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<mods:title id="995B998D1AD6C8FA525CFA5F14956C65">Phylogeny, taxonomy, and zoogeography of the genus Gekko Laurenti, 1768 with the revalidation of G. reevesii Gray, 1831 (Sauria: Gekkonidae)</mods:title>
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<treatment id="19358A1A8B07FFB88980FB73FEC5F7A7" ID-DOI="http://doi.org/10.5281/zenodo.6188386" ID-GBIF-Taxon="119404895" ID-Zenodo-Dep="6188386" LSID="urn:lsid:plazi:treatment:19358A1A8B07FFB88980FB73FEC5F7A7" httpUri="http://treatment.plazi.org/id/19358A1A8B07FFB88980FB73FEC5F7A7" lastPageId="16" lastPageNumber="17" pageId="12" pageNumber="13">
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<paragraph id="91233B0C8B07FFA48980FB73FE12F1D1" blockId="12.[151,435,1184,1244]" box="[151,435,1184,1211]" pageId="12" pageNumber="13">
<heading id="CA6B8C608B07FFA48980FB73FE12F1D1" bold="true" box="[151,435,1184,1211]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<emphasis id="A3E8E71E8B07FFA48980FB73FE12F1D1" bold="true" box="[151,435,1184,1211]" pageId="12" pageNumber="13">
<taxonomicName id="569C408F8B07FFA48980FB73FEC0F1D0" box="[151,353,1185,1211]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="japonicus">
<emphasis id="A3E8E71E8B07FFA48980FB73FEC0F1D0" bold="true" box="[151,353,1185,1211]" italics="true" pageId="12" pageNumber="13">Gekko japonicus</emphasis>
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group
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<paragraph id="91233B0C8B07FFA48980FB16FEE3F1B7" blockId="12.[151,435,1184,1244]" box="[151,322,1220,1244]" pageId="12" pageNumber="13">
(
<figureCitation id="09A727898B07FFA48989FB16FEAAF1B7" box="[158,267,1220,1244]" captionStart="FIGURE 4. A" captionStartId="14.[151,250,1008,1031]" captionTargetBox="[151,1436,193,986]" captionTargetId="figure@14.[151,1436,193,987]" captionTargetPageId="14" captionText="FIGURE 4. A. Female Gekko japonicus (G. japonicus group), photo by J. Boone; B. Male G. hokouensis (G. japonicus group), photo by J. Boone; C. G. palmatus (G. japonicus group), Quang Binh Province, Vietnam, photo by T. Ziegler; D. G. scientiadventura (G. japonicus group), Quang Binh Province, Vietnam, photo by T. Ziegler." httpUri="https://zenodo.org/record/278397/files/figure.png" pageId="12" pageNumber="13">Figures 4</figureCitation>
, 
<figureCitation id="09A727898B07FFA48800FB16FE9BF1B7" box="[279,314,1220,1244]" captionStart="FIGURE 5. A" captionStartId="14.[151,250,1944,1967]" captionTargetBox="[151,1436,1129,1922]" captionTargetId="figure@14.[151,1436,1129,1923]" captionTargetPageId="14" captionText="FIGURE 5. A. Male Gekko vertebralis (G. japonicus group), photo by J. Boone; B. G. monarchus (G. monarchus group), Banggai Islands, Indonesia, photo by A. Koch; C. G. ernstkelleri (G. monarchus group), Panay Island, Philippines, photo by M. Gaulke; D. G. mindorensis (G. monarchus group), Cebu Island, Philippines, photo by M. Gaulke." httpUri="https://zenodo.org/record/278398/files/figure.png" pageId="12" pageNumber="13">5A</figureCitation>
)
</paragraph>
<paragraph id="91233B0C8B07FFA48980FAD9FC84F097" blockId="12.[151,1436,1291,2036]" pageId="12" pageNumber="13">
<emphasis id="A3E8E71E8B07FFA48980FAD9FF54F04F" bold="true" box="[151,245,1291,1316]" pageId="12" pageNumber="13">Species.</emphasis>
<taxonomicName id="569C408F8B07FFA489E9FADFFD01F04F" authority="Zhou &amp; Liu 1982" authorityName="Zhou &amp; Liu" authorityYear="1982" box="[254,672,1292,1316]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="auriverrucosus">
<emphasis id="A3E8E71E8B07FFA489E9FADFFE68F04F" box="[254,457,1293,1316]" italics="true" pageId="12" pageNumber="13">G. auriverrucosus</emphasis>
Zhou &amp; Liu 1982
</taxonomicName>
; 
<taxonomicName id="569C408F8B07FFA48BB9FADFFAA4F04F" authority="Rosler, Nguyen, Doan, Ho &amp; Ziegler 2010" authorityName="Rosler, Nguyen, Doan, Ho &amp; Ziegler" authorityYear="2010" box="[686,1285,1292,1317]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="canhi">
<emphasis id="A3E8E71E8B07FFA48BB9FADFFCACF04F" box="[686,781,1292,1316]" italics="true" pageId="12" pageNumber="13">G. canhi</emphasis>
<bibRefCitation id="F50D46FD8B07FFA48A00FADEFAA4F04F" author="Rosler" box="[791,1285,1292,1317]" pageId="12" pageNumber="13" refString="Rosler, H., Nguyen, Q. T., Doan, V. K., Ho, T. C., Nguyen, T. T. &amp; Ziegler, T. (2010) A new species of the genus Gekko Laurenti, 1768 (Squamata: Sauria: Gekkonidae) from Vietnam with remarks on G. japonicus (Schlegel, 1836). Zootaxa, 2329, 56 - 68." type="journal article" year="2010">Rösler, Nguyen, Doan, Ho &amp; Ziegler 2010</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="569C408F8B07FFA48C04FADFFEB7F02C" authority="Gray, 1842" authorityName="Gray" authorityYear="1842" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="chinensis">
<emphasis id="A3E8E71E8B07FFA48C04FADFFA3AF04F" box="[1299,1435,1292,1316]" italics="true" pageId="12" pageNumber="13">G. chinensis</emphasis>
Gray, 1842
</taxonomicName>
; 
<taxonomicName id="569C408F8B07FFA48836FAE2FDE1F02C" authority="Pope, 1928" authorityName="Pope" authorityYear="1928" box="[289,576,1327,1352]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="hokouensis">
<emphasis id="A3E8E71E8B07FFA48836FAE2FE1AF023" box="[289,443,1328,1352]" italics="true" pageId="12" pageNumber="13">G. hokouensis</emphasis>
Pope, 1928
</taxonomicName>
(
<figureCitation id="09A727898B07FFA48B58FAFDFD11F02C" box="[591,688,1327,1351]" captionStart="FIGURE 4. A" captionStartId="14.[151,250,1008,1031]" captionTargetBox="[151,1436,193,986]" captionTargetId="figure@14.[151,1436,193,987]" captionTargetPageId="14" captionText="FIGURE 4. A. Female Gekko japonicus (G. japonicus group), photo by J. Boone; B. Male G. hokouensis (G. japonicus group), photo by J. Boone; C. G. palmatus (G. japonicus group), Quang Binh Province, Vietnam, photo by T. Ziegler; D. G. scientiadventura (G. japonicus group), Quang Binh Province, Vietnam, photo by T. Ziegler." httpUri="https://zenodo.org/record/278397/files/figure.png" pageId="12" pageNumber="13">Figure 4</figureCitation>
B); 
<taxonomicName id="569C408F8B07FFA48BCEFAE2FB82F02C" authority="Schlegel, 1836" authorityName="Schlegel" authorityYear="1836" box="[729,1059,1327,1352]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="japonicus">
<emphasis id="A3E8E71E8B07FFA48BCEFAE2FCC5F023" box="[729,868,1328,1352]" italics="true" pageId="12" pageNumber="13">G. japonicus</emphasis>
(
<bibRefCitation id="F50D46FD8B07FFA48A65FAFDFBBAF02C" author="Schlegel" box="[882,1051,1327,1351]" pageId="12" pageNumber="13" refString="Schlegel, H. (1836) Le Platydactyle Monarque. Platydactylus Monarchus. In: Dumeril, A. M. C. &amp; Bibron, G. (1836), Erpetologie Generale ou Histoire Naturelle Complete des Reptiles. Vol. 3. Roret, Paris, pp. 335 - 336." type="book chapter" year="1836">Schlegel, 1836</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="09A727898B07FFA48D26FAFDFB08F02C" box="[1073,1193,1327,1351]" captionStart="FIGURE 4. A" captionStartId="14.[151,250,1008,1031]" captionTargetBox="[151,1436,193,986]" captionTargetId="figure@14.[151,1436,193,987]" captionTargetPageId="14" captionText="FIGURE 4. A. Female Gekko japonicus (G. japonicus group), photo by J. Boone; B. Male G. hokouensis (G. japonicus group), photo by J. Boone; C. G. palmatus (G. japonicus group), Quang Binh Province, Vietnam, photo by T. Ziegler; D. G. scientiadventura (G. japonicus group), Quang Binh Province, Vietnam, photo by T. Ziegler." httpUri="https://zenodo.org/record/278397/files/figure.png" pageId="12" pageNumber="13">Figure 4A</figureCitation>
); 
<emphasis id="A3E8E71E8B07FFA48DAAFAE2FA9AF023" box="[1213,1339,1328,1352]" italics="true" pageId="12" pageNumber="13">G. l i b o e n s i s</emphasis>
Zaho &amp; Li, 1982; 
<taxonomicName id="569C408F8B07FFA48812FA87FE41F007" authority="Vogt, 1922" authorityName="Vogt" authorityYear="1922" box="[261,480,1364,1389]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="melli">
<emphasis id="A3E8E71E8B07FFA48812FA87FEFDF007" box="[261,348,1364,1388]" italics="true" pageId="12" pageNumber="13">G. melli</emphasis>
<bibRefCitation id="F50D46FD8B07FFA48874FA87FE41F007" author="Vogt" box="[355,480,1364,1389]" pageId="12" pageNumber="13" refString="Vogt, T. (1922) Zur Reptilien- und Amphibienfauna Sudchinas. Archiv fur Naturgeschichte, 10, 135 - 146." type="journal article" year="1922">Vogt, 1922</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="569C408F8B07FFA488FBFA87FC92F007" authority="Boulenger, 1907" authorityName="Boulenger" authorityYear="1907" box="[492,819,1364,1388]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="palmatus">
<emphasis id="A3E8E71E8B07FFA488FBFA87FDD3F007" box="[492,626,1364,1388]" italics="true" pageId="12" pageNumber="13">G. palmatus</emphasis>
<bibRefCitation id="F50D46FD8B07FFA48B6EFA86FC92F007" author="Boulenger" box="[633,819,1364,1388]" pageId="12" pageNumber="13" refString="Boulenger, G. A. (1907) Description of new lizard in the British Museum. Annals and Magazine of Natural History, 19, 486 - 489." type="journal article" year="1907">Boulenger, 1907</bibRefCitation>
</taxonomicName>
(
<figureCitation id="09A727898B07FFA48A56FA86FC02F007" box="[833,931,1364,1388]" captionStart="FIGURE 4. A" captionStartId="14.[151,250,1008,1031]" captionTargetBox="[151,1436,193,986]" captionTargetId="figure@14.[151,1436,193,987]" captionTargetPageId="14" captionText="FIGURE 4. A. Female Gekko japonicus (G. japonicus group), photo by J. Boone; B. Male G. hokouensis (G. japonicus group), photo by J. Boone; C. G. palmatus (G. japonicus group), Quang Binh Province, Vietnam, photo by T. Ziegler; D. G. scientiadventura (G. japonicus group), Quang Binh Province, Vietnam, photo by T. Ziegler." httpUri="https://zenodo.org/record/278397/files/figure.png" pageId="12" pageNumber="13">Figure 4</figureCitation>
C); 
<taxonomicName id="569C408F8B07FFA48ADBFA87FA8FF007" authority="Liu &amp; Zhou, 1982" authorityName="Liu &amp; Zhou" authorityYear="1982" box="[972,1326,1364,1388]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="scabridus">
<emphasis id="A3E8E71E8B07FFA48ADBFA87FBF9F007" box="[972,1112,1364,1388]" italics="true" pageId="12" pageNumber="13">G. scabridus</emphasis>
Liu &amp; Zhou, 1982
</taxonomicName>
; 
<emphasis id="A3E8E71E8B07FFA48C2DFA87FEB9F0FB" italics="true" pageId="12" pageNumber="13">G. s c i e n - tiadventura</emphasis>
Rösler, Ziegler, Vu, Herrmann &amp; Böhme, 2004 (
<figureCitation id="09A727898B07FFA48A2CFAA5FC3DF0E4" box="[827,924,1399,1423]" captionStart="FIGURE 4. A" captionStartId="14.[151,250,1008,1031]" captionTargetBox="[151,1436,193,986]" captionTargetId="figure@14.[151,1436,193,987]" captionTargetPageId="14" captionText="FIGURE 4. A. Female Gekko japonicus (G. japonicus group), photo by J. Boone; B. Male G. hokouensis (G. japonicus group), photo by J. Boone; C. G. palmatus (G. japonicus group), Quang Binh Province, Vietnam, photo by T. Ziegler; D. G. scientiadventura (G. japonicus group), Quang Binh Province, Vietnam, photo by T. Ziegler." httpUri="https://zenodo.org/record/278397/files/figure.png" pageId="12" pageNumber="13">Figure 4</figureCitation>
D); 
<taxonomicName id="569C408F8B07FFA48AD0FAAAFF72F0DF" authority="Toda, Sengoku, Hikida &amp; Ota, 2008" authorityName="Toda, Sengoku, Hikida &amp; Ota" authorityYear="2008" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="shibatai">
<emphasis id="A3E8E71E8B07FFA48AD0FAAAFBE1F0FB" box="[967,1088,1400,1424]" italics="true" pageId="12" pageNumber="13">G. shibatai</emphasis>
<bibRefCitation id="F50D46FD8B07FFA48D50FAA5FF72F0DF" author="Toda" pageId="12" pageNumber="13" refString="Toda, M., Sengoku, S., Hikida, T. &amp; Ota, H. (2008) Description of two new species of the genus Gekko (Squamata: Gekkonidae) from the Tokara and Amami Island Group in the Ryukyu Archipelago, Japan. Copeia, 2008, 452 - 466." type="journal article" year="2008">Toda, Sengoku, Hikida &amp; Ota, 2008</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="569C408F8B07FFA489CAFA4FFDAAF0DF" authority="Smith, 1923" authorityName="Smith" authorityYear="1923" box="[221,523,1436,1460]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="similignum">
<emphasis id="A3E8E71E8B07FFA489CAFA4FFED9F0DF" box="[221,376,1436,1460]" italics="true" pageId="12" pageNumber="13">G. similignum</emphasis>
<bibRefCitation id="F50D46FD8B07FFA48868FA4EFDAAF0DF" author="Smith" box="[383,523,1436,1460]" pageId="12" pageNumber="13" refString="Smith, M. A. (1923) On a collection of reptiles and batrachians from the island of Hainan. Journal of the Natural History Society of Siam, 6, 195 - 212." type="journal article" year="1923">Smith, 1923</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="569C408F8B07FFA48B01FA4FFCCCF0DF" authority="Gunther, 1864" authorityName="Gunther" authorityYear="1864" box="[534,877,1436,1460]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="subpalmatus">
<emphasis id="A3E8E71E8B07FFA48B01FA4FFD62F0DF" box="[534,707,1436,1460]" italics="true" pageId="12" pageNumber="13">G. subpalmatus</emphasis>
<bibRefCitation id="F50D46FD8B07FFA48BDEFA4EFCCCF0DF" author="Gunther" box="[713,877,1436,1460]" pageId="12" pageNumber="13" refString="Gunther, A. (1864) The Reptiles of British India. The Ray Society London, xxvii + 452 pp., 26 pls." type="book" year="1864">Günther, 1864</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="569C408F8B07FFA48A6FFA4FFB10F0DF" authority="Gunther, 1864" authorityName="Gunther" authorityYear="1864" box="[888,1201,1436,1460]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="swinhonis">
<emphasis id="A3E8E71E8B07FFA48A6FFA4FFBA6F0DF" box="[888,1031,1436,1460]" italics="true" pageId="12" pageNumber="13">G. swinhonis</emphasis>
<bibRefCitation id="F50D46FD8B07FFA48D19FA4EFB10F0DF" author="Gunther" box="[1038,1201,1436,1460]" pageId="12" pageNumber="13" refString="Gunther, A. (1864) The Reptiles of British India. The Ray Society London, xxvii + 452 pp., 26 pls." type="book" year="1864">Günther, 1864</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="569C408F8B07FFA48DAAFA4FFF73F0BC" authority="Song, 1985" authorityName="Song" authorityYear="1985" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="taibaiensis">
<emphasis id="A3E8E71E8B07FFA48DAAFA4FFAF5F0DF" box="[1213,1364,1436,1460]" italics="true" pageId="12" pageNumber="13">G. taibaiensis</emphasis>
<bibRefCitation id="F50D46FD8B07FFA48C4CFA4EFF73F0BC" author="Song" pageId="12" pageNumber="13" refString="Song, M. (1985) A new species of Gekko from Shaanxi. Acta Herpetologica Sinica, 4, 329 - 330." type="journal article" year="1985">Song, 1985</bibRefCitation>
</taxonomicName>
; 
<emphasis id="A3E8E71E8B07FFA489C9FA12FEC9F0B3" box="[222,360,1472,1496]" italics="true" pageId="12" pageNumber="13">G. t a w a e n s i s</emphasis>
<bibRefCitation id="F50D46FD8B07FFA48863FA6DFDA6F0BC" author="Okada" box="[372,519,1471,1495]" pageId="12" pageNumber="13" refString="Okada, Y. (1956) A new species of gekko from Shikoku, Japan. Annotationes Zoologicae Japonensis, 29, 239 - 241." type="journal article" year="1956">Okada, 1956</bibRefCitation>
; 
<taxonomicName id="569C408F8B07FFA48B04FA12FBEAF0BC" authority="Toda, Sengoku, Hikida &amp; Ota, 2008" authorityName="Toda, Sengoku, Hikida &amp; Ota" authorityYear="2008" box="[531,1099,1471,1496]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="vertebralis">
<emphasis id="A3E8E71E8B07FFA48B04FA12FD0AF0B3" box="[531,683,1472,1496]" italics="true" pageId="12" pageNumber="13">G. vertebralis</emphasis>
<bibRefCitation id="F50D46FD8B07FFA48BA4FA6DFBEAF0BC" author="Toda" box="[691,1099,1471,1495]" pageId="12" pageNumber="13" refString="Toda, M., Sengoku, S., Hikida, T. &amp; Ota, H. (2008) Description of two new species of the genus Gekko (Squamata: Gekkonidae) from the Tokara and Amami Island Group in the Ryukyu Archipelago, Japan. Copeia, 2008, 452 - 466." type="journal article" year="2008">Toda, Sengoku, Hikida &amp; Ota, 2008</bibRefCitation>
</taxonomicName>
(
<figureCitation id="09A727898B07FFA48D4DFA6DFB73F0BC" box="[1114,1234,1471,1495]" captionStart="FIGURE 5. A" captionStartId="14.[151,250,1944,1967]" captionTargetBox="[151,1436,1129,1922]" captionTargetId="figure@14.[151,1436,1129,1923]" captionTargetPageId="14" captionText="FIGURE 5. A. Male Gekko vertebralis (G. japonicus group), photo by J. Boone; B. G. monarchus (G. monarchus group), Banggai Islands, Indonesia, photo by A. Koch; C. G. ernstkelleri (G. monarchus group), Panay Island, Philippines, photo by M. Gaulke; D. G. mindorensis (G. monarchus group), Cebu Island, Philippines, photo by M. Gaulke." httpUri="https://zenodo.org/record/278398/files/figure.png" pageId="12" pageNumber="13">Figure 5A</figureCitation>
); 
<taxonomicName id="569C408F8B07FFA48DF0FA12FE21F097" authority="Zhou &amp; Wang, 2008" authorityName="Zhou &amp; Wang" authorityYear="2008" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="wenxianensis">
<emphasis id="A3E8E71E8B07FFA48DF0FA12FA3DF0B3" box="[1255,1436,1472,1496]" italics="true" pageId="12" pageNumber="13">G. wenxianensis</emphasis>
<bibRefCitation id="F50D46FD8B07FFA48980FA36FE21F097" author="Zhou" box="[151,384,1508,1533]" pageId="12" pageNumber="13" refString="Zhou, K. &amp; Wang, Q. (2008) New species of Gekko (Squamata: Sauria: Gekkonidae) from China: morphological and molecular evidence. Zootaxa, 1778, 59 - 68." type="journal article" year="2008">Zhou &amp; Wang, 2008</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="569C408F8B07FFA4889CFA37FC80F097" authority="Matsui &amp; Okada, 1968" authorityName="Matsui &amp; Okada" authorityYear="1968" box="[395,801,1508,1532]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="yakuensis">
<emphasis id="A3E8E71E8B07FFA4889CFA37FDB7F097" box="[395,534,1508,1532]" italics="true" pageId="12" pageNumber="13">G. yakuensis</emphasis>
<bibRefCitation id="F50D46FD8B07FFA48B0AFA36FC80F097" author="Matsui" box="[541,801,1508,1532]" pageId="12" pageNumber="13" refString="Matsui, T. &amp; Okada, Y. (1968) A new species of Gekko found in Yakushima, one of the small island south of Kyushu. Acta Herpetologica Japonica, 3, 1 - 4." type="journal article" year="1968">Matsui &amp; Okada, 1968</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="91233B0C8B07FFA489D2F9D5FC8AF277" blockId="12.[151,1436,1291,2036]" pageId="12" pageNumber="13">
<emphasis id="A3E8E71E8B07FFA489D2F9D5FEF3F34B" bold="true" box="[197,338,1543,1568]" pageId="12" pageNumber="13">Characters.</emphasis>
<quantity id="566496E98B07FFA4884FF9D5FE5BF34B" box="[344,506,1543,1568]" metricMagnitude="-2" metricUnit="m" metricValue="7.904999999999999" metricValueMax="9.92" metricValueMin="5.89" pageId="12" pageNumber="13" unit="mm" value="79.05" valueMax="99.2" valueMin="58.9">58.9–99.2 mm</quantity>
SVL; nares in contact with rostral (except for 
<taxonomicName id="569C408F8B07FFA48AE9F9DAFB67F34B" box="[1022,1222,1544,1568]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="auriverrucosus">
<emphasis id="A3E8E71E8B07FFA48AE9F9DAFB67F34B" box="[1022,1222,1544,1568]" italics="true" pageId="12" pageNumber="13">G. auriverrucosus</emphasis>
</taxonomicName>
); nasals 3 (rarely 
<quantity id="566496E98B07FFA48C99F9D5FF0CF32F" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="12" pageNumber="13" unit="in" value="2.0">2 in</quantity>
<emphasis id="A3E8E71E8B07FFA489A3F9FFFE97F32F" box="[180,310,1580,1604]" italics="true" pageId="12" pageNumber="13">G. c h i n e n s i s</emphasis>
); postmentals relatively small (e.g., 
<taxonomicName id="569C408F8B07FFA48BC5F9FFFCCCF32F" box="[722,877,1580,1604]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="similignum">
<emphasis id="A3E8E71E8B07FFA48BC5F9FFFCCCF32F" box="[722,877,1580,1604]" italics="true" pageId="12" pageNumber="13">G. similignum</emphasis>
</taxonomicName>
), largest in 
<emphasis id="A3E8E71E8B07FFA48AE5F9FFFBEAF32F" box="[1010,1099,1580,1604]" italics="true" pageId="12" pageNumber="13">G. c a n h i</emphasis>
, 
<emphasis id="A3E8E71E8B07FFA48D4AF9FFFB41F32F" box="[1117,1248,1580,1604]" italics="true" pageId="12" pageNumber="13">G. c h i n e n s i s</emphasis>
, 
<taxonomicName id="569C408F8B07FFA48DE6F9FFFAD6F32F" box="[1265,1399,1580,1604]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="palmatus">
<emphasis id="A3E8E71E8B07FFA48DE6F9FFFAD6F32F" box="[1265,1399,1580,1604]" italics="true" pageId="12" pageNumber="13">G. palmatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="569C408F8B07FFA48C93F9FFFEF3F303" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="scientiadventura">
<emphasis id="A3E8E71E8B07FFA48C93F9FFFEF3F303" italics="true" pageId="12" pageNumber="13">G. scientiadventura</emphasis>
</taxonomicName>
; 0–21 dorsal tubercle rows; 0–32 precloacal pores; postcloacal tubercles 1–4; webbing between fingers and toes weakly developed to extensive (
<taxonomicName id="569C408F8B07FFA48BA1F9A7FC9DF3E7" box="[694,828,1652,1676]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="chinensis">
<emphasis id="A3E8E71E8B07FFA48BA1F9A7FC9DF3E7" box="[694,828,1652,1676]" italics="true" pageId="12" pageNumber="13">G. chinensis</emphasis>
</taxonomicName>
, 
<emphasis id="A3E8E71E8B07FFA48A5DF9A7FC3CF3E7" box="[842,925,1652,1676]" italics="true" pageId="12" pageNumber="13">G. m e l l i</emphasis>
, 
<taxonomicName id="569C408F8B07FFA48AB9F9A7FB95F3E7" box="[942,1076,1652,1676]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="palmatus">
<emphasis id="A3E8E71E8B07FFA48AB9F9A7FB95F3E7" box="[942,1076,1652,1676]" italics="true" pageId="12" pageNumber="13">G. palmatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="569C408F8B07FFA48D56F9A7FABBF3E7" box="[1089,1306,1652,1676]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="scientiadventura">
<emphasis id="A3E8E71E8B07FFA48D56F9A7FABBF3E7" box="[1089,1306,1652,1676]" italics="true" pageId="12" pageNumber="13">G. scientiadventura</emphasis>
</taxonomicName>
, 
<taxonomicName id="569C408F8B07FFA48C3FF9A7FF66F3DB" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="similignum">
<emphasis id="A3E8E71E8B07FFA48C3FF9A7FF66F3DB" italics="true" pageId="12" pageNumber="13">G. similignum</emphasis>
</taxonomicName>
, 
<taxonomicName id="569C408F8B07FFA489C3F94AFE21F3DB" box="[212,384,1688,1712]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="subpalmatus">
<emphasis id="A3E8E71E8B07FFA489C3F94AFE21F3DB" box="[212,384,1688,1712]" italics="true" pageId="12" pageNumber="13">G. subpalmatus</emphasis>
</taxonomicName>
); tubercles present on fore and hind limbs, hind limbs only, or lacking all together; lateral fold without tubercles; subcaudals enlarged, in a longitudinal row (in 
<taxonomicName id="569C408F8B07FFA48A6BF96FFBA8F3BF" box="[892,1033,1724,1748]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="yakuensis">
<emphasis id="A3E8E71E8B07FFA48A6BF96FFBA8F3BF" box="[892,1033,1724,1748]" italics="true" pageId="12" pageNumber="13">G. yakuensis</emphasis>
</taxonomicName>
medially subdivided); head pattern present or not, without figure-shape (UU- to W-shaped in 
<emphasis id="A3E8E71E8B07FFA48A33F932FCD6F393" box="[804,887,1760,1784]" italics="true" pageId="12" pageNumber="13">G. m e l l i</emphasis>
and W-shaped in 
<taxonomicName id="569C408F8B07FFA48D51F932FA81F393" box="[1094,1312,1760,1784]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="scientiadventura">
<emphasis id="A3E8E71E8B07FFA48D51F932FA81F393" box="[1094,1312,1760,1784]" italics="true" pageId="12" pageNumber="13">G. scientiadventura</emphasis>
</taxonomicName>
); vertebral region with relatively large, light flecks, blotches or bands.
</paragraph>
</subSubSection>
<subSubSection id="D98668878B07FFA489D2F8F5FF68F20F" pageId="12" pageNumber="13" type="distribution">
<paragraph id="91233B0C8B07FFA489D2F8F5FF68F20F" blockId="12.[151,1436,1291,2036]" pageId="12" pageNumber="13">
<emphasis id="A3E8E71E8B07FFA489D2F8F5FEFFF22B" bold="true" box="[197,350,1831,1856]" pageId="12" pageNumber="13">Distribution.</emphasis>
<collectingCountry id="E98B7B9C8B07FFA4887EF8FAFE0EF22B" box="[361,431,1832,1856]" name="China" pageId="12" pageNumber="13">China</collectingCountry>
, 
<collectingCountry id="E98B7B9C8B07FFA488AAF8FBFDA1F22B" box="[445,512,1833,1856]" name="Japan" pageId="12" pageNumber="13">Japan</collectingCountry>
, 
<collectingCountry id="E98B7B9C8B07FFA48B18F8FBFDF6F22B" box="[527,599,1833,1856]" name="South Korea" pageId="12" pageNumber="13">Korea</collectingCountry>
, 
<collectingCountry id="E98B7B9C8B07FFA48B72F8FAFD1BF22B" box="[613,698,1832,1856]" name="Taiwan" pageId="12" pageNumber="13">Taiwan</collectingCountry>
, and 
<collectingCountry id="E98B7B9C8B07FFA48BEBF8FAFCFAF22B" box="[764,859,1832,1856]" name="Vietnam" pageId="12" pageNumber="13">Vietnam</collectingCountry>
including offshore islands; possibly 
<collectingCountry id="E98B7B9C8B07FFA48C1EF8FBFA9EF22B" box="[1289,1343,1833,1856]" name="Laos" pageId="12" pageNumber="13">Laos</collectingCountry>
(
<figureCitation id="09A727898B07FFA48C46F8FAFF04F20F" captionStart="FIGURE 3" captionStartId="13.[151,250,1859,1882]" captionTargetBox="[151,1436,193,1838]" captionTargetId="figure@13.[151,1436,193,1838]" captionTargetPageId="13" captionText="FIGURE 3. Map of East and Southeast Asia and the Indo-Australian Archipelago showing the approximate distribution of (A) the Gekko japonicus, G. petricolus, and G. monarchus groups and G. a t h y m u s and (B) the G. g e c k o, G. p o ro s u s, and G. vittatus groups and G. vietnamensis." httpUri="https://zenodo.org/record/278396/files/figure.png" pageId="12" pageNumber="13">Figure 3</figureCitation>
A).
</paragraph>
</subSubSection>
<subSubSection id="D98668878B07FFA789D2F8A2FF5BF0F7" lastPageId="15" lastPageNumber="16" pageId="12" pageNumber="13" type="discussion">
<paragraph id="91233B0C8B07FFA489D2F8A2FCA5F29F" blockId="12.[151,1436,1291,2036]" pageId="12" pageNumber="13">
<emphasis id="A3E8E71E8B07FFA489D2F8A2FE98F2E3" bold="true" box="[197,313,1904,1928]" pageId="12" pageNumber="13">Remarks.</emphasis>
The 
<taxonomicName id="569C408F8B07FFA48865F8A2FD8FF2E3" box="[370,558,1904,1928]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="japonicus">
<emphasis id="A3E8E71E8B07FFA48865F8A2FD8FF2E3" box="[370,558,1904,1928]" italics="true" pageId="12" pageNumber="13">Gekko japonicus</emphasis>
</taxonomicName>
group is very complex. Different morphological features of the species of this group (e. g., nares in contact with rostral or not, broad webbing between fingers and toes or not, dorsal tubercles absent or present, tubercles on limbs absent or present, see 
<tableCitation id="DC1E0EB78B07FFA48A22F86AFC28F2BB" box="[821,905,1976,2000]" captionStart="TABLE 2" captionStartId="10.[151,235,752,773]" captionText="TABLE 2. Gekko species group characters. 1 = SVL (&lt;115 mm = 0,&gt; 115 mm = 1); 2 = nostril in contact with rostral but not first supralabial (no = 0, yes = 1); 3 = nasals (three or less = 0, three or more = 1); 4 = postmentals (small = 0, large = 1); 5 = dorsal tubercles (available = 0, missing = 1); 6 = precloacal pores (24 or less = 0, 32 or more = 1); 7 = postcloacal tubercles (one = 0, one or more = 1); 8 = toes webbed (no = 0, yes = 1); 9 = tubercles on limbs (present = 0, absent = 1); 10 = tubercles on lateral fold (absent = 0, present = 1); 11 = subcaudals (enlarged = 0, not enlarged = 1)." pageId="12" pageNumber="13">Table 2</tableCitation>
) demonstrate that this genetically well-defined clade is highly morphologically variable (see 
<figureCitation id="09A727898B07FFA48B81F80EFD52F29F" box="[662,755,2012,2036]" captionStart="FIGURE 1" captionStartId="4.[151,250,1550,1573]" captionTargetBox="[152,1436,448,1524]" captionTargetId="figure@4.[151,1436,447,1528]" captionTargetPageId="4" captionText="FIGURE 1. Bayesian phylogeny of Gekko, based on nucleotide sequences of the mitochondrial ND 2 gene and nuclear RAG- 1 and PDC genes (2881 bp). Support values (Bayesian pp / ML bootstrap / MP bootstrap) are given at nodes. Species groups proposed herein are indicated." httpUri="https://zenodo.org/record/278394/files/figure.png" pageId="12" pageNumber="13">Figure 1</figureCitation>
).
</paragraph>
<caption id="C5E36B848B06FFA58980F891FE10F2FC" httpUri="https://zenodo.org/record/278396/files/figure.png" pageId="13" pageNumber="14" targetBox="[151,1436,193,1838]" targetPageId="13">
<paragraph id="91233B0C8B06FFA58980F891FE10F2FC" blockId="13.[151,1435,1859,1943]" pageId="13" pageNumber="14">
<emphasis id="A3E8E71E8B06FFA58980F891FEADF232" bold="true" box="[151,268,1859,1882]" pageId="13" pageNumber="14">FIGURE 3</emphasis>
. Map of East and Southeast Asia and the Indo-Australian Archipelago showing the approximate distribution of (A) the 
<taxonomicName id="569C408F8B06FFA589AAF8B1FEC6F212" box="[189,359,1891,1913]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="japonicus">
<emphasis id="A3E8E71E8B06FFA589AAF8B1FEC6F212" box="[189,359,1891,1913]" italics="true" pageId="13" pageNumber="14">Gekko japonicus</emphasis>
</taxonomicName>
, 
<taxonomicName id="569C408F8B06FFA58863F8B1FE57F212" box="[372,502,1891,1913]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="petricolus">
<emphasis id="A3E8E71E8B06FFA58863F8B1FE57F212" box="[372,502,1891,1913]" italics="true" pageId="13" pageNumber="14">G. petricolus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="569C408F8B06FFA58B38F8B1FD1BF212" box="[559,698,1891,1913]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="monarchus">
<emphasis id="A3E8E71E8B06FFA58B38F8B1FD1BF212" box="[559,698,1891,1913]" italics="true" pageId="13" pageNumber="14">G. monarchus</emphasis>
</taxonomicName>
groups and 
<emphasis id="A3E8E71E8B06FFA58A2EF8B1FC06F212" box="[825,935,1891,1913]" italics="true" pageId="13" pageNumber="14">G. a t h y m u s</emphasis>
and (B) the 
<emphasis id="A3E8E71E8B06FFA58D3BF8B1FBDEF212" box="[1068,1151,1891,1913]" italics="true" pageId="13" pageNumber="14">G. g e c k o</emphasis>
, 
<emphasis id="A3E8E71E8B06FFA58D87F8B1FB58F212" box="[1168,1273,1891,1913]" italics="true" pageId="13" pageNumber="14">G. p o ro s u s</emphasis>
, and 
<taxonomicName id="569C408F8B06FFA58C22F8B1FA3AF212" box="[1333,1435,1891,1913]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="vittatus">
<emphasis id="A3E8E71E8B06FFA58C22F8B1FA3AF212" box="[1333,1435,1891,1913]" italics="true" pageId="13" pageNumber="14">G. vittatus</emphasis>
</taxonomicName>
groups and 
<taxonomicName id="569C408F8B06FFA5881AF850FE0DF2FC" box="[269,428,1922,1943]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="vietnamensis">
<emphasis id="A3E8E71E8B06FFA5881AF850FE0DF2FC" box="[269,428,1922,1943]" italics="true" pageId="13" pageNumber="14">G. vietnamensis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="C5E36B848B05FFA68980FC22FC7BF128" httpUri="https://zenodo.org/record/278397/files/figure.png" pageId="14" pageNumber="15" targetBox="[151,1436,193,986]" targetPageId="14">
<paragraph id="91233B0C8B05FFA68980FC22FC7BF128" blockId="14.[151,1436,1008,1091]" pageId="14" pageNumber="15">
<emphasis id="A3E8E71E8B05FFA68980FC22FEADF16E" bold="true" box="[151,268,1008,1031]" pageId="14" pageNumber="15">FIGURE 4</emphasis>
. A. Female 
<taxonomicName id="569C408F8B05FFA6889EFC22FD93F16D" box="[393,562,1008,1030]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="japonicus">
<emphasis id="A3E8E71E8B05FFA6889EFC22FD93F16D" box="[393,562,1008,1030]" italics="true" pageId="14" pageNumber="15">Gekko japonicus</emphasis>
</taxonomicName>
(
<taxonomicName id="569C408F8B05FFA68B56FC23FD1EF16D" box="[577,703,1009,1030]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="japonicus">
<emphasis id="A3E8E71E8B05FFA68B56FC23FD1EF16D" box="[577,703,1009,1030]" italics="true" pageId="14" pageNumber="15">G. japonicus</emphasis>
</taxonomicName>
group), photo by J. Boone; B. Male 
<taxonomicName id="569C408F8B05FFA68D24FC23FB1EF16D" box="[1075,1215,1008,1030]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="hokouensis">
<emphasis id="A3E8E71E8B05FFA68D24FC23FB1EF16D" box="[1075,1215,1008,1030]" italics="true" pageId="14" pageNumber="15">G. hokouensis</emphasis>
</taxonomicName>
(
<taxonomicName id="569C408F8B05FFA68DDAFC23FAEAF16D" box="[1229,1355,1009,1030]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="japonicus">
<emphasis id="A3E8E71E8B05FFA68DDAFC23FAEAF16D" box="[1229,1355,1009,1030]" italics="true" pageId="14" pageNumber="15">G. japonicus</emphasis>
</taxonomicName>
group), photo by J. Boone; C. 
<taxonomicName id="569C408F8B05FFA6886AFBDDFE56F14E" box="[381,503,1039,1061]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="palmatus">
<emphasis id="A3E8E71E8B05FFA6886AFBDDFE56F14E" box="[381,503,1039,1061]" italics="true" pageId="14" pageNumber="15">G. palmatus</emphasis>
</taxonomicName>
(
<taxonomicName id="569C408F8B05FFA68B11FBDDFD24F14E" box="[518,645,1039,1061]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="japonicus">
<emphasis id="A3E8E71E8B05FFA68B11FBDDFD24F14E" box="[518,645,1039,1061]" italics="true" pageId="14" pageNumber="15">G. japonicus</emphasis>
</taxonomicName>
group), Quang Binh Province, Vietnam, photo by T. Ziegler; D. 
<taxonomicName id="569C408F8B05FFA68C0BFBDDFF45F128" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="scientiadventura">
<emphasis id="A3E8E71E8B05FFA68C0BFBDDFF45F128" italics="true" pageId="14" pageNumber="15">G. scientiadventura</emphasis>
</taxonomicName>
(
<taxonomicName id="569C408F8B05FFA689E4FBFCFED0F128" box="[243,369,1070,1091]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="japonicus">
<emphasis id="A3E8E71E8B05FFA689E4FBFCFED0F128" box="[243,369,1070,1091]" italics="true" pageId="14" pageNumber="15">G. japonicus</emphasis>
</taxonomicName>
group), Quang Binh Province, Vietnam, photo by T. Ziegler.
</paragraph>
</caption>
<caption id="C5E36B848B05FFA68980F84AFBCAF280" httpUri="https://zenodo.org/record/278398/files/figure.png" pageId="14" pageNumber="15" targetBox="[151,1436,1129,1922]" targetPageId="14">
<paragraph id="91233B0C8B05FFA68980F84AFBCAF280" blockId="14.[151,1436,1944,2028]" pageId="14" pageNumber="15">
<emphasis id="A3E8E71E8B05FFA68980F84AFEAAF2C5" bold="true" box="[151,267,1944,1967]" pageId="14" pageNumber="15">FIGURE 5</emphasis>
. A. Male 
<taxonomicName id="569C408F8B05FFA68867F84AFD85F2C5" box="[368,548,1944,1966]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="vertebralis">
<emphasis id="A3E8E71E8B05FFA68867F84AFD85F2C5" box="[368,548,1944,1966]" italics="true" pageId="14" pageNumber="15">Gekko vertebralis</emphasis>
</taxonomicName>
(
<taxonomicName id="569C408F8B05FFA68B25F84BFD0EF2C4" box="[562,687,1945,1967]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="japonicus">
<emphasis id="A3E8E71E8B05FFA68B25F84BFD0EF2C4" box="[562,687,1945,1967]" italics="true" pageId="14" pageNumber="15">G. japonicus</emphasis>
</taxonomicName>
group), photo by J. Boone; B. 
<taxonomicName id="569C408F8B05FFA68AF2F84BFBD1F2C5" box="[997,1136,1944,1966]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="monarchus">
<emphasis id="A3E8E71E8B05FFA68AF2F84BFBD1F2C5" box="[997,1136,1944,1966]" italics="true" pageId="14" pageNumber="15">G. monarchus</emphasis>
</taxonomicName>
(
<taxonomicName id="569C408F8B05FFA68D69F84BFAA9F2C5" box="[1150,1288,1944,1966]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="monarchus">
<emphasis id="A3E8E71E8B05FFA68D69F84BFAA9F2C5" box="[1150,1288,1944,1966]" italics="true" pageId="14" pageNumber="15">G. monarchus</emphasis>
</taxonomicName>
group), Banggai Islands, Indonesia, photo by A. Koch; C. 
<taxonomicName id="569C408F8B05FFA68B61F86AFCA8F2A6" box="[630,777,1975,1997]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="ernstkelleri">
<emphasis id="A3E8E71E8B05FFA68B61F86AFCA8F2A6" box="[630,777,1975,1997]" italics="true" pageId="14" pageNumber="15">G. ernstkelleri</emphasis>
</taxonomicName>
(
<taxonomicName id="569C408F8B05FFA68A0DF86AFC06F2A6" box="[794,935,1975,1997]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="monarchus">
<emphasis id="A3E8E71E8B05FFA68A0DF86AFC06F2A6" box="[794,935,1975,1997]" italics="true" pageId="14" pageNumber="15">G. monarchus</emphasis>
</taxonomicName>
group), Panay Island, Philippines, photo by M. Gaulke; D. 
<taxonomicName id="569C408F8B05FFA6881BF804FE02F287" box="[268,419,2006,2028]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="mindorensis">
<emphasis id="A3E8E71E8B05FFA6881BF804FE02F287" box="[268,419,2006,2028]" italics="true" pageId="14" pageNumber="15">G. mindorensis</emphasis>
</taxonomicName>
(
<taxonomicName id="569C408F8B05FFA688A6F804FD9CF287" box="[433,573,2006,2028]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="monarchus">
<emphasis id="A3E8E71E8B05FFA688A6F804FD9CF287" box="[433,573,2006,2028]" italics="true" pageId="14" pageNumber="15">G. monarchus</emphasis>
</taxonomicName>
group), Cebu Island, Philippines, photo by M. Gaulke.
</paragraph>
</caption>
<paragraph id="91233B0C8B04FFA789D2FF4AFD33F652" blockId="15.[151,1437,151,2012]" pageId="15" pageNumber="16">
Numerous authors have listed 
<taxonomicName id="569C408F8B04FFA78B31FF4AFD41F5DB" box="[550,736,152,176]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="chinensis">
<emphasis id="A3E8E71E8B04FFA78B31FF4AFD41F5DB" box="[550,736,152,176]" italics="true" pageId="15" pageNumber="16">Gekko chinensis</emphasis>
</taxonomicName>
as a representative of Vietnam’s herpetofauna (e.g., 
<bibRefCitation id="F50D46FD8B04FFA78C51FF4AFF41F5BF" author="Bourret" pageId="15" pageNumber="16" refString="Bourret, R. (1937 b) Notes herpetologiques sur l'Indochine francaise. XV. Lezards et Serpents recus au Laboratoire des Sciences Naturelles de l'Universite au cours de l'annee 1937. Descriptions de deux especes et de deux varietes nouvelles. Bulletin general de l'Instruction publique, 17 (4), annexe, 57 - 80." type="journal article" year="1937" yearSuffix="b">Bourret 1937b</bibRefCitation>
, 
<bibRefCitation id="F50D46FD8B04FFA789FCFF6EFE93F5BF" author="Bourret" box="[235,306,188,212]" pageId="15" pageNumber="16" refString="Bourret, R. (1940 a &quot; 1939 &quot;) Notes herpetologiques sur l'Indochine francaise. XVIII. Reptiles et Batraciens recus au Laboratoire des Sciences Naturelles de l'Universite au cours de l'annee 1939. Descriptions de quatre especes et d'une variete nouvelles. Bulletin general de l'Instruction publique, 18 (4), annexe, 5 - 39, 1 pl." type="journal article" year="1940" yearSuffix="a">1940a</bibRefCitation>
, 
<bibRefCitation id="F50D46FD8B04FFA7882AFF6EFED9F5BF" author="Bourret" box="[317,376,188,212]" pageId="15" pageNumber="16" refString="Bourret, R. (1943) Notes herpetologiques sur l'Indochine francaise. XXIV. Reptiles et Batraciens recus au Laboratoire des Sciences Naturelles de l'Universite au cours de l'annee 1942. Notes et travaux de l'Ecole Superieure des sciences de l'Universite Indochinoise, 2, 1 - 17, 2 pls." type="journal article" year="1943">1943</bibRefCitation>
; 
<bibRefCitation id="F50D46FD8B04FFA78893FF6EFDC3F5BF" author="Zhao" box="[388,610,188,212]" pageId="15" pageNumber="16" refString="Zhao, E. &amp; Adler, K. (1993) Herpetology of China. Society for the Study of Amphibians and Reptiles, Oxford, Ohio, 522 pp., 48 pls., 1 foldout map." type="book" year="1993">Zhao &amp; Adler 1993</bibRefCitation>
; 
<bibRefCitation id="F50D46FD8B04FFA78B79FF6EFD56F5BF" author="Welch" box="[622,759,188,212]" pageId="15" pageNumber="16" refString="Welch, K. R. G. (1994) Lizards of the World. 1. Geckos. R &amp; A Research and Information Limited, Taunton, 165 pp." type="book" year="1994">Welch 1994</bibRefCitation>
; 
<bibRefCitation id="F50D46FD8B04FFA78A15FF6EFC04F5BF" author="Bobrov" box="[770,933,188,212]" pageId="15" pageNumber="16" refString="Bobrov, V. V. (1995 a) Checklist and bibliography of the Lizards of Vietnam. Smithsonian Herpetological Information Service, 105, 1 - 28." type="book chapter" year="1995" yearSuffix="a">Bobrov 1995a</bibRefCitation>
; Rösler 
<emphasis id="A3E8E71E8B04FFA78D17FF6FFB98F5BF" box="[1024,1081,188,212]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
2005). According to 
<bibRefCitation id="F50D46FD8B04FFA78C3FFF6FFEA9F59C" pageId="15" pageNumber="16" refString="Nguyen, V. S., Ho, T. C. &amp; Nguyen, Q. T. (2005) A Checklist of Amphibians and Reptiles of Vietnam [in Vietnamese]. Agriculture Publishing House, Hanoi, 180 pp." type="book">
Nguyen 
<emphasis id="A3E8E71E8B04FFA78C9FFF6FFF15F593" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
(2005)
</bibRefCitation>
, 
<taxonomicName id="569C408F8B04FFA78804FF32FE38F593" box="[275,409,224,248]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="chinensis">
<emphasis id="A3E8E71E8B04FFA78804FF32FE38F593" box="[275,409,224,248]" italics="true" pageId="15" pageNumber="16">G. chinensis</emphasis>
</taxonomicName>
is widely distributed in the north of the country, whereas 
<taxonomicName id="569C408F8B04FFA78D09FF32FB05F593" box="[1054,1188,224,248]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="palmatus">
<emphasis id="A3E8E71E8B04FFA78D09FF32FB05F593" box="[1054,1188,224,248]" italics="true" pageId="15" pageNumber="16">G. palmatus</emphasis>
</taxonomicName>
is only recorded from the two northern provinces of Lang Son and Vinh Phuc (see also 
<bibRefCitation id="F50D46FD8B04FFA78A82FED7FB21F477" author="Nguyen" box="[917,1152,260,285]" pageId="15" pageNumber="16" refString="Nguyen, V. S. &amp; Ho, T. C. (1996) A Checklist of the Reptiles and Amphibians of Vietnam [in Vietnamese]. Science and Technique Publishing House, Hanoi, 264 pp." type="book" year="1996">Nguyen &amp; Ho 1996</bibRefCitation>
). However, according to 
<bibRefCitation id="F50D46FD8B04FFA78980FEF5FED9F454" box="[151,376,295,320]" pageId="15" pageNumber="16" refString="Ziegler, T., Ohler, A., Vu, N. T., Le, K. Q., Nguyen, X. T., Dinh, H. T. &amp; Bui, N. T. (2006) Review of the amphibian and reptile diversity of Phong Nha-Ke Bang National Park and adjacent areas, central Truong Son, Vietnam. In: Vences, M., Kohler, J., Ziegler, T. &amp; Bohme, W. (eds), Herpetologia Bonnensis II. Proceedings of the 13 th Congress of the Societas Europaea Herpetologica, Bonn, pp. 247 - 262." type="proceedings paper">
Ziegler 
<emphasis id="A3E8E71E8B04FFA789F9FEFBFE87F42B" box="[238,294,296,320]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
(2006)
</bibRefCitation>
and based on new locality records (see Appendix 1) the distribution of 
<taxonomicName id="569C408F8B04FFA78D82FEFAFABBF42B" box="[1173,1306,296,320]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="palmatus">
<emphasis id="A3E8E71E8B04FFA78D82FEFAFABBF42B" box="[1173,1306,296,320]" italics="true" pageId="15" pageNumber="16">G. palmatus</emphasis>
</taxonomicName>
in 
<collectingCountry id="E98B7B9C8B04FFA78C2AFEF5FA3DF454" box="[1341,1436,295,319]" name="Vietnam" pageId="15" pageNumber="16">Vietnam</collectingCountry>
in fact extends from several northern provinces southward to central 
<collectingCountry id="E98B7B9C8B04FFA78A8EFE9EFC59F40F" box="[921,1016,332,356]" name="Vietnam" pageId="15" pageNumber="16">Vietnam</collectingCountry>
in Quang Binh Province. Taking into account the records given for 
<emphasis id="A3E8E71E8B04FFA788F1FEA2FDC8F4E3" box="[486,617,368,392]" italics="true" pageId="15" pageNumber="16">G. c h i n e n s i s</emphasis>
by 
<bibRefCitation id="F50D46FD8B04FFA78B8FFEA2FC2AF4EC" author="Nguyen" box="[664,907,367,392]" pageId="15" pageNumber="16" refString="Nguyen, V. S. &amp; Ho, T. C. (1996) A Checklist of the Reptiles and Amphibians of Vietnam [in Vietnamese]. Science and Technique Publishing House, Hanoi, 264 pp." type="book" year="1996">Nguyen &amp; Ho (1996)</bibRefCitation>
, the latter species must occur in sympatry with 
<taxonomicName id="569C408F8B04FFA78980FE47FEBFF4C7" box="[151,286,404,428]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="palmatus">
<emphasis id="A3E8E71E8B04FFA78980FE47FEBFF4C7" box="[151,286,404,428]" italics="true" pageId="15" pageNumber="16">G. palmatus</emphasis>
</taxonomicName>
at least in the northern Vietnamese provinces of Bac Giang, Bac Kan, Cao Bang, Lang Son, Quang Ninh, and Vinh Phuc. However, all 
<taxonomicName id="569C408F8B04FFA78B3AFE6AFDD4F4BB" box="[557,629,440,464]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
<emphasis id="A3E8E71E8B04FFA78B3AFE6AFDD4F4BB" box="[557,629,440,464]" italics="true" pageId="15" pageNumber="16">Gekko</emphasis>
</taxonomicName>
studied by us from these provinces proved to be 
<taxonomicName id="569C408F8B04FFA78DB1FE6AFA8CF4BB" box="[1190,1325,440,464]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="palmatus">
<emphasis id="A3E8E71E8B04FFA78DB1FE6AFA8CF4BB" box="[1190,1325,440,464]" italics="true" pageId="15" pageNumber="16">G. palmatus</emphasis>
</taxonomicName>
and none had the diagnostic characters of 
<emphasis id="A3E8E71E8B04FFA788EAFE0FFD21F49F" box="[509,640,476,500]" italics="true" pageId="15" pageNumber="16">G. c h i n e n s i s</emphasis>
. This also makes the records of 
<taxonomicName id="569C408F8B04FFA78AFDFE0FFBD1F49F" box="[1002,1136,476,500]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="chinensis">
<emphasis id="A3E8E71E8B04FFA78AFDFE0FFBD1F49F" box="[1002,1136,476,500]" italics="true" pageId="15" pageNumber="16">G. chinensis</emphasis>
</taxonomicName>
from Hai Duong and Ninh Binh provinces (see 
<bibRefCitation id="F50D46FD8B04FFA7886BFDD2FDFAF77C" author="Nguyen" box="[380,603,511,536]" pageId="15" pageNumber="16" refString="Nguyen, V. S. &amp; Ho, T. C. (1996) A Checklist of the Reptiles and Amphibians of Vietnam [in Vietnamese]. Science and Technique Publishing House, Hanoi, 264 pp." type="book" year="1996">Nguyen &amp; Ho 1996</bibRefCitation>
) questionable and they are probably based on misidentifications of 
<taxonomicName id="569C408F8B04FFA78C47FDD2FF7BF757" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="palmatus">
<emphasis id="A3E8E71E8B04FFA78C47FDD2FF7BF757" italics="true" pageId="15" pageNumber="16">G. palmatus</emphasis>
</taxonomicName>
. 
<bibRefCitation id="F50D46FD8B04FFA789FFFDF6FDBCF757" author="Bobrov" box="[232,541,548,572]" pageId="15" pageNumber="16" refString="Bobrov, V. V. &amp; Semenov, D. V. (2008) Jastschjerizy Vietnama [Lizards of Vietnam]. Towarristschestowo nautschnyx isdanii KMK, Moscow, 225 pp." type="book" year="2008">Bobrov &amp; Semenov (2008)</bibRefCitation>
listed all previous identifications of 
<taxonomicName id="569C408F8B04FFA78AABFDF7FBE2F757" box="[956,1091,548,572]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="chinensis">
<emphasis id="A3E8E71E8B04FFA78AABFDF7FBE2F757" box="[956,1091,548,572]" italics="true" pageId="15" pageNumber="16">G. chinensis</emphasis>
</taxonomicName>
from 
<collectingCountry id="E98B7B9C8B04FFA78D9EFDF6FB49F757" box="[1161,1256,548,572]" name="Vietnam" pageId="15" pageNumber="16">Vietnam</collectingCountry>
as 
<taxonomicName id="569C408F8B04FFA78C18FDF7FA34F757" box="[1295,1429,548,572]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="palmatus">
<emphasis id="A3E8E71E8B04FFA78C18FDF7FA34F757" box="[1295,1429,548,572]" italics="true" pageId="15" pageNumber="16">G. palmatus</emphasis>
</taxonomicName>
. Ota 
<emphasis id="A3E8E71E8B04FFA789D0FD9BFEA1F70B" box="[199,256,584,608]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
(1995) believed 
<emphasis id="A3E8E71E8B04FFA788D5FD9AFDE4F70B" box="[450,581,584,608]" italics="true" pageId="15" pageNumber="16">G. c h i n e n s i s</emphasis>
and 
<taxonomicName id="569C408F8B04FFA78B95FD9AFCA9F70B" box="[642,776,584,608]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="palmatus">
<emphasis id="A3E8E71E8B04FFA78B95FD9AFCA9F70B" box="[642,776,584,608]" italics="true" pageId="15" pageNumber="16">G. palmatus</emphasis>
</taxonomicName>
to represent allopatric taxa (see also 
<bibRefCitation id="F50D46FD8B04FFA78DA7FD95FA31F734" author="Matsui" box="[1200,1424,583,607]" pageId="15" pageNumber="16" refString="Matsui, M. &amp; Ota, H. (1995) On Chinese herpetology. Herpetologica, 51 (2), 234 - 250." type="journal article" year="1995">Matsui &amp; Ota 1995</bibRefCitation>
). Concordant with the latter authors we herein suggest that the known occurrence of 
<taxonomicName id="569C408F8B04FFA78D2AFDBFFB65F7EF" box="[1085,1220,620,644]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="chinensis">
<emphasis id="A3E8E71E8B04FFA78D2AFDBFFB65F7EF" box="[1085,1220,620,644]" italics="true" pageId="15" pageNumber="16">G. chinensis</emphasis>
</taxonomicName>
is limited to 
<collectingCountry id="E98B7B9C8B04FFA78C4EFDBEFA3DF7EF" box="[1369,1436,620,644]" name="China" pageId="15" pageNumber="16">China</collectingCountry>
(southern provinces of Fujian, Guangdong, Guangxi, Guizhou, 
<collectingCountry id="E98B7B9C8B04FFA78A48FD42FC47F7C3" box="[863,998,656,680]" name="Hong Kong" pageId="15" pageNumber="16">Hong Kong</collectingCountry>
, Sichuan, and Yunnan) whereas that of 
<taxonomicName id="569C408F8B04FFA78980FD67FEBAF7A7" box="[151,283,692,716]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="palmatus">
<emphasis id="A3E8E71E8B04FFA78980FD67FEBAF7A7" box="[151,283,692,716]" italics="true" pageId="15" pageNumber="16">G. palmatus</emphasis>
</taxonomicName>
is only in northern and central 
<collectingCountry id="E98B7B9C8B04FFA78B60FD66FD77F7A7" box="[631,726,692,716]" name="Vietnam" pageId="15" pageNumber="16">Vietnam</collectingCountry>
(Bac Giang, Bac Kan, Cao Bang, Hai Duong, Hai Phong, Lang Son, Ninh Binh, Quang Binh, Quang Ninh, and Vinh Phuc provinces). Further collections or study of museum specimens are needed to clarify the identification of populations along the China-Vietnam border and whether 
<taxonomicName id="569C408F8B04FFA78C93FD2FFF5EF653" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="chinensis">
<emphasis id="A3E8E71E8B04FFA78C93FD2FFF5EF653" italics="true" pageId="15" pageNumber="16">G. chinensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="569C408F8B04FFA78822FCF2FE1BF653" box="[309,442,800,824]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="palmatus">
<emphasis id="A3E8E71E8B04FFA78822FCF2FE1BF653" box="[309,442,800,824]" italics="true" pageId="15" pageNumber="16">G. palmatus</emphasis>
</taxonomicName>
occur in sympatry.
</paragraph>
<paragraph id="91233B0C8B04FFA789D2FC96FA87F18C" blockId="15.[151,1437,151,2012]" pageId="15" pageNumber="16">
The distinguishing character of broad toe webbing in 
<taxonomicName id="569C408F8B04FFA78A31FC97FC0CF637" box="[806,941,836,860]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="palmatus">
<emphasis id="A3E8E71E8B04FFA78A31FC97FC0CF637" box="[806,941,836,860]" italics="true" pageId="15" pageNumber="16">G. palmatus</emphasis>
</taxonomicName>
versus a narrower webbing in 
<emphasis id="A3E8E71E8B04FFA78C1AFC97FA30F637" box="[1293,1425,836,860]" italics="true" pageId="15" pageNumber="16">G. c h i n e n s i s</emphasis>
, which can regularly be found in the older literature (e.g., 
<bibRefCitation id="F50D46FD8B04FFA78A08FCB5FC77F614" author="Boulenger" box="[799,982,871,895]" pageId="15" pageNumber="16" refString="Boulenger, G. A. (1907) Description of new lizard in the British Museum. Annals and Magazine of Natural History, 19, 486 - 489." type="journal article" year="1907">Boulenger 1907</bibRefCitation>
; 
<bibRefCitation id="F50D46FD8B04FFA78AF5FCB5FBC9F614" author="Smith" box="[994,1128,871,895]" pageId="15" pageNumber="16" refString="Smith, M. A. (1935) The Fauna of British India, Including Ceylon and Burma. Reptilia and Amphibia. II. Sauria. Taylor &amp; Francis Ltd., London, xiii + 440 pp., 1 pl., 2 maps." type="book" year="1935">Smith 1935</bibRefCitation>
; 
<bibRefCitation id="F50D46FD8B04FFA78D63FCBBFABAF614" author="Bourret" box="[1140,1307,871,896]" pageId="15" pageNumber="16" refString="Bourret, R. (1937 b) Notes herpetologiques sur l'Indochine francaise. XV. Lezards et Serpents recus au Laboratoire des Sciences Naturelles de l'Universite au cours de l'annee 1937. Descriptions de deux especes et de deux varietes nouvelles. Bulletin general de l'Instruction publique, 17 (4), annexe, 57 - 80." type="journal article" year="1937" yearSuffix="b">Bourret 1937b</bibRefCitation>
), may have conspicuously added to the above listed confusion and misidentifications, because it represents an ambiguous diagnostic character (Ota 
<emphasis id="A3E8E71E8B04FFA7889FFC63FE60F6A3" box="[392,449,944,968]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
1995). 
<bibRefCitation id="F50D46FD8B04FFA78B0FFC7DFD44F6AC" author="Smith" box="[536,741,943,967]" pageId="15" pageNumber="16" refString="Smith, M. A. (1935) The Fauna of British India, Including Ceylon and Burma. Reptilia and Amphibia. II. Sauria. Taylor &amp; Francis Ltd., London, xiii + 440 pp., 1 pl., 2 maps." type="book" year="1935">Smith (1935: 114)</bibRefCitation>
even concluded that 
<taxonomicName id="569C408F8B04FFA78AC2FC62FBFAF6A3" box="[981,1115,944,968]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="palmatus">
<emphasis id="A3E8E71E8B04FFA78AC2FC62FBFAF6A3" box="[981,1115,944,968]" italics="true" pageId="15" pageNumber="16">G. palmatus</emphasis>
</taxonomicName>
is “possibly a racial form of 
<emphasis id="A3E8E71E8B04FFA78980FC07FEB8F687" box="[151,281,980,1004]" italics="true" pageId="15" pageNumber="16">G. c h i n e n s i s</emphasis>
.” However, 
<taxonomicName id="569C408F8B04FFA788BDFC07FD8FF687" box="[426,558,980,1004]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="palmatus">
<emphasis id="A3E8E71E8B04FFA788BDFC07FD8FF687" box="[426,558,980,1004]" italics="true" pageId="15" pageNumber="16">G. palmatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="569C408F8B04FFA78B74FC07FD48F687" box="[611,745,980,1004]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="chinensis">
<emphasis id="A3E8E71E8B04FFA78B74FC07FD48F687" box="[611,745,980,1004]" italics="true" pageId="15" pageNumber="16">G. chinensis</emphasis>
</taxonomicName>
can be easily distinguished by the size of the internasals compared to the nasorostrals (internasals same size or larger than nasorostrals in 
<taxonomicName id="569C408F8B04FFA78AEFFC2AFB21F17B" box="[1016,1152,1016,1040]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="chinensis">
<emphasis id="A3E8E71E8B04FFA78AEFFC2AFB21F17B" box="[1016,1152,1016,1040]" italics="true" pageId="15" pageNumber="16">G. chinensis</emphasis>
</taxonomicName>
versus always smaller in 
<taxonomicName id="569C408F8B04FFA78980FBCFFEBCF15F" box="[151,285,1052,1076]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="palmatus">
<emphasis id="A3E8E71E8B04FFA78980FBCFFEBCF15F" box="[151,285,1052,1076]" italics="true" pageId="15" pageNumber="16">G. palmatus</emphasis>
</taxonomicName>
) and statistically significant differences in the number of midbody scales and the number of lamellae on the first, third and fifth fingers as well as the first toe (Ota 
<emphasis id="A3E8E71E8B04FFA78A4EFB93FC32F133" box="[857,915,1088,1112]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
1995). A key character that allows for proper identification of both aforementioned species is the presence of limb tubercles. These are present in 
<emphasis id="A3E8E71E8B04FFA78C1BFBB7FA30F117" box="[1292,1425,1124,1148]" italics="true" pageId="15" pageNumber="16">G. c h i n e n s i s</emphasis>
, but are completely lacking in 
<taxonomicName id="569C408F8B04FFA788FAFB5AFDD5F1CB" box="[493,628,1160,1184]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="palmatus">
<emphasis id="A3E8E71E8B04FFA788FAFB5AFDD5F1CB" box="[493,628,1160,1184]" italics="true" pageId="15" pageNumber="16">G. palmatus</emphasis>
</taxonomicName>
(our data). Ota 
<emphasis id="A3E8E71E8B04FFA78A3AFB5BFCC6F1CB" box="[813,871,1160,1184]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
(1995) indicated shank tubercles for 
<taxonomicName id="569C408F8B04FFA78C04FB5AFA3DF1CB" box="[1299,1436,1160,1184]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="chinensis">
<emphasis id="A3E8E71E8B04FFA78C04FB5AFA3DF1CB" box="[1299,1436,1160,1184]" italics="true" pageId="15" pageNumber="16">G. chinensis</emphasis>
</taxonomicName>
and the specimen depicted in 
<bibRefCitation id="F50D46FD8B04FFA788F6FB7EFCCFF1AF" author="Zhao" box="[481,878,1196,1220]" pageId="15" pageNumber="16" refString="Zhao, E. &amp; Adler, K. (1993) Herpetology of China. Society for the Study of Amphibians and Reptiles, Oxford, Ohio, 522 pp., 48 pls., 1 foldout map." type="book" year="1993">Zhao &amp; Adler (1993: pl. 21, fig. G.)</bibRefCitation>
has tubercles both on the fore and hind limbs; but the specimen of 
<taxonomicName id="569C408F8B04FFA78858FB02FE74F183" box="[335,469,1232,1256]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="palmatus">
<emphasis id="A3E8E71E8B04FFA78858FB02FE74F183" box="[335,469,1232,1256]" italics="true" pageId="15" pageNumber="16">G. palmatus</emphasis>
</taxonomicName>
depicted by 
<bibRefCitation id="F50D46FD8B04FFA78B73FB02FCE9F18C" author="Nguyen" box="[612,840,1231,1256]" pageId="15" pageNumber="16" refString="Nguyen, V. S., Ho, T. C. &amp; Nguyen, Q. T. (2009) Herpetofauna of Vietnam. Edition Chimaira, Frankfurt am Main, 768 pp." type="book" year="2009">
Nguyen 
<emphasis id="A3E8E71E8B04FFA78BD3FB03FD5DF183" box="[708,764,1232,1256]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
(2009
</bibRefCitation>
: fig. 314) completely lacks such tubercles.
</paragraph>
<paragraph id="91233B0C8B04FFA789D2FB26FF5BF0F7" blockId="15.[151,1437,151,2012]" pageId="15" pageNumber="16">
The taxonomic history of 
<taxonomicName id="569C408F8B04FFA788F2FB27FD21F067" box="[485,640,1268,1292]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="similignum">
<emphasis id="A3E8E71E8B04FFA788F2FB27FD21F067" box="[485,640,1268,1292]" italics="true" pageId="15" pageNumber="16">G. similignum</emphasis>
</taxonomicName>
is controversial. The 
<typeStatus id="4E2785AE8B04FFA78A66FB27FC00F066" box="[881,929,1269,1293]" pageId="15" pageNumber="16">type</typeStatus>
locality of this taxon is Hainan 
<collectingCountry id="E98B7B9C8B04FFA78C12FB26FAE8F067" box="[1285,1353,1268,1292]" name="Iceland" pageId="15" pageNumber="16">Island</collectingCountry>
(
<bibRefCitation id="F50D46FD8B04FFA78C40FB26FF72F044" author="Smith" pageId="15" pageNumber="16" refString="Smith, M. A. (1923) On a collection of reptiles and batrachians from the island of Hainan. Journal of the Natural History Society of Siam, 6, 195 - 212." type="journal article" year="1923">Smith 1923</bibRefCitation>
). 
<bibRefCitation id="F50D46FD8B04FFA789F1FAC5FE00F044" author="Stejneger" box="[230,417,1303,1327]" pageId="15" pageNumber="16" refString="Stejneger, L. (1932) The Chinese lizards of the genus Gekko. Proceedings of the United States National Museum, 132, 1 - 6." type="journal article" year="1932">Stejneger (1932)</bibRefCitation>
synonymized 
<taxonomicName id="569C408F8B04FFA78B53FACAFD7EF05B" box="[580,735,1304,1328]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="similignum">
<emphasis id="A3E8E71E8B04FFA78B53FACAFD7EF05B" box="[580,735,1304,1328]" italics="true" pageId="15" pageNumber="16">G. similignum</emphasis>
</taxonomicName>
with 
<taxonomicName id="569C408F8B04FFA78A08FACAFC07F05B" box="[799,934,1304,1328]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="chinensis">
<emphasis id="A3E8E71E8B04FFA78A08FACAFC07F05B" box="[799,934,1304,1328]" italics="true" pageId="15" pageNumber="16">G. chinensis</emphasis>
</taxonomicName>
(see also 
<bibRefCitation id="F50D46FD8B04FFA78D01FAC5FB70F044" author="Zhao" box="[1046,1233,1303,1328]" pageId="15" pageNumber="16" refString="Zhao, E., Zhao, K. &amp; Zhou, K. (1999) Fauna Sinica. Reptilia Vol. 2. Squamata, Lacertilia [in Chinese]. Science Press, Beijing, xi + 394 pp., 8 pls." type="book" year="1999">
Zhao 
<emphasis id="A3E8E71E8B04FFA78D41FACBFB2EF05B" box="[1110,1167,1304,1328]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
1999
</bibRefCitation>
). However, Ota 
<emphasis id="A3E8E71E8B04FFA78C9FFACBFF15F03F" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
(1995) and 
<bibRefCitation id="F50D46FD8B04FFA7882CFAEEFE73F03F" author="Kluge" box="[315,466,1340,1364]" pageId="15" pageNumber="16" refString="Kluge, A. G. (2001) Gekkotan lizard taxonomy. Hamadryad, 26, 1 - 209." type="journal article" year="2001">Kluge (2001)</bibRefCitation>
listed it as valid at the specific rank. 
<taxonomicName id="569C408F8B04FFA78A65FAEEFB89F03F" box="[882,1064,1340,1364]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="chinensis">
<emphasis id="A3E8E71E8B04FFA78A65FAEEFB89F03F" box="[882,1064,1340,1364]" italics="true" pageId="15" pageNumber="16">Gekko chinensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="569C408F8B04FFA78D48FAEFFB53F03F" box="[1119,1266,1340,1364]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="smilignum">
<emphasis id="A3E8E71E8B04FFA78D48FAEFFB53F03F" box="[1119,1266,1340,1364]" italics="true" pageId="15" pageNumber="16">G. smilignum</emphasis>
</taxonomicName>
differ in maximum SVL, in the number of scales around midbody and in the number of lamellae on the first and fourth toes (see 
<tableCitation id="DC1E0EB78B04FFA78980FA56FF48F0F7" box="[151,233,1412,1436]" captionStart="TABLE 1" captionStartId="5.[151,235,1777,1798]" captionText="TABLE 1. Measurements, scalation, and distribution of Gekko species." pageId="15" pageNumber="16">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D98668878B04FFB889D2FA7AFEC5F7A7" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="reference_group">
<paragraph id="91233B0C8B04FFA789D2FA7AFD5DF243" blockId="15.[151,1437,151,2012]" pageId="15" pageNumber="16">
<taxonomicName id="569C408F8B04FFA789D2FA7AFEDBF0AB" box="[197,378,1448,1472]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="liboensis">
<emphasis id="A3E8E71E8B04FFA789D2FA7AFEDBF0AB" box="[197,378,1448,1472]" italics="true" pageId="15" pageNumber="16">Gekko liboensis</emphasis>
</taxonomicName>
was described by Zhou &amp; Li (1982) based on an adult female specimen collected in 1979. Three further specimens (two females and a juvenile) were collected in 1984 (
<bibRefCitation id="F50D46FD8B04FFA78D08FA1EFB45F08F" author="Zhao" box="[1055,1252,1484,1508]" pageId="15" pageNumber="16" refString="Zhao, E., Zhao, K. &amp; Zhou, K. (1999) Fauna Sinica. Reptilia Vol. 2. Squamata, Lacertilia [in Chinese]. Science Press, Beijing, xi + 394 pp., 8 pls." type="book" year="1999">
Zhao 
<emphasis id="A3E8E71E8B04FFA78D73FA1FFB3EF08F" box="[1124,1183,1484,1508]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
1999
</bibRefCitation>
). 
<bibRefCitation id="F50D46FD8B04FFA78DEDFA1EFF40F36C" author="Zhao" pageId="15" pageNumber="16" refString="Zhao, E. &amp; Adler, K. (1993) Herpetology of China. Society for the Study of Amphibians and Reptiles, Oxford, Ohio, 522 pp., 48 pls., 1 foldout map." type="book" year="1993">Zhao &amp; Adler (1993)</bibRefCitation>
synonymized 
<taxonomicName id="569C408F8B04FFA7889AFA22FDB5F363" box="[397,532,1520,1544]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="liboensis">
<emphasis id="A3E8E71E8B04FFA7889AFA22FDB5F363" box="[397,532,1520,1544]" italics="true" pageId="15" pageNumber="16">G. liboensis</emphasis>
</taxonomicName>
with 
<taxonomicName id="569C408F8B04FFA78B4CFA22FD5BF363" box="[603,762,1520,1544]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="hokouensis">
<emphasis id="A3E8E71E8B04FFA78B4CFA22FD5BF363" box="[603,762,1520,1544]" italics="true" pageId="15" pageNumber="16">G. hokouensis</emphasis>
</taxonomicName>
(see also 
<bibRefCitation id="F50D46FD8B04FFA78A61FA3DFBB9F36C" author="Gunther" box="[886,1048,1519,1543]" pageId="15" pageNumber="16" refString="Gunther, R. (1994) Eine neue Art der Gattung Gekko (Reptilia, Squamata, Gekkonidae) aus dem Suden Vietnams. Zoologischer Anzeiger, 233, 57 - 67." type="journal article" year="1994">Günther 1994</bibRefCitation>
; 
<bibRefCitation id="F50D46FD8B04FFA78D3FFA3DFB13F36C" author="Kluge" box="[1064,1202,1519,1543]" pageId="15" pageNumber="16" refString="Kluge, A. G. (2001) Gekkotan lizard taxonomy. Hamadryad, 26, 1 - 209." type="journal article" year="2001">Kluge 2001</bibRefCitation>
; 
<bibRefCitation id="F50D46FD8B04FFA78DD5FA3DFF75F347" author="Henkel" pageId="15" pageNumber="16" refString="Henkel, F. W. &amp; Schmidt, W. (2003) Praxis Ratgeber Geckos. Edition Chimaira, Frankfurt am Main, 159 pp." type="book" year="2003">Henkel &amp; Schmidt 2003</bibRefCitation>
). In contrast, 
<bibRefCitation id="F50D46FD8B04FFA78865F9C6FDADF347" author="Kluge" box="[370,524,1556,1580]" pageId="15" pageNumber="16" refString="Kluge, A. G. (1993) Gekkonoid Lizard Taxonomy. International Gecko Society, San Diego, 245 pp." type="book" year="1993">Kluge (1993)</bibRefCitation>
, 
<bibRefCitation id="F50D46FD8B04FFA78B0CF9C7FD15F347" author="Bauer" box="[539,692,1556,1580]" pageId="15" pageNumber="16" refString="Bauer, A. M. (1994) Gekkonidae (Reptilia, Sauria), Part I Australia and Oceania. Das Tierreich 109 (Part). Walter de Gruyter, Berlin, XII + 306 pp." type="book" year="1994">Bauer (1994)</bibRefCitation>
, 
<bibRefCitation id="F50D46FD8B04FFA78BD5F9C6FCFEF347" author="Welch" box="[706,863,1556,1580]" pageId="15" pageNumber="16" refString="Welch, K. R. G. (1994) Lizards of the World. 1. Geckos. R &amp; A Research and Information Limited, Taunton, 165 pp." type="book" year="1994">Welch (1994)</bibRefCitation>
, 
<bibRefCitation id="F50D46FD8B04FFA78A79F9C6FBC2F347" author="Matsui" box="[878,1123,1556,1580]" pageId="15" pageNumber="16" refString="Matsui, M. &amp; Ota, H. (1995) On Chinese herpetology. Herpetologica, 51 (2), 234 - 250." type="journal article" year="1995">Matsui &amp; Ota (1995)</bibRefCitation>
, 
<bibRefCitation id="F50D46FD8B04FFA78D66F9C6FAE5F347" box="[1137,1348,1556,1580]" pageId="15" pageNumber="16" refString="Zhao, E., Zhao, K. &amp; Zhou, K. (1999) Fauna Sinica. Reptilia Vol. 2. Squamata, Lacertilia [in Chinese]. Science Press, Beijing, xi + 394 pp., 8 pls." type="book">
Zhao 
<emphasis id="A3E8E71E8B04FFA78DA3F9C7FB4EF347" box="[1204,1263,1556,1580]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
(1999)
</bibRefCitation>
, 
<bibRefCitation id="F50D46FD8B04FFA78C44F9C6FF45F33B" author="Rosler" pageId="15" pageNumber="16" refString="Rosler, H. (2000) Kommentierte Liste der rezent, subrezent und fossil bekannten Gecko-Taxa (Reptilia: Gekkonomorpha). Gekkota, 2, 28 - 153." type="journal article" year="2000">Rösler (2000)</bibRefCitation>
, Rösler 
<emphasis id="A3E8E71E8B04FFA78828F9EBFED6F33B" box="[319,375,1592,1616]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
(2005, 2006) treated 
<taxonomicName id="569C408F8B04FFA78B71F9EAFD48F33B" box="[614,745,1592,1616]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="liboensis">
<emphasis id="A3E8E71E8B04FFA78B71F9EAFD48F33B" box="[614,745,1592,1616]" italics="true" pageId="15" pageNumber="16">G. liboensis</emphasis>
</taxonomicName>
as a valid species. 
<taxonomicName id="569C408F8B04FFA78AAAF9EAFBCEF33B" box="[957,1135,1592,1616]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="liboensis">
<emphasis id="A3E8E71E8B04FFA78AAAF9EAFBCEF33B" box="[957,1135,1592,1616]" italics="true" pageId="15" pageNumber="16">Gekko liboensis</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="569C408F8B04FFA78C16F9EAFA3AF33B" box="[1281,1435,1592,1616]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="hokouensis">
<emphasis id="A3E8E71E8B04FFA78C16F9EAFA3AF33B" box="[1281,1435,1592,1616]" italics="true" pageId="15" pageNumber="16">G. hokouensis</emphasis>
</taxonomicName>
by a larger SVL (85 versus 
<quantity id="566496E98B04FFA788DAF98EFD80F31F" box="[461,545,1628,1652]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="15" pageNumber="16" unit="mm" value="70.0">70 mm</quantity>
), absence of internasals, more interorbitals (40 versus 30–33), flat dorsal tubercles, fewer dorsal tubercle rows (10 versus 12–18) and by the number of lamellae below the fourth toe (9 versus 15–18). However, as adult males are still lacking for 
<emphasis id="A3E8E71E8B04FFA78A12F977FC27F3D7" box="[773,902,1700,1724]" italics="true" pageId="15" pageNumber="16">G. l i b o e n s i s</emphasis>
, important diagnostic characters and possibly additional distinguishing features such as the number of precloacal pores are still missing. The nares of 
<taxonomicName id="569C408F8B04FFA78C35F91AFF4FF26F" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="auriverrucosus">
<emphasis id="A3E8E71E8B04FFA78C35F91AFF4FF26F" italics="true" pageId="15" pageNumber="16">G. auriverrucosus</emphasis>
</taxonomicName>
are not in contact with the first supralabial (Zhou &amp; Liu 1982), which seems to represent a derived character of the 
<taxonomicName id="569C408F8B04FFA78814F8C2FE33F243" box="[259,402,1808,1832]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="gecko">
<emphasis id="A3E8E71E8B04FFA78814F8C2FE33F243" box="[259,402,1808,1832]" italics="true" pageId="15" pageNumber="16">Gekko gecko</emphasis>
</taxonomicName>
group (except for 
<taxonomicName id="569C408F8B04FFA78B74F8C2FD4DF243" box="[611,748,1808,1832]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="verreauxi">
<emphasis id="A3E8E71E8B04FFA78B74F8C2FD4DF243" box="[611,748,1808,1832]" italics="true" pageId="15" pageNumber="16">G. verreauxi</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="91233B0C8B04FFA789D2F8E6FA38F2B7" blockId="15.[151,1437,151,2012]" pageId="15" pageNumber="16">
<bibRefCitation id="F50D46FD8B04FFA789D2F8E6FE1BF227" author="Zhao" box="[197,442,1844,1868]" pageId="15" pageNumber="16" refString="Zhao, E. &amp; Adler, K. (1993) Herpetology of China. Society for the Study of Amphibians and Reptiles, Oxford, Ohio, 522 pp., 48 pls., 1 foldout map." type="book" year="1993">Zhao &amp; Adler (1993)</bibRefCitation>
synonymized 
<taxonomicName id="569C408F8B04FFA78B76F8E7FD4EF227" box="[609,751,1844,1868]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="scabridus">
<emphasis id="A3E8E71E8B04FFA78B76F8E7FD4EF227" box="[609,751,1844,1868]" italics="true" pageId="15" pageNumber="16">G. scabridus</emphasis>
</taxonomicName>
with 
<taxonomicName id="569C408F8B04FFA78A24F8E7FC1DF227" box="[819,956,1844,1868]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="chinensis">
<emphasis id="A3E8E71E8B04FFA78A24F8E7FC1DF227" box="[819,956,1844,1868]" italics="true" pageId="15" pageNumber="16">G. chinensis</emphasis>
</taxonomicName>
(see also 
<bibRefCitation id="F50D46FD8B04FFA78D25F8E6FB1AF227" author="Kluge" box="[1074,1211,1844,1868]" pageId="15" pageNumber="16" refString="Kluge, A. G. (2001) Gekkotan lizard taxonomy. Hamadryad, 26, 1 - 209." type="journal article" year="2001">Kluge 2001</bibRefCitation>
), but 
<bibRefCitation id="F50D46FD8B04FFA78DE9F8E6FF40F21B" author="Matsui" pageId="15" pageNumber="16" refString="Matsui, M. &amp; Ota, H. (1995) On Chinese herpetology. Herpetologica, 51 (2), 234 - 250." type="journal article" year="1995">Matsui &amp; Ota (1995)</bibRefCitation>
and 
<bibRefCitation id="F50D46FD8B04FFA7880EF88AFE45F21B" box="[281,484,1880,1904]" pageId="15" pageNumber="16" refString="Zhao, E., Zhao, K. &amp; Zhou, K. (1999) Fauna Sinica. Reptilia Vol. 2. Squamata, Lacertilia [in Chinese]. Science Press, Beijing, xi + 394 pp., 8 pls." type="book">
Zhao 
<emphasis id="A3E8E71E8B04FFA7884EF88BFE33F21B" box="[345,402,1880,1904]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
(1999)
</bibRefCitation>
again evaluated the former taxon as valid species. Liu &amp; Zhou (1982) pointed to the phenotypic similarity between 
<taxonomicName id="569C408F8B04FFA788F8F8AFFDDBF2FF" box="[495,634,1916,1940]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="scabridus">
<emphasis id="A3E8E71E8B04FFA788F8F8AFFDDBF2FF" box="[495,634,1916,1940]" italics="true" pageId="15" pageNumber="16">G. scabridus</emphasis>
</taxonomicName>
and 
<taxonomicName id="569C408F8B04FFA78BB8F8AFFC9BF2FF" box="[687,826,1917,1940]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="japonicus">
<emphasis id="A3E8E71E8B04FFA78BB8F8AFFC9BF2FF" box="[687,826,1917,1940]" italics="true" pageId="15" pageNumber="16">G. japonicus</emphasis>
</taxonomicName>
. According to Liu &amp; Zhou (1982), 
<taxonomicName id="569C408F8B04FFA78DD5F8AFFAECF2FF" box="[1218,1357,1916,1940]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="scabridus">
<emphasis id="A3E8E71E8B04FFA78DD5F8AFFAECF2FF" box="[1218,1357,1916,1940]" italics="true" pageId="15" pageNumber="16">G. scabridus</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="569C408F8B04FFA789C2F873FEC0F2D3" box="[213,353,1953,1976]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="japonicus">
<emphasis id="A3E8E71E8B04FFA789C2F873FEC0F2D3" box="[213,353,1953,1976]" italics="true" pageId="15" pageNumber="16">G. japonicus</emphasis>
</taxonomicName>
by having tubercles on its back and hind limbs and by a higher count of precloacal pores. From 
<tableCitation id="DC1E0EB78B04FFA78980F816FF4BF2B7" box="[151,234,1988,2012]" captionStart="TABLE 1" captionStartId="5.[151,235,1777,1798]" captionText="TABLE 1. Measurements, scalation, and distribution of Gekko species." pageId="15" pageNumber="16">Table 1</tableCitation>
it is obvious that there exist additional distinguishing features between the two species. Compared with 
<emphasis id="A3E8E71E8B04FFA78C93F817FA38F2B7" box="[1412,1433,1989,2012]" italics="true" pageId="15" pageNumber="16">G.</emphasis>
</paragraph>
<paragraph id="91233B0C8B1BFFB88980FF4BFB8BF42B" blockId="16.[151,1436,151,716]" pageId="16" pageNumber="17">
<taxonomicName id="569C408F8B1BFFB88980FF4BFEA5F5DB" box="[151,260,153,176]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="japonicus">
<emphasis id="A3E8E71E8B1BFFB88980FF4BFEA5F5DB" box="[151,260,153,176]" italics="true" pageId="16" pageNumber="17">japonicus</emphasis>
</taxonomicName>
, 
<taxonomicName id="569C408F8B1BFFB88803FF4AFE00F5DB" box="[276,417,152,176]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="scabridus">
<emphasis id="A3E8E71E8B1BFFB88803FF4AFE00F5DB" box="[276,417,152,176]" italics="true" pageId="16" pageNumber="17">G. scabridus</emphasis>
</taxonomicName>
has fewer interorbitals (30 versus 32–35), fewer lamellae below the first toe (6–9 versus 10–12), and fourth toe (7–9 versus 14–16), more dorsal tubercle rows (17–21 versus 9–14) and a higher number of precloacal pores (10–15 versus 6–9). In comparsion to 
<taxonomicName id="569C408F8B1BFFB88BEBFF32FC23F593" box="[764,898,224,248]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="chinensis">
<emphasis id="A3E8E71E8B1BFFB88BEBFF32FC23F593" box="[764,898,224,248]" italics="true" pageId="16" pageNumber="17">G. chinensis</emphasis>
</taxonomicName>
it has only slightly developed webbing between the fingers and toes, fewer precloacal pores, and more postcloacal tubercles. We, therefore, accept the specific status of 
<taxonomicName id="569C408F8B1BFFB889CAFEFAFEC8F42B" box="[221,361,296,320]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="scabridus">
<emphasis id="A3E8E71E8B1BFFB889CAFEFAFEC8F42B" box="[221,361,296,320]" italics="true" pageId="16" pageNumber="17">G. scabridus</emphasis>
</taxonomicName>
and regard it as a valid member of the 
<taxonomicName id="569C408F8B1BFFB88A08FEFAFC7BF42B" box="[799,986,296,320]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="japonicus">
<emphasis id="A3E8E71E8B1BFFB88A08FEFAFC7BF42B" box="[799,986,296,320]" italics="true" pageId="16" pageNumber="17">Gekko japonicus</emphasis>
</taxonomicName>
group.
</paragraph>
<paragraph id="91233B0C8B1BFFB889D2FE9FFD98F4A4" blockId="16.[151,1436,151,716]" pageId="16" pageNumber="17">
<bibRefCitation id="F50D46FD8B1BFFB889D2FE9FFEEDF40F" author="Vogt" box="[197,332,332,357]" pageId="16" pageNumber="17" refString="Vogt, T. (1922) Zur Reptilien- und Amphibienfauna Sudchinas. Archiv fur Naturgeschichte, 10, 135 - 146." type="journal article" year="1922">Vogt (1922)</bibRefCitation>
originally described 
<emphasis id="A3E8E71E8B1BFFB88B2AFE9FFD30F40F" box="[573,657,332,356]" italics="true" pageId="16" pageNumber="17">G. m e l l i</emphasis>
based on a large series of which now only two specimens still exist (
<bibRefCitation id="F50D46FD8B1BFFB88989FEBDFE05F4EC" author="Bauer" box="[158,420,367,391]" pageId="16" pageNumber="17" refString="Bauer, A. M. &amp; Gunther, R. (1991) An annotated type catalogue of the geckos (Reptilia: Gekkonidae) in the Zoological Museum, Berlin. Mitteilungen aus dem Zoologischen Museum in Berlin, 67, 279 - 310." type="journal article" year="1991">Bauer &amp; Günther 1991</bibRefCitation>
). 
<bibRefCitation id="F50D46FD8B1BFFB888A0FEBDFDD0F4EC" author="Stejneger" box="[439,625,367,391]" pageId="16" pageNumber="17" refString="Stejneger, L. (1932) The Chinese lizards of the genus Gekko. Proceedings of the United States National Museum, 132, 1 - 6." type="journal article" year="1932">Stejneger (1932)</bibRefCitation>
listed 
<emphasis id="A3E8E71E8B1BFFB88BADFEA2FCADF4E3" box="[698,780,368,392]" italics="true" pageId="16" pageNumber="17">G. m e l l i</emphasis>
as a subspecies of 
<taxonomicName id="569C408F8B1BFFB88AF4FEA2FB2EF4E3" box="[995,1167,368,392]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="subpalmatus">
<emphasis id="A3E8E71E8B1BFFB88AF4FEA2FB2EF4E3" box="[995,1167,368,392]" italics="true" pageId="16" pageNumber="17">G. subpalmatus</emphasis>
</taxonomicName>
and 
<bibRefCitation id="F50D46FD8B1BFFB88DD3FEA2FAEDF4EC" author="Pope" box="[1220,1356,367,391]" pageId="16" pageNumber="17" refString="Pope, C. H. (1935) The Reptiles of China. Turtles, Crocodilians, Snakes, Lizards. The American Museum of Natural History, New York, lii + 604 pp." type="book" year="1935">Pope (1935)</bibRefCitation>
finally synonymized the former species with 
<taxonomicName id="569C408F8B1BFFB88B56FE47FD4CF4C7" box="[577,749,404,428]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="subpalmatus">
<emphasis id="A3E8E71E8B1BFFB88B56FE47FD4CF4C7" box="[577,749,404,428]" italics="true" pageId="16" pageNumber="17">G. subpalmatus</emphasis>
</taxonomicName>
. Rösler 
<emphasis id="A3E8E71E8B1BFFB88A5BFE47FC24F4C7" box="[844,901,404,428]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
(2005) revalidated 
<taxonomicName id="569C408F8B1BFFB88D76FE47FB19F4C7" box="[1121,1208,404,428]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="melli">
<emphasis id="A3E8E71E8B1BFFB88D76FE47FB19F4C7" box="[1121,1208,404,428]" italics="true" pageId="16" pageNumber="17">G. melli</emphasis>
</taxonomicName>
as a distinct species (see also 
<bibRefCitation id="F50D46FD8B1BFFB889E8FE65FD8CF4A4" author="Rosler" box="[255,557,439,463]" pageId="16" pageNumber="17" refString="Rosler, H. &amp; Tiedemann, F. (2007) Gekko melli Vogt, 1922 and its types (Reptilia, Sauria, Gekkonidae). Mitteilungen aus dem Museum fur Naturkunde in Berlin, Zoologische Reihe, 83, 105 - 108." type="journal article" year="2007">Rösler &amp; Tiedemann 2007</bibRefCitation>
).
</paragraph>
<paragraph id="91233B0C8B1BFFB889D2FE0EFEC5F7A7" blockId="16.[151,1436,151,716]" pageId="16" pageNumber="17">
With respect to 
<taxonomicName id="569C408F8B1BFFB88861FE0FFD82F49F" box="[374,547,476,500]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="subpalmatus">
<emphasis id="A3E8E71E8B1BFFB88861FE0FFD82F49F" box="[374,547,476,500]" italics="true" pageId="16" pageNumber="17">G. subpalmatus</emphasis>
</taxonomicName>
it is interesting to note that the back of the 
<typeStatus id="4E2785AE8B1BFFB88D1BFE0EFBCFF49F" box="[1036,1134,476,500]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
is slightly banded and that the vertebral region shows four indistinct light blotches (Rösler 
<emphasis id="A3E8E71E8B1BFFB88A74FDD3FC3DF773" box="[867,924,512,536]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
2005: figs. 11–12). However, the ground coloration of the live specimen depicted in 
<bibRefCitation id="F50D46FD8B1BFFB88B41FDF6FBBAF757" author="Zhao" box="[598,1051,548,572]" pageId="16" pageNumber="17" refString="Zhao, E. &amp; Adler, K. (1993) Herpetology of China. Society for the Study of Amphibians and Reptiles, Oxford, Ohio, 522 pp., 48 pls., 1 foldout map." type="book" year="1993">Zhao &amp; Adler (1993: their Pl. 32, fig. B)</bibRefCitation>
is blackish-olive (Black 3U). This specimen, in addition, has a repeatedly interrupted orange (722U) vertebral stripe and tail blotches of the same color; a banded pattern as observable from the 
<typeStatus id="4E2785AE8B1BFFB88BB2FDBEFCA6F7EF" box="[677,775,620,644]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
is not discernible (see also fig. in 
<bibRefCitation id="F50D46FD8B1BFFB88D90FDBFFAD5F7EF" author="Huang" box="[1159,1396,620,645]" pageId="16" pageNumber="17" refString="Huang, Z. Y. &amp; Zong, Y. (1998) Zhongguo Techan de Paxing Dongwu. Fudan University, Shanghai, 116 pp." type="book" year="1998">Huang &amp; Zong 1998</bibRefCitation>
: pl. 29). Whether this reflects intraspecific variation or a persistent confusion over the identity of this species remains to be investigated.
</paragraph>
</subSubSection>
</treatment>
</document>