427 lines
52 KiB
XML
427 lines
52 KiB
XML
<document id="906EEE8C0268A033A882AD0F6397E78E" ID-DOI="10.11646/zootaxa.5396.1.13" ID-ISSN="1175-5326" ID-Zenodo-Dep="10441322" ID-ZooBank="1CCB5F08-CD60-423D-9E32-91480A39465D" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1703840934995" checkinUser="plazi" docAuthor="Hakim, Marina, Azar, Dany & Huang, Di-Ying" docDate="2023" docId="03D64A58D235FFA71CC0FA92667D9807" docLanguage="en" docName="zootaxa.5396.1.13.pdf" docOrigin="Zootaxa 5396 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5396.1.13/52606" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Longiantennum fashengi Liang, Li & Yao 2022" docType="treatment" docVersion="3" lastPageNumber="89" masterDocId="FFEF3220D23FFFA81C57FF8565649E2A" masterDocTitle="First record of fossil psocodeans in copula from mid-Cretaceous Burmese amber" masterLastPageNumber="93" masterPageNumber="74" pageNumber="84" updateTime="1704294611933" updateUser="tatiana" zenodo-license-document="CLOSED">
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<mods:title id="098EFF5E363695CD149D5285D96F0503">First record of fossil psocodeans in copula from mid-Cretaceous Burmese amber</mods:title>
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<mods:namePart id="9F21DD1C77C69BC55118660E493111E8">Hakim, Marina</mods:namePart>
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<mods:affiliation id="382013D4011DAB66881DC5300A7C5943">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China</mods:affiliation>
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<mods:namePart id="ABCD525127135412DEFCE60D6C4A2276">Azar, Dany</mods:namePart>
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<mods:affiliation id="3D571CAB3C25A6E5EF589C824E6E6D65">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China & Lebanese University, Faculty of Science II, Natural Sciences Department, Fanar - El-Matn, PO box 26110217, Lebanon</mods:affiliation>
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<mods:namePart id="92BEAB22EB500067BF862567F6D44C85">Huang, Di-Ying</mods:namePart>
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<mods:affiliation id="A83B1B5C168DC204AF00DD6FCB02FACF">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China</mods:affiliation>
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<mods:date id="1BAC81553BDB26B4CB4C82E7F36CB4AB">2023</mods:date>
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<treatment id="03D64A58D235FFA71CC0FA92667D9807" LSID="urn:lsid:plazi:treatment:03D64A58D235FFA71CC0FA92667D9807" httpUri="http://treatment.plazi.org/id/03D64A58D235FFA71CC0FA92667D9807" lastPageId="15" lastPageNumber="89" pageId="10" pageNumber="84">
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<subSubSection id="C365A8C5D235FFA21CC0FA9267809B1A" box="[151,740,1302,1329]" pageId="10" pageNumber="84" type="nomenclature">
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<paragraph id="8BC0FB4ED235FFA21CC0FA9267809B1A" blockId="10.[151,740,1302,1365]" box="[151,740,1302,1329]" pageId="10" pageNumber="84">
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<heading id="D0884C22D235FFA21CC0FA9267809B1A" bold="true" box="[151,740,1302,1329]" fontSize="11" level="1" pageId="10" pageNumber="84" reason="1">
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<taxonomicName id="4C7F80CDD235FFA21CC0FA9267809B1A" authority="Liang, Li & Yao, 2022" authorityName="Liang, Li & Yao" authorityYear="2022" box="[151,740,1302,1329]" class="Insecta" family="Archaeatropidae" genus="Longiantennum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="84" phylum="Arthropoda" rank="species" species="fashengi">
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<emphasis id="B90B275CD235FFA21CC0FA9267809B1A" bold="true" box="[151,740,1302,1329]" pageId="10" pageNumber="84">
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<emphasis id="B90B275CD235FFA21CC0FA9264AE9B1B" bold="true" box="[151,458,1302,1329]" italics="true" pageId="10" pageNumber="84">Longiantennum fashengi</emphasis>
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<bibRefCitation id="EFEE86BFD235FFA21D85FA9367809B1A" author="Liang, F. Y. & Li, S. & Liu, X. Y. & Bai, M. & Yao, Y. Z." box="[466,740,1302,1329]" pageId="10" pageNumber="84" pagination="105233" refId="ref9702" refString="Liang, F. Y., Li, S., Liu, X. Y., Bai, M. & Yao, Y. Z. (2022) A new genus and species of the family Archaeatropidae (Psocodea: Trogiomorpha) from mid-Cretaceous amber of Northern Myanmar. Cretaceous Research, 138, 105233. https: // doi. org / 10.1016 / j. cretres. 2022.105233" type="journal article" year="2022">Liang, Li & Yao, 2022</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C365A8C5D235FFA21CC0FABE647D9B7F" box="[151,281,1339,1365]" pageId="10" pageNumber="84" type="description">
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<paragraph id="8BC0FB4ED235FFA21CC0FABE647D9B7F" blockId="10.[151,740,1302,1365]" box="[151,281,1339,1365]" pageId="10" pageNumber="84">
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(
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<figureCitation id="1344E7CBD235FFA21CC8FABE64759B7F" box="[159,273,1339,1365]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStart-2="FIGURE 9" captionStart-3="FIGURE 10" captionStartId-0="11.[151,250,1697,1721]" captionStartId-1="12.[151,250,1934,1958]" captionStartId-2="13.[151,250,1943,1967]" captionStartId-3="14.[151,250,1943,1967]" captionTargetBox-0="[151,1436,181,1674]" captionTargetBox-1="[254,1333,597,1911]" captionTargetBox-2="[260,1327,181,1919]" captionTargetBox-3="[246,1345,207,1895]" captionTargetId-0="figure-31@11.[151,1436,181,1674]" captionTargetId-1="figure-267@12.[254,1333,597,1911]" captionTargetId-2="figure-31@13.[260,1327,181,1919]" captionTargetId-3="figure-19@14.[209,1378,181,1919]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionTargetPageId-3="14" captionText-0="FIGURE 7. Longiantennum fashengi, NIGP171866, male (right side) and female (left side) in mating position. A, Habitus in dorsal view. B, Habitus in ventral view. Scale bars = 0.5 mm." captionText-1="FIGURE 8. Longiantennum fashengi. A, Antenna, under green fluorescence. B, Maxillary palpomere. C, Lacinia. D, Fore tibia and tarsus. E, Hind tibial spines. F, Claw. G, Wings, under green fluorescence. Scale bars = 0.2 mm for G, 0.1 mm for A, 0.05 mm for B, D, E, 0.02 mm for C, F." captionText-2="FIGURE 9. Longiantennum fashengi. A, B, Details of genitalia. Scale bars = 0.1 mm." captionText-3="FIGURE 10. Longiantennum fashengi, hand drawings. A, antennae (female). B, forewing (male). C, Hind wing (female). D, Genitalia. Scale bars = 0.2 mm for A–C, 0.1 mm for D." figureDoi-0="http://doi.org/10.5281/zenodo.10441342" figureDoi-1="http://doi.org/10.5281/zenodo.10441344" figureDoi-2="http://doi.org/10.5281/zenodo.10441348" figureDoi-3="http://doi.org/10.5281/zenodo.10441350" httpUri-0="https://zenodo.org/record/10441342/files/figure.png" httpUri-1="https://zenodo.org/record/10441344/files/figure.png" httpUri-2="https://zenodo.org/record/10441348/files/figure.png" httpUri-3="https://zenodo.org/record/10441350/files/figure.png" pageId="10" pageNumber="84">Figs 7–10</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C365A8C5D235FFA21CC0FA0766379807" pageId="10" pageNumber="84" type="materials_examined">
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<paragraph id="8BC0FB4ED235FFA21CC0FA0767CE9BCF" blockId="10.[151,1437,1410,1905]" pageId="10" pageNumber="84">
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<emphasis id="B90B275CD235FFA21CC0FA0764E39BB6" bold="true" box="[151,391,1410,1436]" pageId="10" pageNumber="84">Additional material.</emphasis>
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NIGP171866;
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<specimenCount id="9D7930C7D235FFA21E7AFA0667EB9BB7" box="[557,655,1411,1437]" count="1" pageId="10" pageNumber="84" type="male">one male</specimenCount>
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(NIGP171866-a) and
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<materialsCitation id="3B17F113D235FFA21FD4FA0667C39BCF" country="China" county="Nanjing" location="The" municipality="Chinese Academy of Sciences" pageId="10" pageNumber="84" specimenCount="1" specimenCount-female="1">
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<specimenCount id="9D7930C7D235FFA21FD4FA06669F9BB7" box="[899,1019,1411,1437]" count="1" pageId="10" pageNumber="84" type="female">one female</specimenCount>
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(NIGP171866-b) preserved in mating position (
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<figureCitation id="1344E7CBD235FFA21D57FA2264209BEB" box="[256,324,1447,1473]" captionStart="FIGURE 7" captionStartId="11.[151,250,1697,1721]" captionTargetBox="[151,1436,181,1674]" captionTargetId="figure-31@11.[151,1436,181,1674]" captionTargetPageId="11" captionText="FIGURE 7. Longiantennum fashengi, NIGP171866, male (right side) and female (left side) in mating position. A, Habitus in dorsal view. B, Habitus in ventral view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10441342" httpUri="https://zenodo.org/record/10441342/files/figure.png" pageId="10" pageNumber="84">Fig. 7</figureCitation>
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), no syninclusions.
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<location id="8EA0AD95D235FFA21E77FA2267289BEB" LSID="urn:lsid:plazi:treatment:03D64A58D235FFA71CC0FA92667D9807:8EA0AD95D235FFA21E77FA2267289BEB" box="[544,588,1447,1473]" country="China" county="Nanjing" municipality="Chinese Academy of Sciences" name="The" pageId="10" pageNumber="84">The</location>
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material is deposited at the
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<location id="8EA0AD95D235FFA21FD6FA2261A59BEB" LSID="urn:lsid:plazi:treatment:03D64A58D235FFA71CC0FA92667D9807:8EA0AD95D235FFA21FD6FA2261A59BEB" box="[897,1217,1447,1473]" country="China" county="Nanjing" municipality="Chinese Academy of Sciences" name="Nanjing Institute of Geology" pageId="10" pageNumber="84">Nanjing Institute of Geology</location>
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and
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<location id="8EA0AD95D235FFA218A0FA2260FC9BEB" LSID="urn:lsid:plazi:treatment:03D64A58D235FFA71CC0FA92667D9807:8EA0AD95D235FFA218A0FA2260FC9BEB" box="[1271,1432,1447,1473]" country="China" county="Nanjing" municipality="Chinese Academy of Sciences" name="Palaeontology" pageId="10" pageNumber="84">Palaeontology</location>
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,
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<collectingMunicipality id="6BA46134D235FFA21CC0FA4E64889BCF" box="[151,492,1483,1509]" pageId="10" pageNumber="84">Chinese Academy of Sciences</collectingMunicipality>
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,
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<collectingCounty id="62A183C2D235FFA21DA0FA4E67309BCF" box="[503,596,1483,1509]" pageId="10" pageNumber="84">Nanjing</collectingCounty>
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,
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<collectingCountry id="F368BBDED235FFA21E37FA4E67C39BCF" box="[608,679,1483,1509]" name="China" pageId="10" pageNumber="84">China</collectingCountry>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BC0FB4ED235FFA21C90FA6B66379807" blockId="10.[151,1437,1410,1905]" pageId="10" pageNumber="84">
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<emphasis id="B90B275CD235FFA21C90FA6B64A19822" bold="true" box="[199,453,1518,1545]" pageId="10" pageNumber="84">Locality and horizon.</emphasis>
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Burmese
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<collectingRegion id="49BB35ACD235FFA21E63FA6A67E19823" box="[564,645,1519,1545]" country="Myanmar" name="Kachin" pageId="10" pageNumber="84">Kachin</collectingRegion>
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amber (Burmite), from an amber mine located near Noije Bum Village, Tanai Township, Hukawng Valley,
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<collectingCountry id="F368BBDED235FFA21E48F99667F49807" box="[543,656,1555,1581]" name="Myanmar" pageId="10" pageNumber="84">Myanmar</collectingCountry>
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; mid-Cretaceous.
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</paragraph>
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</subSubSection>
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<subSubSection id="C365A8C5D235FFA71C90F9B367FA9D3F" lastPageId="15" lastPageNumber="89" pageId="10" pageNumber="84" type="description">
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<paragraph id="8BC0FB4ED235FFA21C90F9B367C6995B" blockId="10.[151,1437,1410,1905]" pageId="10" pageNumber="84">
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<emphasis id="B90B275CD235FFA21C90F9B3643D987B" bold="true" box="[199,345,1590,1617]" pageId="10" pageNumber="84">Description.</emphasis>
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<emphasis id="B90B275CD235FFA21D33F9B264C5987B" box="[356,417,1591,1617]" italics="true" pageId="10" pageNumber="84">Head</emphasis>
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: relatively triangular in shape, with sparse setae on vertex and frons; epicranial suture present with coronal suture well pronounced from base of head capsule to ocelli and frontal suture absent, possibly limited to only base of the arms. Compound eyes globulous, half spheres in shape, flattened dorso-ventrally; ommatidia not setose. Three ocelli present, disposed in narrow inverted triangle at middle of vertex. Antennae (
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<figureCitation id="1344E7CBD235FFA21938F92665DF98CB" captionStart="FIGURE 8" captionStartId="12.[151,250,1934,1958]" captionTargetBox="[254,1333,597,1911]" captionTargetId="figure-267@12.[254,1333,597,1911]" captionTargetPageId="12" captionText="FIGURE 8. Longiantennum fashengi. A, Antenna, under green fluorescence. B, Maxillary palpomere. C, Lacinia. D, Fore tibia and tarsus. E, Hind tibial spines. F, Claw. G, Wings, under green fluorescence. Scale bars = 0.2 mm for G, 0.1 mm for A, 0.05 mm for B, D, E, 0.02 mm for C, F." figureDoi="http://doi.org/10.5281/zenodo.10441344" httpUri="https://zenodo.org/record/10441344/files/figure.png" pageId="10" pageNumber="84">Fig. 8A</figureCitation>
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) slightly shorter than body, scarcely setose; scape and pedicel elongate, 2–3× the width of flagellomeres; 32 short flagellomeres (longest preserved antenna in female), no secondary annulations. Postclypeus bulged. Maxillary palpus four-segmented (
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<figureCitation id="1344E7CBD235FFA21DF6F88A64929903" box="[417,502,1807,1833]" captionStart="FIGURE 8" captionStartId="12.[151,250,1934,1958]" captionTargetBox="[254,1333,597,1911]" captionTargetId="figure-267@12.[254,1333,597,1911]" captionTargetPageId="12" captionText="FIGURE 8. Longiantennum fashengi. A, Antenna, under green fluorescence. B, Maxillary palpomere. C, Lacinia. D, Fore tibia and tarsus. E, Hind tibial spines. F, Claw. G, Wings, under green fluorescence. Scale bars = 0.2 mm for G, 0.1 mm for A, 0.05 mm for B, D, E, 0.02 mm for C, F." figureDoi="http://doi.org/10.5281/zenodo.10441344" httpUri="https://zenodo.org/record/10441344/files/figure.png" pageId="10" pageNumber="84">Fig. 8B</figureCitation>
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), with numerous long setae; strong short spur sensillum near base of second maxillary palpomere. Labial palpomeres weakly visible, apical segment setose, lobe shaped. Lacinia present, bicuspid, outer cusp without visible apical denticles (
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<figureCitation id="1344E7CBD235FFA21E6BF8D267F1995B" box="[572,661,1879,1905]" captionStart="FIGURE 8" captionStartId="12.[151,250,1934,1958]" captionTargetBox="[254,1333,597,1911]" captionTargetId="figure-267@12.[254,1333,597,1911]" captionTargetPageId="12" captionText="FIGURE 8. Longiantennum fashengi. A, Antenna, under green fluorescence. B, Maxillary palpomere. C, Lacinia. D, Fore tibia and tarsus. E, Hind tibial spines. F, Claw. G, Wings, under green fluorescence. Scale bars = 0.2 mm for G, 0.1 mm for A, 0.05 mm for B, D, E, 0.02 mm for C, F." figureDoi="http://doi.org/10.5281/zenodo.10441344" httpUri="https://zenodo.org/record/10441344/files/figure.png" pageId="10" pageNumber="84">Fig. 8C</figureCitation>
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).
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</paragraph>
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<caption id="DF00ABC6D234FFA31CC0F924666098F7" ID-DOI="http://doi.org/10.5281/zenodo.10441342" ID-Zenodo-Dep="10441342" httpUri="https://zenodo.org/record/10441342/files/figure.png" pageId="11" pageNumber="85" startId="11.[151,250,1697,1721]" targetBox="[151,1436,181,1674]" targetPageId="11" targetType="figure">
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<paragraph id="8BC0FB4ED234FFA31CC0F924666098F7" blockId="11.[151,1437,1697,1758]" pageId="11" pageNumber="85">
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<emphasis id="B90B275CD234FFA31CC0F92464709890" bold="true" box="[151,276,1697,1722]" pageId="11" pageNumber="85">FIGURE 7.</emphasis>
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<taxonomicName id="4C7F80CDD234FFA31D4CF92767069890" authority=", NIGP" authorityName="NIGP" box="[283,610,1698,1722]" class="Insecta" family="Empheriidae" genus="Longiantennum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Psocodea" pageId="11" pageNumber="85" phylum="Arthropoda" rank="species" species="fashengi">
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<emphasis id="B90B275CD234FFA31D4CF92767739890" box="[283,535,1698,1722]" italics="true" pageId="11" pageNumber="85">Longiantennum fashengi</emphasis>
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, NIGP
|
||
</taxonomicName>
|
||
171866, male (right side) and female (left side) in mating position.
|
||
<emphasis id="B90B275CD234FFA31944F92460419893" bold="true" box="[1299,1317,1697,1721]" pageId="11" pageNumber="85">A</emphasis>
|
||
, Habitus in dorsal view.
|
||
<emphasis id="B90B275CD234FFA31D4FF943644D98F4" bold="true" box="[280,297,1734,1758]" pageId="11" pageNumber="85">B</emphasis>
|
||
, Habitus in ventral view. Scale bars = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC0FB4ED234FFA41C90F8AD60049C17" blockId="11.[151,1437,1832,2038]" lastBlockId="12.[151,1437,151,579]" lastPageId="12" lastPageNumber="86" pageId="11" pageNumber="85">
|
||
<emphasis id="B90B275CD234FFA31C90F8AD647C9968" box="[199,280,1832,1858]" italics="true" pageId="11" pageNumber="85">Thorax</emphasis>
|
||
: Prothorax collar-like, few setae visible near edge; mesothorax well developed, triangular, setose.
|
||
<emphasis id="B90B275CD234FFA31CC0F8C865A8994C" box="[151,204,1869,1894]" italics="true" pageId="11" pageNumber="85">Legs</emphasis>
|
||
: all segments setose; hind tibiae with row of spurs, all tibiae with two longer apical spines (
|
||
<figureCitation id="1344E7CBD234FFA3188EF8C9603C994C" box="[1241,1368,1868,1894]" captionStart="FIGURE 8" captionStartId="12.[151,250,1934,1958]" captionTargetBox="[254,1333,597,1911]" captionTargetId="figure-267@12.[254,1333,597,1911]" captionTargetPageId="12" captionText="FIGURE 8. Longiantennum fashengi. A, Antenna, under green fluorescence. B, Maxillary palpomere. C, Lacinia. D, Fore tibia and tarsus. E, Hind tibial spines. F, Claw. G, Wings, under green fluorescence. Scale bars = 0.2 mm for G, 0.1 mm for A, 0.05 mm for B, D, E, 0.02 mm for C, F." figureDoi="http://doi.org/10.5281/zenodo.10441344" httpUri="https://zenodo.org/record/10441344/files/figure.png" pageId="11" pageNumber="85">Figs 8D–E</figureCitation>
|
||
); tarsi three-segmented, basal tarsomere the longest, fore and middle tarsomeres about equal in length, basal and middle tarsomere bearing two apical spurs; claws sharply curved, with one strong preapical tooth, without pulvillus (
|
||
<figureCitation id="1344E7CBD234FFA31938F81165DC99F8" captionStart="FIGURE 8" captionStartId="12.[151,250,1934,1958]" captionTargetBox="[254,1333,597,1911]" captionTargetId="figure-267@12.[254,1333,597,1911]" captionTargetPageId="12" captionText="FIGURE 8. Longiantennum fashengi. A, Antenna, under green fluorescence. B, Maxillary palpomere. C, Lacinia. D, Fore tibia and tarsus. E, Hind tibial spines. F, Claw. G, Wings, under green fluorescence. Scale bars = 0.2 mm for G, 0.1 mm for A, 0.05 mm for B, D, E, 0.02 mm for C, F." figureDoi="http://doi.org/10.5281/zenodo.10441344" httpUri="https://zenodo.org/record/10441344/files/figure.png" pageId="11" pageNumber="85">Fig. 8F</figureCitation>
|
||
). Forewing (
|
||
<figureCitation id="1344E7CBD234FFA31D11F83D64C499F8" box="[326,416,1976,2002]" captionStart="FIGURE 8" captionStartId="12.[151,250,1934,1958]" captionTargetBox="[254,1333,597,1911]" captionTargetId="figure-267@12.[254,1333,597,1911]" captionTargetPageId="12" captionText="FIGURE 8. Longiantennum fashengi. A, Antenna, under green fluorescence. B, Maxillary palpomere. C, Lacinia. D, Fore tibia and tarsus. E, Hind tibial spines. F, Claw. G, Wings, under green fluorescence. Scale bars = 0.2 mm for G, 0.1 mm for A, 0.05 mm for B, D, E, 0.02 mm for C, F." figureDoi="http://doi.org/10.5281/zenodo.10441344" httpUri="https://zenodo.org/record/10441344/files/figure.png" pageId="11" pageNumber="85">Fig. 8G</figureCitation>
|
||
; measurements are taken from male) membranous, translucent; membrane not setose; wing margin with short fringe only at base of costal margin; all veins except CuP with one row of setae; stems R and M+CuA separating at
|
||
<quantity id="4C8756ABD233FFA41DCFFF12649A9E9B" box="[408,510,151,177]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="12" pageNumber="86" unit="mm" value="0.24">0.24 mm</quantity>
|
||
from wing base; Sc setose, well developed, curving perpendicularly into R
|
||
<subScript id="17FBF90BD233FFA41930FF2C600B9E9D" attach="left" box="[1383,1391,169,183]" fontSize="6" pageId="12" pageNumber="86">1</subScript>
|
||
; no veinlet present between Sc and anterior wing margin; Sc’ directed upward, almost perpendicular to wing margin; R
|
||
<subScript id="17FBF90BD233FFA419C3FF4860F89EF1" attach="left" box="[1428,1436,205,219]" fontSize="6" pageId="12" pageNumber="86">1</subScript>
|
||
straight, connected to Rs by short crossvein; Pt elongated, narrow at base and wide at apex, four-angled (trapezoid in shape), transparent with membrane not setose; Rs and M fused for a short length,
|
||
<quantity id="4C8756ABD233FFA41818FE8661D79F37" box="[1103,1203,259,285]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="86" unit="mm" value="0.06">0.06 mm</quantity>
|
||
long; Rs forked into R
|
||
<subScript id="17FBF90BD233FFA41CFDFEBC65A79F6D" attach="left" box="[170,195,313,327]" fontSize="6" pageId="12" pageNumber="86">2+3</subScript>
|
||
and R
|
||
<subScript id="17FBF90BD233FFA41D5BFEBC64429F6D" attach="left" box="[268,294,313,327]" fontSize="6" pageId="12" pageNumber="86">4+5</subScript>
|
||
at
|
||
<quantity id="4C8756ABD233FFA41D11FEA264C89F6B" box="[326,428,295,321]" metricMagnitude="-4" metricUnit="m" metricValue="9.3" pageId="12" pageNumber="86" unit="mm" value="0.93">0.93 mm</quantity>
|
||
, fork R
|
||
<subScript id="17FBF90BD233FFA41DA8FEBC677C9F6D" attach="left" box="[511,536,313,327]" fontSize="6" pageId="12" pageNumber="86">2+3</subScript>
|
||
and R
|
||
<subScript id="17FBF90BD233FFA41E36FEBC671F9F6D" attach="left" box="[609,635,313,327]" fontSize="6" pageId="12" pageNumber="86">4+5</subScript>
|
||
longer than stem Rs after separation from M; basi-radial cell elongated, four-angled; radial cell seven-angled, short; M+CuA branching into stems M and CuA at
|
||
<quantity id="4C8756ABD233FFA418C4FECF61989F4F" box="[1171,1276,330,357]" metricMagnitude="-4" metricUnit="m" metricValue="5.1" pageId="12" pageNumber="86" unit="mm" value="0.51">0.51 mm</quantity>
|
||
; M with three branches; M
|
||
<subScript id="17FBF90BD233FFA41D73FE0464489FA5" attach="left" box="[292,300,385,399]" fontSize="6" pageId="12" pageNumber="86">3</subScript>
|
||
emerging first at
|
||
<quantity id="4C8756ABD233FFA41DA0FEEA673B9FA3" box="[503,607,367,393]" metricMagnitude="-4" metricUnit="m" metricValue="9.1" pageId="12" pageNumber="86" unit="mm" value="0.91">0.91 mm</quantity>
|
||
, M
|
||
<subScript id="17FBF90BD233FFA41ED3FE0467E89FA5" attach="left" box="[644,652,385,399]" fontSize="6" pageId="12" pageNumber="86">1</subScript>
|
||
and M
|
||
<subScript id="17FBF90BD233FFA41E88FE0467839FA5" attach="left" box="[735,743,385,399]" fontSize="6" pageId="12" pageNumber="86">2</subScript>
|
||
splitting at
|
||
<quantity id="4C8756ABD233FFA41F27FEEA66B39FA3" box="[880,983,367,393]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="12" pageNumber="86" unit="mm" value="1.1">1.10 mm</quantity>
|
||
, fork M
|
||
<subScript id="17FBF90BD233FFA41864FE04615F9FA5" attach="left" box="[1075,1083,385,399]" fontSize="6" pageId="12" pageNumber="86">1</subScript>
|
||
+M
|
||
<subScript id="17FBF90BD233FFA41833FE0461089FA5" attach="left" box="[1124,1132,385,399]" fontSize="6" pageId="12" pageNumber="86">2</subScript>
|
||
longer than common stem M
|
||
<subScript id="17FBF90BD233FFA41CE7FE2065AD9F99" attach="left" box="[176,201,421,435]" fontSize="6" pageId="12" pageNumber="86">1+2</subScript>
|
||
; fork of CuA at
|
||
<quantity id="4C8756ABD233FFA41DD5FE17648D9F87" box="[386,489,402,429]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="12" pageNumber="86" unit="mm" value="0.65">0.65 mm</quantity>
|
||
; CuA
|
||
<subScript id="17FBF90BD233FFA41E7EFE2067559F99" attach="left" box="[553,561,421,435]" fontSize="6" pageId="12" pageNumber="86">1</subScript>
|
||
and CuA
|
||
<subScript id="17FBF90BD233FFA41ECAFE2067C19F99" attach="left" box="[669,677,421,435]" fontSize="6" pageId="12" pageNumber="86">2</subScript>
|
||
long, curved, convex, CuA
|
||
<subScript id="17FBF90BD233FFA41F8DFE2066869F99" attach="left" box="[986,994,421,435]" fontSize="6" pageId="12" pageNumber="86">2</subScript>
|
||
half the length of CuA
|
||
<subScript id="17FBF90BD233FFA418B2FE2061899F99" attach="left" box="[1253,1261,421,435]" fontSize="6" pageId="12" pageNumber="86">1</subScript>
|
||
; AP free, semielliptic in shape, relatively short; CuP thinner than other veins; A clearly visible; CuP and A meeting at wing margin; in-flight wing-coupling structures present, composed of separate and curved setae. Hind wing (
|
||
<figureCitation id="1344E7CBD233FFA418E2FE5E606F9FDF" box="[1205,1291,475,501]" captionStart="FIGURE 8" captionStartId="12.[151,250,1934,1958]" captionTargetBox="[254,1333,597,1911]" captionTargetId="figure-267@12.[254,1333,597,1911]" captionTargetPageId="12" captionText="FIGURE 8. Longiantennum fashengi. A, Antenna, under green fluorescence. B, Maxillary palpomere. C, Lacinia. D, Fore tibia and tarsus. E, Hind tibial spines. F, Claw. G, Wings, under green fluorescence. Scale bars = 0.2 mm for G, 0.1 mm for A, 0.05 mm for B, D, E, 0.02 mm for C, F." figureDoi="http://doi.org/10.5281/zenodo.10441344" httpUri="https://zenodo.org/record/10441344/files/figure.png" pageId="12" pageNumber="86">Fig. 8G</figureCitation>
|
||
) translucent, without setae on membrane, wing margin or veins; Sc short, not reaching margin; basi-radial cell present, four-angled; R
|
||
<subScript id="17FBF90BD233FFA41D54FDB0646F9C69" attach="left" box="[259,267,565,579]" fontSize="6" pageId="12" pageNumber="86">1</subScript>
|
||
and Rs separating before fusion of Rs with M; Rs, M and A forked; CuA simple; CuP thin, simple.
|
||
</paragraph>
|
||
<caption id="DF00ABC6D233FFA41CC0F80B649899C4" ID-DOI="http://doi.org/10.5281/zenodo.10441344" ID-Zenodo-Dep="10441344" httpUri="https://zenodo.org/record/10441344/files/figure.png" pageId="12" pageNumber="86" startId="12.[151,250,1934,1958]" targetBox="[254,1333,597,1911]" targetPageId="12" targetType="figure">
|
||
<paragraph id="8BC0FB4ED233FFA41CC0F80B649899C4" blockId="12.[151,1436,1934,2031]" pageId="12" pageNumber="86">
|
||
<emphasis id="B90B275CD233FFA41CC0F80B6476998D" bold="true" box="[151,274,1934,1959]" pageId="12" pageNumber="86">FIGURE 8.</emphasis>
|
||
<taxonomicName id="4C7F80CDD233FFA41D4FF80A6777998D" box="[280,531,1935,1959]" class="Insecta" family="Empheriidae" genus="Longiantennum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Psocodea" pageId="12" pageNumber="86" phylum="Arthropoda" rank="species" species="fashengi">
|
||
<emphasis id="B90B275CD233FFA41D4FF80A6777998D" box="[280,531,1935,1959]" italics="true" pageId="12" pageNumber="86">Longiantennum fashengi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B90B275CD233FFA41E48F80B6755998C" bold="true" box="[543,561,1934,1958]" pageId="12" pageNumber="86">A</emphasis>
|
||
, Antenna, under green fluorescence.
|
||
<emphasis id="B90B275CD233FFA41FFFF80A66DD998D" bold="true" box="[936,953,1935,1959]" pageId="12" pageNumber="86">B</emphasis>
|
||
, Maxillary palpomere.
|
||
<emphasis id="B90B275CD233FFA418F1F80B61DC998C" bold="true" box="[1190,1208,1934,1958]" pageId="12" pageNumber="86">C</emphasis>
|
||
, Lacinia.
|
||
<emphasis id="B90B275CD233FFA4194BF80A604A998D" bold="true" box="[1308,1326,1935,1959]" pageId="12" pageNumber="86">D</emphasis>
|
||
, Fore tibia and tarsus.
|
||
<emphasis id="B90B275CD233FFA41D5DF836647F99E1" bold="true" box="[266,283,1971,1995]" pageId="12" pageNumber="86">E</emphasis>
|
||
, Hind tibial spines.
|
||
<emphasis id="B90B275CD233FFA41DB0F836649299E1" bold="true" box="[487,502,1971,1995]" pageId="12" pageNumber="86">F</emphasis>
|
||
, Claw.
|
||
<emphasis id="B90B275CD233FFA41E13F837673C99E0" bold="true" box="[580,600,1970,1994]" pageId="12" pageNumber="86">G</emphasis>
|
||
, Wings, under green fluorescence. Scale bars = 0.2 mm for
|
||
<emphasis id="B90B275CD233FFA418EFF83761A899E0" bold="true" box="[1208,1228,1970,1994]" pageId="12" pageNumber="86">G</emphasis>
|
||
, 0.1 mm for
|
||
<emphasis id="B90B275CD233FFA41906F837600799E0" bold="true" box="[1361,1379,1970,1994]" pageId="12" pageNumber="86">A</emphasis>
|
||
, 0.05 mm for
|
||
<emphasis id="B90B275CD233FFA41CBEF852659E99C5" bold="true" box="[233,250,2007,2031]" pageId="12" pageNumber="86">B</emphasis>
|
||
,
|
||
<emphasis id="B90B275CD233FFA41D51F852647C99C5" bold="true" box="[262,280,2007,2031]" pageId="12" pageNumber="86">D</emphasis>
|
||
,
|
||
<emphasis id="B90B275CD233FFA41D72F852645299C5" bold="true" box="[293,310,2007,2031]" pageId="12" pageNumber="86">E</emphasis>
|
||
, 0.02 mm for
|
||
<emphasis id="B90B275CD233FFA41D90F85364BD99C4" bold="true" box="[455,473,2006,2030]" pageId="12" pageNumber="86">C</emphasis>
|
||
,
|
||
<emphasis id="B90B275CD233FFA41DB1F852649199C5" bold="true" box="[486,501,2007,2031]" pageId="12" pageNumber="86">F</emphasis>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF00ABC6D232FFA51CC0F81261669985" ID-DOI="http://doi.org/10.5281/zenodo.10441348" ID-Zenodo-Dep="10441348" box="[151,1026,1943,1968]" httpUri="https://zenodo.org/record/10441348/files/figure.png" pageId="13" pageNumber="87" startId="13.[151,250,1943,1967]" targetBox="[260,1327,181,1919]" targetPageId="13" targetType="figure">
|
||
<paragraph id="8BC0FB4ED232FFA51CC0F81261669985" blockId="13.[151,1026,1943,1968]" box="[151,1026,1943,1968]" pageId="13" pageNumber="87">
|
||
<emphasis id="B90B275CD232FFA51CC0F81264779985" bold="true" box="[151,275,1943,1967]" pageId="13" pageNumber="87">FIGURE 9.</emphasis>
|
||
<taxonomicName id="4C7F80CDD232FFA51D4EF81D67719985" box="[281,533,1943,1967]" class="Insecta" family="Empheriidae" genus="Longiantennum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Psocodea" pageId="13" pageNumber="87" phylum="Arthropoda" rank="species" species="fashengi">
|
||
<emphasis id="B90B275CD232FFA51D4EF81D67719985" box="[281,533,1943,1967]" italics="true" pageId="13" pageNumber="87">Longiantennum fashengi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B90B275CD232FFA51E76F81267579985" bold="true" box="[545,563,1943,1967]" pageId="13" pageNumber="87">A</emphasis>
|
||
,
|
||
<emphasis id="B90B275CD232FFA51E17F81D6735999A" bold="true" box="[576,593,1944,1968]" pageId="13" pageNumber="87">B</emphasis>
|
||
, Details of genitalia. Scale bars = 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF00ABC6D231FFA61CC0F81267AC99F9" ID-DOI="http://doi.org/10.5281/zenodo.10441350" ID-Zenodo-Dep="10441350" httpUri="https://zenodo.org/record/10441350/files/figure.png" pageId="14" pageNumber="88" startId="14.[151,250,1943,1967]" targetBox="[246,1345,207,1895]" targetPageId="14" targetType="figure">
|
||
<paragraph id="8BC0FB4ED231FFA61CC0F81267AC99F9" blockId="14.[151,1436,1943,2004]" pageId="14" pageNumber="88">
|
||
<emphasis id="B90B275CD231FFA61CC0F81264459985" bold="true" box="[151,289,1943,1967]" pageId="14" pageNumber="88">FIGURE 10.</emphasis>
|
||
<taxonomicName id="4C7F80CDD231FFA61D7FF81D6741999A" box="[296,549,1944,1968]" class="Insecta" family="Empheriidae" genus="Longiantennum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Psocodea" pageId="14" pageNumber="88" phylum="Arthropoda" rank="species" species="fashengi">
|
||
<emphasis id="B90B275CD231FFA61D7FF81D6741999A" box="[296,549,1944,1968]" italics="true" pageId="14" pageNumber="88">Longiantennum fashengi</emphasis>
|
||
</taxonomicName>
|
||
, hand drawings.
|
||
<emphasis id="B90B275CD231FFA61E81F812678C9985" bold="true" box="[726,744,1943,1967]" pageId="14" pageNumber="88">A</emphasis>
|
||
, antennae (female).
|
||
<emphasis id="B90B275CD231FFA61FECF81D66A8999A" bold="true" box="[955,972,1944,1968]" pageId="14" pageNumber="88">B</emphasis>
|
||
, forewing (male).
|
||
<emphasis id="B90B275CD231FFA618DBF81261FA9985" bold="true" box="[1164,1182,1943,1967]" pageId="14" pageNumber="88">C</emphasis>
|
||
, Hind wing (female).
|
||
<emphasis id="B90B275CD231FFA619D3F81D60F2999A" bold="true" box="[1412,1430,1944,1968]" pageId="14" pageNumber="88">D</emphasis>
|
||
, Genitalia. Scale bars = 0.2 mm for
|
||
<emphasis id="B90B275CD231FFA61DACF83E674899F9" bold="true" box="[507,556,1979,2003]" pageId="14" pageNumber="88">A–C</emphasis>
|
||
, 0.1 mm for
|
||
<emphasis id="B90B275CD231FFA61EE7F83967A699FE" bold="true" box="[688,706,1980,2004]" pageId="14" pageNumber="88">D</emphasis>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC0FB4ED230FFA71C90FF1265969F6B" blockId="15.[151,1437,151,1581]" pageId="15" pageNumber="89">
|
||
<emphasis id="B90B275CD230FFA71C90FF1264559E9B" box="[199,305,151,177]" italics="true" pageId="15" pageNumber="89">Abdomen</emphasis>
|
||
: Partially obstructed in ventral view by male hind leg and several fractures over male and female. Epiproct and paraprocts setose; paraproct with apical spine. Male genitalia: hypandrium simple, apically rounded, setose; female genitalia: ovipositor with well-developed external valvulae, membranous, elongated and very broad with multiple very thin long setae; one broad membranous structure visible between paraprocts and external valvulae (
|
||
<figureCitation id="1344E7CBD230FFA71CC8FEA265809F6B" box="[159,228,295,321]" captionStart="FIGURE 9" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[260,1327,181,1919]" captionTargetId="figure-31@13.[260,1327,181,1919]" captionTargetPageId="13" captionText="FIGURE 9. Longiantennum fashengi. A, B, Details of genitalia. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10441348" httpUri="https://zenodo.org/record/10441348/files/figure.png" pageId="15" pageNumber="89">Fig. 9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC0FB4ED230FFA71C90FECE61849FDF" blockId="15.[151,1437,151,1581]" pageId="15" pageNumber="89">
|
||
<emphasis id="B90B275CD230FFA71C90FECE641A9F4F" bold="true" box="[199,382,331,357]" pageId="15" pageNumber="89">Measurements.</emphasis>
|
||
Male. Head length
|
||
<quantity id="4C8756ABD230FFA71E0DFECE67A49F4F" box="[602,704,331,357]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="15" pageNumber="89" unit="mm" value="0.44">0.44 mm</quantity>
|
||
, width
|
||
<quantity id="4C8756ABD230FFA71F45FECE66129F4F" box="[786,886,331,357]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="15" pageNumber="89" unit="mm" value="0.41">0.41 mm</quantity>
|
||
(compound eyes included),
|
||
<quantity id="4C8756ABD230FFA718E4FECE60729F4F" box="[1203,1302,331,357]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="15" pageNumber="89" unit="mm" value="0.24">0.24 mm</quantity>
|
||
(compound eyes excluded); compound eyes length
|
||
<quantity id="4C8756ABD230FFA71E0EFEEA67A59FA3" box="[601,705,367,393]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="15" pageNumber="89" unit="mm" value="0.18">0.18 mm</quantity>
|
||
, width
|
||
<quantity id="4C8756ABD230FFA71F41FEEA661B9FA3" box="[790,895,367,393]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="15" pageNumber="89" unit="mm" value="0.09">0.09 mm</quantity>
|
||
. Thorax length
|
||
<quantity id="4C8756ABD230FFA71864FEEA61FF9FA3" box="[1075,1179,367,393]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="15" pageNumber="89" unit="mm" value="0.42">0.42 mm</quantity>
|
||
, maximum width
|
||
<quantity id="4C8756ABD230FFA7193CFEEA65A29F87" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="15" pageNumber="89" unit="mm" value="0.32">0.32 mm</quantity>
|
||
; prothorax length
|
||
<quantity id="4C8756ABD230FFA71DCDFE1667679F87" box="[410,515,403,429]" metricMagnitude="-5" metricUnit="m" metricValue="3.0" pageId="15" pageNumber="89" unit="mm" value="0.03">0.03 mm</quantity>
|
||
, mesothorax length
|
||
<quantity id="4C8756ABD230FFA71EBAFE1666319F87" box="[749,853,403,429]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="15" pageNumber="89" unit="mm" value="0.17">0.17 mm</quantity>
|
||
, metathorax length
|
||
<quantity id="4C8756ABD230FFA7186CFE1661C09F87" box="[1083,1188,403,429]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="15" pageNumber="89" unit="mm" value="0.22">0.22 mm</quantity>
|
||
; Fw length
|
||
<quantity id="4C8756ABD230FFA71978FE1660FC9F87" box="[1327,1432,403,429]" metricMagnitude="-3" metricUnit="m" metricValue="1.49" pageId="15" pageNumber="89" unit="mm" value="1.49">1.49 mm</quantity>
|
||
, width
|
||
<quantity id="4C8756ABD230FFA71C8AFE3364209FFB" box="[221,324,438,465]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="15" pageNumber="89" unit="mm" value="0.55">0.55 mm</quantity>
|
||
. Abdomen length
|
||
<quantity id="4C8756ABD230FFA71E59FE3267179FFB" box="[526,627,439,465]" metricMagnitude="-4" metricUnit="m" metricValue="8.1" pageId="15" pageNumber="89" unit="mm" value="0.81">0.81 mm</quantity>
|
||
, maximum width
|
||
<quantity id="4C8756ABD230FFA71F6AFE3366C79FFB" box="[829,931,438,465]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="15" pageNumber="89" unit="mm" value="0.35">0.35 mm</quantity>
|
||
. Tarsi length: Ft1
|
||
<quantity id="4C8756ABD230FFA7183CFE3261AB9FFB" box="[1131,1231,439,465]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="15" pageNumber="89" unit="mm" value="0.11">0.11 mm</quantity>
|
||
, Ft2
|
||
<quantity id="4C8756ABD230FFA71951FE3260089FFB" box="[1286,1388,439,465]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="15" pageNumber="89" unit="mm" value="0.04">0.04 mm</quantity>
|
||
, Ft3
|
||
<quantity id="4C8756ABD230FFA71CC0FE5E65999FDF" box="[151,253,475,501]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="15" pageNumber="89" unit="mm" value="0.04">0.04 mm</quantity>
|
||
, Mt1
|
||
<quantity id="4C8756ABD230FFA71D69FE5E64C09FDF" box="[318,420,475,501]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="15" pageNumber="89" unit="mm" value="0.13">0.13 mm</quantity>
|
||
, Mt2
|
||
<quantity id="4C8756ABD230FFA71DB1FE5E67289FDF" box="[486,588,475,501]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="15" pageNumber="89" unit="mm" value="0.04">0.04 mm</quantity>
|
||
, Mt3
|
||
<quantity id="4C8756ABD230FFA71EDAFE5E67979FDF" box="[653,755,475,501]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="15" pageNumber="89" unit="mm" value="0.04">0.04 mm</quantity>
|
||
, Ht1
|
||
<quantity id="4C8756ABD230FFA71F78FE5E66F19FDF" box="[815,917,475,501]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="15" pageNumber="89" unit="mm" value="0.17">0.17 mm</quantity>
|
||
, Ht2
|
||
<quantity id="4C8756ABD230FFA71F85FE5E615C9FDF" box="[978,1080,475,501]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="15" pageNumber="89" unit="mm" value="0.04">0.04 mm</quantity>
|
||
, Ht3
|
||
<quantity id="4C8756ABD230FFA71822FE5F61B89FDF" box="[1141,1244,474,501]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="89" unit="mm" value="0.05">0.05 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC0FB4ED230FFA71C90FE7A67E49CAF" blockId="15.[151,1437,151,1581]" pageId="15" pageNumber="89">
|
||
Female. Head width
|
||
<quantity id="4C8756ABD230FFA71DFAFE7A67749C33" box="[429,528,511,537]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="15" pageNumber="89" unit="mm" value="0.43">0.43 mm</quantity>
|
||
wide (compound eyes included),
|
||
<quantity id="4C8756ABD230FFA71FD2FE7A668C9C33" box="[901,1000,511,537]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="15" pageNumber="89" unit="mm" value="0.3">0.30 mm</quantity>
|
||
(compound eyes excluded), compound eyes length
|
||
<quantity id="4C8756ABD230FFA71D77FDA664EC9C17" box="[288,392,547,573]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="15" pageNumber="89" unit="mm" value="0.18">0.18 mm</quantity>
|
||
, width
|
||
<quantity id="4C8756ABD230FFA71D8AFDA667229C17" box="[477,582,547,573]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="89" unit="mm" value="0.1">0.10 mm</quantity>
|
||
. Thorax length
|
||
<quantity id="4C8756ABD230FFA71EADFDA666069C17" box="[762,866,547,573]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="15" pageNumber="89" unit="mm" value="0.43">0.43 mm</quantity>
|
||
, maximum width
|
||
<quantity id="4C8756ABD230FFA71866FDA661FE9C17" box="[1073,1178,547,573]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="15" pageNumber="89" unit="mm" value="0.31">0.31 mm</quantity>
|
||
; prothorax length
|
||
<quantity id="4C8756ABD230FFA7193CFDA665A19C4B" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="89" unit="mm" value="0.02">0.02 mm</quantity>
|
||
, mesothorax length
|
||
<quantity id="4C8756ABD230FFA71DE7FDC2677D9C4B" box="[432,537,583,609]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="89" unit="mm" value="0.2">0.20 mm</quantity>
|
||
, metathorax length
|
||
<quantity id="4C8756ABD230FFA71F57FDC2660E9C4B" box="[768,874,583,609]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="15" pageNumber="89" unit="mm" value="0.21">0.21 mm</quantity>
|
||
; Fw length
|
||
<quantity id="4C8756ABD230FFA71FA1FDC2613B9C4B" box="[1014,1119,583,609]" metricMagnitude="-3" metricUnit="m" metricValue="1.42" pageId="15" pageNumber="89" unit="mm" value="1.42">1.42 mm</quantity>
|
||
, width
|
||
<quantity id="4C8756ABD230FFA718EFFDC360469C4B" box="[1208,1314,582,609]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="89" unit="mm" value="0.5">0.50 mm</quantity>
|
||
. Abdomen length
|
||
<quantity id="4C8756ABD230FFA71CB3FDEE642E9CAF" box="[228,330,619,645]" metricMagnitude="-4" metricUnit="m" metricValue="6.4" pageId="15" pageNumber="89" unit="mm" value="0.64">0.64 mm</quantity>
|
||
, maximum width
|
||
<quantity id="4C8756ABD230FFA71E42FDEE67189CAF" box="[533,636,619,645]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="15" pageNumber="89" unit="mm" value="0.3">0.30 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC0FB4ED230FFA71C90FD0B67FA9D3F" blockId="15.[151,1437,151,1581]" pageId="15" pageNumber="89">
|
||
<emphasis id="B90B275CD230FFA71C90FD0B64BF9C82" bold="true" box="[199,475,654,680]" pageId="15" pageNumber="89">Additional description.</emphasis>
|
||
Two individuals are encapsulated within the same amber piece, preserved in a mating position. They both appear similar in size, facing opposite directions, with their heads oriented away from each other and their abdomens directed towards one another at terminalia level, but not in direct contact. The male’s genital organs remain completely internalised.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C365A8C5D230FFA71C90FC9A667D9807" pageId="15" pageNumber="89" type="discussion">
|
||
<paragraph id="8BC0FB4ED230FFA71C90FC9A67F09DAB" blockId="15.[151,1437,151,1581]" pageId="15" pageNumber="89">
|
||
<emphasis id="B90B275CD230FFA71C90FC9A645F9D13" bold="true" box="[199,315,799,825]" pageId="15" pageNumber="89">Remarks.</emphasis>
|
||
In specimen NIGP171866-b, one forewing has a very short common stem M
|
||
<subScript id="17FBF90BD230FFA718E5FCB461A89D15" attach="left" box="[1202,1228,817,831]" fontSize="6" pageId="15" pageNumber="89">1+2</subScript>
|
||
(
|
||
<figureCitation id="1344E7CBD230FFA7188AFC9A605C9D13" box="[1245,1336,799,825]" captionStart="FIGURE 8" captionStartId="12.[151,250,1934,1958]" captionTargetBox="[254,1333,597,1911]" captionTargetId="figure-267@12.[254,1333,597,1911]" captionTargetPageId="12" captionText="FIGURE 8. Longiantennum fashengi. A, Antenna, under green fluorescence. B, Maxillary palpomere. C, Lacinia. D, Fore tibia and tarsus. E, Hind tibial spines. F, Claw. G, Wings, under green fluorescence. Scale bars = 0.2 mm for G, 0.1 mm for A, 0.05 mm for B, D, E, 0.02 mm for C, F." figureDoi="http://doi.org/10.5281/zenodo.10441344" httpUri="https://zenodo.org/record/10441344/files/figure.png" pageId="15" pageNumber="89">Fig. 8G</figureCitation>
|
||
), though it is not observed in the second wing or in the male. Wing venation teratisms were also reported in the
|
||
<typeStatus id="54C445ECD230FFA7194CFCC660199D77" box="[1307,1405,835,861]" pageId="15" pageNumber="89" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C7F80CDD230FFA71CC0FCE267F49DAB" authority="(Liang et al., 2022)" baseAuthorityName="Liang" baseAuthorityYear="2022" box="[151,656,871,897]" class="Insecta" family="Empheriidae" genus="Longiantennum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Psocodea" pageId="15" pageNumber="89" phylum="Arthropoda" rank="species" species="fashengi">
|
||
<emphasis id="B90B275CD230FFA71CC0FCE264C99DAB" box="[151,429,871,897]" italics="true" pageId="15" pageNumber="89">Longiantennum fashengi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFEE86BFD230FFA71DECFCE267EC9DAB" author="Liang, F. Y. & Li, S. & Liu, X. Y. & Bai, M. & Yao, Y. Z." box="[443,648,871,897]" pageId="15" pageNumber="89" pagination="105233" refId="ref9702" refString="Liang, F. Y., Li, S., Liu, X. Y., Bai, M. & Yao, Y. Z. (2022) A new genus and species of the family Archaeatropidae (Psocodea: Trogiomorpha) from mid-Cretaceous amber of Northern Myanmar. Cretaceous Research, 138, 105233. https: // doi. org / 10.1016 / j. cretres. 2022.105233" type="journal article" year="2022">
|
||
Liang
|
||
<emphasis id="B90B275CD230FFA71E52FCE2675A9DAB" box="[517,574,871,897]" italics="true" pageId="15" pageNumber="89">et al.</emphasis>
|
||
, 2022
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC0FB4ED230FFA71C90FC0E64E09B1B" blockId="15.[151,1437,151,1581]" pageId="15" pageNumber="89">
|
||
The discovery of a male individual belonging to
|
||
<taxonomicName id="4C7F80CDD230FFA71F4BFC0E66A89D8E" authorityName="Liang, Li & Yao" authorityYear="2022" box="[796,972,907,932]" class="Insecta" family="Empheriidae" genus="Longiantennum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Psocodea" pageId="15" pageNumber="89" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90B275CD230FFA71F4BFC0E66A89D8E" box="[796,972,907,932]" italics="true" pageId="15" pageNumber="89">Longiantennum</emphasis>
|
||
</taxonomicName>
|
||
suggests that male genitalia is mainly internalised—the only external structure being the hypandrium—, as it is usually found in non-parasitic
|
||
<taxonomicName id="4C7F80CDD230FFA7197BFC2A60FD9DE3" authorityName="Hennig" authorityYear="1966" box="[1324,1433,943,969]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Psocodea" pageId="15" pageNumber="89" phylum="Arthropoda" rank="order">Psocodea</taxonomicName>
|
||
, which warrants a hypothesis that both
|
||
<typeStatus id="54C445ECD230FFA71E05FC5667D09DC7" box="[594,692,979,1005]" pageId="15" pageNumber="89" type="holotype">holotype</typeStatus>
|
||
CNU-PSO-MA-2016020 and
|
||
<typeStatus id="54C445ECD230FFA71845FC5661169DC7" box="[1042,1138,979,1005]" pageId="15" pageNumber="89" type="paratype">paratype</typeStatus>
|
||
CAU-BA-LFY-21003 are actually female specimens, possibility of two different species based on the descriptions of
|
||
<bibRefCitation id="EFEE86BFD230FFA718ECFC7260FD9A3B" author="Liang, F. Y. & Li, S. & Liu, X. Y. & Bai, M. & Yao, Y. Z." box="[1211,1433,1015,1041]" pageId="15" pageNumber="89" pagination="105233" refId="ref9702" refString="Liang, F. Y., Li, S., Liu, X. Y., Bai, M. & Yao, Y. Z. (2022) A new genus and species of the family Archaeatropidae (Psocodea: Trogiomorpha) from mid-Cretaceous amber of Northern Myanmar. Cretaceous Research, 138, 105233. https: // doi. org / 10.1016 / j. cretres. 2022.105233" type="journal article" year="2022">
|
||
Liang
|
||
<emphasis id="B90B275CD230FFA71951FC7260259A3B" box="[1286,1345,1015,1041]" italics="true" pageId="15" pageNumber="89">et al.</emphasis>
|
||
(2022)
|
||
</bibRefCitation>
|
||
. We examined the
|
||
<typeStatus id="54C445ECD230FFA71D34FB9E64A19A1F" box="[355,453,1051,1077]" pageId="15" pageNumber="89" type="holotype">holotype</typeStatus>
|
||
CNU-PSO-MA-2016020, but could not observe any structures to suggest the presence of the hypandrium. In fact, no genitalia structures are clearly visible in ventral view; two membranous rod-like lobes are distinguished in lateral view. We believe that the structures identified originally as parameres in
|
||
<typeStatus id="54C445ECD230FFA7196DFBE660F89A57" box="[1338,1436,1123,1149]" pageId="15" pageNumber="89" type="holotype">holotype</typeStatus>
|
||
CNU-PSO-MA-2016020 could be the setose external valvulae and the darkish edge of the originally identified hypandrium is the apex of the female subgenital plate. We note that the external valvulae in NIGP171866-b appear to be wider and longer than those of the
|
||
<typeStatus id="54C445ECD230FFA71E01FB4A67DF9AC3" box="[598,699,1231,1257]" pageId="15" pageNumber="89" type="holotype">holotype</typeStatus>
|
||
, though we cannot confirm whether it is a diagnostic difference or an artifact of preservation in the resin. The collection of more material belonging to
|
||
<taxonomicName id="4C7F80CDD230FFA7181DFB76619E9B26" authorityName="Liang, Li & Yao" authorityYear="2022" box="[1098,1274,1267,1292]" class="Insecta" family="Empheriidae" genus="Longiantennum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Psocodea" pageId="15" pageNumber="89" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90B275CD230FFA7181DFB76619E9B26" box="[1098,1274,1267,1292]" italics="true" pageId="15" pageNumber="89">Longiantennum</emphasis>
|
||
</taxonomicName>
|
||
would help to clarify this character.
|
||
</paragraph>
|
||
<paragraph id="8BC0FB4ED230FFA71C90FABE667D9807" blockId="15.[151,1437,151,1581]" pageId="15" pageNumber="89">
|
||
The only other difference between CNU-PSO-MA-2016020 and specimens of NIGP171866 according to the description by
|
||
<bibRefCitation id="EFEE86BFD230FFA71D3BFADA67279B53" author="Liang, F. Y. & Li, S. & Liu, X. Y. & Bai, M. & Yao, Y. Z." box="[364,579,1375,1401]" pageId="15" pageNumber="89" pagination="105233" refId="ref9702" refString="Liang, F. Y., Li, S., Liu, X. Y., Bai, M. & Yao, Y. Z. (2022) A new genus and species of the family Archaeatropidae (Psocodea: Trogiomorpha) from mid-Cretaceous amber of Northern Myanmar. Cretaceous Research, 138, 105233. https: // doi. org / 10.1016 / j. cretres. 2022.105233" type="journal article" year="2022">
|
||
Liang et
|
||
<emphasis id="B90B275CD230FFA71D85FADA64949B53" box="[466,496,1375,1401]" italics="true" pageId="15" pageNumber="89">al.</emphasis>
|
||
(2022)
|
||
</bibRefCitation>
|
||
is the absence/presence of a sensillum on the second maxillary palpomere. However, the sensillum is a very delicate character to observe—especially without an optical microscope—as the palpomeres are not always well positioned for an optimal observation. It is present in both the male and female in NIGP171866, and we could vaguely see what might be a sensillum in CNU-PSO-MA-2016020. Thus, we tentatively assign the new individuals to
|
||
<taxonomicName id="4C7F80CDD230FFA71D8AFA6A67969823" box="[477,754,1519,1545]" class="Insecta" family="Empheriidae" genus="Longiantennum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Psocodea" pageId="15" pageNumber="89" phylum="Arthropoda" rank="species" species="fashengi">
|
||
<emphasis id="B90B275CD230FFA71D8AFA6A67969823" box="[477,754,1519,1545]" italics="true" pageId="15" pageNumber="89">Longiantennum fashengi</emphasis>
|
||
</taxonomicName>
|
||
, and refrain from establishing a new species—until definitive diagnostic traits are discovered on future new specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |