292 lines
34 KiB
XML
292 lines
34 KiB
XML
<document id="63CB6EC03A1DDBCFA26B049A3125A97C" ID-DOI="10.5281/zenodo.178388" ID-GBIF-Dataset="5468dffa-e7d5-45e3-9935-7d45a69f1de3" ID-ISSN="1175-5326" ID-Zenodo-Dep="178388" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459892172593" checkinUser="plazi" docAuthor="Haase, Martin, Wilke, Thomas & Mildner, Paul" docDate="2007" docId="03E2F349FF9C7403FF0CCF6FFF44FE80" docLanguage="en" docName="zt01563p016.pdf" docOrigin="Zootaxa 1563" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Bythinella angelitae" docType="treatment" docVersion="9" lastPageNumber="11" masterDocId="FFDB8B31FF947409FF9BCE58FFA6FFAA" masterDocTitle="Identifying species of Bythinella (Caenogastropoda: Rissooidea): A plea for an integrative approach" masterLastPageNumber="16" masterPageNumber="1" pageNumber="9" updateTime="1698228136151" updateUser="plazi">
|
||
<mods:mods id="B397ECC81809C84A8189BE2BF25ACC00" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="8447928D05C503ACE6BACCBD6AE73AC3">
|
||
<mods:title id="E294130CA5A3A47962B3310E0D17870C">Identifying species of Bythinella (Caenogastropoda: Rissooidea): A plea for an integrative approach</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="3A8ECCE1C13BD02C55B9536CB8BCFEB5" type="personal">
|
||
<mods:role id="0117EE1E54715DDF2A6FE1A6C2BB2F83">
|
||
<mods:roleTerm id="A45F810090FCD100F155CE8BC21E3E65">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="3162DE5C02EF5C5BC1B73DBF76B3C912">Haase, Martin</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="32BCBC1CA3346924102765663ACFA5D6" type="personal">
|
||
<mods:role id="BE69C66298252FD86FA56B4758EFCB78">
|
||
<mods:roleTerm id="0158320D7FA30B2BDFFC87662D532996">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="D8E1FFD061594F43E105424232CD3FDF">Wilke, Thomas</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="3375B714D3229A0DFFD27927F958D105" type="personal">
|
||
<mods:role id="032F38466CA4C36577F8EF488C3E2456">
|
||
<mods:roleTerm id="8615E65C629ECE53D06A9696FCD5E746">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="8924CFA7058A7EA710B2677F9E1EE671">Mildner, Paul</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="1485AE40EA7F80A8486A930EFCD783CD">text</mods:typeOfResource>
|
||
<mods:relatedItem id="A08990A5B87B62AFA845171CC6BA0611" type="host">
|
||
<mods:titleInfo id="8C34BB58EF87FACD5913407EA3069B9A">
|
||
<mods:title id="0F89DC8E824F34EA9282736D758F3147">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="4CE1FB47C62AAFF33091A68FA4DF0C11">
|
||
<mods:date id="75E8A35810B102FA2BF3595DC3406A64">2007</mods:date>
|
||
<mods:detail id="3222B0485550FD78E6D872CE69A05077" type="volume">
|
||
<mods:number id="960EC0FA3881B7FF34DF61842AB90926">1563</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="033ACE8F5993D50962180AE043E6D314" unit="page">
|
||
<mods:start id="3AED9903072DE3CC426C92BB188AA80E">1</mods:start>
|
||
<mods:end id="C63E9C5A2DF071257FC7C9B194030AFB">16</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="97AE097413F3F504F2D21C2E45764ED7">journal article</mods:classification>
|
||
<mods:identifier id="739DCDD45EE67506CF3830AF6807D774" type="DOI">10.5281/zenodo.178388</mods:identifier>
|
||
<mods:identifier id="3E410DE8F625670AD19BFFF511366B13" type="GBIF-Dataset">5468dffa-e7d5-45e3-9935-7d45a69f1de3</mods:identifier>
|
||
<mods:identifier id="C6EB95B68D1018F26AD016046E7CCA07" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="09B7ABEE0558A4EC1902D737526B67EB" type="Zenodo-Dep">178388</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03E2F349FF9C7403FF0CCF6FFF44FE80" ID-DOI="http://doi.org/10.5281/zenodo.5629037" ID-GBIF-Taxon="119372116" ID-Zenodo-Dep="5629037" LSID="urn:lsid:plazi:treatment:03E2F349FF9C7403FF0CCF6FFF44FE80" httpUri="http://treatment.plazi.org/id/03E2F349FF9C7403FF0CCF6FFF44FE80" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9">
|
||
<subSubSection id="C35111D4FF9C7401FF0CCF6FFB7CFE33" pageId="8" pageNumber="9" type="nomenclature">
|
||
<paragraph id="8BF4425FFF9C7401FF0CCF6FFDACFEFB" blockId="8.[151,522,311,337]" box="[151,522,311,337]" pageId="8" pageNumber="9">
|
||
<heading id="D0BCF533FF9C7401FF0CCF6FFDACFEFB" bold="true" box="[151,522,311,337]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
|
||
<emphasis id="B93F9E4DFF9C7401FF0CCF6FFDACFEFB" bold="true" box="[151,522,311,337]" pageId="8" pageNumber="9">
|
||
<taxonomicName id="4C4B39DCFF9C7401FF0CCF6FFE2FFEFB" box="[151,393,311,337]" class="Gastropoda" family="Hydrobiidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="angelitae">
|
||
<emphasis id="B93F9E4DFF9C7401FF0CCF6FFE2FFEFB" bold="true" box="[151,393,311,337]" italics="true" pageId="8" pageNumber="9">Bythinella angelitae</emphasis>
|
||
</taxonomicName>
|
||
nom. nov.
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BF4425FFF9C7401FF0CCFDAFB7CFE33" blockId="8.[151,1242,386,409]" box="[151,1242,386,409]" pageId="8" pageNumber="9">
|
||
Synonymy:
|
||
<taxonomicName id="4C4B39DCFF9C7401FE83CFDAFD96FE33" authority="Ziegler" authorityName="Ziegler" box="[280,560,386,409]" class="Gastropoda" family="Assimineidae" genus="Paludinella" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="opaca">
|
||
<emphasis id="B93F9E4DFF9C7401FE83CFDAFE7BFE33" box="[280,477,386,409]" italics="true" pageId="8" pageNumber="9">Paludinella opaca</emphasis>
|
||
Ziegler
|
||
</taxonomicName>
|
||
—
|
||
<bibRefCitation id="EFDA3FAEFF9C7401FDC3CFDAFCB4FE33" author="Frauenfeld" box="[600,786,386,409]" pageId="8" pageNumber="15" refString="Frauenfeld, G. v. (1857) Uber die Paludinen aus der Gruppe der Pal. viridis Poir. Sitzungsberichte der Akademie der Wissenschaften Wien, 22, 569 - 578." type="journal article" year="1857">Frauenfeld, 1857</bibRefCitation>
|
||
: 576, fig. 6, non
|
||
<bibRefCitation id="EFDA3FAEFF9C7401FC5DCFDAFB20FE33" author="Gallenstein" box="[966,1158,386,409]" pageId="8" pageNumber="15" refString="Gallenstein, M. v. (1848) Systematisches Verzeichnis der in der Provinz Karnten bisher entdeckten Land- & Susswasser- Conchylien, mit Angabe der wichtigsten Fundorte, nebst einer kurzen Anleitung fur angehende Conchylien-Sammler. Joseph Blasnik, Laibach, 105 pp." type="book" year="1848">Gallenstein, 1848</bibRefCitation>
|
||
: 18–19.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35111D4FF9C7401FF0CCF92FDAAFCE6" pageId="8" pageNumber="9" type="materials_examined">
|
||
<paragraph id="8BF4425FFF9C7401FF0CCF92FEFBFD2E" blockId="8.[151,1437,458,1288]" pageId="8" pageNumber="9">
|
||
<emphasis id="B93F9E4DFF9C7401FF0CCF92FEFEFE4E" bold="true" box="[151,344,458,484]" pageId="8" pageNumber="9">
|
||
<typeStatus id="54F0FCFDFF9C7401FF0CCF92FF72FE4E" box="[151,212,458,484]" pageId="8" pageNumber="9">Type</typeStatus>
|
||
material:
|
||
</emphasis>
|
||
<typeStatus id="54F0FCFDFF9C7401FEF1CF92FE79FE4E" box="[362,479,458,484]" pageId="8" pageNumber="9" type="lectotype">Lectotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4B39DCFF9C7401FD81CF92FC47FE4E" authority="Frauenfeld, 1857" authorityName="Frauenfeld" authorityYear="1857" box="[538,993,458,484]" class="Gastropoda" family="Assimineidae" genus="Paludinella" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="opaca">
|
||
<emphasis id="B93F9E4DFF9C7401FD81CF92FD5AFE49" box="[538,764,458,483]" italics="true" pageId="8" pageNumber="9">Paludinella opaca</emphasis>
|
||
<bibRefCitation id="EFDA3FAEFF9C7401FC97CF92FC47FE4E" author="Frauenfeld" box="[780,993,458,484]" pageId="8" pageNumber="15" refString="Frauenfeld, G. v. (1857) Uber die Paludinen aus der Gruppe der Pal. viridis Poir. Sitzungsberichte der Akademie der Wissenschaften Wien, 22, 569 - 578." type="journal article" year="1857">Frauenfeld, 1857</bibRefCitation>
|
||
</taxonomicName>
|
||
, here designated, and
|
||
<typeStatus id="54F0FCFDFF9C7401FA97CF92FAD4FE4E" box="[1292,1394,458,484]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4B39DCFF9C7401FF0CCFAAFE25FDA1" box="[151,387,498,523]" class="Gastropoda" family="Hydrobiidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="angelitae">
|
||
<emphasis id="B93F9E4DFF9C7401FF0CCFAAFE25FDA1" box="[151,387,498,523]" italics="true" pageId="8" pageNumber="9">Bythinella angelitae</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<collectionCode id="ED5ADA9AFF9C7401FE08CFAAFE55FDA6" box="[403,499,498,524]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="9">NHMW</collectionCode>
|
||
(Naturhistorisches Museum Wien) 103753 (Veldes See = Lake Bled,
|
||
<collectingCountry id="F35C02CFFF9C7401FAB8CFAAFA29FDA6" box="[1315,1423,498,524]" name="Slovenia" pageId="8" pageNumber="9">Slovenia</collectingCountry>
|
||
);
|
||
<typeStatus id="54F0FCFDFF9C7401FF0CCC42FEE6FD9E" box="[151,320,538,564]" pageId="8" pageNumber="9" type="paralectotype">paralectotypes</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4B39DCFF9C7401FEF2CC42FD06FD9E" authority="Frauenfeld, 1857" authorityName="Frauenfeld" authorityYear="1857" box="[361,672,538,564]" class="Gastropoda" family="Assimineidae" genus="Paludinella" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="opaca">
|
||
<emphasis id="B93F9E4DFF9C7401FEF2CC42FE68FD99" box="[361,462,538,563]" italics="true" pageId="8" pageNumber="9">P. opaca</emphasis>
|
||
<bibRefCitation id="EFDA3FAEFF9C7401FE4DCC42FD06FD9E" author="Frauenfeld" box="[470,672,538,564]" pageId="8" pageNumber="15" refString="Frauenfeld, G. v. (1857) Uber die Paludinen aus der Gruppe der Pal. viridis Poir. Sitzungsberichte der Akademie der Wissenschaften Wien, 22, 569 - 578." type="journal article" year="1857">Frauenfeld, 1857</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<typeStatus id="54F0FCFDFF9C7401FD41CC42FCECFD9E" box="[730,842,538,564]" pageId="8" pageNumber="9" type="paratype">paratypes</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4B39DCFF9C7401FCE8CC42FBA4FD99" box="[883,1026,538,563]" class="Gastropoda" family="Hydrobiidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="angelitae">
|
||
<emphasis id="B93F9E4DFF9C7401FCE8CC42FBA4FD99" box="[883,1026,538,563]" italics="true" pageId="8" pageNumber="9">B. angelitae</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<collectionCode id="ED5ADA9AFF9C7401FB89CC42FBD4FD9E" box="[1042,1138,538,564]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="9">NHMW</collectionCode>
|
||
103754 (4, Veldes See = Lake Bled,
|
||
<collectingCountry id="F35C02CFFF9C7401FE84CC1AFE2DFDF6" box="[287,395,578,604]" name="Slovenia" pageId="8" pageNumber="9">Slovenia</collectingCountry>
|
||
);
|
||
<collectionCode id="ED5ADA9AFF9C7401FE04CC1AFE59FDF6" box="[415,511,578,604]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="9">NHMW</collectionCode>
|
||
103758 (three specimens, no locality data);
|
||
<collectionCode id="ED5ADA9AFF9C7401FB93CC1AFBCEFDF6" box="[1032,1128,578,604]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="9">NHMW</collectionCode>
|
||
103759 (three specimens, no locality data).
|
||
</paragraph>
|
||
<paragraph id="8BF4425FFF9C7401FF5DCCCAFDC7FD56" blockId="8.[151,1437,458,1288]" pageId="8" pageNumber="9">
|
||
<emphasis id="B93F9E4DFF9C7401FF5DCCCAFED7FD06" bold="true" box="[198,369,658,684]" pageId="8" pageNumber="9">
|
||
<typeStatus id="54F0FCFDFF9C7401FF5DCCCAFEA2FD06" box="[198,260,658,684]" pageId="8" pageNumber="9">Type</typeStatus>
|
||
locality:
|
||
</emphasis>
|
||
Through our designation of the
|
||
<typeStatus id="54F0FCFDFF9C7401FD63CCCAFCC5FD06" box="[760,867,658,684]" pageId="8" pageNumber="9" type="lectotype">lectotype</typeStatus>
|
||
from the Slovenian sample, the
|
||
<typeStatus id="54F0FCFDFF9C7401FB77CCCAFAB9FD06" box="[1260,1311,658,684]" pageId="8" pageNumber="9">type</typeStatus>
|
||
locality is restricted to Veldes See = Lake Bled, which, since the species is not lacustrine (
|
||
<bibRefCitation id="EFDA3FAEFF9C7401FBA8CCE2FB54FD7E" author="Radoman" box="[1075,1266,698,724]" pageId="8" pageNumber="16" refString="Radoman, P. (1976) Speciation within the family Bythinellidae on the Balkans and Asia Minor. Zeitschrift fur zoologische Systematik und Evolutionsforschung, 14, 130 - 152." type="journal article" year="1976">Radoman, 1976</bibRefCitation>
|
||
), must be conceived as “the area around Lake Bled”.
|
||
</paragraph>
|
||
<paragraph id="8BF4425FFF9C7401FF5DCD52FDAAFCE6" blockId="8.[151,1437,458,1288]" pageId="8" pageNumber="9">
|
||
<emphasis id="B93F9E4DFF9C7401FF5DCD52FE60FC8E" bold="true" box="[198,454,778,804]" pageId="8" pageNumber="9">Additional material:</emphasis>
|
||
spring in Tscheppaschlucht, southern Carinthia,
|
||
<geoCoordinate id="EE7F2498FF9C7401FB9ACD52FB23FC8E" box="[1025,1157,778,804]" direction="north" orientation="latitude" pageId="8" pageNumber="9" precision="5" value="46.4837">46.4837°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE7F2498FF9C7401FB0BCD52FAB6FC8E" box="[1168,1296,778,804]" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="5" value="14.2632">14.2632°E</geoCoordinate>
|
||
,
|
||
<quantity id="4CB3EFBAFF9C7401FA80CD52FAC3FC8E" box="[1307,1381,778,804]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="8" pageNumber="9" unit="m" value="680.0">680 m</quantity>
|
||
asl., leg. P. Mildner,
|
||
<date id="FFF5649FFF9C7401FECACD6AFDA1FCE6" box="[337,519,818,844]" pageId="8" pageNumber="9" value="2006-03-30">30 March 2006</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35111D4FF9C7401FF5DCD02FAE2FCDE" box="[198,1348,858,884]" pageId="8" pageNumber="9" type="etymology">
|
||
<paragraph id="8BF4425FFF9C7401FF5DCD02FAE2FCDE" blockId="8.[151,1437,458,1288]" box="[198,1348,858,884]" pageId="8" pageNumber="9">
|
||
<emphasis id="B93F9E4DFF9C7401FF5DCD02FEF1FCDE" bold="true" box="[198,343,858,884]" pageId="8" pageNumber="9">Etymology:</emphasis>
|
||
This species is dedicated to the first author’s wife Angela “Angelita” Schmitz-Ornés.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35111D4FF9C7400FF5DCDDAFCC8FAE1" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="description">
|
||
<paragraph id="8BF4425FFF9C7401FF5DCDDAFB98FBCE" blockId="8.[151,1437,458,1288]" pageId="8" pageNumber="9">
|
||
<emphasis id="B93F9E4DFF9C7401FF5DCDDAFEC7FC36" bold="true" box="[198,353,898,924]" pageId="8" pageNumber="9">Description:</emphasis>
|
||
Shell (
|
||
<figureCitation id="13705EDAFF9C7401FE22CDDAFDAEFC36" box="[441,520,898,924]" captionStart="FIGURE 2" captionStartId="8.[151,255,1863,1887]" captionTargetBox="[151,1435,1344,1840]" captionTargetId="figure@8.[151,1436,1344,1842]" captionTargetPageId="8" captionText="FIGURE 2. Shells. A. B. angelitae, holotype (and lectotype of Paludinella opaca Frauenfeld, 1857); B, C. B. angelitae, paratypes (and paralectotypes of P. opaca Frauenfeld, 1857); D. B. robiciana, topotype; E. B. opaca, (topotype of Paludina schmidtii Küster, 1852). Scale bar = 2 mm." httpUri="https://zenodo.org/record/178390/files/figure.png" pageId="8" pageNumber="9">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation id="13705EDAFF9C7401FD8DCDDAFD81FC36" box="[534,551,898,924]" captionStart="FIGURE 3" captionStartId="9.[151,255,745,769]" captionTargetBox="[264,1322,163,722]" captionTargetId="figure@9.[264,1323,163,723]" captionTargetPageId="9" captionText="FIGURE 3. Shells of Carinthian populations. A – G. B. opaca; H. B. angelitae, Tscheppaschlucht. A. Bärental; B. Farchtnersee; C. Granitztal St. Paul; D. Sand Grafenstein; E. Oberdrauburger Bach; F. Niedermarkt; G. Podkraj." httpUri="https://zenodo.org/record/178391/files/figure.png" pageId="8" pageNumber="9">3</figureCitation>
|
||
;
|
||
<tableCitation id="C6C977E4FF9C7401FDACCDDAFD37FC36" box="[567,657,898,924]" captionStart="TABLE 2" captionStartId="4.[151,239,394,418]" captionTargetBox="[159,1430,675,911]" captionTargetPageId="4" captionText="TABLE 2. Shell dimensions. ah, aperture height; aw, aperture width; bww, width of body (= penultimate) whorl; cv, coefficient of variation corrected for sample size; f, females, m, males; max, maximum; min, minimum; N, number of specimens measured (not necessarily identical with number of sexed specimens); sd, standard deviation, sh, shell height; sw, shell width; w, number of whorls. Measurements in mm." httpUri="http://table.plazi.org/id/DF3412D7FF90740DFF0CCFD2FCBAFDA0" pageId="8" pageNumber="9" tableUuid="DF3412D7FF90740DFF0CCFD2FCBAFDA0">Table 2</tableCitation>
|
||
) conical to subcylindrical with typical blunt apex,
|
||
<typeStatus id="54F0FCFDFF9C7401FB63CDDAFAF8FC36" box="[1272,1374,898,924]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
|
||
<quantity id="4CB3EFBAFF9C7401FAF2CDDAFF63FC6E" metricMagnitude="-3" metricUnit="m" metricValue="3.18" pageId="8" pageNumber="9" unit="mm" value="3.18">3.18 mm</quantity>
|
||
high and
|
||
<quantity id="4CB3EFBAFF9C7401FEA6CDF2FE01FC6E" box="[317,423,938,964]" metricMagnitude="-3" metricUnit="m" metricValue="1.71" pageId="8" pageNumber="9" unit="mm" value="1.71">1.71 mm</quantity>
|
||
wide, with 4 whorls, shells from the Tscheppaschlucht 2.61 ±
|
||
<quantity id="4CB3EFBAFF9C7401FB0FCDF2FB58FC6E" box="[1172,1278,938,964]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="9" unit="mm" value="0.08">0.08 mm</quantity>
|
||
(range
|
||
<quantity id="4CB3EFBAFF9C7401FAC1CDF2FEA5FC46" metricMagnitude="-3" metricUnit="m" metricValue="2.62" metricValueMax="2.76" metricValueMin="2.48" pageId="8" pageNumber="9" unit="mm" value="2.62" valueMax="2.76" valueMin="2.48">2.48– 2.76 mm</quantity>
|
||
) high and 1.67 ±
|
||
<quantity id="4CB3EFBAFF9C7401FE40CD8AFDE0FC46" box="[475,582,978,1004]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="9" unit="mm" value="0.06">0.06 mm</quantity>
|
||
(range
|
||
<quantity id="4CB3EFBAFF9C7401FD38CD8AFCF7FC46" box="[675,849,978,1004]" metricMagnitude="-3" metricUnit="m" metricValue="1.68" metricValueMax="1.8" metricValueMin="1.56" pageId="8" pageNumber="9" unit="mm" value="1.68" valueMax="1.8" valueMin="1.56">1.56–1.80 mm</quantity>
|
||
) wide, with 3 3/8–3 7/8 convex whorls of which protoconch comprises 1.1–1.2 whorls with delicately pitted spiral lines, teleoconch without structure apart from growth lines; aperture ovate with typically pronounced angle posteriorly; umbilicus largely covered; operculum corneous, yellow, thin, ellipsoidal, paucispiral, nucleus submarginal.
|
||
</paragraph>
|
||
<paragraph id="8BF4425FFF9C7401FF5DCA2AFC6DFAAE" blockId="8.[151,1437,458,1288]" pageId="8" pageNumber="9">Tentacles without conspicuous ciliation; epidermis black except mantle edge; ctenidium with 21–26 gill filaments, connected to pericardium with short vessel, osphradium under or slightly behind middle of ctenidium, elongate, about 1/3 or somewhat less of gill length, kidney may penetrate slightly into the mantle roof; radula formula R: 4–5 1 4–5/1-2 1–2, L: 3 1 4, M1: 17–23, M2: 27–31.</paragraph>
|
||
<caption id="DF3412D7FF9C7401FF0CC91FFD3EF809" httpUri="https://zenodo.org/record/178390/files/figure.png" pageId="8" pageNumber="9" targetBox="[151,1435,1344,1840]" targetPageId="8">
|
||
<paragraph id="8BF4425FFF9C7401FF0CC91FFD3EF809" blockId="8.[151,1436,1863,1956]" pageId="8" pageNumber="9">
|
||
<emphasis id="B93F9E4DFF9C7401FF0CC91FFEBFF8F5" bold="true" box="[151,281,1863,1887]" pageId="8" pageNumber="9">FIGURE 2.</emphasis>
|
||
Shells. A.
|
||
<taxonomicName id="4C4B39DCFF9C7401FE14C91FFDB6F8F4" box="[399,528,1863,1886]" class="Gastropoda" family="Hydrobiidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="angelitae">
|
||
<emphasis id="B93F9E4DFF9C7401FE14C91FFDB6F8F4" box="[399,528,1863,1886]" italics="true" pageId="8" pageNumber="9">B. angelitae</emphasis>
|
||
</taxonomicName>
|
||
, holotype (and lectotype of
|
||
<taxonomicName id="4C4B39DCFF9C7401FCA6C91FFB64F8F4" authority="Frauenfeld, 1857" authorityName="Frauenfeld" authorityYear="1857" box="[829,1218,1863,1886]" class="Gastropoda" family="Assimineidae" genus="Paludinella" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="opaca">
|
||
<emphasis id="B93F9E4DFF9C7401FCA6C91FFBA4F8F4" box="[829,1026,1863,1886]" italics="true" pageId="8" pageNumber="9">Paludinella opaca</emphasis>
|
||
Frauenfeld, 1857
|
||
</taxonomicName>
|
||
); B, C.
|
||
<taxonomicName id="4C4B39DCFF9C7401FA8FC91FFA33F8F4" box="[1300,1429,1863,1886]" class="Gastropoda" family="Hydrobiidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="angelitae">
|
||
<emphasis id="B93F9E4DFF9C7401FA8FC91FFA33F8F4" box="[1300,1429,1863,1886]" italics="true" pageId="8" pageNumber="9">B. angelitae</emphasis>
|
||
</taxonomicName>
|
||
, paratypes (and paralectotypes of
|
||
<taxonomicName id="4C4B39DCFF9C7401FE62C932FCB1F82B" authority="Frauenfeld, 1857" authorityName="Frauenfeld" authorityYear="1857" box="[505,791,1898,1921]" class="Gastropoda" family="Assimineidae" genus="Paludinella" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="opaca">
|
||
<emphasis id="B93F9E4DFF9C7401FE62C932FDF3F82B" box="[505,597,1898,1921]" italics="true" pageId="8" pageNumber="9">P. opaca</emphasis>
|
||
Frauenfeld, 1857
|
||
</taxonomicName>
|
||
); D.
|
||
<taxonomicName id="4C4B39DCFF9C7401FCD7C932FC77F82B" box="[844,977,1898,1921]" class="Gastropoda" family="Hydrobiidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="robiciana">
|
||
<emphasis id="B93F9E4DFF9C7401FCD7C932FC77F82B" box="[844,977,1898,1921]" italics="true" pageId="8" pageNumber="9">B. robiciana</emphasis>
|
||
</taxonomicName>
|
||
, topotype; E.
|
||
<taxonomicName id="4C4B39DCFF9C7401FBF2C932FB6EF82B" box="[1129,1224,1898,1921]" class="Gastropoda" family="Assimineidae" genus="Paludinella" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="opaca">
|
||
<emphasis id="B93F9E4DFF9C7401FBF2C932FB6EF82B" box="[1129,1224,1898,1921]" italics="true" pageId="8" pageNumber="9">B. opaca</emphasis>
|
||
</taxonomicName>
|
||
, (topotype of
|
||
<taxonomicName id="4C4B39DCFF9C7401FAFAC932FE67F809" authority="Kuster, 1852" authorityName="Kuster" authorityYear="1852" class="Gastropoda" family="Hydrobiidae" genus="Paludina" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="schmidtii">
|
||
<emphasis id="B93F9E4DFF9C7401FAFAC932FE89F80E" italics="true" pageId="8" pageNumber="9">Paludina schmidtii</emphasis>
|
||
Küster, 1852
|
||
</taxonomicName>
|
||
). Scale bar = 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3412D7FF9D7400FF0CCCB1FB5CFC8A" httpUri="https://zenodo.org/record/178391/files/figure.png" pageId="9" pageNumber="10" targetBox="[264,1322,163,722]" targetPageId="9">
|
||
<paragraph id="8BF4425FFF9D7400FF0CCCB1FB5CFC8A" blockId="9.[151,1436,745,800]" pageId="9" pageNumber="10">
|
||
<emphasis id="B93F9E4DFF9D7400FF0CCCB1FEBFFCAB" bold="true" box="[151,281,745,769]" pageId="9" pageNumber="10">FIGURE 3.</emphasis>
|
||
Shells of Carinthian populations. A–G.
|
||
<taxonomicName id="4C4B39DCFF9D7400FD59CCB1FC87FCAA" box="[706,801,745,768]" class="Gastropoda" family="Assimineidae" genus="Paludinella" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="opaca">
|
||
<emphasis id="B93F9E4DFF9D7400FD59CCB1FC87FCAA" box="[706,801,745,768]" italics="true" pageId="9" pageNumber="10">B. opaca</emphasis>
|
||
</taxonomicName>
|
||
; H.
|
||
<taxonomicName id="4C4B39DCFF9D7400FCD4CCB1FA83FCAA" authority="Tscheppaschlucht. A. Barental" authorityName="Tscheppaschlucht. A. Barental" box="[847,1317,745,768]" class="Gastropoda" family="Hydrobiidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="angelitae">
|
||
<emphasis id="B93F9E4DFF9D7400FCD4CCB1FC76FCAA" box="[847,976,745,768]" italics="true" pageId="9" pageNumber="10">B. angelitae</emphasis>
|
||
, Tscheppaschlucht. A. Bärental
|
||
</taxonomicName>
|
||
; B. Farchtnersee; C. Granitztal St. Paul; D. Sand Grafenstein; E. Oberdrauburger Bach; F. Niedermarkt; G. Podkraj.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF4425FFF9D7400FF5DCD09FE27FB51" blockId="9.[151,1437,849,2035]" pageId="9" pageNumber="10">
|
||
Stomach without appendix, rectum with shallow loop along pallial genital glands; ovary small sac with wide lobes comprising 0.25 whorls, starting about 1 whorl below apex, distal end close to but not reaching posterior end of stomach, albumen gland comprising three fifth length of pallial oviduct, renal oviduct describing wide loop of 270°, seminal receptacle small, tubular, arising close to junction with bursal duct, receptacle and bursal duct may be embedded into albumen gland tissue, bursa copulatrix a long sac arching around posterior end of albumen gland, bursal duct entering ventrally (
|
||
<figureCitation id="13705EDAFF9D7400FC4DCA41FBB8FB99" box="[982,1054,1049,1075]" captionStart="FIGURE 6" captionStartId="12.[151,255,747,771]" captionTargetBox="[189,1389,227,694]" captionTargetId="figure@12.[151,1436,194,725]" captionTargetPageId="12" captionText="FIGURE 6. Distal genitalia of three females of B. angelitae from the Tscheppaschlucht. ag, albumen gland; bc, bursa copulatrix; bd, bursal duct; cg, capsule gland; go, genital opening; od, oviduct; rs, receptaculum seminis. Scale bar = 400 µm." httpUri="https://zenodo.org/record/178394/files/figure.png" pageId="9" pageNumber="10">Fig. 6</figureCitation>
|
||
); testis lobate, comprising about 1 whorl, starting about 0.5 whorls below apex, covering posterior chamber of stomach, seminal vesicle comprising 0.25 whorls, proximal vas deferens entering kidney-shaped prostate just in front of posterior wall of mantle cavity, distal vas deferens emerging at distal tip of prostate becoming muscular ductus ejaculatorius coiling in proximal third of resting penis, penis longer than tubular accessory gland, anterior third of penis with black pigment.
|
||
</paragraph>
|
||
<paragraph id="8BF4425FFF9D7400FF5DCB50FCC8FAE1" blockId="9.[151,1437,849,2035]" pageId="9" pageNumber="10">
|
||
<emphasis id="B93F9E4DFF9D7400FF5DCB50FDA2FA88" bold="true" box="[198,516,1288,1314]" pageId="9" pageNumber="10">Habitat and distribution:</emphasis>
|
||
It is only known from springs in the area of Lake Bled in
|
||
<collectingCountry id="F35C02CFFF9D7400FB23CB51FAB8FA89" box="[1208,1310,1289,1315]" name="Slovenia" pageId="9" pageNumber="10">Slovenia</collectingCountry>
|
||
and in the Tscheppaschlucht just across the Austrian border in Carinthia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35111D4FF9D7403FF5DCB00FF44FE80" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="discussion">
|
||
<paragraph id="8BF4425FFF9D7400FF5DCB00FCE4F8F9" blockId="9.[151,1437,849,2035]" pageId="9" pageNumber="10">
|
||
<emphasis id="B93F9E4DFF9D7400FF5DCB00FEE4FAD8" bold="true" box="[198,322,1368,1394]" pageId="9" pageNumber="10">Remarks:</emphasis>
|
||
Morphologically and anatomically,
|
||
<taxonomicName id="4C4B39DCFF9D7400FD72CB01FCD0FAD8" box="[745,886,1369,1394]" class="Gastropoda" family="Hydrobiidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="angelitae">
|
||
<emphasis id="B93F9E4DFF9D7400FD72CB01FCD0FAD8" box="[745,886,1369,1394]" italics="true" pageId="9" pageNumber="10">B. angelitae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93F9E4DFF9D7400FCE7CB00FC53FAD8" bold="true" box="[892,1013,1368,1394]" pageId="9" pageNumber="10">nom. nov.</emphasis>
|
||
falls well into the range of variation of its immediate sister species,
|
||
<taxonomicName id="4C4B39DCFF9D7400FD9DCBD9FDC9FA30" box="[518,623,1409,1434]" class="Gastropoda" family="Assimineidae" genus="Paludinella" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="opaca">
|
||
<emphasis id="B93F9E4DFF9D7400FD9DCBD9FDC9FA30" box="[518,623,1409,1434]" italics="true" pageId="9" pageNumber="10">B. opaca</emphasis>
|
||
</taxonomicName>
|
||
, which is also stem species of
|
||
<taxonomicName id="4C4B39DCFF9D7400FC42CBD9FBCDFA30" box="[985,1131,1409,1434]" class="Gastropoda" family="Hydrobiidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="robiciana">
|
||
<emphasis id="B93F9E4DFF9D7400FC42CBD9FBCDFA30" box="[985,1131,1409,1434]" italics="true" pageId="9" pageNumber="10">B. robiciana</emphasis>
|
||
</taxonomicName>
|
||
characterized by a globular, derived shell shape (see below). The only distinguishing feature of the new species is the denticulation of the marginal teeth of the radula, with the fewest number of denticles on the inner marginal and the highest number on the outer one. Although there is some overlap with the remaining populations — hardly though in case of the outer marginal —, the shift towards the extreme ends of the overall distribution in opposite directions must be considered distinctive. There are reports of phenotypic plasticity of radula tooth shape depending on diet (
|
||
<bibRefCitation id="EFDA3FAEFF9D7400FEABC829FE6BF921" author="Padilla" box="[304,461,1649,1675]" pageId="9" pageNumber="16" refString="Padilla, D. K. (1998) Inducible phenotypic plasticity in the radula of Lacuna (Gastropoda: Littorinidae). Veli ger, 41, 201 - 204." type="journal article" year="1998">Padilla 1998</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFDA3FAEFF9D7400FE46C829FC87F921" author="Andrade" box="[477,801,1649,1675]" pageId="9" pageNumber="15" refString="Andrade, S. C. S. & Solferini, V. N. (2006) Transfer experiment suggests environmental effects on the radula of Littoraria flava (Gastropoda: Littorinidae). Journal of Molluscan Studies, 72, 111 - 116." type="journal article" year="2006">Andrade & Solferini 2006</bibRefCitation>
|
||
), but we are not aware of any case of phenotypical plasticity in numbers of denticles. The numbers representing the extreme ends of the total variation observed among the populations investigated suggest that probably more than one controlling genetic locus is involved. In addition,
|
||
<taxonomicName id="4C4B39DCFF9D7400FEBEC8B1FE15F8A8" box="[293,435,1769,1794]" class="Gastropoda" family="Hydrobiidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="angelitae">
|
||
<emphasis id="B93F9E4DFF9D7400FEBEC8B1FE15F8A8" box="[293,435,1769,1794]" italics="true" pageId="9" pageNumber="10">B. angelitae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93F9E4DFF9D7400FE20C8B0FD92F8A8" bold="true" box="[443,564,1768,1794]" pageId="9" pageNumber="10">nom. nov.</emphasis>
|
||
turned out to be sister species of
|
||
<taxonomicName id="4C4B39DCFF9D7400FC26C8B1FB80F8A8" box="[957,1062,1769,1794]" class="Gastropoda" family="Assimineidae" genus="Paludinella" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="opaca">
|
||
<emphasis id="B93F9E4DFF9D7400FC26C8B1FB80F8A8" box="[957,1062,1769,1794]" italics="true" pageId="9" pageNumber="10">B. opaca</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4B39DCFF9D7400FBC4C8B1FB56F8A8" box="[1119,1264,1769,1794]" class="Gastropoda" family="Hydrobiidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="robiciana">
|
||
<emphasis id="B93F9E4DFF9D7400FBC4C8B1FB56F8A8" box="[1119,1264,1769,1794]" italics="true" pageId="9" pageNumber="10">B. robiciana</emphasis>
|
||
</taxonomicName>
|
||
with HKY + I + Γ distances>
|
||
<quantity id="4CB3EFBAFF9D7400FECEC949FE10F881" box="[341,438,1809,1835]" metricMagnitude="-3" metricUnit="m" metricValue="1.2192" pageId="9" pageNumber="10" unit="in" value="0.048">0.048 in</quantity>
|
||
the phylogenetic analysis based on the 638 bp long fragment of COI. The combination of these facts justifies recognition as separate species.
|
||
</paragraph>
|
||
<paragraph id="8BF4425FFF9D7400FF5CC939FE51F859" blockId="9.[151,1437,849,2035]" pageId="9" pageNumber="10">
|
||
According to paragraph 72.7 of the ICZN the
|
||
<typeStatus id="54F0FCFDFF9D7400FD71C939FC8EF8D1" box="[746,808,1889,1915]" pageId="9" pageNumber="10">types</typeStatus>
|
||
of the species whose name is to be replaced are also
|
||
<typeStatus id="54F0FCFDFF9D7400FF0CC9D1FF73F809" box="[151,213,1929,1955]" pageId="9" pageNumber="10">types</typeStatus>
|
||
of the new name regardless of any restriction of the applicability of that name as in the present case. Therefore, the samples from
|
||
<collectingCountry id="F35C02CFFF9D7400FE76C9E9FD85F861" box="[493,547,1969,1995]" name="Italy" pageId="9" pageNumber="10">Italy</collectingCountry>
|
||
preserved their status as
|
||
<typeStatus id="54F0FCFDFF9D7400FCD5C9E9FC2AF861" box="[846,908,1969,1995]" pageId="9" pageNumber="10">types</typeStatus>
|
||
although they most likely represent a different species (see Introduction).
|
||
</paragraph>
|
||
<paragraph id="8BF4425FFF9E7403FF5CCEC0FF44FE80" blockId="10.[151,1436,152,298]" pageId="10" pageNumber="11">
|
||
Frauenfeld’s (1857) Lake Bled-sample apparently consisted of representatives of two populations judging from the color of the periostracum and the different ways of original treatment. The three lighter shells had been glued onto black cardboard (see
|
||
<figureCitation id="13705EDAFF9E7403FDC0CEB0FD05FEA8" box="[603,675,232,258]" captionStart="FIGURE 2" captionStartId="8.[151,255,1863,1887]" captionTargetBox="[151,1435,1344,1840]" captionTargetId="figure@8.[151,1436,1344,1842]" captionTargetPageId="8" captionText="FIGURE 2. Shells. A. B. angelitae, holotype (and lectotype of Paludinella opaca Frauenfeld, 1857); B, C. B. angelitae, paratypes (and paralectotypes of P. opaca Frauenfeld, 1857); D. B. robiciana, topotype; E. B. opaca, (topotype of Paludina schmidtii Küster, 1852). Scale bar = 2 mm." httpUri="https://zenodo.org/record/178390/files/figure.png" pageId="10" pageNumber="11">Fig. 2</figureCitation>
|
||
), whereas the two darker shells were removed from white cardboard.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |