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<document id="F6369047ACE1A6204A2001FA2CD27D22" ID-CLB-Dataset="30030" ID-DOI="10.11646/zootaxa.4422.4.3" ID-GBIF-Dataset="d2c5d8c4-d2e2-44a7-add4-3a94812d2581" ID-ISSN="1175-5326" ID-Zenodo-Dep="1253219" ID-ZooBank="45E16185-7EE6-4768-88DF-6ACA1D29DCE4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1527266432226" checkinUser="plazi" docAuthor="Yang, Ho Jin, Seo, Ji Eun &amp; Gordon, Dennis P." docDate="2018" docId="03B387C39C67FFAE76DCC59052CFFAC1" docLanguage="en" docName="zootaxa.4422.4.3.pdf.imf" docOrigin="Zootaxa 4422 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Calloporina hayamiae Arakawa 1995" docType="treatment" docVersion="6" lastPageNumber="507" masterDocId="FF8AFFBB9C69FFA0764BC508565FFFC1" masterDocTitle="Sixteen new generic records of Korean Bryozoa from southern coastal waters and Jeju Island, East China Sea: evidence of tropical affinities" masterLastPageNumber="518" masterPageNumber="493" pageNumber="507" updateTime="1698545350699" updateUser="ExternalLinkService">
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<mods:title id="329D06BC19F76AEFC30526F2EC924A0A">Sixteen new generic records of Korean Bryozoa from southern coastal waters and Jeju Island, East China Sea: evidence of tropical affinities</mods:title>
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<mods:namePart id="ABBBD3CCC2E718C89078E93F04694ED9">Yang, Ho Jin</mods:namePart>
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<mods:namePart id="0A74A2E4A7CD32723515CCAF2D265322">Seo, Ji Eun</mods:namePart>
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<mods:namePart id="858985387942422AF7E2AD547660FA58">Gordon, Dennis P.</mods:namePart>
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<mods:title id="0E15F8AC5281E7118A2049813562C2A8">Zootaxa</mods:title>
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<mods:date id="A3ED3B2C272F6A7505F66D7B71A03B29">2018</mods:date>
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<treatment id="03B387C39C67FFAE76DCC59052CFFAC1" ID-DOI="http://doi.org/10.5281/zenodo.5964317" ID-GBIF-Taxon="144128916" ID-Zenodo-Dep="5964317" LSID="urn:lsid:plazi:treatment:03B387C39C67FFAE76DCC59052CFFAC1" httpUri="http://treatment.plazi.org/id/03B387C39C67FFAE76DCC59052CFFAC1" lastPageNumber="507" pageId="14" pageNumber="507">
<subSubSection id="C300655E9C67FFAE76DCC5905778FF12" pageId="14" pageNumber="507" type="nomenclature">
<paragraph id="8BA536D59C67FFAE76DCC5905436FF73" blockId="14.[151,617,152,211]" box="[151,617,152,178]" pageId="14" pageNumber="507">
<heading id="D0ED81B99C67FFAE76DCC5905436FF73" bold="true" box="[151,617,152,178]" fontSize="11" level="1" pageId="14" pageNumber="507" reason="1">
<taxonomicName id="4C1A4D569C67FFAE76DCC5905436FF73" authority="Arakawa, 1995" authorityName="Arakawa" authorityYear="1995" box="[151,617,152,178]" class="Gymnolaemata" family="Microporellidae" genus="Calloporina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="14" pageNumber="507" phylum="Bryozoa" rank="species" species="hayamiae">
<emphasis id="B96EEAC79C67FFAE76DCC5905436FF73" bold="true" box="[151,617,152,178]" pageId="14" pageNumber="507">
<emphasis id="B96EEAC79C67FFAE76DCC59057FBFF73" bold="true" box="[151,420,152,178]" italics="true" pageId="14" pageNumber="507">Calloporina hayamiae</emphasis>
<bibRefCitation id="EF8B4B249C67FFAE77E7C5905436FF73" box="[428,617,152,178]" pageId="14" pageNumber="507" refId="ref11662" refString="Arakawa, S. (1995) Bryozoan fauna in the Jizodo Formation (Pleistocene), Boso Peninsula, Honshu, Japan. Natural History Research, 3, 75 - 110." type="journal article">Arakawa, 1995</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BA536D59C67FFAE76DCC5B25778FF12" blockId="14.[151,617,152,211]" box="[151,295,186,211]" pageId="14" pageNumber="507">(Figs 26, 27)</paragraph>
</subSubSection>
<subSubSection id="C300655E9C67FFAE76DCC40A54C1FEF6" pageId="14" pageNumber="507" type="reference_group">
<paragraph id="8BA536D59C67FFAE76DCC40A52EAFED9" blockId="14.[151,1205,258,311]" box="[151,1205,258,280]" pageId="14" pageNumber="507">
<treatmentCitationGroup id="AB0A11FB9C67FFAE76DCC40A52EAFED9" box="[151,1205,258,280]" pageId="14" pageNumber="507">
<taxonomicName id="4C1A4D569C67FFAE76DCC40A5417FED9" ID-CoL="PVXW" authority=": Nishizawa 1990: 119" authorityName="Nishizawa" authorityPageNumber="119" authorityYear="1990" box="[151,584,258,280]" class="Gymnolaemata" family="Microporellidae" genus="Calloporina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="14" pageNumber="507" phylum="Bryozoa" rank="species" species="sculpta">
<emphasis id="B96EEAC79C67FFAE76DCC40A573DFED9" box="[151,354,258,280]" italics="true" pageId="14" pageNumber="507">Calloporina sculpta</emphasis>
:
<treatmentCitation id="0ABB10C49C67FFAE7724C40A5417FED9" author="" box="[367,584,258,280]" page="119" pageId="14" pageNumber="507">
<bibRefCitation id="EF8B4B249C67FFAE7724C40A5448FED9" box="[367,535,258,280]" pageId="14" pageNumber="507" refId="ref14849" refString="Nishizawa, Y. (1990) Illustrated introduction of fossils in the Niigata Prefecture 3: Cheilostomata, Bryozoa. Journal of the Association of Earth Sciences Education Niigata Prefecture, 25, 107 - 133." type="journal article">Nishizawa 1990</bibRefCitation>
: 119
</treatmentCitation>
</taxonomicName>
, pl. 4, fig. 3.
<emphasis id="B96EEAC79C67FFAE7485C40B54A7FED9" box="[718,760,259,280]" italics="true" pageId="14" pageNumber="507">Non</emphasis>
<taxonomicName id="4C1A4D569C67FFAE74B5C40A52EDFED9" ID-CoL="PVXW" authority="Canu &amp; Bassler, 1929" authorityName="Canu &amp; Bassler" authorityYear="1929" box="[766,1202,258,280]" class="Gymnolaemata" family="Microporellidae" genus="Calloporina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="14" pageNumber="507" phylum="Bryozoa" rank="species" species="sculpta">
<emphasis id="B96EEAC79C67FFAE74B5C40A5596FED9" box="[766,969,258,280]" italics="true" pageId="14" pageNumber="507">Calloporina sculpta</emphasis>
<bibRefCitation id="EF8B4B249C67FFAE759BC40A52EDFED9" box="[976,1202,258,280]" pageId="14" pageNumber="507" refId="ref12255" refString="Canu, F. &amp; Bassler, R. S. (1929) Bryozoa of the Philippine region. Bulletin of the United States National Museum, 100, 1 - 685, 94 pls." type="journal article">Canu &amp; Bassler, 1929</bibRefCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA536D59C67FFAE76DCC42954C1FEF6" blockId="14.[151,1205,258,311]" box="[151,670,289,311]" pageId="14" pageNumber="507">
<treatmentCitationGroup id="AB0A11FB9C67FFAE76DCC42954C1FEF6" box="[151,670,289,311]" pageId="14" pageNumber="507">
<taxonomicName id="4C1A4D569C67FFAE76DCC429541AFEF6" authority="Arakawa, 1995: 98" authorityName="Arakawa" authorityPageNumber="98" authorityYear="1995" box="[151,581,289,311]" class="Gymnolaemata" family="Microporellidae" genus="Calloporina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="14" pageNumber="507" phylum="Bryozoa" rank="species" species="hayamiae">
<emphasis id="B96EEAC79C67FFAE76DCC4295724FEF6" box="[151,379,289,311]" italics="true" pageId="14" pageNumber="507">Calloporina hayamiae</emphasis>
<treatmentCitation id="0ABB10C49C67FFAE77C9C429541AFEF6" author="" box="[386,581,289,311]" page="98" pageId="14" pageNumber="507">
<bibRefCitation id="EF8B4B249C67FFAE77C9C429547FFEF6" box="[386,544,289,311]" pageId="14" pageNumber="507" refId="ref11662" refString="Arakawa, S. (1995) Bryozoan fauna in the Jizodo Formation (Pleistocene), Boso Peninsula, Honshu, Japan. Natural History Research, 3, 75 - 110." type="journal article">Arakawa, 1995</bibRefCitation>
: 98
</treatmentCitation>
</taxonomicName>
, fig. 9A.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C300655E9C67FFAE76DCC46C55CDFE61" pageId="14" pageNumber="507" type="materials_examined">
<paragraph id="8BA536D59C67FFAE76DCC46C55CDFE61" blockId="14.[151,1437,356,1280]" pageId="14" pageNumber="507">
<materialsCitation id="3B723C889C67FFAE76DCC46C55CDFE61" ID-GBIF-Occurrence="1845321967" collectingDate="2016-07-30" collectorName="RV Kuklipgongwon-yeoungu" county="South Sea" location="Material" municipality="Hajo Island" pageId="14" pageNumber="507" specimenCount="1">
<emphasis id="B96EEAC79C67FFAE76DCC46C57DFFEBC" bold="true" box="[151,384,356,381]" pageId="14" pageNumber="507">
<location id="8EC5600E9C67FFAE76DCC46C56A0FEBC" LSID="urn:lsid:plazi:treatment:03B387C39C67FFAE76DCC59052CFFAC1:8EC5600E9C67FFAE76DCC46C56A0FEBC" box="[151,255,356,381]" county="South Sea" municipality="Hajo Island" name="Material" pageId="14" pageNumber="507">Material</location>
examined.
</emphasis>
<location id="8EC5600E9C67FFAE77CCC46C5786FEBC" LSID="urn:lsid:plazi:treatment:03B387C39C67FFAE76DCC59052CFFAC1:8EC5600E9C67FFAE77CCC46C5786FEBC" box="[391,473,356,381]" county="South Sea" municipality="Hajo Island" name="Unique" pageId="14" pageNumber="507">Unique</location>
colony (NIBRIV00000805890), from
<collectingMunicipality id="6BC1ACAF9C67FFAE75C7C46C524FFEBC" box="[908,1040,356,381]" pageId="14" pageNumber="507">Hajo Island</collectingMunicipality>
,
<collectingCounty id="62C44E599C67FFAE7257C46C52D0FEBD" box="[1052,1167,356,381]" pageId="14" pageNumber="507">South Sea</collectingCounty>
,
<date id="FFA410159C67FFAE72D0C46C5371FEBC" box="[1179,1326,356,381]" pageId="14" pageNumber="507" value="2016-07-30">
<collectingDate id="EFE0E9FD9C67FFAE72D0C46C5371FEBC" box="[1179,1326,356,381]" pageId="14" pageNumber="507" value="2016-07-30">
30 July
<quantity id="4CE29B309C67FFAE72B8C46C5371FEBC" box="[1267,1326,356,381]" metricMagnitude="3" metricUnit="m" metricValue="2.01633" pageId="14" pageNumber="507" unit="m" value="2016.33">2016</quantity>
</collectingDate>
</date>
, 33 m, on oyster shell, collected by grab from
<collectorName id="26EF53039C67FFAE7461C4805529FE61" box="[554,886,391,416]" pageId="14" pageNumber="507">RV Kuklipgongwon-yeoungu</collectorName>
1.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C300655E9C67FFAE768CC4A45348FBE8" pageId="14" pageNumber="507" type="description">
<paragraph id="8BA536D59C67FFAE768CC4A454E3FDB9" blockId="14.[151,1437,356,1280]" pageId="14" pageNumber="507">
<emphasis id="B96EEAC79C67FFAE768CC4A45706FE04" bold="true" box="[199,345,428,453]" pageId="14" pageNumber="507">Description.</emphasis>
Colony encrusting, multiserial, unilaminar, large, up to
<quantity id="4CE29B309C67FFAE7246C4A5523BFE05" box="[1037,1124,429,452]" metricMagnitude="-2" metricUnit="m" metricValue="7.7" pageId="14" pageNumber="507" unit="mm" value="77.0">77 mm</quantity>
across. Autozooids fairly regularly rhomboidal, rounded distally and proximally, more acute laterally, a little longer than wide. Lepralioid frontal shield with 1217 conspicuous areolar-septular pores around margin, with slight ribbing between pores that converges to low frontal-shield ridges of irregular disposition. Orifice high-arched; weakly concave proximal margin with low, sloping condyles in corners. Oral spines 56, stout. Ascopore at some distance from orifice, incised, semi-lunate, very thin, lacking denticles.
</paragraph>
<paragraph id="8BA536D59C67FFAE768CC78C5218FD24" blockId="14.[151,1437,356,1280]" pageId="14" pageNumber="507">Avicularia mostly single, on left or right sides of autozooids, less often paired, rarely absent; rostrum moderately long, acute, directed mostly obliquely distolaterally, tip often projecting just beyond lateral margin, crossbar well developed; half rostral length with palatal shelf, no opesial cryptocyst.</paragraph>
<paragraph id="8BA536D59C67FFAE768CC7E75527FC59" blockId="14.[151,1437,356,1280]" pageId="14" pageNumber="507">Developing ooecium with extensive membranous area of proximofrontal ectooecium that is calcified around periphery and has apical transverse rimmed cap; endooecium skeletally complex, initially with transverse creases proximofrontally and honeycomb of calcification frontally; as development proceeds, lateral calcification of ectooecium covers most of endooecium, leaving bean-shaped or bifenestrate window showing endooecial pits, distal of which is rimmed ectooecial cap, itself sometimes etched.</paragraph>
<paragraph id="8BA536D59C67FFAE768CC6AC57ABFC1E" blockId="14.[151,1437,356,1280]" pageId="14" pageNumber="507">Well-developed basal pore-chambers at colony margin, two distolateral, one distal, each subdivided into paired chambers. Ancestrula not seen.</paragraph>
<paragraph id="8BA536D59C67FFAE768CC6E45348FBE8" blockId="14.[151,1437,356,1280]" pageId="14" pageNumber="507">
<emphasis id="B96EEAC79C67FFAE768CC6E45722FBC4" bold="true" box="[199,381,1004,1029]" pageId="14" pageNumber="507">Measurements.</emphasis>
ZL 388524 (432) µm; ZW 295551 (378) µm; OrL 87108 (99) µm; OrW 91110 (100) µm; AvL 101158 (138) µm; AvW 3967 (55) µm; OoL 137166 (156) µm; OoW 153186 (172) µm.
</paragraph>
</subSubSection>
<subSubSection id="C300655E9C67FFAE768CC13C52B2FB1C" pageId="14" pageNumber="507" type="discussion">
<paragraph id="8BA536D59C67FFAE768CC13C52B2FB1C" blockId="14.[151,1437,356,1280]" pageId="14" pageNumber="507">
<emphasis id="B96EEAC79C67FFAE768CC13C5764FB8C" bold="true" box="[199,315,1076,1101]" pageId="14" pageNumber="507">Remarks.</emphasis>
<taxonomicName id="4C1A4D569C67FFAE7709C13C5460FB8C" box="[322,575,1076,1101]" class="Gymnolaemata" family="Microporellidae" genus="Calloporina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="14" pageNumber="507" phylum="Bryozoa" rank="species" species="hayamiae">
<emphasis id="B96EEAC79C67FFAE7709C13C5460FB8C" box="[322,575,1076,1101]" italics="true" pageId="14" pageNumber="507">Calloporina hayamiae</emphasis>
</taxonomicName>
was previously known only as a Pleistocene fossil from the Boso Peninsula, on the east coast of central Honshu,
<collectingCountry id="F30D76459C67FFAE741AC15154CBFBB1" box="[593,660,1113,1136]" name="Japan" pageId="14" pageNumber="507">Japan</collectingCountry>
. The characters of our material are identical to that described by
<bibRefCitation id="EF8B4B249C67FFAE76DCC174570EFB54" box="[151,337,1148,1173]" pageId="14" pageNumber="507" refId="ref11662" refString="Arakawa, S. (1995) Bryozoan fauna in the Jizodo Formation (Pleistocene), Boso Peninsula, Honshu, Japan. Natural History Research, 3, 75 - 110." type="journal article">Arakawa (1995)</bibRefCitation>
, especially the distinctive semi-lunate smile-like ascopore incised within sloping margins, and the form of the mature ooecium.
<bibRefCitation id="EF8B4B249C67FFAE744CC19754E1FB79" box="[519,702,1183,1208]" pageId="14" pageNumber="507" refId="ref11662" refString="Arakawa, S. (1995) Bryozoan fauna in the Jizodo Formation (Pleistocene), Boso Peninsula, Honshu, Japan. Natural History Research, 3, 75 - 110." type="journal article">Arakawa (1995)</bibRefCitation>
noted a range of 46 oral-spine bases.
<taxonomicName id="4C1A4D569C67FFAE7238C197532FFB79" box="[1139,1392,1183,1208]" class="Gymnolaemata" family="Microporellidae" genus="Calloporina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="14" pageNumber="507" phylum="Bryozoa" rank="species" species="hayamiae">
<emphasis id="B96EEAC79C67FFAE7238C197532FFB79" box="[1139,1392,1183,1208]" italics="true" pageId="14" pageNumber="507">Calloporina hayamiae</emphasis>
</taxonomicName>
has not yet been reported from modern Japanese waters but it is highly likely that it will be discovered.
</paragraph>
</subSubSection>
<subSubSection id="C300655E9C67FFAE768CC1EF52CFFAC1" box="[199,1168,1255,1280]" pageId="14" pageNumber="507" type="distribution">
<paragraph id="8BA536D59C67FFAE768CC1EF52CFFAC1" blockId="14.[151,1437,356,1280]" box="[199,1168,1255,1280]" pageId="14" pageNumber="507">
<emphasis id="B96EEAC79C67FFAE768CC1EF573FFAC1" bold="true" box="[199,352,1255,1280]" pageId="14" pageNumber="507">Distribution.</emphasis>
<collectingCountry id="F30D76459C67FFAE772CC1E157F4FAC1" box="[359,427,1257,1280]" name="Japan" pageId="14" pageNumber="507">Japan</collectingCountry>
: Pleistocene, east-central Honshu.
<collectingCountry id="F30D76459C67FFAE757BC1E15526FAC1" box="[816,889,1257,1280]" name="South Korea" pageId="14" pageNumber="507">Korea</collectingCountry>
:
<collectingRegion id="49DEF8379C67FFAE75C8C1EF55EFFAC1" box="[899,944,1255,1280]" country="South Korea" name="Jeju" pageId="14" pageNumber="507">Jeju</collectingRegion>
Island,
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depth.
</paragraph>
</subSubSection>
</treatment>
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