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<mods:title id="D4B79F8DC5180A6C44B9955B536EE435">Taxonomic notes on the tribe Dryptini Bonelli, 1810 with description of a new genus and species from China (Coleoptera: Carabidae: Dryptini)</mods:title>
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<treatment id="03D77E1287116B1DFF0EFF0D60FCFC60" ID-DOI="http://doi.org/10.5281/zenodo.3665225" ID-GBIF-Taxon="162074184" ID-Zenodo-Dep="3665225" LSID="urn:lsid:plazi:treatment:03D77E1287116B1DFF0EFF0D60FCFC60" httpUri="http://treatment.plazi.org/id/03D77E1287116B1DFF0EFF0D60FCFC60" lastPageNumber="525" pageId="3" pageNumber="525">
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<taxonomicName id="4C7EB48787116B1DFF0EFF0D60D1FF71" ID-CoL="8KZNN" authority="Jeannel, 1949" authorityName="Jeannel" authorityYear="1949" box="[151,473,153,180]" class="Insecta" family="Carabidae" genus="Nesiodrypta" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="525" phylum="Arthropoda" rank="genus">
<emphasis id="B90A131687116B1DFF0EFF0D60D1FF71" bold="true" box="[151,473,153,180]" pageId="3" pageNumber="525">
<emphasis id="B90A131687116B1DFF0EFF0D6020FF76" bold="true" box="[151,296,153,180]" italics="true" pageId="3" pageNumber="525">Nesiodrypta</emphasis>
<bibRefCitation id="EFEFB2F587116B1DFEA9FF0E60D1FF71" author="Jeannel, R." box="[304,473,153,180]" pageId="3" pageNumber="525" pagination="767 - 1146" refId="ref5904" refString="Jeannel, R. (1949) Faune de l'empire francais. XI. Coleopteres carabiques de la region Malgache. Troisieme Partie. Libriarie Larose Paris, 380 pp. [pp. 767 - 1146]" type="book chapter" year="1949">Jeannel, 1949</bibRefCitation>
</emphasis>
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<paragraph id="8BC1CF0487116B1DFF0EFF766395FF39" blockId="3.[151,669,225,252]" box="[151,669,225,252]" pageId="3" pageNumber="525">
<typeStatus id="54C571A687116B1DFF0EFF7661C7FF3E" box="[151,207,226,252]" pageId="3" pageNumber="525">Type</typeStatus>
species:
<taxonomicName id="4C7EB48787116B1DFEACFF766395FF39" authority="Fairmaire, 1834" authorityName="Fairmaire" authorityYear="1834" box="[309,669,225,251]" class="Insecta" family="Carabidae" genus="Drypta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="525" phylum="Arthropoda" rank="species" species="perrieri">
<emphasis id="B90A131687116B1DFEACFF7660EBFF39" box="[309,483,226,251]" italics="true" pageId="3" pageNumber="525">Drypta perrieri</emphasis>
Fairmaire, 1834
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<paragraph id="8BC1CF0487116B1DFF0EFEB26279FEFC" blockId="3.[151,881,293,318]" box="[151,881,293,318]" pageId="3" pageNumber="525">
<emphasis id="B90A131687116B1DFF0EFEB261A0FEFF" box="[151,168,294,317]" italics="true" pageId="3" pageNumber="525">=</emphasis>
<taxonomicName id="4C7EB48787116B1DFF36FEB260CAFEFC" authority="Jeannel, 1949" authorityName="Jeannel" authorityYear="1949" box="[175,450,293,318]" class="Insecta" genus="Goniodrypta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="525" phylum="Arthropoda" rank="genus">
<emphasis id="B90A131687116B1DFF36FEB26038FEFC" box="[175,304,294,318]" italics="true" pageId="3" pageNumber="525">Goniodrypta</emphasis>
<bibRefCitation id="EFEFB2F587116B1DFEAFFEB160CAFEFC" author="Jeannel, R." box="[310,450,293,318]" pageId="3" pageNumber="525" pagination="767 - 1146" refId="ref5904" refString="Jeannel, R. (1949) Faune de l'empire francais. XI. Coleopteres carabiques de la region Malgache. Troisieme Partie. Libriarie Larose Paris, 380 pp. [pp. 767 - 1146]" type="book chapter" year="1949">Jeannel, 1949</bibRefCitation>
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species:
<taxonomicName id="4C7EB48787116B1DFDCEFEB26279FEFC" ID-CoL="8LDPH" authority="Castelnau, 1840" authorityName="Castelnau" authorityYear="1840" box="[599,881,293,318]" class="Insecta" family="Carabidae" genus="Drypta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="525" phylum="Arthropoda" rank="species" species="iris">
<emphasis id="B90A131687116B1DFDCEFEB263CFFEFF" box="[599,711,294,317]" italics="true" pageId="3" pageNumber="525">Drypta iris</emphasis>
Castelnau, 1840
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<paragraph id="8BC1CF0487116B1DFF0EFEF863CFFDD5" blockId="3.[151,1437,364,931]" pageId="3" pageNumber="525">
This genus, described by
<bibRefCitation id="EFEFB2F587116B1DFE2AFEF86351FE44" author="Jeannel, R." box="[435,601,364,390]" pageId="3" pageNumber="525" pagination="767 - 1146" refId="ref5904" refString="Jeannel, R. (1949) Faune de l'empire francais. XI. Coleopteres carabiques de la region Malgache. Troisieme Partie. Libriarie Larose Paris, 380 pp. [pp. 767 - 1146]" type="book chapter" year="1949">Jeannel (1949)</bibRefCitation>
at the same time as the preceding one, has been regarded as systematically close to it because of its well developed pronotal bead.
<bibRefCitation id="EFEFB2F587116B1DFC86FE0462C0FE68" author="Jeannel, R." box="[799,968,400,426]" pageId="3" pageNumber="525" pagination="767 - 1146" refId="ref5904" refString="Jeannel, R. (1949) Faune de l'empire francais. XI. Coleopteres carabiques de la region Malgache. Troisieme Partie. Libriarie Larose Paris, 380 pp. [pp. 767 - 1146]" type="book chapter" year="1949">Jeannel (1949)</bibRefCitation>
also described a subgenus
<taxonomicName id="4C7EB48787116B1DFA9FFE04649DFE68" authorityName="Jeannel" authorityYear="1949" box="[1286,1429,400,426]" class="Insecta" genus="Goniodrypta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="525" phylum="Arthropoda" rank="genus">
<emphasis id="B90A131687116B1DFA9FFE04649DFE68" box="[1286,1429,400,426]" italics="true" pageId="3" pageNumber="525">Goniodrypta</emphasis>
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, distinct on the basis of an evident angle in the middle of the lateral pronotal margins. Although very evident, this character does not seem to us sufficient for a subgeneric distinction, so we agree with the opinion of
<bibRefCitation id="EFEFB2F587116B1DFB6CFE4D6490FE30" author="Lorenz, W." box="[1269,1432,471,499]" pageId="3" pageNumber="525" refId="ref6110" refString="Lorenz, W. (2005) A systematic list of extant ground beetles of the World. (Insecta, Coleoptera, Adephaga: Trachypachidae &amp; Carabidae incl. Paussinae, Cicindelinae, Rhysodinae), 2 nd edition. W. Lorenz, Tutzing, 530 pp." type="book" year="2005">Lorenz (2005)</bibRefCitation>
, who regards it as a synonym of
<taxonomicName id="4C7EB48787116B1DFE65FE68638DFDD4" authorityName="Jeannel" authorityYear="1949" box="[508,645,508,534]" class="Insecta" family="Carabidae" genus="Nesiodrypta" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="525" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B90A131687116B1DFE65FE68638DFDD4" box="[508,645,508,534]" italics="true" pageId="3" pageNumber="525">Nesiodrypta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A239AE6D87116B1DFD15FE6963CFFDD5" box="[652,711,509,535]" pageId="3" pageNumber="525" sensu="stricto">s. str.</taxonomicNameLabel>
</paragraph>
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<paragraph id="8BC1CF0487116B1DFF5EFDB460DEFD40" blockId="3.[151,1437,364,931]" pageId="3" pageNumber="525">
This genus contains 18 species primarily occuring in
<collectingCountry id="F3698F9487116B1DFC81FDB46297FDF8" box="[792,927,544,570]" name="Madagascar" pageId="3" pageNumber="525">Madagascar</collectingCountry>
(15 species), but marginally also in continental Africa (3 species), with two species in East Africa (
<collectingCountry id="F3698F9487116B1DFC99FDD06243FD9C" box="[768,843,580,606]" name="Kenya" pageId="3" pageNumber="525">Kenya</collectingCountry>
and
<collectingCountry id="F3698F9487116B1DFC1BFDD062E1FD9D" box="[898,1001,580,607]" name="Tanzania" pageId="3" pageNumber="525">Tanzania</collectingCountry>
) and one in
<collectingCountry id="F3698F9487116B1DFBEFFDD065CBFD9C" box="[1142,1219,580,606]" name="Gabon" pageId="3" pageNumber="525">Gabon</collectingCountry>
, recently described (
<bibRefCitation id="EFEFB2F587116B1DFF06FDFC60C2FD40" author="Facchini, S. &amp; Czeppel, R." box="[159,458,616,642]" pageId="3" pageNumber="525" pagination="287 - 291" refId="ref5821" refString="Facchini, S. &amp; Czeppel, R. (2011) Una nuova specie di Drypta Latreille, 1796 (Nesiodrypta Jeannel, 1949) dell'Africa occiden- tale (Coleoptera Carabidae, Dryptinae). Giornale italiano di Entomologia, 12, 287 - 291." type="journal article" year="2011">Facchini &amp; Czeppel, 2011</bibRefCitation>
).
</paragraph>
<paragraph id="8BC1CF0487116B1DFF5EFD1860FCFC60" blockId="3.[151,1437,364,931]" pageId="3" pageNumber="525">
The unique combination of characters distinguishing this genus is: pronotal bead well marked, with sides not serrate (fig. 13); punctuation on head sparse, often leaving the frons smooth, on pronotum often limited to the lateral parts, the disc covered with transversal wrinkles; elytral microsculpture well developed; elytral pubescence limited to some small and sparse punctures, only in few cases some more regular punctures are aligned along the striae (except
<taxonomicName id="4C7EB48787116B1DFF69FC896062FCF4" baseAuthorityName="Gerstaecker" baseAuthorityYear="1867" box="[240,362,797,822]" class="Insecta" family="Carabidae" genus="Nesiodrypta" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="525" phylum="Arthropoda" rank="species" species="setigera">
<emphasis id="B90A131687116B1DFF69FC896062FCF4" box="[240,362,797,822]" italics="true" pageId="3" pageNumber="525">N. setigera</emphasis>
</taxonomicName>
, that shows a punctuation on head, pronotum and elytra almost as dense as in
<taxonomicName id="4C7EB48787116B1DFB57FC88646DFCF4" authorityName="Jeannel" authorityYear="1949" box="[1230,1381,796,822]" class="Insecta" family="Carabidae" genus="Prionodrypta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="525" phylum="Arthropoda" rank="genus">
<emphasis id="B90A131687116B1DFB57FC88646DFCF4" box="[1230,1381,796,822]" italics="true" pageId="3" pageNumber="525">Prionodrypta</emphasis>
</taxonomicName>
; it is possible that future studies will show the necessity to isolate this species in a separate genus); scutellar pore single; intervals rather convex; tarsal claws thick, strongly curved, smooth on inner side; no setae, or a rudimental one, on outer side of stylomere (
<figureCitation id="1345D38187116B1DFE31FC1C60EEFC60" box="[424,486,903,930]" captionStart="FIGS" captionStartId="5.[151,208,1725,1750]" captionTargetBox="[350,1235,1327,1678]" captionTargetId="figure@5.[340,1247,1314,1700]" captionText="FIGS. 48. Stylomere structure of: 4—Megadrypta mirabilis gen. n. sp. n.; 5—Nesiodrypta waterhousei (Oberthür, 1881); 6—Prionodrypta lugens (Schmidt-Goebel, 1846); 7—Drypta lineola W. S. MacLeay, 1825; 8—Dendrocellus australis (Peringuey, 1896)." figureDoi="http://doi.org/10.5281/zenodo.3690687" httpUri="https://zenodo.org/record/3690687/files/figure.png" pageId="3" pageNumber="525">fig. 5</figureCitation>
).
</paragraph>
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