304 lines
38 KiB
XML
304 lines
38 KiB
XML
<document id="5A395F15C4B200EFB0417E2E6350E2A5" ID-DOI="10.5281/zenodo.5392079" ID-ISSN="1638-9387" ID-Zenodo-Dep="5392079" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630316032044" checkinUser="marcus" docAuthor="Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul & Spiegel, Didier Van Den" docDate="2007" docId="03F4C378A2213058FF450270FB77F935" docLanguage="en" docName="Zoosystema.2007.29.1.7-49.pdf" docOrigin="Zoosystema 29 (1)" docStyle="DocumentStyle:2A9C6F9EA5F90D98170D9E0D9CA29375.4:Zoosystema.2000-2008.journal_article.type1" docStyleId="2A9C6F9EA5F90D98170D9E0D9CA29375" docStyleName="Zoosystema.2000-2008.journal_article.type1" docStyleVersion="4" docTitle="Glyphiulus subgranulatus Golovatch & Geoffroy & Mauriès & Spiegel 2007, n. sp." docType="treatment" docVersion="4" lastPageNumber="23" masterDocId="FFCDBB00A22F3048FF94075DFFE6FF91" masterDocTitle="Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group" masterLastPageNumber="49" masterPageNumber="7" pageNumber="21" updateTime="1699223690750" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="5637C35837D1EF9B01A04EDEDB87F9A7">Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group</mods:title>
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<mods:name id="9BE9A8F4F215677A1B7D5551DC3B9BAA" type="personal">
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<mods:roleTerm id="E1C9E51499E01474930CCBDAA5F50193">Author</mods:roleTerm>
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<mods:namePart id="39C54D7CA221413E25445EB7A6AF5177">Golovatch, Sergei I.</mods:namePart>
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<mods:affiliation id="6A50224631060C945B1D1910E3CBD94D">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071 (Russia) sgol @ orc. ru</mods:affiliation>
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<mods:nameIdentifier id="26336F12D0B944148C181789884BEFC7" type="email">sgol@orc.ru</mods:nameIdentifier>
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<mods:roleTerm id="E86CE4C2FF505460E985427917540040">Author</mods:roleTerm>
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<mods:namePart id="438CF9A7A5D0C159A23B800FCC9D2C76">Geoffroy, Jean-Jacques</mods:namePart>
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<mods:affiliation id="29A3786676AF54C1F804127E2A57F43A">Muséum national d’Histoire naturelle, Département Écologie et Gestion de la Biodiversité, USM 306 Réseaux trophiques du sol, 4 avenue du Petit Château, F- 91800 Brunoy (France) geoffroy @ mnhn. fr</mods:affiliation>
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<mods:nameIdentifier id="245F5CAA51493F6C51C02891ADAACA59" type="email">geoffroy@mnhn.fr</mods:nameIdentifier>
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<mods:roleTerm id="C88BDEF7E37DDFEB33DE21A96A34B125">Author</mods:roleTerm>
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<mods:namePart id="0ADC3FB5C627B8DFDE39E2BCB90BAC91">Mauriès, Jean-Paul</mods:namePart>
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<mods:affiliation id="7E3F075618B90729CF48D9C66648FF99">Muséum national d’Histoire naturelle, Département Systématique et Évolution, USM 602, case postale 53, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) collmill @ mnhn. fr</mods:affiliation>
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<mods:nameIdentifier id="6363CDCD0185210A01C0E6EFED269810" type="email">collmill@mnhn.fr</mods:nameIdentifier>
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<mods:namePart id="5402A77E15BA07E859B89232B30148B0">Spiegel, Didier Van Den</mods:namePart>
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<mods:affiliation id="C333FAFF02758DA190AB13FCA0BFAC1F">Musée royal de l’Afrique centrale, Leuvensesteenweg 13, B- 3080 Tervuren (Belgium) didier. van. den. spiegel @ africamuseum. be</mods:affiliation>
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<mods:title id="4DA5DA1E2624C5F242BBB410F62DE4F5">Zoosystema</mods:title>
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<mods:part id="D19D92CE7FDEA4A2715445AAC4F92823">
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<mods:date id="15AF0328E3C036E0ADB3D81658EF89C5">2007</mods:date>
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<mods:number id="900C52BF25B68B2DE0B4DA9594F12A14">29</mods:number>
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<treatment id="03F4C378A2213058FF450270FB77F935" ID-DOI="http://doi.org/10.5281/zenodo.5468294" ID-Zenodo-Dep="5468294" LSID="urn:lsid:plazi:treatment:03F4C378A2213058FF450270FB77F935" httpUri="http://treatment.plazi.org/id/03F4C378A2213058FF450270FB77F935" lastPageId="16" lastPageNumber="23" pageId="14" pageNumber="21">
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<subSubSection id="C34721E5A2213046FF450270FDC1FAD6" box="[209,551,1325,1351]" pageId="14" pageNumber="21" type="nomenclature">
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<paragraph id="8BE2726EA2213046FF450270FDC1FAD6" blockId="14.[209,551,1325,1383]" box="[209,551,1325,1351]" pageId="14" pageNumber="21">
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<heading id="D0AAC502A2213046FF450270FDC1FAD6" box="[209,551,1325,1351]" centered="true" fontSize="11" level="2" pageId="14" pageNumber="21" reason="2">
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<taxonomicName id="4C5D09EDA2213046FF450270FE03FAD6" ID-CoL="3GGL6" authority="Golovatch & Geoffroy & Mauriès & Spiegel, 2007" authorityName="Golovatch & Geoffroy & Mauriès & Spiegel" authorityYear="2007" box="[209,485,1325,1351]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="14" pageNumber="21" phylum="Arthropoda" rank="species" species="subgranulatus" status="sp. nov.">
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<emphasis id="B929AE7CA2213046FF450270FE03FAD6" bold="true" box="[209,485,1325,1351]" italics="true" pageId="14" pageNumber="21">Glyphiulus subgranulatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21A1307A2213046FE780270FDC1FAD6" box="[492,551,1325,1351]" pageId="14" pageNumber="21" rank="species">n. sp.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<subSubSection id="C34721E5A2213046FED30210FE57FAF6" box="[327,433,1357,1383]" pageId="14" pageNumber="21" type="description">
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<paragraph id="8BE2726EA2213046FED30210FE57FAF6" blockId="14.[209,551,1325,1383]" box="[327,433,1357,1383]" pageId="14" pageNumber="21">
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(
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<figureCitation id="13666EEBA2213046FEDB0210FE41FAF6" box="[335,423,1357,1383]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="12.[113,124,1318,1334]" captionStartId-1="13.[113,124,1024,1040]" captionStartId-2="14.[113,124,963,979]" captionTargetBox-0="[121,1200,215,1260]" captionTargetBox-1="[120,1200,215,977]" captionTargetBox-2="[113,1209,218,907]" captionTargetId-0="figure-36@12.[390,661,595,786]" captionTargetId-1="figure-32@13.[390,663,596,787]" captionTargetId-2="figure-18@14.[74,1220,182,921]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="14" captionText-0="FIG. 7. — Glyphiulus subgranulatus n. sp., ♀ paratype:A, head, ventral view; B, collum, dorsal view; C, middle part of body, dorsal view; D, same, lateral view; E, caudal part of body, lateral view; F, same, dorsal view. Scale bars: A, 0.1 mm; B, 0.2 mm, C-F, 0.5 mm." captionText-1="FIG. 8. — Glyphiulus subgranulatus n. sp., ♂ and ♀ paratypes: A, anterior part of body, lateral view; B, caudal part of body, ventral view; C, midbody segment section (a little damaged), caudal view; D, male legs 1, caudal view. Scale bars: A, C, 0.5 mm; B, 0.2 mm; D, 0.1 mm." captionText-2="FIG. 9. — Glyphiulus subgranulatus n. sp., ♂ paratype: A, antenna; B, gnathochilarium, ventral view; C, legs 1, caudal view; D, legs 2, caudal view; E, anterior gonopods, caudal view; F, posterior gonopods, frontal view; G, same, caudal view. Scale bar: A, B, 0.3 mm; C-G, 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5392095" figureDoi-1="http://doi.org/10.5281/zenodo.5392097" figureDoi-2="http://doi.org/10.5281/zenodo.5392099" httpUri-0="https://zenodo.org/record/5392095/files/figure.png" httpUri-1="https://zenodo.org/record/5392097/files/figure.png" httpUri-2="https://zenodo.org/record/5392099/files/figure.png" pageId="14" pageNumber="21">Figs 7-9</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C34721E5A2213046FFE502DAFE53F9C6" pageId="14" pageNumber="21" type="materials_examined">
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<materialsCitation id="3B357833A2213046FFE502DAFEB4FA42" collectingDate="1989-01-05" collectionCode="NMNHS" collectorName="P. Beron" country="China" county="Mengzi County" location="Mengzi County" pageId="14" pageNumber="21" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
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TYPE MATERIAL. —
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.
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<collectingRegion id="4999BC8CA2213046FEFE02DAFE0FFA0F" box="[362,489,1415,1438]" country="China" name="Yunnan" pageId="14" pageNumber="21">Yunnan Prov.</collectingRegion>
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,
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<collectingCounty id="62830AE2A2213046FE6402DAFD65FA0F" box="[496,643,1415,1438]" pageId="14" pageNumber="21">Mengzi County</collectingCounty>
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, cave near footpath to plateau,
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<date id="FFE354AEA2213046FE0B02FFFE10FA28" box="[415,502,1442,1465]" pageId="14" pageNumber="21" value="1989-01-05">
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<collectingDate id="EFA7AD46A2213046FE0B02FFFE10FA28" box="[415,502,1442,1465]" pageId="14" pageNumber="21" value="1989-01-05">5.I.1989</collectingDate>
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</date>
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, leg.
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<collectorName id="26A817B8A2213046FDBF02FFFD67FA28" box="[555,641,1442,1465]" pageId="14" pageNumber="21">P. Beron</collectorName>
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,
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<typeStatus id="54E6CCCCA2213046FFE702E1FF21FA42" box="[115,199,1468,1491]" pageId="14" pageNumber="21" type="holotype">holotype</typeStatus>
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<specimenCount id="9D5BB9E7A2213046FF5E02E2FF38FA42" box="[202,222,1471,1491]" pageId="14" pageNumber="21" type="male">♂</specimenCount>
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(
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<collectionCode id="ED4CEAABA2213046FF7D02E1FEADFA42" box="[233,331,1468,1491]" pageId="14" pageNumber="21">NMNHS</collectionCode>
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)
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</materialsCitation>
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;
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<materialsCitation id="3B357833A2213046FECE02E1FF04FA7F" collectingDate="1989-01-05" collectionCode="NMNHS" collectorName="P. Beron" country="China" county="Mengzi County" isTailEnumeration="true" location="Mengzi County" pageId="14" pageNumber="21" specimenCount="32" specimenCount-female="14" specimenCount-male="7" stateProvince="Yunnan" typeStatus="paratype">
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<typeStatus id="54E6CCCCA2213046FECE02E1FE52FA42" box="[346,436,1468,1491]" pageId="14" pageNumber="21" type="paratype">paratypes</typeStatus>
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,
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<specimenCount id="9D5BB9E7A2213046FE6F02E1FDDCFA42" box="[507,570,1468,1491]" pageId="14" pageNumber="21" type="female">14 ♀♀</specimenCount>
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,
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<specimenCount id="9D5BB9E7A2213046FDD102E1FD6EFA42" box="[581,648,1468,1491]" pageId="14" pageNumber="21" type="generic">11 juv.</specimenCount>
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(
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;
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<specimenCount id="9D5BB9E7A2213046FF7A028AFECAFA7C" box="[238,300,1495,1518]" pageId="14" pageNumber="21" type="male">2 ♂♂</specimenCount>
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,
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(
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<collectionCode id="ED4CEAABA2213046FE15028AFE33FA7F" box="[385,469,1495,1518]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="21">MNHN</collectionCode>
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<specimenCode id="DBFBDA15A2213046FE49028AFDD6FA7F" box="[477,560,1495,1518]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="21">GA 035</specimenCode>
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)
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</materialsCitation>
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;
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<materialsCitation id="3B357833A2213046FDD7028AFEECF999" collectionCode="ZMUM" country="China" pageId="14" pageNumber="21" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Yunnan" typeStatus="paratype">
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<specimenCount id="9D5BB9E7A2213046FDD7028AFD66FA7C" box="[579,640,1495,1518]" pageId="14" pageNumber="21" type="male">2 ♂♂</specimenCount>
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,
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<specimenCount id="9D5BB9E7A2213046FFE502ACFF40F996" box="[113,166,1521,1544]" pageId="14" pageNumber="21" type="female">2 ♀♀</specimenCount>
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(
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<collectionCode id="ED4CEAABA2213046FF2402ACFEE5F999" box="[176,259,1521,1544]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="14" pageNumber="21">ZMUM</collectionCode>
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)
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</materialsCitation>
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;
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<materialsCitation id="3B357833A2213046FE8502ACFE59F999" box="[273,447,1521,1544]" collectionCode="ZMUC" country="China" pageId="14" pageNumber="21" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
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<specimenCount id="9D5BB9E7A2213046FE8502ACFED3F996" box="[273,309,1521,1544]" pageId="14" pageNumber="21" type="male">1 ♂</specimenCount>
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,
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<specimenCount id="9D5BB9E7A2213046FEAB02ACFE86F996" box="[319,352,1521,1544]" pageId="14" pageNumber="21" type="female">1 ♀</specimenCount>
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(
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<collectionCode id="ED4CEAABA2213046FEFF02ACFE5EF999" box="[363,440,1521,1544]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="14" pageNumber="21">ZMUC</collectionCode>
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)
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</materialsCitation>
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;
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<materialsCitation id="3B357833A2213046FE5302ACFD65F999" box="[455,643,1521,1544]" collectionCode="MCSNV" country="China" pageId="14" pageNumber="21" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
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<specimenCount id="9D5BB9E7A2213046FE5302ACFE0CF996" box="[455,490,1521,1544]" pageId="14" pageNumber="21" type="male">1 ♂</specimenCount>
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,
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<specimenCount id="9D5BB9E7A2213046FE6002ACFDF0F996" box="[500,534,1521,1544]" pageId="14" pageNumber="21" type="female">1 ♀</specimenCount>
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(
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<collectionCode id="ED4CEAABA2213046FDB402ACFD9AF999" box="[544,636,1521,1544]" pageId="14" pageNumber="21">MCSNV</collectionCode>
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)
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</materialsCitation>
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;
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<materialsCitation id="3B357833A2213046FFFB0156FE56F9C6" collectingDate="1989-01-06" collectionCode="SEM, NMNHS" collectorName="P. Beron" country="China" county="Mengzi County" location="Pothole No." municipality="Pothole No." pageId="14" pageNumber="21" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
|
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<specimenCount id="9D5BB9E7A2213046FFFB0156FF7DF9B3" box="[111,155,1547,1570]" pageId="14" pageNumber="21" type="male">1 ♂</specimenCount>
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,
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<specimenCount id="9D5BB9E7A2213046FF3E0156FF37F9B3" box="[170,209,1547,1570]" pageId="14" pageNumber="21" type="female">1 ♀</specimenCount>
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(
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<collectionCode id="ED4CEAABA2213046FF740156FEFFF9B3" box="[224,281,1547,1570]" pageId="14" pageNumber="21">SEM</collectionCode>
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). —
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<collectingRegion id="4999BC8CA2213046FEDF0156FE31F9B3" box="[331,471,1547,1570]" country="China" name="Yunnan" pageId="14" pageNumber="21">Yunnan Prov.</collectingRegion>
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,
|
||
<collectingCounty id="62830AE2A2213046FE760156FD64F9B3" box="[482,642,1547,1570]" pageId="14" pageNumber="21">Mengzi County</collectingCounty>
|
||
,
|
||
<location id="8E8224B5A2213046FFE7017BFF09F9AC" LSID="urn:lsid:plazi:treatment:03F4C378A2213058FF450270FB77F935:8E8224B5A2213046FFE7017BFF09F9AC" box="[115,239,1574,1597]" country="China" county="Mengzi County" municipality="Pothole No." name="Pothole No." pageId="14" pageNumber="21" stateProvince="Yunnan">
|
||
<collectingMunicipality id="6B86E814A2213046FFE7017BFF09F9AC" box="[115,239,1574,1597]" pageId="14" pageNumber="21">Pothole No.</collectingMunicipality>
|
||
</location>
|
||
2 (
|
||
<location id="8E8224B5A2213046FE87017BFE34F9AC" LSID="urn:lsid:plazi:treatment:03F4C378A2213058FF450270FB77F935:8E8224B5A2213046FE87017BFE34F9AC" box="[275,466,1574,1597]" country="China" county="Mengzi County" municipality="Pothole No." name="Ma Fa Tiao Dong" pageId="14" pageNumber="21" stateProvince="Yunnan">Ma Fa Tiao Dong</location>
|
||
),
|
||
<date id="FFE354AEA2213046FE72017BFDD9F9AC" box="[486,575,1574,1597]" pageId="14" pageNumber="21" value="1989-01-06">
|
||
<collectingDate id="EFA7AD46A2213046FE72017BFDD9F9AC" box="[486,575,1574,1597]" pageId="14" pageNumber="21" value="1989-01-06">6.I.1989</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26A817B8A2213046FDEC017BFF57F9C6" pageId="14" pageNumber="21">P. Beron</collectorName>
|
||
,
|
||
<typeStatus id="54E6CCCCA2213046FF2F011DFEE8F9C6" box="[187,270,1600,1623]" pageId="14" pageNumber="21" type="paratype">paratype</typeStatus>
|
||
<specimenCount id="9D5BB9E7A2213046FE81011DFED1F9C6" box="[277,311,1600,1623]" pageId="14" pageNumber="21" type="female">1 ♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED4CEAABA2213046FED1011DFE4FF9C6" box="[325,425,1600,1623]" pageId="14" pageNumber="21">NMNHS</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34721E5A2213046FFE5012CFEF4F932" pageId="14" pageNumber="21" type="etymology">
|
||
<paragraph id="8BE2726EA2213046FFE5012CFEF4F932" blockId="14.[113,646,1649,1699]" pageId="14" pageNumber="21">
|
||
ETYMOLOGY. — To emphasize the obvious similarity to
|
||
<taxonomicName id="4C5D09EDA2213046FF1801D1FEEAF932" baseAuthorityName="Gervais" baseAuthorityYear="1847" box="[140,268,1676,1699]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="14" pageNumber="21" phylum="Arthropoda" rank="species" species="granulatus">
|
||
<emphasis id="B929AE7CA2213046FF1801D1FEEAF932" box="[140,268,1676,1699]" italics="true" pageId="14" pageNumber="21">G. granulatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34721E5A2213046FD310313FCD9FB5E" pageId="14" pageNumber="21" type="diagnosis">
|
||
<paragraph id="8BE2726EA2213046FD310313FCD9FB5E" blockId="14.[675,1210,1102,1231]" pageId="14" pageNumber="21">DIAGNOSIS. — Differs from the most similar congeners by the crests on the collum being underdeveloped (especially so anteriorly), the body segments somewhat compressed laterally, the anterior gonopods with unusually slender telopodites, etc.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34721E5A2213058FD3103B3FED6F935" lastPageId="16" lastPageNumber="23" pageId="14" pageNumber="21" type="description">
|
||
<paragraph id="8BE2726EA2213046FD3103B3FCD3FA97" blockId="14.[674,1212,1262,1700]" box="[677,821,1262,1288]" pageId="14" pageNumber="21">DESCRIPTION</paragraph>
|
||
<paragraph id="8BE2726EA2213046FD310253FCD8F9D4" blockId="14.[674,1212,1262,1700]" pageId="14" pageNumber="21">
|
||
Length of adults of both sexes
|
||
<quantity id="4CA5DF8BA2213046FC620253FB88FAB9" box="[1014,1134,1294,1320]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" metricValueMax="2.8" metricValueMin="1.9" pageId="14" pageNumber="21" unit="mm" value="23.5" valueMax="28.0" valueMin="19.0">19-28 mm</quantity>
|
||
, width 1.0-
|
||
<quantity id="4CA5DF8BA2213046FD400270FCD5FAD6" box="[724,819,1325,1351]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="14" pageNumber="21" unit="mm" value="1.3">1.3 mm</quantity>
|
||
; males usually a little shorter and more slender than females. Coloration much as in
|
||
<taxonomicName id="4C5D09EDA2213046FD510230FCBFFA16" baseAuthorityName="Gervais" baseAuthorityYear="1847" box="[709,857,1389,1415]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="14" pageNumber="21" phylum="Arthropoda" rank="species" species="granulatus">
|
||
<emphasis id="B929AE7CA2213046FD510230FCBFFA16" box="[709,857,1389,1415]" italics="true" pageId="14" pageNumber="21">G. granulatus</emphasis>
|
||
</taxonomicName>
|
||
, variegate, yellow-brown, with a dark brown vertex, blackish ocellaria, mainly brown crests on collum and grey-brownish lateral longitudinal stripes running down from dark brown ozoporiferous tubercles; dorsum generally light, but a thin axial line mostly present due to darker median crests.
|
||
</paragraph>
|
||
<paragraph id="8BE2726EA2213047FD2F0117FDB5F994" blockId="14.[674,1212,1262,1700]" lastBlockId="15.[112,648,1484,1700]" lastPageId="15" lastPageNumber="22" pageId="14" pageNumber="21">
|
||
Adults with 38-52p+5-2a+T in males, 39-63p+5- 1a+T in females; largest juvenile
|
||
<specimenCount id="9D5BB9E7A2213046FC620130FBEAF912" box="[1014,1036,1645,1667]" pageId="14" pageNumber="21" type="male">♂</specimenCount>
|
||
with 31p+3a+T; two smallest juveniles with 17 or 18 haplopodous segments after 6th, plus 2a+T.
|
||
<typeStatus id="54E6CCCCA2203047FE470291FDDBFA77" box="[467,573,1484,1510]" pageId="15" pageNumber="22" type="holotype">Holotype</typeStatus>
|
||
about
|
||
<quantity id="4CA5DF8BA2203047FFE702B6FF24F994" box="[115,194,1515,1541]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="15" pageNumber="22" unit="mm" value="23.0">23 mm</quantity>
|
||
long, 1.0 mm wide, with 47p+2a+T.
|
||
</paragraph>
|
||
<caption id="DF2222E6A2203047FFE50209FEE8FA00" ID-DOI="http://doi.org/10.5281/zenodo.5392101" ID-Zenodo-Dep="5392101" httpUri="https://zenodo.org/record/5392101/files/figure.png" pageId="15" pageNumber="22" startId="15.[113,124,1364,1380]" targetBox="[119,1198,221,1320]" targetPageId="15">
|
||
<paragraph id="8BE2726EA2203047FFE50209FEE8FA00" blockId="15.[113,1210,1363,1426]" pageId="15" pageNumber="22">
|
||
FIG. 10. —
|
||
<taxonomicName id="4C5D09EDA2203047FF5E020EFE47FAF5" authority="Golovatch & Geoffroy & Mauriès & Spiegel, 2007" authorityName="Golovatch & Geoffroy & Mauriès & Spiegel" authorityYear="2007" box="[202,417,1363,1380]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="15" pageNumber="22" phylum="Arthropoda" rank="species" species="paragranulatus" status="sp. nov.">
|
||
<emphasis id="B929AE7CA2203047FF5E020EFE47FAF5" box="[202,417,1363,1380]" italics="true" pageId="15" pageNumber="22">Glyphiulus paragranulatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A1307A2203047FE320209FE30FAF5" box="[422,470,1364,1380]" pageId="15" pageNumber="22" rank="species">n. sp.</taxonomicNameLabel>
|
||
, ♂ and ♀ paratypes:
|
||
<emphasis id="B929AE7CA2203047FD10020EFD77FAF5" bold="true" box="[644,657,1363,1380]" pageId="15" pageNumber="22">A</emphasis>
|
||
, anterior part of body, lateral view;
|
||
<emphasis id="B929AE7CA2203047FC27020EFC26FAF2" bold="true" box="[947,960,1363,1379]" pageId="15" pageNumber="22">B</emphasis>
|
||
, same, dorsal view;
|
||
<emphasis id="B929AE7CA2203047FBFD020EFB91FAF5" bold="true" box="[1129,1143,1363,1380]" pageId="15" pageNumber="22">C</emphasis>
|
||
, middle part of body, dorsal view;
|
||
<emphasis id="B929AE7CA2203047FEDD0237FEB1FAEB" bold="true" box="[329,343,1386,1402]" pageId="15" pageNumber="22">D</emphasis>
|
||
, same, lateral view;
|
||
<emphasis id="B929AE7CA2203047FD950237FDEBFAEB" bold="true" box="[513,525,1386,1402]" pageId="15" pageNumber="22">E</emphasis>
|
||
, caudal part of body, dorsal view;
|
||
<emphasis id="B929AE7CA2203047FCB80237FCD1FAEB" bold="true" box="[812,823,1386,1402]" pageId="15" pageNumber="22">F</emphasis>
|
||
, same, caudal view. Scale bars: A, D, 0.5 mm; B, C, E, F, 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE2726EA2203058FF1C0156FED6F935" blockId="15.[112,648,1484,1700]" lastBlockId="16.[113,647,1610,1700]" lastPageId="16" lastPageNumber="23" pageId="15" pageNumber="22">
|
||
Other characters as in
|
||
<taxonomicName id="4C5D09EDA2203047FE140156FDF6F9B4" baseAuthorityName="Gervais" baseAuthorityYear="1847" box="[384,528,1547,1573]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="15" pageNumber="22" phylum="Arthropoda" rank="species" species="granulatus">
|
||
<emphasis id="B929AE7CA2203047FE140156FDF6F9B4" box="[384,528,1547,1573]" italics="true" pageId="15" pageNumber="22">G. granulatus</emphasis>
|
||
</taxonomicName>
|
||
, including mentum either divided (n = 1) (
|
||
<figureCitation id="13666EEBA2203047FE480176FDD6F9D4" box="[476,560,1579,1605]" captionStart="FIG" captionStartId="14.[113,124,963,979]" captionTargetBox="[113,1209,218,907]" captionTargetId="figure-18@14.[74,1220,182,921]" captionTargetPageId="14" captionText="FIG. 9. — Glyphiulus subgranulatus n. sp., ♂ paratype: A, antenna; B, gnathochilarium, ventral view; C, legs 1, caudal view; D, legs 2, caudal view; E, anterior gonopods, caudal view; F, posterior gonopods, frontal view; G, same, caudal view. Scale bar: A, B, 0.3 mm; C-G, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5392099" httpUri="https://zenodo.org/record/5392099/files/figure.png" pageId="15" pageNumber="22">Fig. 9B</figureCitation>
|
||
) or undivided (n = 1) (
|
||
<figureCitation id="13666EEBA2203047FEBC0117FE98F9F5" box="[296,382,1610,1636]" captionStart="FIG" captionStartId="12.[113,124,1318,1334]" captionTargetBox="[121,1200,215,1260]" captionTargetId="figure-36@12.[390,661,595,786]" captionTargetPageId="12" captionText="FIG. 7. — Glyphiulus subgranulatus n. sp., ♀ paratype:A, head, ventral view; B, collum, dorsal view; C, middle part of body, dorsal view; D, same, lateral view; E, caudal part of body, lateral view; F, same, dorsal view. Scale bars: A, 0.1 mm; B, 0.2 mm, C-F, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5392095" httpUri="https://zenodo.org/record/5392095/files/figure.png" pageId="15" pageNumber="22">Fig. 7A</figureCitation>
|
||
), 11-18 ocelli, antennae (
|
||
<figureCitation id="13666EEBA2203047FFEE0137FF29F915" box="[122,207,1642,1668]" captionStart="FIG" captionStartId="13.[113,124,1024,1040]" captionTargetBox="[120,1200,215,977]" captionTargetId="figure-32@13.[390,663,596,787]" captionTargetPageId="13" captionText="FIG. 8. — Glyphiulus subgranulatus n. sp., ♂ and ♀ paratypes: A, anterior part of body, lateral view; B, caudal part of body, ventral view; C, midbody segment section (a little damaged), caudal view; D, male legs 1, caudal view. Scale bars: A, C, 0.5 mm; B, 0.2 mm; D, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5392097" httpUri="https://zenodo.org/record/5392097/files/figure.png" pageId="15" pageNumber="22">Figs 8A</figureCitation>
|
||
;
|
||
<figureCitation id="13666EEBA2203047FF4C0137FF1DF915" box="[216,251,1642,1668]" captionStart="FIG" captionStartId="14.[113,124,963,979]" captionTargetBox="[113,1209,218,907]" captionTargetId="figure-18@14.[74,1220,182,921]" captionTargetPageId="14" captionText="FIG. 9. — Glyphiulus subgranulatus n. sp., ♂ paratype: A, antenna; B, gnathochilarium, ventral view; C, legs 1, caudal view; D, legs 2, caudal view; E, anterior gonopods, caudal view; F, posterior gonopods, frontal view; G, same, caudal view. Scale bar: A, B, 0.3 mm; C-G, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5392099" httpUri="https://zenodo.org/record/5392099/files/figure.png" pageId="15" pageNumber="22">9A</figureCitation>
|
||
), the carinotaxy formulae (
|
||
<figureCitation id="13666EEBA2203047FD870137FDA9F915" box="[531,591,1642,1668]" captionStart="FIG" captionStartId="12.[113,124,1318,1334]" captionTargetBox="[121,1200,215,1260]" captionTargetId="figure-36@12.[390,661,595,786]" captionTargetPageId="12" captionText="FIG. 7. — Glyphiulus subgranulatus n. sp., ♀ paratype:A, head, ventral view; B, collum, dorsal view; C, middle part of body, dorsal view; D, same, lateral view; E, caudal part of body, lateral view; F, same, dorsal view. Scale bars: A, 0.1 mm; B, 0.2 mm, C-F, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5392095" httpUri="https://zenodo.org/record/5392095/files/figure.png" pageId="15" pageNumber="22">Fig. 7</figureCitation>
|
||
B-F), claw, posterior gonopod structure (
|
||
<figureCitation id="13666EEBA2203047FE4301D7FDA6F935" box="[471,576,1674,1700]" captionStart="FIG" captionStartId="14.[113,124,963,979]" captionTargetBox="[113,1209,218,907]" captionTargetId="figure-18@14.[74,1220,182,921]" captionTargetPageId="14" captionText="FIG. 9. — Glyphiulus subgranulatus n. sp., ♂ paratype: A, antenna; B, gnathochilarium, ventral view; C, legs 1, caudal view; D, legs 2, caudal view; E, anterior gonopods, caudal view; F, posterior gonopods, frontal view; G, same, caudal view. Scale bar: A, B, 0.3 mm; C-G, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5392099" httpUri="https://zenodo.org/record/5392099/files/figure.png" pageId="15" pageNumber="22">Fig. 9F, G</figureCitation>
|
||
), male legs 1 and 2 (
|
||
<figureCitation id="13666EEBA2203047FCBC0291FC98FA77" box="[808,894,1484,1510]" captionStart="FIG" captionStartId="13.[113,124,1024,1040]" captionTargetBox="[120,1200,215,977]" captionTargetId="figure-32@13.[390,663,596,787]" captionTargetPageId="13" captionText="FIG. 8. — Glyphiulus subgranulatus n. sp., ♂ and ♀ paratypes: A, anterior part of body, lateral view; B, caudal part of body, ventral view; C, midbody segment section (a little damaged), caudal view; D, male legs 1, caudal view. Scale bars: A, C, 0.5 mm; B, 0.2 mm; D, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5392097" httpUri="https://zenodo.org/record/5392097/files/figure.png" pageId="15" pageNumber="22">Figs 8D</figureCitation>
|
||
;
|
||
<figureCitation id="13666EEBA2203047FC1C0291FC2AFA77" box="[904,972,1484,1510]" captionStart="FIG" captionStartId="14.[113,124,963,979]" captionTargetBox="[113,1209,218,907]" captionTargetId="figure-18@14.[74,1220,182,921]" captionTargetPageId="14" captionText="FIG. 9. — Glyphiulus subgranulatus n. sp., ♂ paratype: A, antenna; B, gnathochilarium, ventral view; C, legs 1, caudal view; D, legs 2, caudal view; E, anterior gonopods, caudal view; F, posterior gonopods, frontal view; G, same, caudal view. Scale bar: A, B, 0.3 mm; C-G, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5392099" httpUri="https://zenodo.org/record/5392099/files/figure.png" pageId="15" pageNumber="22">9C, D</figureCitation>
|
||
), telson (
|
||
<figureCitation id="13666EEBA2203047FBBE0291FB9CFA77" box="[1066,1146,1484,1510]" captionStart="FIG" captionStartId="13.[113,124,1024,1040]" captionTargetBox="[120,1200,215,977]" captionTargetId="figure-32@13.[390,663,596,787]" captionTargetPageId="13" captionText="FIG. 8. — Glyphiulus subgranulatus n. sp., ♂ and ♀ paratypes: A, anterior part of body, lateral view; B, caudal part of body, ventral view; C, midbody segment section (a little damaged), caudal view; D, male legs 1, caudal view. Scale bars: A, C, 0.5 mm; B, 0.2 mm; D, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5392097" httpUri="https://zenodo.org/record/5392097/files/figure.png" pageId="15" pageNumber="22">Fig. 8B</figureCitation>
|
||
), etc., but body size much larger (see above); segments compressed laterally (
|
||
<figureCitation id="13666EEBA2203047FC020156FC08F9B4" box="[918,1006,1547,1573]" captionStart="FIG" captionStartId="13.[113,124,1024,1040]" captionTargetBox="[120,1200,215,977]" captionTargetId="figure-32@13.[390,663,596,787]" captionTargetPageId="13" captionText="FIG. 8. — Glyphiulus subgranulatus n. sp., ♂ and ♀ paratypes: A, anterior part of body, lateral view; B, caudal part of body, ventral view; C, midbody segment section (a little damaged), caudal view; D, male legs 1, caudal view. Scale bars: A, C, 0.5 mm; B, 0.2 mm; D, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5392097" httpUri="https://zenodo.org/record/5392097/files/figure.png" pageId="15" pageNumber="22">Fig. 8C</figureCitation>
|
||
); crests on collum (7+m+7 at caudal margin) evidently reduced in size, almost obsolete anteromedially (
|
||
<figureCitation id="13666EEBA2203047FBBB0117FB64F9F5" box="[1071,1154,1610,1636]" captionStart="FIG" captionStartId="12.[113,124,1318,1334]" captionTargetBox="[121,1200,215,1260]" captionTargetId="figure-36@12.[390,661,595,786]" captionTargetPageId="12" captionText="FIG. 7. — Glyphiulus subgranulatus n. sp., ♀ paratype:A, head, ventral view; B, collum, dorsal view; C, middle part of body, dorsal view; D, same, lateral view; E, caudal part of body, lateral view; F, same, dorsal view. Scale bars: A, 0.1 mm; B, 0.2 mm, C-F, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5392095" httpUri="https://zenodo.org/record/5392095/files/figure.png" pageId="15" pageNumber="22">Figs 7B</figureCitation>
|
||
;
|
||
<figureCitation id="13666EEBA2203047FB180117FB48F9F5" box="[1164,1198,1610,1636]" captionStart="FIG" captionStartId="13.[113,124,1024,1040]" captionTargetBox="[120,1200,215,977]" captionTargetId="figure-32@13.[390,663,596,787]" captionTargetPageId="13" captionText="FIG. 8. — Glyphiulus subgranulatus n. sp., ♂ and ♀ paratypes: A, anterior part of body, lateral view; B, caudal part of body, ventral view; C, midbody segment section (a little damaged), caudal view; D, male legs 1, caudal view. Scale bars: A, C, 0.5 mm; B, 0.2 mm; D, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5392097" httpUri="https://zenodo.org/record/5392097/files/figure.png" pageId="15" pageNumber="22">8A</figureCitation>
|
||
); legs somewhat longer (about 4/5 as long as body height); anterior gonopods with telopodites more slender, while the coxosternum has much shorter distomedian outgrowths and acute distolateral corners (
|
||
<figureCitation id="13666EEBA23F3058FF4401D7FEC2F935" box="[208,292,1674,1700]" captionStart="FIG" captionStartId="14.[113,124,963,979]" captionTargetBox="[113,1209,218,907]" captionTargetId="figure-18@14.[74,1220,182,921]" captionTargetPageId="14" captionText="FIG. 9. — Glyphiulus subgranulatus n. sp., ♂ paratype: A, antenna; B, gnathochilarium, ventral view; C, legs 1, caudal view; D, legs 2, caudal view; E, anterior gonopods, caudal view; F, posterior gonopods, frontal view; G, same, caudal view. Scale bar: A, B, 0.3 mm; C-G, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5392099" httpUri="https://zenodo.org/record/5392099/files/figure.png" pageId="16" pageNumber="23">Fig. 9E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF2222E6A23F3058FFE502BAFD52F9B4" ID-DOI="http://doi.org/10.5281/zenodo.5392103" ID-Zenodo-Dep="5392103" httpUri="https://zenodo.org/record/5392103/files/figure.png" pageId="16" pageNumber="23" startId="16.[113,124,1511,1527]" targetBox="[123,1201,216,1478]" targetPageId="16">
|
||
<paragraph id="8BE2726EA23F3058FFE502BAFD52F9B4" blockId="16.[113,1210,1510,1573]" pageId="16" pageNumber="23">
|
||
FIG. 11. —
|
||
<taxonomicName id="4C5D09EDA23F3058FF5902BBFE43FA66" authority="Golovatch & Geoffroy & Mauriès & Spiegel, 2007" authorityName="Golovatch & Geoffroy & Mauriès & Spiegel" authorityYear="2007" box="[205,421,1510,1527]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="16" pageNumber="23" phylum="Arthropoda" rank="species" species="paragranulatus" status="sp. nov.">
|
||
<emphasis id="B929AE7CA23F3058FF5902BBFE43FA66" box="[205,421,1510,1527]" italics="true" pageId="16" pageNumber="23">Glyphiulus paragranulatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A1307A23F3058FE3F02BAFE3DFA66" box="[427,475,1511,1527]" pageId="16" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
|
||
, ♂ and ♀ paratypes:
|
||
<emphasis id="B929AE7CA23F3058FD1B02BBFD7AFA66" bold="true" box="[655,668,1510,1527]" pageId="16" pageNumber="23">A</emphasis>
|
||
, caudal part of body, ventral view;
|
||
<emphasis id="B929AE7CA23F3058FC5702BAFC36FA66" bold="true" box="[963,976,1511,1527]" pageId="16" pageNumber="23">B</emphasis>
|
||
, midbody segment section, caudal view;
|
||
<emphasis id="B929AE7CA23F3058FF4B02A0FF0BF99F" bold="true" box="[223,237,1533,1550]" pageId="16" pageNumber="23">C</emphasis>
|
||
, midbody leg;
|
||
<emphasis id="B929AE7CA23F3058FEFE02A3FE9EF99F" bold="true" box="[362,376,1534,1550]" pageId="16" pageNumber="23">D</emphasis>
|
||
, claw;
|
||
<emphasis id="B929AE7CA23F3058FE2702A3FE59F99F" bold="true" box="[435,447,1534,1550]" pageId="16" pageNumber="23">E</emphasis>
|
||
, posterior gonopods
|
||
<emphasis id="B929AE7CA23F3058FDE002A3FD43F99F" box="[628,677,1534,1550]" italics="true" pageId="16" pageNumber="23">in situ</emphasis>
|
||
in gonopod opening, frontal view;
|
||
<emphasis id="B929AE7CA23F3058FC5C02A3FC35F99F" bold="true" box="[968,979,1534,1550]" pageId="16" pageNumber="23">F</emphasis>
|
||
, anterior gonopods, caudal view. Scale bars: A, 0.2 mm; B, 0.5 mm; C, E, F, 0.1 mm; D, 0.02 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C34721E5A23F3058FD310117FB77F935" pageId="16" pageNumber="23" type="discussion">
|
||
<paragraph id="8BE2726EA23F3058FD310117FCEFF9F3" blockId="16.[675,1209,1610,1700]" box="[677,777,1610,1636]" pageId="16" pageNumber="23">REMARKS</paragraph>
|
||
<paragraph id="8BE2726EA23F3058FD310137FB77F935" blockId="16.[675,1209,1610,1700]" pageId="16" pageNumber="23">In the absence of evident troglomorphic traits, this species can only be considered as troglophilic.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |