FF78FF6AFA4DFF18" box="[166,1417,227,251]" gridrow="0" pageId="11" pageNumber="200" rowspan-0="1" rowspan-3="1" rowspan-6="1">
|
||
<th id="B0025DB9C82CA7C4FD2CFF6AFC16FF18" box="[754,978,227,251]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="11" pageNumber="200">
|
||
<taxonomicName id="8AE34D04C82C5830FCE5FF6AFC09FF18" authorityName=", ZM-CBSU H" authorityYear="1789" box="[827,973,227,251]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="200" phylum="Chordata" rank="species" species="karunensis">
|
||
<emphasis id="7F97EA95C82C5830FCE5FF6AFC09FF18" bold="true" box="[827,973,227,251]" italics="true" pageId="11" pageNumber="200">
|
||
O.
|
||
<emphasis id="7F97EA95C82C5830FC84FF6AFC09FF18" bold="true" box="[858,973,227,251]" italics="true" pageId="11" pageNumber="200">karunensis</emphasis>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="B0025DB9C82CA7C4FB84FF6AFAFEFF18" box="[1114,1338,227,251]" colspan="2" colspanRight="1" gridcol="4" gridrow="0" pageId="11" pageNumber="200">
|
||
<taxonomicName id="8AE34D04C82C5830FB41FF6AFAFCFF18" authorityName="ZM-CBSU H" authorityYear="1998" box="[1183,1336,227,251]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="200" phylum="Chordata" rank="species" species="marunensis">
|
||
<emphasis id="7F97EA95C82C5830FB41FF6AFAFCFF18" bold="true" box="[1183,1336,227,251]" italics="true" pageId="11" pageNumber="200">O. marunensis</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
</tr>
|
||
<tr id="F3D334C5C82CA7C4FF78FE82FA4DFEC0" box="[166,1417,267,291]" gridrow="1" pageId="11" pageNumber="200" rowspan-0="1">
|
||
<td id="B0025DB9C82CA7C4FD2CFE82FC9CFEC0" box="[754,856,267,291]" gridcol="1" gridrow="1" pageId="11" pageNumber="200">
|
||
<emphasis id="7F97EA95C82C5830FCDDFE82FC8CFEC0" bold="true" box="[771,840,267,291]" pageId="11" pageNumber="200">Range</emphasis>
|
||
</td>
|
||
<td id="B0025DB9C82CA7C4FC4AFE82FC16FEC0" box="[916,978,267,291]" gridcol="2" gridrow="1" pageId="11" pageNumber="200">
|
||
<emphasis id="7F97EA95C82C5830FC4AFE82FC16FEC0" bold="true" box="[916,978,267,291]" pageId="11" pageNumber="200">Mean</emphasis>
|
||
</td>
|
||
<td id="B0025DB9C82CA7C4FBDCFE82FBE6FEC0" box="[1026,1058,267,291]" gridcol="3" gridrow="1" pageId="11" pageNumber="200">
|
||
<emphasis id="7F97EA95C82C5830FBDCFE82FBE6FEC0" bold="true" box="[1026,1058,267,291]" pageId="11" pageNumber="200">SD</emphasis>
|
||
</td>
|
||
<td id="B0025DB9C82CA7C4FB84FE82FB04FEC0" box="[1114,1216,267,291]" gridcol="4" gridrow="1" pageId="11" pageNumber="200">
|
||
<emphasis id="7F97EA95C82C5830FBB5FE82FB74FEC0" bold="true" box="[1131,1200,267,291]" pageId="11" pageNumber="200">Range</emphasis>
|
||
</td>
|
||
<td id="B0025DB9C82CA7C4FB22FE82FAFEFEC0" box="[1276,1338,267,291]" gridcol="5" gridrow="1" pageId="11" pageNumber="200">
|
||
<emphasis id="7F97EA95C82C5830FB22FE82FAFEFEC0" bold="true" box="[1276,1338,267,291]" pageId="11" pageNumber="200">Mean</emphasis>
|
||
</td>
|
||
<td id="B0025DB9C82CA7C4FAB7FE82FA4DFEC0" box="[1385,1417,267,291]" gridcol="6" gridrow="1" pageId="11" pageNumber="200">
|
||
<emphasis id="7F97EA95C82C5830FAB7FE82FA4DFEC0" bold="true" box="[1385,1417,267,291]" pageId="11" pageNumber="200">SD</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr id="F3D334C5C82CA7C4FF78FEBAFA4DFEAF" box="[166,1417,307,332]" gridrow="2" pageId="11" pageNumber="200" rowspan-3="1" rowspan-6="1">
|
||
<th id="B0025DB9C82CA7C4FF78FEBAFD5AFEAF" box="[166,670,307,332]" gridcol="0" gridrow="2" pageId="11" pageNumber="200">Standard length (mm)</th>
|
||
<td id="B0025DB9C82CA7C4FD2CFEBAFC9CFEAF" box="[754,856,307,332]" gridcol="1" gridrow="2" pageId="11" pageNumber="200">33–58</td>
|
||
<td id="B0025DB9C82CA7C4FC4AFEBAFC16FEAF" box="[916,978,307,332]" gridcol="2" gridrow="2" pageId="11" pageNumber="200">49.2</td>
|
||
<td id="B0025DB9C82CA7C4FB84FEBAFB04FEAF" box="[1114,1216,307,332]" gridcol="4" gridrow="2" pageId="11" pageNumber="200">27–43</td>
|
||
<td id="B0025DB9C82CA7C4FB22FEBAFAFEFEAF" box="[1276,1338,307,332]" gridcol="5" gridrow="2" pageId="11" pageNumber="200">34.8</td>
|
||
</tr>
|
||
<tr id="F3D334C5C82CA7C4FF78FED2FA4DFE90" box="[166,1417,347,371]" gridrow="3" pageId="11" pageNumber="200">
|
||
<th id="B0025DB9C82CA7C4FF78FED2FA4DFE90" box="[166,1417,347,371]" colspan="7" colspanRight="6" gridcol="0" gridrow="3" pageId="11" pageNumber="200">
|
||
<emphasis id="7F97EA95C82C5830FF78FED2FE26FE90" bold="true" box="[166,482,347,371]" pageId="11" pageNumber="200">In percent of standard length</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="F3D334C5C82CA7C4FF78FE0AFA4DFE7F" box="[166,1417,387,412]" gridrow="4" pageId="11" pageNumber="200">
|
||
<th id="B0025DB9C82CA7C4FF78FE0AFD5AFE7F" box="[166,670,387,412]" gridcol="0" gridrow="4" pageId="11" pageNumber="200">Head length</th>
|
||
<td id="B0025DB9C82CA7C4FD2CFE0AFC9CFE7F" box="[754,856,387,412]" gridcol="1" gridrow="4" pageId="11" pageNumber="200">22.5–25.1</td>
|
||
<td id="B0025DB9C82CA7C4FC4AFE0AFC16FE7F" box="[916,978,387,412]" gridcol="2" gridrow="4" pageId="11" pageNumber="200">23.7</td>
|
||
<td id="B0025DB9C82CA7C4FBDCFE0AFBE6FE7F" box="[1026,1058,387,412]" gridcol="3" gridrow="4" pageId="11" pageNumber="200">5.6</td>
|
||
<td id="B0025DB9C82CA7C4FB84FE0AFB04FE7F" box="[1114,1216,387,412]" gridcol="4" gridrow="4" pageId="11" pageNumber="200">24.7–27.4</td>
|
||
<td id="B0025DB9C82CA7C4FB22FE0AFAFEFE7F" box="[1276,1338,387,412]" gridcol="5" gridrow="4" pageId="11" pageNumber="200">25.5</td>
|
||
<td id="B0025DB9C82CA7C4FAB7FE0AFA4DFE7F" box="[1385,1417,387,412]" gridcol="6" gridrow="4" pageId="11" pageNumber="200">0.6</td>
|
||
</tr>
|
||
<tr id="F3D334C5C82CA7C4FF78FE25FA4DFE27" box="[166,1417,428,452]" gridrow="5" pageId="11" pageNumber="200">
|
||
<th id="B0025DB9C82CA7C4FF78FE25FD5AFE27" box="[166,670,428,452]" gridcol="0" gridrow="5" pageId="11" pageNumber="200">Body depth at dorsal-fin origin</th>
|
||
<td id="B0025DB9C82CA7C4FD2CFE25FC9CFE27" box="[754,856,428,452]" gridcol="1" gridrow="5" pageId="11" pageNumber="200">15.1–18.7</td>
|
||
<td id="B0025DB9C82CA7C4FC4AFE25FC16FE27" box="[916,978,428,452]" gridcol="2" gridrow="5" pageId="11" pageNumber="200">17.2</td>
|
||
<td id="B0025DB9C82CA7C4FBDCFE25FBE6FE27" box="[1026,1058,428,452]" gridcol="3" gridrow="5" pageId="11" pageNumber="200">0.7</td>
|
||
<td id="B0025DB9C82CA7C4FB84FE25FB04FE27" box="[1114,1216,428,452]" gridcol="4" gridrow="5" pageId="11" pageNumber="200">13.7–18.3</td>
|
||
<td id="B0025DB9C82CA7C4FB22FE25FAFEFE27" box="[1276,1338,428,452]" gridcol="5" gridrow="5" pageId="11" pageNumber="200">16.0</td>
|
||
<td id="B0025DB9C82CA7C4FAB7FE25FA4DFE27" box="[1385,1417,428,452]" gridcol="6" gridrow="5" pageId="11" pageNumber="200">1.2</td>
|
||
</tr>
|
||
<tr id="F3D334C5C82CA7C4FF78FE5DFA4DFE0F" box="[166,1417,468,492]" gridrow="6" pageId="11" pageNumber="200">
|
||
<th id="B0025DB9C82CA7C4FF78FE5DFD5AFE0F" box="[166,670,468,492]" gridcol="0" gridrow="6" pageId="11" pageNumber="200">Body width at dorsal-fin origin</th>
|
||
<td id="B0025DB9C82CA7C4FD2CFE5DFC9CFE0F" box="[754,856,468,492]" gridcol="1" gridrow="6" pageId="11" pageNumber="200">11.5–14.0</td>
|
||
<td id="B0025DB9C82CA7C4FC4AFE5DFC16FE0F" box="[916,978,468,492]" gridcol="2" gridrow="6" pageId="11" pageNumber="200">13.0</td>
|
||
<td id="B0025DB9C82CA7C4FBDCFE5DFBE6FE0F" box="[1026,1058,468,492]" gridcol="3" gridrow="6" pageId="11" pageNumber="200">1.0</td>
|
||
<td id="B0025DB9C82CA7C4FB84FE5DFB04FE0F" box="[1114,1216,468,492]" gridcol="4" gridrow="6" pageId="11" pageNumber="200">9.7–13.7</td>
|
||
<td id="B0025DB9C82CA7C4FB22FE5DFAFEFE0F" box="[1276,1338,468,492]" gridcol="5" gridrow="6" pageId="11" pageNumber="200">11.6</td>
|
||
<td id="B0025DB9C82CA7C4FAB7FE5DFA4DFE0F" box="[1385,1417,468,492]" gridcol="6" gridrow="6" pageId="11" pageNumber="200">1.2</td>
|
||
</tr>
|
||
<tr id="F3D334C5C82CA7C4FF78FE75FA4DFDF7" box="[166,1417,508,532]" gridrow="7" pageId="11" pageNumber="200">
|
||
<th id="B0025DB9C82CA7C4FF78FE75FD5AFDF7" box="[166,670,508,532]" gridcol="0" gridrow="7" pageId="11" pageNumber="200">Pre-dorsal length</th>
|
||
<td id="B0025DB9C82CA7C4FD2CFE75FC9CFDF7" box="[754,856,508,532]" gridcol="1" gridrow="7" pageId="11" pageNumber="200">49.6–50.9</td>
|
||
<td id="B0025DB9C82CA7C4FC4AFE75FC16FDF7" box="[916,978,508,532]" gridcol="2" gridrow="7" pageId="11" pageNumber="200">50.4</td>
|
||
<td id="B0025DB9C82CA7C4FBDCFE75FBE6FDF7" box="[1026,1058,508,532]" gridcol="3" gridrow="7" pageId="11" pageNumber="200">0.6</td>
|
||
<td id="B0025DB9C82CA7C4FB84FE75FB04FDF7" box="[1114,1216,508,532]" gridcol="4" gridrow="7" pageId="11" pageNumber="200">47.1–53.1</td>
|
||
<td id="B0025DB9C82CA7C4FB22FE75FAFEFDF7" box="[1276,1338,508,532]" gridcol="5" gridrow="7" pageId="11" pageNumber="200">50.4</td>
|
||
<td id="B0025DB9C82CA7C4FAB7FE75FA4DFDF7" box="[1385,1417,508,532]" gridcol="6" gridrow="7" pageId="11" pageNumber="200">1.3</td>
|
||
</tr>
|
||
<tr id="F3D334C5C82CA7C4FF78FDADFA4DFDDE" box="[166,1417,548,573]" gridrow="8" pageId="11" pageNumber="200">
|
||
<th id="B0025DB9C82CA7C4FF78FDADFD5AFDDE" box="[166,670,548,573]" gridcol="0" gridrow="8" pageId="11" pageNumber="200">Post-dorsal length</th>
|
||
<td id="B0025DB9C82CA7C4FD2CFDADFC9CFDDE" box="[754,856,548,573]" gridcol="1" gridrow="8" pageId="11" pageNumber="200">33.8–35.8</td>
|
||
<td id="B0025DB9C82CA7C4FC4AFDADFC16FDDE" box="[916,978,548,573]" gridcol="2" gridrow="8" pageId="11" pageNumber="200">35.0</td>
|
||
<td id="B0025DB9C82CA7C4FBDCFDADFBE6FDDE" box="[1026,1058,548,573]" gridcol="3" gridrow="8" pageId="11" pageNumber="200">0.5</td>
|
||
<td id="B0025DB9C82CA7C4FB84FDADFB04FDDE" box="[1114,1216,548,573]" gridcol="4" gridrow="8" pageId="11" pageNumber="200">31.9–36.9</td>
|
||
<td id="B0025DB9C82CA7C4FB22FDADFAFEFDDE" box="[1276,1338,548,573]" gridcol="5" gridrow="8" pageId="11" pageNumber="200">34.2</td>
|
||
<td id="B0025DB9C82CA7C4FAB7FDADFA4DFDDE" box="[1385,1417,548,573]" gridcol="6" gridrow="8" pageId="11" pageNumber="200">1.3</td>
|
||
</tr>
|
||
<tr id="F3D334C5C82CA7C4FF78FDC4FA4DFD86" box="[166,1417,589,613]" gridrow="9" pageId="11" pageNumber="200">
|
||
<th id="B0025DB9C82CA7C4FF78FDC4FD5AFD86" box="[166,670,589,613]" gridcol="0" gridrow="9" pageId="11" pageNumber="200">Pre-pelvic length</th>
|
||
<td id="B0025DB9C82CA7C4FD2CFDC4FC9CFD86" box="[754,856,589,613]" gridcol="1" gridrow="9" pageId="11" pageNumber="200">51.1–53.8</td>
|
||
<td id="B0025DB9C82CA7C4FC4AFDC4FC16FD86" box="[916,978,589,613]" gridcol="2" gridrow="9" pageId="11" pageNumber="200">52.6</td>
|
||
<td id="B0025DB9C82CA7C4FBDCFDC4FBE6FD86" box="[1026,1058,589,613]" gridcol="3" gridrow="9" pageId="11" pageNumber="200">0.7</td>
|
||
<td id="B0025DB9C82CA7C4FB84FDC4FB04FD86" box="[1114,1216,589,613]" gridcol="4" gridrow="9" pageId="11" pageNumber="200">51.2–54.1</td>
|
||
<td id="B0025DB9C82CA7C4FB22FDC4FAFEFD86" box="[1276,1338,589,613]" gridcol="5" gridrow="9" pageId="11" pageNumber="200">52.6</td>
|
||
<td id="B0025DB9C82CA7C4FAB7FDC4FA4DFD86" box="[1385,1417,589,613]" gridcol="6" gridrow="9" pageId="11" pageNumber="200">0.8</td>
|
||
</tr>
|
||
<tr id="F3D334C5C82CA7C4FF78FDFCFA4DFD6E" box="[166,1417,629,653]" gridrow="10" pageId="11" pageNumber="200">
|
||
<th id="B0025DB9C82CA7C4FF78FDFCFD5AFD6E" box="[166,670,629,653]" gridcol="0" gridrow="10" pageId="11" pageNumber="200">Pre-anal length</th>
|
||
<td id="B0025DB9C82CA7C4FD2CFDFCFC9CFD6E" box="[754,856,629,653]" gridcol="1" gridrow="10" pageId="11" pageNumber="200">74.0–76.5</td>
|
||
<td id="B0025DB9C82CA7C4FC4AFDFCFC16FD6E" box="[916,978,629,653]" gridcol="2" gridrow="10" pageId="11" pageNumber="200">74.6</td>
|
||
<td id="B0025DB9C82CA7C4FBDCFDFCFBE6FD6E" box="[1026,1058,629,653]" gridcol="3" gridrow="10" pageId="11" pageNumber="200">0.9</td>
|
||
<td id="B0025DB9C82CA7C4FB84FDFCFB04FD6E" box="[1114,1216,629,653]" gridcol="4" gridrow="10" pageId="11" pageNumber="200">73.1–76.2</td>
|
||
<td id="B0025DB9C82CA7C4FB22FDFCFAFEFD6E" box="[1276,1338,629,653]" gridcol="5" gridrow="10" pageId="11" pageNumber="200">74.7</td>
|
||
<td id="B0025DB9C82CA7C4FAB7FDFCFA4DFD6E" box="[1385,1417,629,653]" gridcol="6" gridrow="10" pageId="11" pageNumber="200">1.0</td>
|
||
</tr>
|
||
<tr id="F3D334C5C82CA7C4FF78FD14FA4DFD56" box="[166,1417,669,693]" gridrow="11" pageId="11" pageNumber="200">
|
||
<th id="B0025DB9C82CA7C4FF78FD14FD5AFD56" box="[166,670,669,693]" gridcol="0" gridrow="11" pageId="11" pageNumber="200">Distance between pectoral and pelvic-fin origins</th>
|
||
<td id="B0025DB9C82CA7C4FD2CFD14FC9CFD56" box="[754,856,669,693]" gridcol="1" gridrow="11" pageId="11" pageNumber="200">25.5–30.7</td>
|
||
<td id="B0025DB9C82CA7C4FC4AFD14FC16FD56" box="[916,978,669,693]" gridcol="2" gridrow="11" pageId="11" pageNumber="200">28.8</td>
|
||
<td id="B0025DB9C82CA7C4FBDCFD14FBE6FD56" box="[1026,1058,669,693]" gridcol="3" gridrow="11" pageId="11" pageNumber="200">0.7</td>
|
||
<td id="B0025DB9C82CA7C4FB84FD14FB04FD56" box="[1114,1216,669,693]" gridcol="4" gridrow="11" pageId="11" pageNumber="200">25.2–28.0</td>
|
||
<td id="B0025DB9C82CA7C4FB22FD14FAFEFD56" box="[1276,1338,669,693]" gridcol="5" gridrow="11" pageId="11" pageNumber="200">26.3</td>
|
||
<td id="B0025DB9C82CA7C4FAB7FD14FA4DFD56" box="[1385,1417,669,693]" gridcol="6" gridrow="11" pageId="11" pageNumber="200">0.8</td>
|
||
</tr>
|
||
<tr id="F3D334C5C82CA7C4FF78FD4CFA4DFD3D" box="[166,1417,709,734]" gridrow="12" pageId="11" pageNumber="200">
|
||
<th id="B0025DB9C82CA7C4FF78FD4CFD5AFD3D" box="[166,670,709,734]" gridcol="0" gridrow="12" pageId="11" pageNumber="200">Distance between pelvic and anal-fin origins</th>
|
||
<td id="B0025DB9C82CA7C4FD2CFD4CFC9CFD3D" box="[754,856,709,734]" gridcol="1" gridrow="12" pageId="11" pageNumber="200">19.5–22.3</td>
|
||
<td id="B0025DB9C82CA7C4FC4AFD4CFC16FD3D" box="[916,978,709,734]" gridcol="2" gridrow="12" pageId="11" pageNumber="200">21.0</td>
|
||
<td id="B0025DB9C82CA7C4FBDCFD4CFBE6FD3D" box="[1026,1058,709,734]" gridcol="3" gridrow="12" pageId="11" pageNumber="200">1.0</td>
|
||
<td id="B0025DB9C82CA7C4FB84FD4CFB04FD3D" box="[1114,1216,709,734]" gridcol="4" gridrow="12" pageId="11" pageNumber="200">22.3–23.8</td>
|
||
<td id="B0025DB9C82CA7C4FB22FD4CFAFEFD3D" box="[1276,1338,709,734]" gridcol="5" gridrow="12" pageId="11" pageNumber="200">23.1</td>
|
||
<td id="B0025DB9C82CA7C4FAB7FD4CFA4DFD3D" box="[1385,1417,709,734]" gridcol="6" gridrow="12" pageId="11" pageNumber="200">0.4</td>
|
||
</tr>
|
||
<tr id="F3D334C5C82CA7C4FF78FD67FA4DFCE5" box="[166,1417,750,774]" gridrow="13" pageId="11" pageNumber="200">
|
||
<th id="B0025DB9C82CA7C4FF78FD67FD5AFCE5" box="[166,670,750,774]" gridcol="0" gridrow="13" pageId="11" pageNumber="200">Depth of caudal peduncle</th>
|
||
<td id="B0025DB9C82CA7C4FD2CFD67FC9CFCE5" box="[754,856,750,774]" gridcol="1" gridrow="13" pageId="11" pageNumber="200">7.7–10.9</td>
|
||
<td id="B0025DB9C82CA7C4FC4AFD67FC16FCE5" box="[916,978,750,774]" gridcol="2" gridrow="13" pageId="11" pageNumber="200">8.5</td>
|
||
<td id="B0025DB9C82CA7C4FBDCFD67FBE6FCE5" box="[1026,1058,750,774]" gridcol="3" gridrow="13" pageId="11" pageNumber="200">0.8</td>
|
||
<td id="B0025DB9C82CA7C4FB84FD67FB04FCE5" box="[1114,1216,750,774]" gridcol="4" gridrow="13" pageId="11" pageNumber="200">9.0–11.3</td>
|
||
<td id="B0025DB9C82CA7C4FB22FD67FAFEFCE5" box="[1276,1338,750,774]" gridcol="5" gridrow="13" pageId="11" pageNumber="200">10.2</td>
|
||
<td id="B0025DB9C82CA7C4FAB7FD67FA4DFCE5" box="[1385,1417,750,774]" gridcol="6" gridrow="13" pageId="11" pageNumber="200">0.6</td>
|
||
</tr>
|
||
<tr id="F3D334C5C82CA7C4FF78FC9FFA4DFCCD" box="[166,1417,790,814]" gridrow="14" pageId="11" pageNumber="200">
|
||
<th id="B0025DB9C82CA7C4FF78FC9FFD5AFCCD" box="[166,670,790,814]" gridcol="0" gridrow="14" pageId="11" pageNumber="200">Length of caudal peduncle</th>
|
||
<td id="B0025DB9C82CA7C4FD2CFC9FFC9CFCCD" box="[754,856,790,814]" gridcol="1" gridrow="14" pageId="11" pageNumber="200">17.2–19.6</td>
|
||
<td id="B0025DB9C82CA7C4FC4AFC9FFC16FCCD" box="[916,978,790,814]" gridcol="2" gridrow="14" pageId="11" pageNumber="200">19.1</td>
|
||
<td id="B0025DB9C82CA7C4FBDCFC9FFBE6FCCD" box="[1026,1058,790,814]" gridcol="3" gridrow="14" pageId="11" pageNumber="200">0.8</td>
|
||
<td id="B0025DB9C82CA7C4FB84FC9FFB04FCCD" box="[1114,1216,790,814]" gridcol="4" gridrow="14" pageId="11" pageNumber="200">15.6–19.3</td>
|
||
<td id="B0025DB9C82CA7C4FB22FC9FFAFEFCCD" box="[1276,1338,790,814]" gridcol="5" gridrow="14" pageId="11" pageNumber="200">17.7</td>
|
||
<td id="B0025DB9C82CA7C4FAB7FC9FFA4DFCCD" box="[1385,1417,790,814]" gridcol="6" gridrow="14" pageId="11" pageNumber="200">0.8</td>
|
||
</tr>
|
||
<tr id="F3D334C5C82CA7C4FF78FCB7FA4DFCB5" box="[166,1417,830,854]" gridrow="15" pageId="11" pageNumber="200">
|
||
<th id="B0025DB9C82CA7C4FF78FCB7FD5AFCB5" box="[166,670,830,854]" gridcol="0" gridrow="15" pageId="11" pageNumber="200">Dorsal-fin length</th>
|
||
<td id="B0025DB9C82CA7C4FD2CFCB7FC9CFCB5" box="[754,856,830,854]" gridcol="1" gridrow="15" pageId="11" pageNumber="200">20.3–23.1</td>
|
||
<td id="B0025DB9C82CA7C4FC4AFCB7FC16FCB5" box="[916,978,830,854]" gridcol="2" gridrow="15" pageId="11" pageNumber="200">21.5</td>
|
||
<td id="B0025DB9C82CA7C4FBDCFCB7FBE6FCB5" box="[1026,1058,830,854]" gridcol="3" gridrow="15" pageId="11" pageNumber="200">0.7</td>
|
||
<td id="B0025DB9C82CA7C4FB84FCB7FB04FCB5" box="[1114,1216,830,854]" gridcol="4" gridrow="15" pageId="11" pageNumber="200">22.5–27.2</td>
|
||
<td id="B0025DB9C82CA7C4FB22FCB7FAFEFCB5" box="[1276,1338,830,854]" gridcol="5" gridrow="15" pageId="11" pageNumber="200">24.4</td>
|
||
<td id="B0025DB9C82CA7C4FAB7FCB7FA4DFCB5" box="[1385,1417,830,854]" gridcol="6" gridrow="15" pageId="11" pageNumber="200">1.2</td>
|
||
</tr>
|
||
<tr id="F3D334C5C82CA7C4FF78FCEFFA4DFC9C" box="[166,1417,870,895]" gridrow="16" pageId="11" pageNumber="200">
|
||
<th id="B0025DB9C82CA7C4FF78FCEFFD5AFC9C" box="[166,670,870,895]" gridcol="0" gridrow="16" pageId="11" pageNumber="200">Pectoral-fin length</th>
|
||
<td id="B0025DB9C82CA7C4FD2CFCEFFC9CFC9C" box="[754,856,870,895]" gridcol="1" gridrow="16" pageId="11" pageNumber="200">19.6–25.2</td>
|
||
<td id="B0025DB9C82CA7C4FC4AFCEFFC16FC9C" box="[916,978,870,895]" gridcol="2" gridrow="16" pageId="11" pageNumber="200">23.1</td>
|
||
<td id="B0025DB9C82CA7C4FBDCFCEFFBE6FC9C" box="[1026,1058,870,895]" gridcol="3" gridrow="16" pageId="11" pageNumber="200">0.8</td>
|
||
<td id="B0025DB9C82CA7C4FB84FCEFFB04FC9C" box="[1114,1216,870,895]" gridcol="4" gridrow="16" pageId="11" pageNumber="200">24.5–28.3</td>
|
||
<td id="B0025DB9C82CA7C4FB22FCEFFAFEFC9C" box="[1276,1338,870,895]" gridcol="5" gridrow="16" pageId="11" pageNumber="200">26.5</td>
|
||
<td id="B0025DB9C82CA7C4FAB7FCEFFA4DFC9C" box="[1385,1417,870,895]" gridcol="6" gridrow="16" pageId="11" pageNumber="200">1.1</td>
|
||
</tr>
|
||
<tr id="F3D334C5C82CA7C4FF78FC07FA4DFC44" box="[166,1417,910,935]" gridrow="17" pageId="11" pageNumber="200">
|
||
<th id="B0025DB9C82CA7C4FF78FC07FD5AFC44" box="[166,670,910,935]" gridcol="0" gridrow="17" pageId="11" pageNumber="200">Pelvic-fin length</th>
|
||
<td id="B0025DB9C82CA7C4FD2CFC07FC9CFC44" box="[754,856,910,935]" gridcol="1" gridrow="17" pageId="11" pageNumber="200">15.5–19.2</td>
|
||
<td id="B0025DB9C82CA7C4FC4AFC07FC16FC44" box="[916,978,910,935]" gridcol="2" gridrow="17" pageId="11" pageNumber="200">17.3</td>
|
||
<td id="B0025DB9C82CA7C4FBDCFC07FBE6FC44" box="[1026,1058,910,935]" gridcol="3" gridrow="17" pageId="11" pageNumber="200">1.6</td>
|
||
<td id="B0025DB9C82CA7C4FB84FC07FB04FC44" box="[1114,1216,910,935]" gridcol="4" gridrow="17" pageId="11" pageNumber="200">17.8–20.7</td>
|
||
<td id="B0025DB9C82CA7C4FB22FC07FAFEFC44" box="[1276,1338,910,935]" gridcol="5" gridrow="17" pageId="11" pageNumber="200">19.3</td>
|
||
<td id="B0025DB9C82CA7C4FAB7FC07FA4DFC44" box="[1385,1417,910,935]" gridcol="6" gridrow="17" pageId="11" pageNumber="200">0.7</td>
|
||
</tr>
|
||
<tr id="F3D334C5C82CA7C4FF78FC3FFA4DFC2C" box="[166,1417,950,975]" gridrow="18" pageId="11" pageNumber="200">
|
||
<th id="B0025DB9C82CA7C4FF78FC3FFA4DFC2C" box="[166,1417,950,975]" colspan="7" colspanRight="6" gridcol="0" gridrow="18" pageId="11" pageNumber="200">
|
||
<emphasis id="7F97EA95C82C5830FF78FC3EFE71FC2C" bold="true" box="[166,437,950,975]" pageId="11" pageNumber="200">In percent of head length</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="F3D334C5C82CA7C4FF78FC56FA4DFC14" box="[166,1417,991,1015]" gridrow="19" pageId="11" pageNumber="200">
|
||
<th id="B0025DB9C82CA7C4FF78FC56FD5AFC14" box="[166,670,991,1015]" gridcol="0" gridrow="19" pageId="11" pageNumber="200">Head depth at nape</th>
|
||
<td id="B0025DB9C82CA7C4FD2CFC56FC9CFC14" box="[754,856,991,1015]" gridcol="1" gridrow="19" pageId="11" pageNumber="200">55–61</td>
|
||
<td id="B0025DB9C82CA7C4FC4AFC56FC16FC14" box="[916,978,991,1015]" gridcol="2" gridrow="19" pageId="11" pageNumber="200">59.3</td>
|
||
<td id="B0025DB9C82CA7C4FBDCFC56FBE6FC14" box="[1026,1058,991,1015]" gridcol="3" gridrow="19" pageId="11" pageNumber="200">2.0</td>
|
||
<td id="B0025DB9C82CA7C4FB84FC56FB04FC14" box="[1114,1216,991,1015]" gridcol="4" gridrow="19" pageId="11" pageNumber="200">50–61</td>
|
||
<td id="B0025DB9C82CA7C4FB22FC56FAFEFC14" box="[1276,1338,991,1015]" gridcol="5" gridrow="19" pageId="11" pageNumber="200">55.6</td>
|
||
<td id="B0025DB9C82CA7C4FAB7FC56FA4DFC14" box="[1385,1417,991,1015]" gridcol="6" gridrow="19" pageId="11" pageNumber="200">2.9</td>
|
||
</tr>
|
||
<tr id="F3D334C5C82CA7C4FF78FB8EFA4DFBFC" box="[166,1417,1031,1055]" gridrow="20" pageId="11" pageNumber="200">
|
||
<th id="B0025DB9C82CA7C4FF78FB8EFD5AFBFC" box="[166,670,1031,1055]" gridcol="0" gridrow="20" pageId="11" pageNumber="200">Head depth at eye</th>
|
||
<td id="B0025DB9C82CA7C4FD2CFB8EFC9CFBFC" box="[754,856,1031,1055]" gridcol="1" gridrow="20" pageId="11" pageNumber="200">49–55</td>
|
||
<td id="B0025DB9C82CA7C4FC4AFB8EFC16FBFC" box="[916,978,1031,1055]" gridcol="2" gridrow="20" pageId="11" pageNumber="200">51.6</td>
|
||
<td id="B0025DB9C82CA7C4FBDCFB8EFBE6FBFC" box="[1026,1058,1031,1055]" gridcol="3" gridrow="20" pageId="11" pageNumber="200">1.8</td>
|
||
<td id="B0025DB9C82CA7C4FB84FB8EFB04FBFC" box="[1114,1216,1031,1055]" gridcol="4" gridrow="20" pageId="11" pageNumber="200">45–56</td>
|
||
<td id="B0025DB9C82CA7C4FB22FB8EFAFEFBFC" box="[1276,1338,1031,1055]" gridcol="5" gridrow="20" pageId="11" pageNumber="200">51.4</td>
|
||
<td id="B0025DB9C82CA7C4FAB7FB8EFA4DFBFC" box="[1385,1417,1031,1055]" gridcol="6" gridrow="20" pageId="11" pageNumber="200">2.9</td>
|
||
</tr>
|
||
<tr id="F3D334C5C82CA7C4FF78FBA6FA4DFBA4" box="[166,1417,1071,1095]" gridrow="21" pageId="11" pageNumber="200">
|
||
<th id="B0025DB9C82CA7C4FF78FBA6FD5AFBA4" box="[166,670,1071,1095]" gridcol="0" gridrow="21" pageId="11" pageNumber="200">Snout length</th>
|
||
<td id="B0025DB9C82CA7C4FD2CFBA6FC9CFBA4" box="[754,856,1071,1095]" gridcol="1" gridrow="21" pageId="11" pageNumber="200">39–42</td>
|
||
<td id="B0025DB9C82CA7C4FC4AFBA6FC16FBA4" box="[916,978,1071,1095]" gridcol="2" gridrow="21" pageId="11" pageNumber="200">40.7</td>
|
||
<td id="B0025DB9C82CA7C4FBDCFBA6FBE6FBA4" box="[1026,1058,1071,1095]" gridcol="3" gridrow="21" pageId="11" pageNumber="200">0.7</td>
|
||
<td id="B0025DB9C82CA7C4FB84FBA6FB04FBA4" box="[1114,1216,1071,1095]" gridcol="4" gridrow="21" pageId="11" pageNumber="200">37–44</td>
|
||
<td id="B0025DB9C82CA7C4FB22FBA6FAFEFBA4" box="[1276,1338,1071,1095]" gridcol="5" gridrow="21" pageId="11" pageNumber="200">40.8</td>
|
||
<td id="B0025DB9C82CA7C4FAB7FBA6FA4DFBA4" box="[1385,1417,1071,1095]" gridcol="6" gridrow="21" pageId="11" pageNumber="200">1.5</td>
|
||
</tr>
|
||
<tr id="F3D334C5C82CA7C4FF78FBDEFA4DFB93" box="[166,1417,1111,1136]" gridrow="22" pageId="11" pageNumber="200">
|
||
<th id="B0025DB9C82CA7C4FF78FBDEFD5AFB93" box="[166,670,1111,1136]" gridcol="0" gridrow="22" pageId="11" pageNumber="200">Eye diameter</th>
|
||
<td id="B0025DB9C82CA7C4FD2CFBDEFC9CFB93" box="[754,856,1111,1136]" gridcol="1" gridrow="22" pageId="11" pageNumber="200">15–19</td>
|
||
<td id="B0025DB9C82CA7C4FC4AFBDEFC16FB93" box="[916,978,1111,1136]" gridcol="2" gridrow="22" pageId="11" pageNumber="200">16.8</td>
|
||
<td id="B0025DB9C82CA7C4FBDCFBDEFBE6FB93" box="[1026,1058,1111,1136]" gridcol="3" gridrow="22" pageId="11" pageNumber="200">1.2</td>
|
||
<td id="B0025DB9C82CA7C4FB84FBDEFB04FB93" box="[1114,1216,1111,1136]" gridcol="4" gridrow="22" pageId="11" pageNumber="200">18–25</td>
|
||
<td id="B0025DB9C82CA7C4FB22FBDEFAFEFB93" box="[1276,1338,1111,1136]" gridcol="5" gridrow="22" pageId="11" pageNumber="200">20.6</td>
|
||
<td id="B0025DB9C82CA7C4FAB7FBDEFA4DFB93" box="[1385,1417,1111,1136]" gridcol="6" gridrow="22" pageId="11" pageNumber="200">2.2</td>
|
||
</tr>
|
||
<tr id="F3D334C5C82CA7C4FF78FB09FA4DFB7B" box="[166,1417,1152,1176]" gridrow="23" pageId="11" pageNumber="200">
|
||
<th id="B0025DB9C82CA7C4FF78FB09FD5AFB7B" box="[166,670,1152,1176]" gridcol="0" gridrow="23" pageId="11" pageNumber="200">Postorbital distance</th>
|
||
<td id="B0025DB9C82CA7C4FD2CFB09FC9CFB7B" box="[754,856,1152,1176]" gridcol="1" gridrow="23" pageId="11" pageNumber="200">42–46</td>
|
||
<td id="B0025DB9C82CA7C4FC4AFB09FC16FB7B" box="[916,978,1152,1176]" gridcol="2" gridrow="23" pageId="11" pageNumber="200">43.1</td>
|
||
<td id="B0025DB9C82CA7C4FBDCFB09FBE6FB7B" box="[1026,1058,1152,1176]" gridcol="3" gridrow="23" pageId="11" pageNumber="200">1.5</td>
|
||
<td id="B0025DB9C82CA7C4FB84FB09FB04FB7B" box="[1114,1216,1152,1176]" gridcol="4" gridrow="23" pageId="11" pageNumber="200">40–45</td>
|
||
<td id="B0025DB9C82CA7C4FB22FB09FAFEFB7B" box="[1276,1338,1152,1176]" gridcol="5" gridrow="23" pageId="11" pageNumber="200">43.1</td>
|
||
<td id="B0025DB9C82CA7C4FAB7FB09FA4DFB7B" box="[1385,1417,1152,1176]" gridcol="6" gridrow="23" pageId="11" pageNumber="200">1.2</td>
|
||
</tr>
|
||
<tr id="F3D334C5C82CA7C4FF78FB21FA4DFB23" box="[166,1417,1192,1216]" gridrow="24" pageId="11" pageNumber="200">
|
||
<th id="B0025DB9C82CA7C4FF78FB21FD5AFB23" box="[166,670,1192,1216]" gridcol="0" gridrow="24" pageId="11" pageNumber="200">Maximum head width</th>
|
||
<td id="B0025DB9C82CA7C4FD2CFB21FC9CFB23" box="[754,856,1192,1216]" gridcol="1" gridrow="24" pageId="11" pageNumber="200">62–67</td>
|
||
<td id="B0025DB9C82CA7C4FC4AFB21FC16FB23" box="[916,978,1192,1216]" gridcol="2" gridrow="24" pageId="11" pageNumber="200">64.4</td>
|
||
<td id="B0025DB9C82CA7C4FBDCFB21FBE6FB23" box="[1026,1058,1192,1216]" gridcol="3" gridrow="24" pageId="11" pageNumber="200">1.3</td>
|
||
<td id="B0025DB9C82CA7C4FB84FB21FB04FB23" box="[1114,1216,1192,1216]" gridcol="4" gridrow="24" pageId="11" pageNumber="200">50–66</td>
|
||
<td id="B0025DB9C82CA7C4FB22FB21FAFEFB23" box="[1276,1338,1192,1216]" gridcol="5" gridrow="24" pageId="11" pageNumber="200">58.3</td>
|
||
<td id="B0025DB9C82CA7C4FAB7FB21FA4DFB23" box="[1385,1417,1192,1216]" gridcol="6" gridrow="24" pageId="11" pageNumber="200">3.8</td>
|
||
</tr>
|
||
<tr id="F3D334C5C82CA7C4FF78FB59FA4DFB0B" box="[166,1417,1232,1256]" gridrow="25" pageId="11" pageNumber="200">
|
||
<th id="B0025DB9C82CA7C4FF78FB59FD5AFB0B" box="[166,670,1232,1256]" gridcol="0" gridrow="25" pageId="11" pageNumber="200">Interorbital width</th>
|
||
<td id="B0025DB9C82CA7C4FD2CFB59FC9CFB0B" box="[754,856,1232,1256]" gridcol="1" gridrow="25" pageId="11" pageNumber="200">26–29</td>
|
||
<td id="B0025DB9C82CA7C4FC4AFB59FC16FB0B" box="[916,978,1232,1256]" gridcol="2" gridrow="25" pageId="11" pageNumber="200">27.1</td>
|
||
<td id="B0025DB9C82CA7C4FBDCFB59FBE6FB0B" box="[1026,1058,1232,1256]" gridcol="3" gridrow="25" pageId="11" pageNumber="200">0.8</td>
|
||
<td id="B0025DB9C82CA7C4FB84FB59FB04FB0B" box="[1114,1216,1232,1256]" gridcol="4" gridrow="25" pageId="11" pageNumber="200">26–31</td>
|
||
<td id="B0025DB9C82CA7C4FB22FB59FAFEFB0B" box="[1276,1338,1232,1256]" gridcol="5" gridrow="25" pageId="11" pageNumber="200">28.3</td>
|
||
<td id="B0025DB9C82CA7C4FAB7FB59FA4DFB0B" box="[1385,1417,1232,1256]" gridcol="6" gridrow="25" pageId="11" pageNumber="200">1.4</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="4D5C3687C82C5830FF19FA91FC81FA20" blockId="11.[151,1437,1304,1475]" pageId="11" pageNumber="200">
|
||
Among congeners studied for molecular characters,
|
||
<taxonomicName id="8AE34D04C82C5830FCCDFA91FC72FAD1" authorityName="ZM-CBSU H" authorityYear="1998" box="[787,950,1304,1330]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="200" phylum="Chordata" rank="species" species="marunensis">
|
||
<emphasis id="7F97EA95C82C5830FCCDFA91FCE3FAD1" box="[787,807,1304,1330]" italics="true" pageId="11" pageNumber="200">O</emphasis>
|
||
.
|
||
<emphasis id="7F97EA95C82C5830FCEBFA90FC72FAD1" box="[821,950,1305,1330]" italics="true" pageId="11" pageNumber="200">marunensis</emphasis>
|
||
</taxonomicName>
|
||
is characterised by a K2P nearest neighbor distance of 2% to
|
||
<taxonomicName id="8AE34D04C82C5830FE80FAB5FE1FFAB5" baseAuthorityName="Heckel" baseAuthorityYear="1843" box="[350,475,1340,1366]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="200" phylum="Chordata" rank="species" species="frenatus">
|
||
<emphasis id="7F97EA95C82C5830FE80FAB5FE1FFAB5" box="[350,475,1340,1366]" italics="true" pageId="11" pageNumber="200">O. frenatus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<tableCitation id="0061033CC82C5830FE36FAB5FDF9FAB5" box="[488,573,1340,1366]" captionStart="TABLE 3" captionStartId="4.[151,239,1123,1147]" captionTargetPageId="4" captionText="TABLE 3. Estimates of evolutionary divergence (%) over sequence pairs between species found in the COI barcode region of the Iranian Oxynoemacheilus species studied." httpUri="http://table.plazi.org/id/199C660FC823583FFF49FBEAFD21FB7C" pageId="11" pageNumber="200" tableUuid="199C660FC823583FFF49FBEAFD21FB7C">Table 3</tableCitation>
|
||
), whereas, they are largely diverged based on morphological characters.
|
||
<taxonomicName id="8AE34D04C82C5830FAB8FAB5FE7EFA99" authorityName="ZM-CBSU H" authorityYear="1998" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="200" phylum="Chordata" rank="species" species="marunensis">
|
||
<emphasis id="7F97EA95C82C5830FAB8FAB5FE7EFA99" italics="true" pageId="11" pageNumber="200">Oxynoemacheilus marunensis</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="8AE34D04C82C5830FD66FAE9FCF2FA99" baseAuthorityName="Heckel" baseAuthorityYear="1843" box="[696,822,1376,1402]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="200" phylum="Chordata" rank="species" species="frenatus">
|
||
<emphasis id="7F97EA95C82C5830FD66FAE9FCF2FA99" box="[696,822,1376,1402]" italics="true" pageId="11" pageNumber="200">O. frenatus</emphasis>
|
||
</taxonomicName>
|
||
by having a suborbital groove in males (vs. absent), a deeply emarginate or forked caudal fin (vs. slightly emarginate), complete lateral line (vs. incomplete), and having two prominent black spots at the caudal-fin base (vs. absent).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |