166 lines
20 KiB
XML
166 lines
20 KiB
XML
<document id="9781540EFBB3DA08F54FAD2FDC77393B" ID-CLB-Dataset="9312" ID-DOI="10.11646/zootaxa.4808.2.9" ID-GBIF-Dataset="93d8b512-f031-4345-a6fe-83d6f563ced8" ID-ISSN="1175-5326" ID-Zenodo-Dep="3933634" ID-ZooBank="8EDEC876-C608-46F4-91FE-13AB3F0DC029" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1594125075647" checkinUser="plazi" docAuthor="Frolov, Andrey V., Akhmetova, Lilia A. & Vishnevskaya, Maria S." docDate="2020" docId="03D45649FF84F174FF612B21FC43FC1D" docLanguage="en" docName="zootaxa.4808.2.9.pdf" docOrigin="Zootaxa 4808 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Madecorphnus grebennikovi Frolov, Akhmetova & Vishnevskaya 2020, new species" docType="treatment" docVersion="6" lastPageNumber="376" masterDocId="FFED2E31FF85F177FFF62F63FFD8FFD6" masterDocTitle="A new species of the endemic Madagascan scarab beetle genus, Madecorphnus grebennikovi (Coleoptera: Scarabaeidae: Orphninae): morphological description and DNA barcode" masterLastPageNumber="378" masterPageNumber="373" pageNumber="374" updateTime="1698843121469" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="44C9ADEBF8B2BA7852FC0F3E1E331968" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B5FD2A41B5279ADB120A0FBBC279CE71">
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<mods:title id="4907588D0E6499174B8A610BC97ABDCE">A new species of the endemic Madagascan scarab beetle genus, Madecorphnus grebennikovi (Coleoptera: Scarabaeidae: Orphninae): morphological description and DNA barcode</mods:title>
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</mods:titleInfo>
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<mods:name id="25E5358A175AF67198CA432532B4F99C" type="personal">
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<mods:role id="83F1C2768B202CE6C49C4DC2402E282E">
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<mods:roleTerm id="01398E6EBEBF07BA79FAF13464200BA4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="1B59CB9FD5E563E02E2EE1D0FC44C531">Frolov, Andrey V.</mods:namePart>
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<mods:affiliation id="80B6404C174B7CF75FF67C016B3DF3FF">Zoological Institute, Russian Academy of Sciences, Universitetskaya nab., 1, Saint-Petersburg 199034, Russia. frolov @ scarabaeoidea. com; https: // orcid. org / 0000 - 0002 - 6724 - 6828</mods:affiliation>
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</mods:name>
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<mods:name id="400CA1ED3EFEB40448441172F5E91BEA" type="personal">
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<mods:roleTerm id="A525A9AE66633A01957F34C3249B1CF7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="EC4AAE1E226B664A899F6200E3345617">Akhmetova, Lilia A.</mods:namePart>
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<mods:affiliation id="BE1EB8B029E1252DCB97DCA7B14FCF31">Zoological Institute, Russian Academy of Sciences, Universitetskaya nab., 1, Saint-Petersburg 199034, Russia. frolov @ scarabaeoidea. com; https: // orcid. org / 0000 - 0002 - 6724 - 6828</mods:affiliation>
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</mods:name>
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<mods:name id="B46F1C9FB1E6CD365B06C9B0135487B0" type="personal">
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<mods:roleTerm id="619A12F94995E8715CDCD4FAE36B4FFA">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6697E881A564F23CC579CD932C72F473">Vishnevskaya, Maria S.</mods:namePart>
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<mods:affiliation id="DB5DF7A4EC2F6EA1ECFEB9F63B75BB45">Chromas Research Resource Center, Saint-Petersburg State University, Oranienbaumskoe hwy. 2 - 5, Saint-Petersburg 198504, Russia.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="0F21198D1F5C4957701B813E1AB707AB">text</mods:typeOfResource>
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<mods:title id="240E30005DAB407CC10128C06EC8C75B">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="03B08394A45CBAF4743C8284C865FDEC">
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<mods:date id="C9B52715AED3154CD2F17C76E5C475AD">2020</mods:date>
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<mods:number id="0D2D4F6E92FB8EB56300DC7A6A9B11D5">2020-07-02</mods:number>
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<mods:number id="72F5BC7A4E4FA743A676BB097691FE6A">4808</mods:number>
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<mods:start id="4BEEDAB17DD9CBB63B3CB480BD6C1234">373</mods:start>
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<mods:identifier id="3D572471A19AB54F9DD2E061318B8A0D" type="DOI">10.11646/zootaxa.4808.2.9</mods:identifier>
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<mods:identifier id="6E9803AC9597F7F519CD616BADC3825C" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="8D25E96FBAEF689BC5991EC1FEFCD528" type="Zenodo-Dep">3933634</mods:identifier>
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<mods:identifier id="0FC9E7291B4C55760613D9176731ED89" type="ZooBank">8EDEC876-C608-46F4-91FE-13AB3F0DC029</mods:identifier>
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</mods:mods>
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<treatment id="03D45649FF84F174FF612B21FC43FC1D" ID-GBIF-Taxon="176086483" LSID="urn:lsid:plazi:treatment:03D45649FF84F174FF612B21FC43FC1D" httpUri="http://treatment.plazi.org/id/03D45649FF84F174FF612B21FC43FC1D" lastPageId="3" lastPageNumber="376" pageId="1" pageNumber="374">
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<subSubSection id="C367B4D4FF84F176FF612B21FECBFBA9" pageId="1" pageNumber="374" type="nomenclature">
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<paragraph id="8BC2E75FFF84F176FF612B21FBBFFB8D" blockId="1.[151,1127,1089,1151]" box="[151,1127,1089,1116]" pageId="1" pageNumber="374">
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<heading id="D08A5033FF84F176FF612B21FBBFFB8D" bold="true" box="[151,1127,1089,1116]" fontSize="11" level="1" pageId="1" pageNumber="374" reason="1">
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<emphasis id="B9093B4DFF84F176FF612B21FBBFFB8D" bold="true" box="[151,1127,1089,1116]" pageId="1" pageNumber="374">
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<taxonomicName id="4C7D9CDCFF84F176FF612B21FC13FB8D" authority="Frolov, Akhmetova & Vishnevskaya" authorityName="Frolov, Akhmetova & Vishnevskaya" authorityYear="2020" box="[151,971,1089,1116]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="374" phylum="Arthropoda" rank="species" species="grebennikovi" status="sp. nov.">Madecorphnus grebennikovi Frolov, Akhmetova & Vishnevskaya</taxonomicName>
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,
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<taxonomicNameLabel id="A23A8636FF84F176FC212B21FBBFFB8D" box="[983,1127,1089,1116]" pageId="1" pageNumber="374" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC2E75FFF84F176FF612B06FECBFBA9" blockId="1.[151,1127,1089,1151]" box="[151,275,1125,1151]" pageId="1" pageNumber="374">(Figs. 1, 2)</paragraph>
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</subSubSection>
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<subSubSection id="C367B4D4FF84F176FF612BCDFE55FB3D" pageId="1" pageNumber="374" type="materials_examined">
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<paragraph id="8BC2E75FFF84F176FF612BCDFE55FB3D" blockId="1.[151,1437,1197,2016]" pageId="1" pageNumber="374">
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<emphasis id="B9093B4DFF84F176FF612BCDFE9DFB1E" bold="true" box="[151,325,1197,1224]" pageId="1" pageNumber="374">Type material.</emphasis>
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<materialsCitation id="3B15ED02FF84F176FEBA2BCEFE55FB3D" ID-GBIF-Occurrence="2995049301" collectingDate="2018-12-10" collectionCode="ZIN, V" country="Madagascar" elevation="1321" latitude="-14.4367" location="Marojejy" longLatPrecision="7" longitude="49.7435" pageId="1" pageNumber="374" specimenCode="MD17" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54C659FDFF84F176FEBA2BCEFE61FB11" box="[332,441,1197,1223]" pageId="1" pageNumber="374" type="holotype">Holotype</typeStatus>
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: male at
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<collectionCode id="ED6C7F9AFF84F176FDEA2BCDFD93FB1E" box="[540,587,1198,1224]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="1" pageNumber="374" type="Museum">ZIN</collectionCode>
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labeled “
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<collectingCountry id="F36AA7CFFF84F176FD452BCEFCE5FB11" box="[691,829,1197,1223]" name="Madagascar" pageId="1" pageNumber="374">Madagascar</collectingCountry>
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<location id="8EA2B184FF84F176FCB22BCEFC73FB11" LSID="urn:lsid:plazi:treatment:03D45649FF84F174FF612B21FC43FC1D:8EA2B184FF84F176FCB22BCEFC73FB11" box="[836,939,1197,1223]" country="Madagascar" latitude="-14.4367" longLatPrecision="7" longitude="49.7435" name="Marojejy" pageId="1" pageNumber="374">Marojejy</location>
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<geoCoordinate id="EE498198FF84F176FC472BCEFBCEFB11" box="[945,1046,1197,1223]" degrees="14.4367" direction="south" orientation="latitude" pageId="1" pageNumber="374" precision="5" value="-14.4367">-14.4367</geoCoordinate>
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<geoCoordinate id="EE498198FF84F176FBEB2BCEFBA0FB1E" box="[1053,1144,1197,1224]" degrees="49.7435" direction="east" orientation="longitude" pageId="1" pageNumber="374" precision="5" value="49.7435">49.7435</geoCoordinate>
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<quantity id="4C854ABAFF84F176FB892BCEFB14FB11" box="[1151,1228,1197,1223]" metricMagnitude="3" metricUnit="m" metricValue="1.321" pageId="1" pageNumber="374" unit="m" value="1321.0">
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<elevation id="0050006CFF84F176FB892BCEFB14FB11" box="[1151,1228,1197,1223]" metricMagnitude="3" metricUnit="m" metricValue="1.321" pageId="1" pageNumber="374" unit="m" value="1321.0">1321m</elevation>
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</quantity>
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<date id="FFC3C19FFF84F176FB222BCEFA8BFB11" box="[1236,1363,1197,1223]" pageId="1" pageNumber="374" value="2018-12-10">
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<collectingDate id="EF873877FF84F176FB222BCEFA8BFB11" box="[1236,1363,1197,1223]" pageId="1" pageNumber="374" value="2018-12-10">10.xii.2018</collectingDate>
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</date>
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sift
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<collectionCode id="ED6C7F9AFF84F176FA732BCDFA41FB1E" box="[1413,1433,1198,1224]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="374" type="Museum">V</collectionCode>
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. Grebennikov
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<specimenCode id="DBDB4F24FF84F176FEC62BB2FEA6FB3D" box="[304,382,1233,1259]" collectionCode="MD" country="Germany" name="Museum Donaueschingen" pageId="1" pageNumber="374">MD17</specimenCode>
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”.
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C367B4D4FF84F175FF312B96FF16F823" lastPageId="2" lastPageNumber="375" pageId="1" pageNumber="374" type="description">
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<paragraph id="8BC2E75FFF84F176FF312B96FC46FAE5" blockId="1.[151,1437,1197,2016]" pageId="1" pageNumber="374">
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<emphasis id="B9093B4DFF84F176FF312B96FE81FAC6" bold="true" box="[199,345,1269,1296]" pageId="1" pageNumber="374">Description.</emphasis>
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<typeStatus id="54C659FDFF84F176FEA92B96FE12FAD9" box="[351,458,1269,1295]" pageId="1" pageNumber="374" type="holotype">Holotype</typeStatus>
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, male (Fig. 1). Body length
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<quantity id="4C854ABAFF84F176FCF22B96FC82FAC6" box="[772,858,1269,1296]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="1" pageNumber="374" unit="mm" value="5.3">5.3 mm</quantity>
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(with mandibles). Color of head and pronotum uniformly brown, disc of elytra and anterior part of head slightly lighter.
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</paragraph>
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<paragraph id="8BC2E75FFF84F176FF312A5EFF27FA31" blockId="1.[151,1437,1197,2016]" pageId="1" pageNumber="374">Right mandible as long as left, without tooth behind apex. Labrum subtrapezoidal, with slightly rounded sides, length about 1/8 width (in dorsal view). Clypeus symmetrical, apically obtuse, with two long and one shorter setae on the apical margin. Canthus and frontal suture indistinct. Clypeus slightly depressed apicomedially. Head without traces of frontoclypeal suture, finely punctate with minute punctures separated by greater than four times their diameter.</paragraph>
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<paragraph id="8BC2E75FFF84F176FF312A91FDC9F9A1" blockId="1.[151,1437,1197,2016]" pageId="1" pageNumber="374">Pronotum approximately 1.6 times wider than long, widest medially. Disc of pronotum convex, without any depressions, tubercles, or ridges. Punctation on pronotum similar to that on head. Margins with relatively wide border, lateral margins with four long setae: one seta on basal angle, one seta approximately in the middle of lateral margin, and two setae on the apical angle.</paragraph>
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<paragraph id="8BC2E75FFF84F176FF3129E2FC04F94D" blockId="1.[151,1437,1197,2016]" box="[199,988,1665,1691]" pageId="1" pageNumber="374">Scutellum subtriangular, angulate apically, about 1/12 length of elytra.</paragraph>
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<paragraph id="8BC2E75FFF84F176FF3129C6FB34F8FD" blockId="1.[151,1437,1197,2016]" pageId="1" pageNumber="374">Elytra convex, with distinct humeral and apical umbones, widest at middle. Stria I distinct and reaching the apex of elytron, other striae indistinct. Elytra with double punctation: entire surface with minute punctures similar to those on head and pronotum; disc with larger, sparse, elongate, setigerous punctures. Epipleura with long, sparse, brown setae. Base of elytron with border from scutellum to humeral callus. Wings fully developed.</paragraph>
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<paragraph id="8BC2E75FFF84F176FF312856FEBBF86D" blockId="1.[151,1437,1197,2016]" pageId="1" pageNumber="374">Protibiae with three outer teeth, lateral margin basad of outer teeth not crenulate. Apex with robust, spur-like seta and a few smaller setae basally. Mesothoracic and metathoracic legs similar in shape to each other. Longer tibial spur slightly shorter than mesotarsomeres I–II in mesothoracic legs and as long as metatarsomeres I–II in metathoracic legs.</paragraph>
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<paragraph id="8BC2E75FFF84F175FF3128A5FD6CFF2D" blockId="1.[151,1437,1197,2016]" lastBlockId="2.[151,1436,153,287]" lastPageId="2" lastPageNumber="375" pageId="1" pageNumber="374">Parameres 0.6 length of phallobase, narrowly rounded in lateral view (Fig. 1F), their apical part as wide as basal part in dorsal view, with small but distinct lateral teeth (Fig. 1D). Endophallus with 1) a long, almost straight sclerite with attached to its end a 2/3 shorter, somewhat curved sclerite, 2) a separate smaller, elongate sclerite, and 3) a rather large area of microspinules (Fig. 1E).</paragraph>
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<paragraph id="8BC2E75FFF87F175FF312E66FE4DFEC9" blockId="2.[151,1436,153,287]" box="[199,405,261,287]" pageId="2" pageNumber="375">
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<emphasis id="B9093B4DFF87F175FF312E66FEC6FEC9" bold="true" box="[199,286,261,287]" pageId="2" pageNumber="375">Female</emphasis>
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unknown.
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</paragraph>
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<paragraph id="8BC2E75FFF87F175FF6128F0FF16F823" blockId="2.[151,1436,1939,2037]" pageId="2" pageNumber="375">
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<emphasis id="B9093B4DFF87F175FF6128F0FECFF87B" bold="true" box="[151,279,1939,1965]" pageId="2" pageNumber="375">FIGURE 1.</emphasis>
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<typeStatus id="54C659FDFF87F175FED628F7FEA6F87B" box="[288,382,1940,1965]" pageId="2" pageNumber="375" type="holotype">Holotype</typeStatus>
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of
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<taxonomicName id="4C7D9CDCFF87F175FE5028F6FB95F87B" authority="Frolov, Akhmetova & Vishnevskaya" authorityName="Frolov, Akhmetova & Vishnevskaya" authorityYear="2020" box="[422,1101,1940,1965]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="375" phylum="Arthropoda" rank="species" species="grebennikovi" status="sp. nov.">
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<emphasis id="B9093B4DFF87F175FE5028F6FD12F87B" box="[422,714,1941,1965]" italics="true" pageId="2" pageNumber="375">Madecorphnus grebennikovi</emphasis>
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Frolov, Akhmetova & Vishnevskaya
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</taxonomicName>
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,
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<taxonomicNameLabel id="A23A8636FF87F175FBAC28F6FB01F87B" box="[1114,1241,1940,1965]" pageId="2" pageNumber="375" rank="species">new species</taxonomicNameLabel>
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. A, habitus, dorsal view; B, habitus, dorsolateral view; C, habitus, lateral view; D, parameres, dorsal view; E, endophallus; F, aedeagus, lateral view.
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</paragraph>
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</subSubSection>
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<subSubSection id="C367B4D4FF86F174FF312FFBFF29FCED" pageId="3" pageNumber="376" type="diagnosis">
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<paragraph id="8BC2E75FFF86F174FF312FFBFD90FE94" blockId="3.[151,1436,152,971]" pageId="3" pageNumber="376">
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<emphasis id="B9093B4DFF86F174FF312FFBFE9AFF65" bold="true" box="[199,322,152,179]" pageId="3" pageNumber="376">Diagnosis.</emphasis>
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The new species is distinguished from other
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<taxonomicName id="4C7D9CDCFF86F174FCB32FFBFC35FF64" authorityName="Paulian" authorityYear="1992" box="[837,1005,152,178]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="376" phylum="Arthropoda" rank="genus">
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<emphasis id="B9093B4DFF86F174FCB32FFBFC35FF64" box="[837,1005,152,178]" italics="true" pageId="3" pageNumber="376">Madecorphnus</emphasis>
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</taxonomicName>
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species by the unique combination of the shape of the parameres and armature of the endophallus, namely the parameres being narrowly rounded in lateral view and having a small but distinct lateral teeth, and the endophallic armature consisting of 1) a long straight sclerite with attached to its end a 2/3 shorted, somewhat curved sclerite, 2) separate smaller, elongate sclerite, and 3) a rather large area of microspinules.
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</paragraph>
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<paragraph id="8BC2E75FFF86F174FF312E2FFF29FCED" blockId="3.[151,1436,152,971]" pageId="3" pageNumber="376">
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<emphasis id="B9093B4DFF86F174FF312E2FFEA8FEB1" bold="true" box="[199,368,332,359]" pageId="3" pageNumber="376">DNA barcode.</emphasis>
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GenBank accession number
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<accessionNumber id="942E7ABCFF86F174FD412E2FFCE0FEB0" box="[695,824,332,358]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT361868" pageId="3" pageNumber="376">MT361868</accessionNumber>
|
||
, length 811 base pairs: TAAAGATATTGGAACATTGTATTTCCTATTTGGAAGTTGAGCAGGAATAGTAGGAACCTCCCTA- AGCCTATTAATCCGTGCCGAACTAGGAAACCCTGGAACTTTAATTGGTGACGATCAAATTTATAAC- GTAATCGTAACAGCTCATGCATTTGTAATAATTTTTTTTATAGTAATACCTATTATAATTGGTGGGTTTG- GAAACTGACTTGTTCCATTAATGCTAGGTGCTCCCGACATAGCATTCCCTCGAATAAACAACATA- AGGTTTTGGCTTCTCCCCCCATCCTTAACACTTTTATTAACTAGAAGAATAGTTGAAAGTGGTGCAG- GTACAGGTTGAACGGTTTATCCTCCCCTTTCAGCTAACATTGCCCATAGAGGTGCATCTGTTGACCTA GCTATTTTTAGCCTTCACCTAGCAGGAATCTCTTCAATTCTAGGGGCTGTAAATTTTATTACTACAGTA- ATCAATATACGATCCACTGGAATAACTTTTGATCGAATACCTTTATTTGTATGGTCCGTAGCATTAACTG CTTTATTACTCTTGCTCTCYCTGCCTGTTTTAGCTGGTGCAATTACCATGTTATTAACAGATCGAAATT- TAAACACTACTTTTTTTGACCCAGCCGGAGGAGGCGATCCAATTCTATACCAACACCTATTTT- GATTCTTTGGACACCCAGAAGTATACATTTTAATTCTACCTGGATTTGGTATAATCTCCCATATTATT AGGCAAGAAAGAAGAAAAAAGGAAACTTTTGGTACTTTAGGTATAATTTATGCTATACTAGCAATTG- GATTA
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C367B4D4FF86F174FF312C27FDCAFC55" pageId="3" pageNumber="376" type="distribution">
|
||
<paragraph id="8BC2E75FFF86F174FF312C27FDCAFC55" blockId="3.[151,1436,152,971]" pageId="3" pageNumber="376">
|
||
<emphasis id="B9093B4DFF86F174FF312C27FEB9FC89" bold="true" box="[199,353,836,863]" pageId="3" pageNumber="376">Distribution.</emphasis>
|
||
The species is known from the
|
||
<typeStatus id="54C659FDFF86F174FD2B2C26FCD5FC89" box="[733,781,837,863]" pageId="3" pageNumber="376">type</typeStatus>
|
||
locality only, the Marojejy National Park, Sava Region, northeastern
|
||
<collectingCountry id="F36AA7CFFF86F174FEDF2C0BFE68FC54" box="[297,432,872,898]" name="Madagascar" pageId="3" pageNumber="376">Madagascar</collectingCountry>
|
||
(
|
||
<figureCitation id="1346FBDAFF86F174FE492C0BFDDDFC55" box="[447,517,872,899]" captionStart="FIGURE 2" captionStartId="3.[151,250,1776,1801]" captionTargetBox="[404,1184,994,1747]" captionTargetId="figure@3.[401,1187,990,1748]" captionTargetPageId="3" captionText="FIGURE 2. Locality map of Madecorphnus Paulian, 1992 species in northern Madagascar." figureDoi="http://doi.org/10.5281/zenodo.3933636" httpUri="https://zenodo.org/record/3933636/files/figure.png" pageId="3" pageNumber="376">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C367B4D4FF86F174FF312CEEFC43FC1D" pageId="3" pageNumber="376" type="etymology">
|
||
<paragraph id="8BC2E75FFF86F174FF312CEEFC43FC1D" blockId="3.[151,1436,152,971]" pageId="3" pageNumber="376">
|
||
<emphasis id="B9093B4DFF86F174FF312CEEFE96FC71" bold="true" box="[199,334,909,935]" pageId="3" pageNumber="376">Etymology.</emphasis>
|
||
The new species is named after Vasily V. Grebennikov (Canadian Food Inspection Agency, Ottawa,
|
||
<collectingCountry id="F36AA7CFFF86F174FF612CD3FF35FC1D" box="[151,237,944,971]" name="Canada" pageId="3" pageNumber="376">Canada</collectingCountry>
|
||
) who collected the
|
||
<typeStatus id="54C659FDFF86F174FE3E2CD2FE20FC1D" box="[456,504,945,971]" pageId="3" pageNumber="376">type</typeStatus>
|
||
specimen. Noun in the genitive case.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |