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<document id="66742A7B917E283ED0B5429CD9553919" ID-CLB-Dataset="8695" ID-DOI="10.11646/zootaxa.4829.1.1" ID-GBIF-Dataset="04b19d7f-1d74-41ec-b657-9b57f47826f9" ID-ISSN="1175-5326" ID-Zenodo-Dep="4402757" ID-ZooBank="7B34E058-03B4-44D0-AC4E-065B010172E1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1609343828179" checkinUser="plazi" docAuthor="Jaschhof, Mathias &amp; Jaschhof, Catrin" docDate="2020" docId="03C00F49FF876E35FF57F8909827F9DD" docLanguage="en" docName="zootaxa.4829.1.1.pdf" docOrigin="Zootaxa 4829 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Winnertzia imbecilla Jaschhof &amp; Jaschhof 2020, sp. nov." docType="treatment" docVersion="8" lastPageNumber="14" masterDocId="FFF97731FF8B6E38FFC0FFBC9D54FF8C" masterDocTitle="Reevaluation of species richness in Winnertzia (Diptera, Cecidomyiidae, Winnertziinae), with descriptions of 37 new species from Sweden, Peru and Australia" masterLastPageNumber="72" masterPageNumber="1" pageNumber="13" updateTime="1698849859117" updateUser="ExternalLinkService">
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<mods:title id="6F5A7AB31F681D46F888771720DB9775">Reevaluation of species richness in Winnertzia (Diptera, Cecidomyiidae, Winnertziinae), with descriptions of 37 new species from Sweden, Peru and Australia</mods:title>
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<mods:namePart id="3EF78172A6388A20FD984EBB01F03D96">Jaschhof, Mathias</mods:namePart>
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<mods:namePart id="68A0BF51C98ED601E31D373B364145BA">Jaschhof, Catrin</mods:namePart>
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<treatment id="03C00F49FF876E35FF57F8909827F9DD" ID-DOI="http://doi.org/10.5281/zenodo.4402412" ID-GBIF-Taxon="177020951" ID-Zenodo-Dep="4402412" LSID="urn:lsid:plazi:treatment:03C00F49FF876E35FF57F8909827F9DD" httpUri="http://treatment.plazi.org/id/03C00F49FF876E35FF57F8909827F9DD" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<subSubSection id="C373EDD4FF876E34FF57F8909CA3F8CA" box="[151,503,1835,1862]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="8BD6BE5FFF876E34FF57F8909CA3F8CA" blockId="12.[151,503,1835,1898]" box="[151,503,1835,1862]" pageId="12" pageNumber="13">
<heading id="D09E0933FF876E34FF57F8909CA3F8CA" bold="true" box="[151,503,1835,1862]" fontSize="11" level="2" pageId="12" pageNumber="13" reason="4">
<emphasis id="B91D624DFF876E34FF57F8909CA3F8CA" bold="true" box="[151,503,1835,1862]" pageId="12" pageNumber="13">
<taxonomicName id="4C69C5DCFF876E34FF57F8909CC6F8CA" ID-CoL="8FMHS" authority="Jaschhof &amp; Jaschhof, 2020" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2020" box="[151,402,1835,1862]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="imbecilla" status="sp. nov.">
<emphasis id="B91D624DFF876E34FF57F8909CC6F8CA" bold="true" box="[151,402,1835,1862]" italics="true" pageId="12" pageNumber="13">Winnertzia imbecilla</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EDF36FF876E34FE59F8909CA3F8CA" box="[409,503,1836,1862]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C373EDD4FF876E35FF57F8EC9CAAFF59" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="description">
<paragraph id="8BD6BE5FFF876E34FF57F8EC9C40F8E6" blockId="12.[151,503,1835,1898]" box="[151,276,1872,1898]" pageId="12" pageNumber="13">
<figureCitation id="1352A2DAFF876E34FF57F8EC9C40F8E6" box="[151,276,1872,1898]" captionStart="FIGURES 713" captionStartId="12.[151,264,1621,1645]" captionTargetBox="[154,1433,188,1588]" captionTargetId="figure-25@12.[151,1436,181,1598]" captionTargetPageId="12" captionText="FIGURES 713. Male morphology of Winnertzia brachytarsus (79) and Winnertzia imbecilla (1013), holotypes. 7: Fourth flagellomere, lateral. 8: Ditto, medial. 9: Genitalia, ventral. 10: Genitalia, ventral. 11: Palpus, lateral. 12: Fourth flagellomere, lateral. 13: Ditto, medial. Scales for 710 and 1213, 0.05 mm; for 11, 0.025 mm. Arrows refer to characters described in the diagnoses." figureDoi="http://doi.org/10.5281/zenodo.4402763" httpUri="https://zenodo.org/record/4402763/files/figure.png" pageId="12" pageNumber="13">Figs 1013</figureCitation>
</paragraph>
<paragraph id="8BD6BE5FFF876E35FF57F8249CAAFF59" blockId="12.[151,1437,1944,2042]" lastBlockId="13.[151,1437,150,1619]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
The specimen designated here as the
<typeStatus id="54D200FDFF876E34FDF6F8249FCCF83E" box="[566,664,1944,1970]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C69C5DCFF876E34FD7DF8259E1CF83E" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2020" box="[701,840,1944,1970]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="imbecilla">
<emphasis id="B91D624DFF876E34FD7DF8259E1CF83E" box="[701,840,1944,1970]" italics="true" pageId="12" pageNumber="13">W. imbecilla</emphasis>
</taxonomicName>
was published in error as the first Swedish find of
<taxonomicName id="4C69C5DCFF876E34FA40F8259F30F85A" authority="(Jaschhof &amp; Jaschhof 2017)" baseAuthorityName="Jaschhof &amp; Jaschhof" baseAuthorityYear="2017" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="brachypalpa">
<emphasis id="B91D624DFF876E34FA40F8259C71F85A" italics="true" pageId="12" pageNumber="13">W. brachypalpa</emphasis>
(
<bibRefCitation id="EFF8C3AEFF876E34FEF2F8009F08F85A" author="Jaschhof, M. &amp; Jaschhof, C." box="[306,604,1980,2006]" pageId="12" pageNumber="13" pagination="546 - 570" refId="ref56260" refString="Jaschhof, M. &amp; Jaschhof, C. (2017) Mycophagous gall midges (Diptera, Cecidomyiidae: Lestremiinae, Micromyinae, Winnertziinae, Porricondylinae): first records in Sweden and descriptions of closely related new species from elsewhere. Zootaxa, 4226 (4), 546 - 570. https: // doi. org / 10.11646 / zootaxa. 4226.4.6" type="journal article" year="2017">Jaschhof &amp; Jaschhof 2017</bibRefCitation>
)
</taxonomicName>
. Another specimen of
<taxonomicName id="4C69C5DCFF876E34FC9AF8019EB0F85A" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2020" box="[858,996,1980,2006]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="imbecilla">
<emphasis id="B91D624DFF876E34FC9AF8019EB0F85A" box="[858,996,1980,2006]" italics="true" pageId="12" pageNumber="13">W. imbecilla</emphasis>
</taxonomicName>
from
<collectingCountry id="F37EFECFFF876E34FBE6F800992BF85A" box="[1062,1151,1980,2006]" name="Sweden" pageId="12" pageNumber="13">Sweden</collectingCountry>
we detected in the NHRS collection, again misidentified as
<taxonomicName id="4C69C5DCFF876E34FDD0F85D9F94F876" baseAuthorityName="Jaschhof &amp; Jaschhof" baseAuthorityYear="2017" box="[528,704,2016,2042]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="brachypalpa">
<emphasis id="B91D624DFF876E34FDD0F85D9F94F876" box="[528,704,2016,2042]" italics="true" pageId="12" pageNumber="13">W. brachypalpa</emphasis>
</taxonomicName>
. This leaves the genuine
<taxonomicName id="4C69C5DCFF876E34FC1CF85D99D8F876" baseAuthorityName="Jaschhof &amp; Jaschhof" baseAuthorityYear="2017" box="[988,1164,2016,2042]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="brachypalpa">
<emphasis id="B91D624DFF876E34FC1CF85D99D8F876" box="[988,1164,2016,2042]" italics="true" pageId="12" pageNumber="13">W. brachypalpa</emphasis>
</taxonomicName>
, a species originally described from the Russian Far East (
<bibRefCitation id="EFF8C3AEFF866E35FDE4FF2B9F9CFF3D" author="Mamaev, B. M." box="[548,712,150,177]" pageId="13" pageNumber="14" pagination="81 - 88" refId="ref56620" refString="Mamaev, B. M. (1975) Some regularities of infestation of Betula costata and Betula manshurica by xylophagous insects. Trudy Biologo-pochvennogo Instituta, Novaya, Seriya, 28, 81 - 88. [in Russian]" type="journal article" year="1975">Mamaev 1975</bibRefCitation>
) and subsequently reported to occur in
<collectingCountry id="F37EFECFFF866E35FB47FF2B99B5FF3D" box="[1159,1249,151,177]" name="Ukraine" pageId="13" pageNumber="14">Ukraine</collectingCountry>
(
<bibRefCitation id="EFF8C3AEFF866E35FB31FF2B98C4FF3D" author="Spungis, V." box="[1265,1424,151,177]" pageId="13" pageNumber="14" pagination="1 - 38" refId="ref57075" refString="Spungis, V. (1992) A revision of the European gall midges of the tribe Winnertziini. Latvijas Entomologs Supplementum, 5, 1 - 38." type="journal article" year="1992">Spungis 1992</bibRefCitation>
), without valid record in
<collectingCountry id="F37EFECFFF866E35FE5EFF079CAEFF59" box="[414,506,187,213]" name="Sweden" pageId="13" pageNumber="14">Sweden</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C373EDD4FF866E35FF07FF629C37FA81" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="8BD6BE5FFF866E35FF07FF629C96FDED" blockId="13.[151,1437,150,1619]" pageId="13" pageNumber="14">
<emphasis id="B91D624DFF866E35FF07FF629C16FF74" bold="true" box="[199,322,222,248]" pageId="13" pageNumber="14">Diagnosis.</emphasis>
A medium-sized, brown
<taxonomicName id="4C69C5DCFF866E35FDABFF639FB0FF74" authorityName="Rondani" authorityYear="1860" box="[619,740,223,248]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B91D624DFF866E35FDABFF639FB0FF74" box="[619,740,223,248]" italics="true" pageId="13" pageNumber="14">Winnertzia</emphasis>
</taxonomicName>
with vestigial mouthparts. A combination of male genitalic characters is diagnostic of
<taxonomicName id="4C69C5DCFF866E35FE01FEBF9F18FE91" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2020" box="[449,588,259,285]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="imbecilla">
<emphasis id="B91D624DFF866E35FE01FEBF9F18FE91" box="[449,588,259,285]" italics="true" pageId="13" pageNumber="14">W. imbecilla</emphasis>
</taxonomicName>
, as follows (
<figureCitation id="1352A2DAFF866E35FD1CFEBF9E7BFE91" box="[732,815,259,285]" captionStart="FIGURES 713" captionStartId="12.[151,264,1621,1645]" captionTargetBox="[154,1433,188,1588]" captionTargetId="figure-25@12.[151,1436,181,1598]" captionTargetPageId="12" captionText="FIGURES 713. Male morphology of Winnertzia brachytarsus (79) and Winnertzia imbecilla (1013), holotypes. 7: Fourth flagellomere, lateral. 8: Ditto, medial. 9: Genitalia, ventral. 10: Genitalia, ventral. 11: Palpus, lateral. 12: Fourth flagellomere, lateral. 13: Ditto, medial. Scales for 710 and 1213, 0.05 mm; for 11, 0.025 mm. Arrows refer to characters described in the diagnoses." figureDoi="http://doi.org/10.5281/zenodo.4402763" httpUri="https://zenodo.org/record/4402763/files/figure.png" pageId="13" pageNumber="14">Fig. 10</figureCitation>
). The compact gonostylus has dense, large microtrichia apically and medially (↓
<superScript id="7C1C1317FF866E35FE69FE9A9CE5FEB8" attach="right" box="[425,433,294,308]" fontSize="6" pageId="13" pageNumber="14">4</superScript>
); the base of the gonocoxal emargination is angular rather than rounded (↓
<superScript id="7C1C1317FF866E35FB3FFE9A9853FEB8" attach="none" box="[1279,1287,294,308]" fontSize="6" pageId="13" pageNumber="14">5</superScript>
); the tegmen is markedly tapered towards the blunt-ended, microtrichose apex (↓
<superScript id="7C1C1317FF866E35FC57FEF69ECBFED4" attach="none" box="[919,927,330,344]" fontSize="6" pageId="13" pageNumber="14">6</superScript>
); and the parameral apodemes are unusually short and broad (↓
<superScript id="7C1C1317FF866E35FEAAFED39C26FEF1" attach="none" box="[362,370,367,381]" fontSize="6" pageId="13" pageNumber="14">7</superScript>
).
<taxonomicName id="4C69C5DCFF866E35FE4CFED39FC8FE05" baseAuthorityName="Jaschhof &amp; Jaschhof" baseAuthorityYear="2017" box="[396,668,367,393]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="brachypalpa">
<emphasis id="B91D624DFF866E35FE4CFED39FC8FE05" box="[396,668,367,393]" italics="true" pageId="13" pageNumber="14">Winnertzia brachypalpa</emphasis>
</taxonomicName>
, a generally similar species whose
<typeStatus id="54D200FDFF866E35FBF4FED399C2FE05" box="[1076,1174,367,393]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
we studied in 2012, is distinguished from
<taxonomicName id="4C69C5DCFF866E35FEB6FE2F9F57FE21" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2020" box="[374,515,403,429]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="imbecilla">
<emphasis id="B91D624DFF866E35FEB6FE2F9F57FE21" box="[374,515,403,429]" italics="true" pageId="13" pageNumber="14">W. imbecilla</emphasis>
</taxonomicName>
by the subtriangular tegmen with pointed apex and by the small, inconspicuous parameral apodemes. Also, the gonostylar apex of
<taxonomicName id="4C69C5DCFF866E35FD0DFE0B9E2AFE5D" baseAuthorityName="Jaschhof &amp; Jaschhof" baseAuthorityYear="2017" box="[717,894,439,465]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="brachypalpa">
<emphasis id="B91D624DFF866E35FD0DFE0B9E2AFE5D" box="[717,894,439,465]" italics="true" pageId="13" pageNumber="14">W. brachypalpa</emphasis>
</taxonomicName>
seems to be more slender compared with
<taxonomicName id="4C69C5DCFF866E35FA94FE0B9DB7FE79" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2020" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="imbecilla">
<emphasis id="B91D624DFF866E35FA94FE0B9DB7FE79" italics="true" pageId="13" pageNumber="14">W. imbecilla</emphasis>
</taxonomicName>
, a possible distinction that is difficult to assess in the
<typeStatus id="54D200FDFF866E35FC85FE679EF3FE79" box="[837,935,475,501]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
specimen.
<taxonomicName id="4C69C5DCFF866E35FBE7FE67986DFE79" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2020" box="[1063,1337,475,501]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="angustistylus">
<emphasis id="B91D624DFF866E35FBE7FE67986DFE79" box="[1063,1337,475,501]" italics="true" pageId="13" pageNumber="14">Winnertzia angustistylus</emphasis>
</taxonomicName>
, another species of the
<taxonomicName id="4C69C5DCFF866E35FEF9FE439CC6FD95" authorityName="Mohrig &amp; Mamaev" authorityYear="1970" box="[313,402,511,537]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="discreta">
<emphasis id="B91D624DFF866E35FEF9FE439CC6FD95" box="[313,402,511,537]" italics="true" pageId="13" pageNumber="14">discreta</emphasis>
</taxonomicName>
group with vestigial mouthparts, differs from both
<taxonomicName id="4C69C5DCFF866E35FC19FE439931FD95" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2020" box="[985,1125,511,537]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="imbecilla">
<emphasis id="B91D624DFF866E35FC19FE439931FD95" box="[985,1125,511,537]" italics="true" pageId="13" pageNumber="14">W. imbecilla</emphasis>
</taxonomicName>
and
<taxonomicName id="4C69C5DCFF866E35FB5DFE43981BFD95" baseAuthorityName="Jaschhof &amp; Jaschhof" baseAuthorityYear="2017" box="[1181,1359,511,537]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="brachypalpa">
<emphasis id="B91D624DFF866E35FB5DFE43981BFD95" box="[1181,1359,511,537]" italics="true" pageId="13" pageNumber="14">W. brachypalpa</emphasis>
</taxonomicName>
by the conspicuously slender gonostylus (
<figureCitation id="1352A2DAFF866E35FDE1FD9F9F32FDB1" box="[545,614,547,573]" captionStart="FIGURES 16" captionStartId="10.[151,264,1742,1766]" captionTargetBox="[159,1430,385,1670]" captionTargetId="figure-111@10.[151,1436,378,1676]" captionTargetPageId="10" captionText="FIGURES 16. Male morphology of Winnertzia angustistylus. 1: Genitalia, ventral, holotype. 2: Gonostylus, ventral, paratype. 3: Fourth flagellomere, lateral, holotype. 4: Ditto, medial. 5: Palpus, lateral, paratype. 6: Wing, dorsal, specimen from Södermanland. Scales for 1, 3 and 4, 0.05 mm; for 2 and 5, 0.025 mm; for 6, 0.50 mm. Arrows refer to characters described in the diagnosis." figureDoi="http://doi.org/10.5281/zenodo.4402759" httpUri="https://zenodo.org/record/4402759/files/figure.png" pageId="13" pageNumber="14">Fig. 2</figureCitation>
), to name just one of the distinctions. Females and preimaginal stages of
<taxonomicName id="4C69C5DCFF866E35FF57FDFB9C76FDED" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2020" box="[151,290,583,609]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="imbecilla">
<emphasis id="B91D624DFF866E35FF57FDFB9C76FDED" box="[151,290,583,609]" italics="true" pageId="13" pageNumber="14">W. imbecilla</emphasis>
</taxonomicName>
are unknown.
</paragraph>
<paragraph id="8BD6BE5FFF866E35FF07FDD69C37FA81" blockId="13.[151,1437,150,1619]" pageId="13" pageNumber="14">
<emphasis id="B91D624DFF866E35FF07FDD69C88FD09" bold="true" box="[199,476,618,645]" pageId="13" pageNumber="14">Other male characters.</emphasis>
Body size
<quantity id="4C9113BAFF866E35FD9BFDD79FE1FD09" box="[603,693,619,645]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="13" pageNumber="14" unit="mm" value="1.7">1.7 mm</quantity>
.
<emphasis id="B91D624DFF866E35FD00FDD79E53FD09" bold="true" box="[704,775,619,645]" pageId="13" pageNumber="14">Head.</emphasis>
Eye bridge 34 ommatidia long dorsally. Antenna slightly longer than half body. Scape and pedicel same size, both concolorous with flagellum. 11 flagellomeres, apical flagellomere long, composed of two nodes; translucent sensilla present on flagellomeres 110. Fourth flagellomere: neck 0.60.7 times as long as node; node 1.5 times as long as broad; sensory hairs sparse; both lateral and medial translucent sensilla long, filiform, linear to variously bent (
<figureCitation id="1352A2DAFF866E35FCF8FD479EE9FC99" box="[824,957,763,789]" captionStart="FIGURES 713" captionStartId="12.[151,264,1621,1645]" captionTargetBox="[154,1433,188,1588]" captionTargetId="figure-25@12.[151,1436,181,1598]" captionTargetPageId="12" captionText="FIGURES 713. Male morphology of Winnertzia brachytarsus (79) and Winnertzia imbecilla (1013), holotypes. 7: Fourth flagellomere, lateral. 8: Ditto, medial. 9: Genitalia, ventral. 10: Genitalia, ventral. 11: Palpus, lateral. 12: Fourth flagellomere, lateral. 13: Ditto, medial. Scales for 710 and 1213, 0.05 mm; for 11, 0.025 mm. Arrows refer to characters described in the diagnoses." figureDoi="http://doi.org/10.5281/zenodo.4402763" httpUri="https://zenodo.org/record/4402763/files/figure.png" pageId="13" pageNumber="14">Figs 1213</figureCitation>
). Palpus much shorter than head height, 3 segments with mostly long setae; apical segment napiform, longer than preceding two segments together (
<figureCitation id="1352A2DAFF866E35FAFCFCA398DAFCB5" box="[1340,1422,799,825]" captionStart="FIGURES 713" captionStartId="12.[151,264,1621,1645]" captionTargetBox="[154,1433,188,1588]" captionTargetId="figure-25@12.[151,1436,181,1598]" captionTargetPageId="12" captionText="FIGURES 713. Male morphology of Winnertzia brachytarsus (79) and Winnertzia imbecilla (1013), holotypes. 7: Fourth flagellomere, lateral. 8: Ditto, medial. 9: Genitalia, ventral. 10: Genitalia, ventral. 11: Palpus, lateral. 12: Fourth flagellomere, lateral. 13: Ditto, medial. Scales for 710 and 1213, 0.05 mm; for 11, 0.025 mm. Arrows refer to characters described in the diagnoses." figureDoi="http://doi.org/10.5281/zenodo.4402763" httpUri="https://zenodo.org/record/4402763/files/figure.png" pageId="13" pageNumber="14">Fig. 11</figureCitation>
). Labella vestigial.
<emphasis id="B91D624DFF866E35FEA7FCFF9C93FCD1" bold="true" box="[359,455,835,861]" pageId="13" pageNumber="14">Thorax.</emphasis>
Pronotal setae 6. Anepimeral setae absent. Lateral mediotergal microtrichia slightly enlarged. Parascutellar area inconspicuous, vaguely contoured.
<emphasis id="B91D624DFF866E35FCA4FCDA9EF1FC0C" bold="true" box="[868,933,870,896]" pageId="13" pageNumber="14">Wing</emphasis>
slightly shorter than body, 2.6 times as long as broad. Costal cell slightly reinforced. M
<subScript id="17EDBC1AFF866E35FDBEFC219FD2FC27" attach="left" box="[638,646,925,939]" fontSize="6" pageId="13" pageNumber="14">4</subScript>
long, straight, CuA moderately bent, both veins extending to edge of wing.
<emphasis id="B91D624DFF866E35FF1EFC139C49FC45" bold="true" box="[222,285,943,969]" pageId="13" pageNumber="14">Legs.</emphasis>
Scales pointed. Basitarsal spines absent. Fore tibia 1.4 times length T
<subScript id="17EDBC1AFF866E35FBF1FC7D996DFC43" attach="left" box="[1073,1081,961,975]" fontSize="6" pageId="13" pageNumber="14">2</subScript>
. Acropods: claws slightly bent, basal tooth large; empodia vestigial.
<emphasis id="B91D624DFF866E35FD86FC6E9F96FC60" bold="true" box="[582,706,978,1004]" pageId="13" pageNumber="14">Abdomen.</emphasis>
Pleural membrane devoid of setae.
<emphasis id="B91D624DFF866E35FBABFC6E998DFC60" bold="true" box="[1131,1241,978,1004]" pageId="13" pageNumber="14">Genitalia</emphasis>
(
<figureCitation id="1352A2DAFF866E35FB2DFC6F9810FC61" box="[1261,1348,979,1005]" captionStart="FIGURES 713" captionStartId="12.[151,264,1621,1645]" captionTargetBox="[154,1433,188,1588]" captionTargetId="figure-25@12.[151,1436,181,1598]" captionTargetPageId="12" captionText="FIGURES 713. Male morphology of Winnertzia brachytarsus (79) and Winnertzia imbecilla (1013), holotypes. 7: Fourth flagellomere, lateral. 8: Ditto, medial. 9: Genitalia, ventral. 10: Genitalia, ventral. 11: Palpus, lateral. 12: Fourth flagellomere, lateral. 13: Ditto, medial. Scales for 710 and 1213, 0.05 mm; for 11, 0.025 mm. Arrows refer to characters described in the diagnoses." figureDoi="http://doi.org/10.5281/zenodo.4402763" httpUri="https://zenodo.org/record/4402763/files/figure.png" pageId="13" pageNumber="14">Fig. 10</figureCitation>
). Ninth tergite about half gonopodal length; setae confined to posterior portion; posterior edge emarginated medially, the emargination darkly pigmented; anterior edge straight, indistinct. Gonocoxal synsclerite slightly broader than long; a short portion ventrobasally non-setose; ventral emargination with broad, darkly pigmented border, membranous basally; ventroanterior edge straight; ventroposterior portions enlarged, markedly protruding beyond dorsoposterior portions; dorsal apodemes moderately long, subtriangular. Gonostylus twice as long as broad; basolateral apophysis normal size, angulated. Aedeagal apodeme gently broadened shortly before the apex, then slightly constricted; solid basal portion long. Aedeagal bulge apparently devoid of microtrichia. Tegmen slightly reinforced apically; flaps narrow, indistinct.
</paragraph>
</subSubSection>
<subSubSection id="C373EDD4FF866E35FF07FAAB9C0DFAD9" pageId="13" pageNumber="14" type="etymology">
<paragraph id="8BD6BE5FFF866E35FF07FAAB9C0DFAD9" blockId="13.[151,1437,150,1619]" pageId="13" pageNumber="14">
<emphasis id="B91D624DFF866E35FF07FAAB9C1AFABD" bold="true" box="[199,334,1303,1329]" pageId="13" pageNumber="14">Etymology.</emphasis>
The Latin adjective
<taxonomicName id="4C69C5DCFF866E35FDF6FAAB9FCAFABD" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2020" box="[566,670,1303,1329]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="imbecilla">
<emphasis id="B91D624DFF866E35FDF6FAAB9FCAFABD" box="[566,670,1303,1329]" italics="true" pageId="13" pageNumber="14">imbecilla</emphasis>
</taxonomicName>
means weak, an allusion to the fact that the morphology of males is partly regressive.
</paragraph>
</subSubSection>
<subSubSection id="C373EDD4FF866E35FF07FAE39F15F986" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="8BD6BE5FFF866E35FF07FAE39945FA11" blockId="13.[151,1437,150,1619]" pageId="13" pageNumber="14">
<emphasis id="B91D624DFF866E35FF07FAE39C22FAF4" bold="true" box="[199,374,1374,1401]" pageId="13" pageNumber="14">Type material.</emphasis>
<materialsCitation id="3B01B402FF866E35FEBDFAE39945FA11" ID-GBIF-Occurrence="3023445511" collectingDate="2015-07-22" collectingDateMax="2015-08-23" collectingDateMin="2015-07-22" collectionCode="CEC, NHRS" collectorName="M. &amp; C. Jaschhof" country="Sweden" county="Skepparsang Nature Reserve" location="Male" municipality="Borgholm" pageId="13" pageNumber="14" specimenCount="1" typeStatus="holotype">
<emphasis id="B91D624DFF866E35FEBDFAE39CA5FAF5" bold="true" box="[381,497,1375,1401]" pageId="13" pageNumber="14">
<typeStatus id="54D200FDFF866E35FEBDFAE39CB8FAF5" box="[381,492,1375,1401]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
.
</emphasis>
<location id="8EB6E884FF866E35FE38FAE39F60FAF5" LSID="urn:lsid:plazi:treatment:03C00F49FF876E35FF57F8909827F9DD:8EB6E884FF866E35FE38FAE39F60FAF5" box="[504,564,1375,1401]" country="Sweden" county="Skepparsang Nature Reserve" municipality="Borgholm" name="Male" pageId="13" pageNumber="14">Male</location>
,
<collectingCountry id="F37EFECFFF866E35FD80FAE39FCFFAF5" box="[576,667,1375,1401]" name="Sweden" pageId="13" pageNumber="14">Sweden</collectingCountry>
,
<location id="8EB6E884FF866E35FD68FADC9FBBFAF7" LSID="urn:lsid:plazi:treatment:03C00F49FF876E35FF57F8909827F9DD:8EB6E884FF866E35FD68FADC9FBBFAF7" box="[680,751,1376,1403]" country="Sweden" county="Skepparsang Nature Reserve" municipality="Borgholm" name="Oland" pageId="13" pageNumber="14">Öland</location>
,
<collectingMunicipality id="6BB22425FF866E35FD3BFAE39E3BFAF5" box="[763,879,1375,1401]" pageId="13" pageNumber="14">Borgholm</collectingMunicipality>
,
<collectingCounty id="62B7C6D3FF866E35FCBAFAE39996FAF5" box="[890,1218,1375,1401]" pageId="13" pageNumber="14">Skepparsäng Nature Reserve</collectingCounty>
, dry pine forest,
<date id="FFD7989FFF866E35FA40FAE39CDCFA11" pageId="13" pageNumber="14" value="2015-07-22" valueMax="2015-08-23" valueMin="2015-07-22">
<collectingDate id="EF936177FF866E35FA40FAE39CDCFA11" pageId="13" pageNumber="14" value="2015-07-22" valueMax="2015-08-23" valueMin="2015-07-22">22 July23 August 2015</collectingDate>
</date>
,
<collectingMethod id="5228C648FF866E35FE53FA3F9F75FA11" box="[403,545,1411,1437]" pageId="13" pageNumber="14">Malaise trap</collectingMethod>
,
<collectorName id="269CDB89FF866E35FDECFA3F9FA7FA11" box="[556,755,1411,1437]" pageId="13" pageNumber="14">M. &amp; C. Jaschhof</collectorName>
(spn
<collectionCode id="ED78269AFF866E35FCF1FA3F9E3CFA11" box="[817,872,1411,1437]" pageId="13" pageNumber="14">CEC</collectionCode>
<quantity id="4C9113BAFF866E35FCA8FA3F9EFAFA11" box="[872,942,1411,1437]" metricMagnitude="0" metricUnit="m" metricValue="6.858" pageId="13" pageNumber="14" unit="in" value="270.0">270 in</quantity>
<collectionCode id="ED78269AFF866E35FC75FA3F9951FA11" box="[949,1029,1411,1437]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="13" pageNumber="14">NHRS</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BD6BE5FFF866E35FF07FA1A9F15F986" blockId="13.[151,1437,150,1619]" pageId="13" pageNumber="14">
<emphasis id="B91D624DFF866E35FF07FA1A9CB0FA4C" bold="true" box="[199,484,1446,1472]" pageId="13" pageNumber="14">Other material studied.</emphasis>
<materialsCitation id="3B01B402FF866E35FE2EFA1B9F69F986" ID-GBIF-Occurrence="3023445367" collectingDate="2002-05" collectingDateMax="2002-06-17" collectingDateMin="2002-05" collectionCode="MT, NHRS, V" collectorName="Tyresta NR &amp; B. Viklund" country="Sweden" location="Haningealpen Station" pageId="13" pageNumber="14" specimenCode="GULI000021001" specimenCount="1" specimenCount-male="1" stateProvince="Sodermanland">
<collectingCountry id="F37EFECFFF866E35FE2EFA1B9F1FFA4D" box="[494,587,1447,1473]" name="Sweden" pageId="13" pageNumber="14">Sweden</collectingCountry>
:
<specimenCount id="9D6F75D6FF866E35FD97FA1B9FFCFA4D" box="[599,680,1447,1473]" pageId="13" pageNumber="14" type="male">1 male</specimenCount>
,
<collectingRegion id="49AD70BDFF866E35FD75FA1B9E0EFA4D" box="[693,858,1447,1473]" country="Sweden" name="Sodermanland" pageId="13" pageNumber="14">Södermanland</collectingRegion>
,
<collectorName id="269CDB89FF866E35FCA7FA1B9EB8FA4D" box="[871,1004,1447,1473]" pageId="13" pageNumber="14">Tyresta NR</collectorName>
,
<date id="FFD7989FFF866E35FC3AFA1B993AFA4D" box="[1018,1134,1447,1473]" pageId="13" pageNumber="14" value="2002-05">
<collectingDate id="EF936177FF866E35FC3AFA1B993AFA4D" box="[1018,1134,1447,1473]" pageId="13" pageNumber="14" value="2002-05">May 2002</collectingDate>
</date>
firesite north of
<location id="8EB6E884FF866E35FAF4FA1B9C7AFA69" LSID="urn:lsid:plazi:treatment:03C00F49FF876E35FF57F8909827F9DD:8EB6E884FF866E35FAF4FA1B9C7AFA69" country="Sweden" name="Haningealpen Station" pageId="13" pageNumber="14" stateProvince="Sodermanland">Haningealpen Station</location>
,
<date id="FFD7989FFF866E35FEFBFA779F68FA69" box="[315,572,1483,1509]" pageId="13" pageNumber="14" value="2002-05-31" valueMax="2002-06-17" valueMin="2002-05-31">
<collectingDate id="EF936177FF866E35FEFBFA779F68FA69" box="[315,572,1483,1509]" pageId="13" pageNumber="14" value="2002-05-31" valueMax="2002-06-17" valueMin="2002-05-31">31 May17 June 2002</collectingDate>
</date>
,
<collectionCode id="ED78269AFF866E35FD88FA779F27FA69" box="[584,627,1483,1509]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="13" pageNumber="14">MT</collectionCode>
,
<collectorName id="269CDB89FF866E35FDBFFA779FAFFA69" box="[639,763,1483,1509]" pageId="13" pageNumber="14">B. Viklund</collectorName>
(spn
<specimenCode id="DBCF1624FF866E35FCFCFA779EA9FA69" box="[828,1021,1483,1509]" collectionCode="GULI" pageId="13" pageNumber="14">GULI000021001</specimenCode>
in
<collectionCode id="ED78269AFF866E35FBE3FA77993BFA69" box="[1059,1135,1483,1509]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="13" pageNumber="14">NHRS</collectionCode>
) (previously labeled as
<taxonomicName id="4C69C5DCFF866E35FA40FA779CDAF985" authority="Mamaev" authorityName="Mamaev" authorityYear="1975" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="brachypalpa">
<emphasis id="B91D624DFF866E35FA40FA779C71F985" italics="true" pageId="13" pageNumber="14">W. brachypalpa</emphasis>
Mamaev
</taxonomicName>
by
<collectionCode id="ED78269AFF866E35FE77FA539C9FF985" box="[439,459,1519,1545]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="13" pageNumber="14" type="Museum">V</collectionCode>
. Spungis)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C373EDD4FF866E35FF07F9AE9827F9DD" pageId="13" pageNumber="14" type="distribution">
<paragraph id="8BD6BE5FFF866E35FF07F9AE9827F9DD" blockId="13.[151,1437,150,1619]" pageId="13" pageNumber="14">
<emphasis id="B91D624DFF866E35FF07F9AE9F42F9A1" bold="true" box="[199,534,1554,1581]" pageId="13" pageNumber="14">Distribution and phenology.</emphasis>
Both specimens known of
<taxonomicName id="4C69C5DCFF866E35FC8BF9AF9E82F9A1" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2020" box="[843,982,1555,1581]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="imbecilla">
<emphasis id="B91D624DFF866E35FC8BF9AF9E82F9A1" box="[843,982,1555,1581]" italics="true" pageId="13" pageNumber="14">W. imbecilla</emphasis>
</taxonomicName>
were collected in dry pine forests in the southern half of
<collectingCountry id="F37EFECFFF866E35FE8FF98B9CFCF9DD" box="[335,424,1591,1617]" name="Sweden" pageId="13" pageNumber="14">Sweden</collectingCountry>
(Öland,
<collectingRegion id="49AD70BDFF866E35FDCBF98B9FE6F9DE" box="[523,690,1591,1618]" country="Sweden" name="Sodermanland" pageId="13" pageNumber="14">Södermanland</collectingRegion>
). Available data suggest the adult flight period is JuneAugust.
</paragraph>
</subSubSection>
</treatment>
</document>