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<document id="4C4E601E3B6C15360373DEB0642D4E4C" ID-ISSN="1175­5334" ID-ZooBank="505937B0-9F57-4068-82E6-8553826DD5AA" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629822157238" checkinUser="felipe" docAuthor="Judson, Sarah W. &amp; Nelson, C. Riley" docDate="2012" docId="03B387E7154F811FFF5AFB11FE9F57FB" docLanguage="en" docName="zt03541p118.pdf" docOrigin="Zootaxa 3541" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Diura majuscula" docType="treatment" docVersion="2" lastPageNumber="42" masterDocId="FF8AFF9F15668136FFCDFFA7FFAF5015" masterDocTitle="3541" masterLastPageNumber="118" masterPageNumber="1" pageNumber="42" updateTime="1699117090469" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:namePart id="395A5D8E6B529C1D7A1E6D7715BAE2A2">Judson, Sarah W.</mods:namePart>
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<mods:namePart id="5F51199773E721F7D3BB151D7BC14E8A">Nelson, C. Riley</mods:namePart>
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<treatment id="03B387E7154F811FFF5AFB11FE9F57FB" LSID="urn:lsid:plazi:treatment:03B387E7154F811FFF5AFB11FE9F57FB" httpUri="http://treatment.plazi.org/id/03B387E7154F811FFF5AFB11FE9F57FB" lastPageNumber="42" pageId="41" pageNumber="42">
<subSubSection id="C300657A154F811FFF5AFB11FE935503" pageId="41" pageNumber="42" type="nomenclature">
<paragraph id="8BA536F1154F811FFF5AFB11FD9654C5" blockId="41.[151,569,1206,1232]" box="[151,569,1206,1232]" pageId="41" pageNumber="42">
<heading id="D0ED819D154F811FFF5AFB11FD9654C5" bold="true" box="[151,569,1206,1232]" fontSize="11" level="1" pageId="41" pageNumber="42" reason="1">
<taxonomicName id="4C1A4D72154F811FFF5AFB11FD9654C5" ID-CoL="36ZL3" authority="(Klapalek, 1912)" baseAuthorityName="Klapalek" baseAuthorityYear="1912" box="[151,569,1206,1232]" class="Insecta" family="Perlodidae" genus="Diura" kingdom="Animalia" order="Plecoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="majuscula">
<emphasis id="B96EEAE3154F811FFF5AFB11FD9654C5" bold="true" box="[151,569,1206,1232]" pageId="41" pageNumber="42">
<emphasis id="B96EEAE3154F811FFF5AFB11FECC54C5" bold="true" box="[151,355,1206,1232]" italics="true" pageId="41" pageNumber="42">Diura majuscula</emphasis>
(Klapalek, 1912)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BA536F1154F811FFF5AFB5AFE935503" blockId="41.[151,591,1277,1427]" box="[151,316,1277,1302]" pageId="41" pageNumber="42">
<heading id="D0ED819D154F811FFF5AFB5AFE935503" bold="true" box="[151,316,1277,1302]" fontSize="10" level="2" pageId="41" pageNumber="42" reason="2">
<emphasis id="B96EEAE3154F811FFF5AFB5AFE935503" bold="true" box="[151,316,1277,1302]" pageId="41" pageNumber="42">SYNONYMY</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C300657A154F811FFF5AFA86FE475586" pageId="41" pageNumber="42" type="description">
<paragraph id="8BA536F1154F811FFF5AFA86FDE05522" blockId="41.[151,591,1277,1427]" box="[151,591,1313,1335]" pageId="41" pageNumber="42">
<taxonomicName id="4C1A4D72154F811FFF5AFA86FDE05522" authority="Klapalek, 1912" authorityName="Klapalek" authorityYear="1912" box="[151,591,1313,1335]" class="Insecta" family="Perlodidae" genus="Dictyopterygella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="majuscule">
<emphasis id="B96EEAE3154F811FFF5AFA86FE025522" box="[151,429,1313,1335]" italics="true" pageId="41" pageNumber="42">Dictyopterygella majuscule</emphasis>
Klapálek, 1912
</taxonomicName>
</paragraph>
<paragraph id="8BA536F1154F811FFF5AFAE6FDAA5543" blockId="41.[151,591,1277,1427]" box="[151,517,1344,1366]" pageId="41" pageNumber="42">
<emphasis id="B96EEAE3154F811FFF5AFAE6FED55543" box="[151,378,1344,1366]" italics="true" pageId="41" pageNumber="42">
<taxonomicName id="4C1A4D72154F811FFF5AFAE6FE865543" baseAuthorityName="Kempny" baseAuthorityYear="1900" box="[151,297,1345,1366]" class="Insecta" family="Perlodidae" genus="Diura" kingdom="Animalia" order="Plecoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="nanseni">Diura nanseni</taxonomicName>
(males)
</emphasis>
Brinck, 1949
</paragraph>
<paragraph id="8BA536F1154F811FFF5AFAF8FD815561" blockId="41.[151,591,1277,1427]" box="[151,558,1374,1397]" pageId="41" pageNumber="42">
<emphasis id="B96EEAE3154F811FFF5AFAF8FE0C5560" box="[151,419,1375,1397]" italics="true" pageId="41" pageNumber="42">
<taxonomicName id="4C1A4D72154F811FFF5AFAF8FE905560" authorityName="Ricker" authorityYear="1952" box="[151,319,1375,1397]" class="Insecta" family="Perlodidae" genus="Diura" kingdom="Animalia" order="Plecoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="bicaudata">Diura bicaudata</taxonomicName>
(females)
</emphasis>
Brinck, 1949
</paragraph>
<paragraph id="8BA536F1154F811FFF5AFAD9FE475586" blockId="41.[151,591,1277,1427]" box="[151,488,1405,1427]" pageId="41" pageNumber="42">
<taxonomicName id="4C1A4D72154F811FFF5AFAD9FE475586" authority="Zhiltzova, 1975" authorityName="Zhiltzova" authorityYear="1975" box="[151,488,1405,1427]" class="Insecta" family="Perlodidae" genus="Diura" kingdom="Animalia" order="Plecoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="majuscula">
<emphasis id="B96EEAE3154F811FFF5AFAD9FEED5586" box="[151,322,1405,1427]" italics="true" pageId="41" pageNumber="42">Diura majuscula</emphasis>
Zhiltzova, 1975
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C300657A154F811FFF5AFA1CFB9F55C7" box="[151,1072,1466,1491]" pageId="41" pageNumber="42" type="materials_examined">
<paragraph id="8BA536F1154F811FFF5AFA1CFB9F55C7" blockId="41.[151,1436,1466,2030]" box="[151,1072,1466,1491]" pageId="41" pageNumber="42">
<materialsCitation id="3B723CAC154F811FFF5AFA1CFB8355C7" box="[151,1068,1466,1491]" country="Russia" county="Lena River" location="Ust-Kutsk" municipality="Bazaikha" pageId="41" pageNumber="42" specimenCount="1" stateProvince="Krasnoyarsk" typeStatus="holotype">
<emphasis id="B96EEAE3154F811FFF5AFA1CFE2B55C6" bold="true" box="[151,388,1466,1491]" pageId="41" pageNumber="42">
<typeStatus id="54A18853154F811FFF5AFA1CFF4D55C6" box="[151,226,1467,1491]" pageId="41" pageNumber="42">TYPE</typeStatus>
LOCALITY:
</emphasis>
<collectingCountry id="F30D7661154F811FFE41FA1DFE7455C7" box="[396,475,1466,1490]" name="Russia" pageId="41" pageNumber="42">Russia</collectingCountry>
:
<collectingCounty id="62C44E7D154F811FFE28FA1CFDCB55C7" box="[485,612,1466,1490]" pageId="41" pageNumber="42">Lena River</collectingCounty>
,
<collectingMunicipality id="6BC1AC8B154F811FFDA2FA1DFD7855C7" box="[623,727,1466,1490]" pageId="41" pageNumber="42">Bazaikha</collectingMunicipality>
(
<collectingRegion id="49DEF813154F811FFD28FA1DFCD955C7" box="[741,886,1466,1490]" country="Russia" name="Krasnoyarsk" pageId="41" pageNumber="42">Krasnoyarsk</collectingRegion>
) and
<location id="8EC5602A154F811FFC78FA1DFB8355C7" LSID="urn:lsid:plazi:treatment:03B387E7154F811FFF5AFB11FE9F57FB:8EC5602A154F811FFC78FA1DFB8355C7" box="[949,1068,1466,1490]" country="Russia" county="Lena River" municipality="Bazaikha" name="Ust-Kutsk" pageId="41" pageNumber="42" stateProvince="Krasnoyarsk">Ust-Kutsk</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C300657A154F811FFF0AFA79FE8F56E7" pageId="41" pageNumber="42" type="diagnosis">
<paragraph id="8BA536F1154F811FFF0AFA79FE8F56E7" blockId="41.[151,1436,1466,2030]" pageId="41" pageNumber="42">
<emphasis id="B96EEAE3154F811FFF0AFA79FEC255E2" bold="true" box="[199,365,1502,1527]" pageId="41" pageNumber="42">DIAGNOSIS:</emphasis>
Males are brachypterous, with the wing tips reaching the middle of the abdomen (
<figureCitation id="13212A74154F811FFACEFA79FACA55E3" box="[1283,1381,1502,1526]" captionStart="FIGURES 268271" captionStartId="85.[151,263,895,918]" captionTargetBox="[153,1420,159,885]" captionTargetId="figure-73@85.[153,1438,159,928]" captionTargetPageId="85" captionText="FIGURES 268271. Diura majuscula. 268: Habitus (preserved); 269: Male paraprocts; 270: Female terminalia, ventral; 271: Predicted ENM map with documented occurrences and range estimate." pageId="41" pageNumber="42">Fig. 268</figureCitation>
), but rarely to the apex, Comparatively, the wing tips of
<taxonomicName id="4C1A4D72154F811FFCCAF9A4FC3E560E" box="[775,913,1539,1563]" class="Insecta" family="Perlodidae" genus="Diura" kingdom="Animalia" order="Plecoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="bicaudat">
<emphasis id="B96EEAE3154F811FFCCAF9A4FC3E560E" box="[775,913,1539,1563]" italics="true" pageId="41" pageNumber="42">D. bicaudat</emphasis>
</taxonomicName>
a only extend slightly beyond the thorax to approximately tergum 2. Adult head of both sexes has a round pale interocellar mark that is closed posteriorly (
<figureCitation id="13212A74154F811FFAA3F981FF6B5677" captionStart="FIGURES 268271" captionStartId="85.[151,263,895,918]" captionTargetBox="[153,1420,159,885]" captionTargetId="figure-73@85.[153,1438,159,928]" captionTargetPageId="85" captionText="FIGURES 268271. Diura majuscula. 268: Habitus (preserved); 269: Male paraprocts; 270: Female terminalia, ventral; 271: Predicted ENM map with documented occurrences and range estimate." pageId="41" pageNumber="42">Fig. 268</figureCitation>
), in contrast to
<taxonomicName id="4C1A4D72154F811FFEBBF9ECFDA55676" authorityName="Ricker" authorityYear="1952" box="[374,522,1611,1635]" class="Insecta" family="Perlodidae" genus="Diura" kingdom="Animalia" order="Plecoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="bicaudata">
<emphasis id="B96EEAE3154F811FFEBBF9ECFDA55676" box="[374,522,1611,1635]" italics="true" pageId="41" pageNumber="42">D. bicaudata</emphasis>
</taxonomicName>
which has a pointed mark that is open posteriorly (
<figureCitation id="13212A74154F811FFB98F9EDFB165677" box="[1109,1209,1610,1634]" captionStart="FIGURES 263267" captionStartId="84.[151,264,1966,1989]" captionTargetBox="[156,1436,1184,1951]" captionTargetId="figure-68@84.[156,1441,1184,1953]" captionTargetPageId="84" captionText="FIGURES 263267. Diura bicaudata. 263: Habitus (preserved); 264: Male paraprocts; 265: Male, lateral; 266: Female terminalia, ventral; 267: Predicted ENM map with documented occurrences and range estimate." pageId="41" pageNumber="42">Fig. 263</figureCitation>
). The pronotum has pale median stripe which is thicker than that of
<taxonomicName id="4C1A4D72154F811FFD74F9C8FCE25693" authorityName="Ricker" authorityYear="1952" box="[697,845,1646,1670]" class="Insecta" family="Perlodidae" genus="Diura" kingdom="Animalia" order="Plecoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="bicaudata">
<emphasis id="B96EEAE3154F811FFD74F9C8FCE25693" box="[697,845,1646,1670]" italics="true" pageId="41" pageNumber="42">D. bicaudata</emphasis>
</taxonomicName>
. The male subanal lobes are pilous (
<figureCitation id="13212A74154F811FFB3CF9C9FAFB5693" box="[1265,1364,1646,1670]" captionStart="FIGURES 268271" captionStartId="85.[151,263,895,918]" captionTargetBox="[153,1420,159,885]" captionTargetId="figure-73@85.[153,1438,159,928]" captionTargetPageId="85" captionText="FIGURES 268271. Diura majuscula. 268: Habitus (preserved); 269: Male paraprocts; 270: Female terminalia, ventral; 271: Predicted ENM map with documented occurrences and range estimate." pageId="41" pageNumber="42">Fig. 269</figureCitation>
), with long, dense setation, whereas
<taxonomicName id="4C1A4D72154F811FFE21F934FDD056BE" authorityName="Ricker" authorityYear="1952" box="[492,639,1683,1707]" class="Insecta" family="Perlodidae" genus="Diura" kingdom="Animalia" order="Plecoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="bicaudata">
<emphasis id="B96EEAE3154F811FFE21F934FDD056BE" box="[492,639,1683,1707]" italics="true" pageId="41" pageNumber="42">D. bicaudata</emphasis>
</taxonomicName>
has entirely glabrous subanal lobes (
<figureCitation id="13212A74154F811FFBEAF935FB2456BF" box="[1063,1163,1682,1706]" captionStart="FIGURES 263267" captionStartId="84.[151,264,1966,1989]" captionTargetBox="[156,1436,1184,1951]" captionTargetId="figure-68@84.[156,1441,1184,1953]" captionTargetPageId="84" captionText="FIGURES 263267. Diura bicaudata. 263: Habitus (preserved); 264: Male paraprocts; 265: Male, lateral; 266: Female terminalia, ventral; 267: Predicted ENM map with documented occurrences and range estimate." pageId="41" pageNumber="42">Fig. 264</figureCitation>
). The female subgenital plate is broadly rounded at the apex (
<figureCitation id="13212A74154F811FFDFCF911FD3C56DB" box="[561,659,1718,1742]" captionStart="FIGURES 268271" captionStartId="85.[151,263,895,918]" captionTargetBox="[153,1420,159,885]" captionTargetId="figure-73@85.[153,1438,159,928]" captionTargetPageId="85" captionText="FIGURES 268271. Diura majuscula. 268: Habitus (preserved); 269: Male paraprocts; 270: Female terminalia, ventral; 271: Predicted ENM map with documented occurrences and range estimate." pageId="41" pageNumber="42">Fig. 270</figureCitation>
), as opposed to
<taxonomicName id="4C1A4D72154F811FFC8EF910FC7A56DB" authorityName="Ricker" authorityYear="1952" box="[835,981,1718,1742]" class="Insecta" family="Perlodidae" genus="Diura" kingdom="Animalia" order="Plecoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="bicaudata">
<emphasis id="B96EEAE3154F811FFC8EF910FC7A56DB" box="[835,981,1718,1742]" italics="true" pageId="41" pageNumber="42">D. bicaudata</emphasis>
</taxonomicName>
which is truncate (
<figureCitation id="13212A74154F811FFB65F911FAA556DB" box="[1192,1290,1718,1742]" captionStart="FIGURES 263267" captionStartId="84.[151,264,1966,1989]" captionTargetBox="[156,1436,1184,1951]" captionTargetId="figure-68@84.[156,1441,1184,1953]" captionTargetPageId="84" captionText="FIGURES 263267. Diura bicaudata. 263: Habitus (preserved); 264: Male paraprocts; 265: Male, lateral; 266: Female terminalia, ventral; 267: Predicted ENM map with documented occurrences and range estimate." pageId="41" pageNumber="42">Fig. 266</figureCitation>
). The nymph is unknown.
</paragraph>
</subSubSection>
<subSubSection id="C300657A154F811FFF0AF959FEFB572F" pageId="41" pageNumber="42" type="distribution">
<paragraph id="8BA536F1154F811FFF0AF959FEFB572F" blockId="41.[151,1436,1466,2030]" pageId="41" pageNumber="42">
<emphasis id="B96EEAE3154F811FFF0AF959FDA35702" bold="true" box="[199,524,1790,1815]" pageId="41" pageNumber="42">DISTRIBUTION—Global:</emphasis>
East Palearctic—
<emphasis id="B96EEAE3154F811FFD2DF959FCFC5702" bold="true" box="[736,851,1790,1815]" pageId="41" pageNumber="42">Regional:</emphasis>
AOB, IDB—
<emphasis id="B96EEAE3154F811FFC36F959FBFC5702" bold="true" box="[1019,1107,1790,1815]" pageId="41" pageNumber="42">Aimag:</emphasis>
AR, BO, KhD, KhG, SE*, TO^, UB*, ZA^.
</paragraph>
</subSubSection>
<subSubSection id="C300657A154F811FFF0AF8E1FE9F57FB" pageId="41" pageNumber="42" type="discussion">
<paragraph id="8BA536F1154F811FFF0AF8E1FE9F57FB" blockId="41.[151,1436,1466,2030]" pageId="41" pageNumber="42">
<emphasis id="B96EEAE3154F811FFF0AF8E1FED4574A" bold="true" box="[199,379,1862,1887]" pageId="41" pageNumber="42">DISCUSSION:</emphasis>
Historically,
<taxonomicName id="4C1A4D72154F811FFDD9F8E0FD05574B" authorityName="Zhiltzova" authorityYear="1975" box="[532,682,1862,1886]" class="Insecta" family="Perlodidae" genus="Diura" kingdom="Animalia" order="Plecoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="majuscula">
<emphasis id="B96EEAE3154F811FFDD9F8E0FD05574B" box="[532,682,1862,1886]" italics="true" pageId="41" pageNumber="42">D. majuscula</emphasis>
</taxonomicName>
males and females have been assigned to separate species (Brinck 1949). Currently, it is currently unclear if
<taxonomicName id="4C1A4D72154F811FFDB9F8CCFCA45796" authorityName="Zhiltzova" authorityYear="1975" box="[628,779,1899,1923]" class="Insecta" family="Perlodidae" genus="Diura" kingdom="Animalia" order="Plecoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="majuscula">
<emphasis id="B96EEAE3154F811FFDB9F8CCFCA45796" box="[628,779,1899,1923]" italics="true" pageId="41" pageNumber="42">D. majuscula</emphasis>
</taxonomicName>
and the very similar
<taxonomicName id="4C1A4D72154F811FFC33F8CCFBD65797" baseAuthorityName="Kempny" baseAuthorityYear="1900" box="[1022,1145,1899,1922]" class="Insecta" family="Perlodidae" genus="Diura" kingdom="Animalia" order="Plecoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="nanseni">
<emphasis id="B96EEAE3154F811FFC33F8CCFBD65797" box="[1022,1145,1899,1922]" italics="true" pageId="41" pageNumber="42">D. nanseni</emphasis>
</taxonomicName>
are the same species. All specimens from
<collectingCountry id="F30D7661154F811FFE91F829FE6757B3" box="[348,456,1934,1958]" name="Mongolia" pageId="41" pageNumber="42">Mongolia</collectingCountry>
have been traditionally identified as
<taxonomicName id="4C1A4D72154F811FFC41F828FB8857B3" authorityName="Zhiltzova" authorityYear="1975" box="[908,1063,1934,1958]" class="Insecta" family="Perlodidae" genus="Diura" kingdom="Animalia" order="Plecoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="majuscula">
<emphasis id="B96EEAE3154F811FFC41F828FB8857B3" box="[908,1063,1934,1958]" italics="true" pageId="41" pageNumber="42">D. majuscula</emphasis>
</taxonomicName>
, but this may be for historical (Zhiltzova 1972) and not taxonomic or morphological reasons.
<taxonomicName id="4C1A4D72154F811FFCA7F814FBA257DF" baseAuthorityName="Kempny" baseAuthorityYear="1900" box="[874,1037,1971,1994]" class="Insecta" family="Perlodidae" genus="Diura" kingdom="Animalia" order="Plecoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="nanseni">
<emphasis id="B96EEAE3154F811FFCA7F814FBA257DF" box="[874,1037,1971,1994]" italics="true" pageId="41" pageNumber="42">Diura nanseni</emphasis>
</taxonomicName>
may occur sympatrically or be the same species.
</paragraph>
</subSubSection>
</treatment>
</document>