406 lines
49 KiB
XML
406 lines
49 KiB
XML
<document id="6F05D0DA826ABB3588A33ECE8F0B9590" ID-DOI="10.1649/0010-065X-72.mo4.1" ID-ISSN="1938-4394" ID-Zenodo-Dep="5102840" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626277672172" checkinUser="carolina" docAuthor="Mullen, Logan J., Campbell, J. M. & Sikes, Derek S." docDate="2018" docId="03971078FFD9BC68FD06FB17FE28C8F4" docLanguage="en" docName="TheColeopBull.mo.72.16.1-54.pdf" docOrigin="The Coleopterists Bulletin 72" docSource="http://dx.doi.org/10.1649/0010-065x-72.mo4.1" docStyle="DocumentStyle:A431C8F32F3D86E670C18C13DA9B5F2B.4:TheColeopBull.2000-2009.monograph" docStyleId="A431C8F32F3D86E670C18C13DA9B5F2B" docStyleName="TheColeopBull.2000-2009.monograph" docStyleVersion="4" docTitle="Phlaeopterus obsoletus Mullen and Campbell 2018, new species" docType="treatment" docUuid="D2D70CE0-DDEC-4834-857E-DC6D80FFD18D" docUuidSource="ZooBank" docVersion="3" lastPageNumber="45" masterDocId="FFAE6800FFF0BC44FFDAFFA6FFBDCE36" masterDocTitle="Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)" masterLastPageNumber="54" masterPageNumber="1" pageNumber="42" updateTime="1699007718751" updateUser="plazi">
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<mods:title id="EAE7A51800807281896CFF97D2CDF7A5">Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)</mods:title>
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<mods:namePart id="031E86972351409986761281661F4496">Mullen, Logan J.</mods:namePart>
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<mods:namePart id="B2F6DB4E34085802ED4BA800FBD8E8C5">Campbell, J. M.</mods:namePart>
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<mods:namePart id="9CCAD2FE649F558E907C8637869F5E2B">Sikes, Derek S.</mods:namePart>
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<mods:typeOfResource id="7AD143492CC4CE681C28C3D3DD8CB9E3">text</mods:typeOfResource>
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<mods:title id="15DF05CCE2243792869F1FCE88E0758B">The Coleopterists Bulletin</mods:title>
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<mods:part id="D59A2F6203C1F696C6CE543A98B0B9CA">
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<mods:date id="53F7E1637CF34AE85C6A7843807AFE2F">2018</mods:date>
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<mods:number id="B1EFDA0A7451AE7AA47553D1D3FDDE21">2018-12-28</mods:number>
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<mods:number id="4627168EDDA13708CA7D197E1A7F0897">72</mods:number>
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<mods:url id="30B491340FA066F99BE40930E7C520C7">http://dx.doi.org/10.1649/0010-065x-72.mo4.1</mods:url>
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<mods:classification id="7A5828750B95D181492D538713A8AD74">journal article</mods:classification>
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<subSubSection id="C324F2E5FFD9BC6DFD06FB17FBAECAD7" pageId="41" pageNumber="42" type="nomenclature">
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<paragraph id="8B81A16EFFD9BC6DFD06FB17FBAECAD7" blockId="41.[690,1168,1201,1355]" pageId="41" pageNumber="42">
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<heading id="D0C91602FFD9BC6DFD06FB17FBAECAD7" centered="true" fontSize="10" level="2" pageId="41" pageNumber="42" reason="2">
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<emphasis id="B94A7D7CFFD9BC6DFD06FB17FBAECAD7" bold="true" pageId="41" pageNumber="42">
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15.
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<taxonomicName id="4C3EDAEDFFD9BC6DFCD8FB17FC2CCAD7" ID-CoL="8Y8D2" authority="Mullen and Campbell" authorityName="Mullen and Campbell" authorityYear="2018" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="obsoletus" status="sp. nov.">
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<emphasis id="B94A7D7CFFD9BC6DFCD8FB17FC55CAF0" bold="true" box="[770,1000,1201,1222]" italics="true" pageId="41" pageNumber="42">Phlaeopterus obsoletus</emphasis>
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Mullen and Campbell
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</taxonomicName>
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,
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<taxonomicNameLabel id="A279C007FFD9BC6DFC44FB6AFBAECAD7" box="[926,1043,1228,1249]" pageId="41" pageNumber="42" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<caption id="DF41F1E6FFD9BC6DFF75FB72FEC1CB21" ID-DOI="http://doi.org/10.5281/zenodo.5102897" ID-Zenodo-Dep="5102897" httpUri="https://zenodo.org/record/5102897/files/figure.png" pageId="41" pageNumber="42" startId="41.[175,206,1236,1255]" targetBox="[154,634,195,1220]" targetPageId="41">
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<paragraph id="8B81A16EFFD9BC6DFF75FB72FEC1CB21" blockId="41.[151,639,1236,1303]" pageId="41" pageNumber="42">
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<emphasis id="B94A7D7CFFD9BC6DFF75FB72FF46CAD1" bold="true" box="[175,251,1236,1255]" pageId="41" pageNumber="42">Fig. 34.</emphasis>
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<taxonomicName id="4C3EDAEDFFD9BC6DFECBFB72FE72CAD1" box="[273,463,1236,1255]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="B94A7D7CFFD9BC6DFECBFB72FE3CCAD1" box="[273,385,1236,1255]" italics="true" pageId="41" pageNumber="42">Phlaeopterus</emphasis>
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species
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</taxonomicName>
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, head, dorsal and dorsal oblique views. A–B)
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<taxonomicName id="4C3EDAEDFFD9BC6DFE7BFB4AFD9CCAC9" authorityName="Hatch" authorityYear="1957" box="[417,545,1260,1279]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="lagrandeuri">
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<emphasis id="B94A7D7CFFD9BC6DFE7BFB4AFD9CCAC9" box="[417,545,1260,1279]" italics="true" pageId="41" pageNumber="42">P. lagrandeuri</emphasis>
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</taxonomicName>
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, C–D)
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<taxonomicName id="4C3EDAEDFFD9BC6DFDB4FB4AFF68CB21" authorityName="Hatch" authorityYear="1957" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="houkae">
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<emphasis id="B94A7D7CFFD9BC6DFDB4FB4AFF68CB21" italics="true" pageId="41" pageNumber="42">P. houkae</emphasis>
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</taxonomicName>
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, E–F)
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<taxonomicName id="4C3EDAEDFFD9BC6DFECBFAA2FECACB21" authorityName="Mullen and Campbell" authorityYear="2018" box="[273,375,1284,1303]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="obsoletus">
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<emphasis id="B94A7D7CFFD9BC6DFECBFAA2FECACB21" box="[273,375,1284,1303]" italics="true" pageId="41" pageNumber="42">P. obsoletus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</caption>
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<subSubSection id="C324F2E5FFD9BC6DFD68FB41FB29CB48" pageId="41" pageNumber="42" type="description">
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<paragraph id="8B81A16EFFD9BC6DFD68FB41FB32CB7D" blockId="41.[690,1168,1201,1355]" pageId="41" pageNumber="42">
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(
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<figureCitation id="1305BDEBFFD9BC6DFD63FB41FCB3CACA" box="[697,782,1255,1276]" captionStart="Fig" captionStartId="13.[175,206,1742,1761]" captionTargetBox="[157,1166,196,1725]" captionTargetId="figure-1@13.[155,1168,194,1727]" captionTargetPageId="13" captionText="Fig. 4. Phlaeopterus species, habitus. A) P. loganensis, female (UAMObs:Ento:233072), B) P. longipennis, male" figureDoi="http://doi.org/10.5281/zenodo.5102849" httpUri="https://zenodo.org/record/5102849/files/figure.png" pageId="41" pageNumber="42">Figs. 4C</figureCitation>
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,
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<figureCitation id="1305BDEBFFD9BC6DFCC3FB41FCF8CACA" box="[793,837,1255,1276]" captionStart="Fig" captionStartId="19.[175,206,1027,1046]" captionText="Fig. 13. Distributions of Phlaeopterus species based on all georeferenced specimen records. A) P. obsoletus, B) P. occidentalis. Grey dots represent localities where at least one Phlaeopterus species, but not the species of interest, was collected. Red dots represent localities where the species of interest was collected." pageId="41" pageNumber="42">13A</figureCitation>
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,
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<figureCitation id="1305BDEBFFD9BC6DFC8AFB41FCC6CACA" box="[848,891,1255,1276]" captionStart="Fig" captionStartId="21.[175,206,1672,1691]" captionTargetBox="[157,1166,199,1650]" captionTargetId="figure-1@21.[154,1169,194,1656]" captionTargetPageId="21" captionText="Fig. 15. Pronota, dorsal view, of A) Lesteva pallipes LeConte, 1863, B) Lesteva longoelytrata (Goeze, 1777), C) Unamis sp. undescribed, D) Unamis sp., E) Phlaeopterus lagrandeuri, F) Phlaeopterus houkae, G) Phlaeopterus longipennis, H) Phlaeopterus obsoletus." figureDoi="http://doi.org/10.5281/zenodo.5102865" httpUri="https://zenodo.org/record/5102865/files/figure.png" pageId="41" pageNumber="42">15H</figureCitation>
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,
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||
<figureCitation id="1305BDEBFFD9BC6DFC5DFB41FC0FCACA" box="[903,946,1255,1276]" captionStart="Fig" captionStartId="23.[175,206,1547,1566]" captionTargetBox="[168,1155,197,1527]" captionTargetId="figure-55@23.[164,1159,194,1532]" captionTargetPageId="23" captionText="Fig. 17. Phlaeopterus species. Pronota, dorsal view, of A) P. fusconiger, B) P. frosti, C) P. filicornis, D) P. hatchi, E) P. elongatus. Abdominal segments 4–6, dorsal view, of F) P. houkae, G) P. obsoletus, H) P. elongatus." figureDoi="http://doi.org/10.5281/zenodo.5102867" httpUri="https://zenodo.org/record/5102867/files/figure.png" pageId="41" pageNumber="42">17G</figureCitation>
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,
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<figureCitation id="1305BDEBFFD9BC6DFC64FB41FC55CACA" box="[958,1000,1255,1276]" captionStart="Fig" captionStartId="25.[709,740,1185,1204]" captionTargetBox="[689,1169,195,1168]" captionTargetId="figure-245@25.[688,1170,194,1169]" captionTargetPageId="25" captionText="Fig. 20. Phlaeopterus species, mesosterna, ventral oblique view. A) P. longipennis, B) P. obsoletus, C) P. olympicus, D) P. frosti, E) P. occidentalis, F) P. elongatus." figureDoi="http://doi.org/10.5281/zenodo.5102871" httpUri="https://zenodo.org/record/5102871/files/figure.png" pageId="41" pageNumber="42">20B</figureCitation>
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, 21D, 29A, 31G, 33G, 34E–F, 37H–I, 39C–D, 41A) Zoobank.org/
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<uri id="FFAFAD6CFFD9BC6DFCAAFABAFB32CB7D" pageId="41" pageNumber="42">
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urn:lsid:zoobank.org:act:
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<uuid id="FF989BBBFFD9BC6DFD68FA90FB32CB7D" box="[690,1167,1334,1355]" pageId="41" pageNumber="42">D2D70CE0-DDEC-4834-857E-DC6D80FFD18D</uuid>
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</uri>
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</paragraph>
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<paragraph id="8B81A16EFFD9BC6DFD77FACFFB29CB48" blockId="41.[685,1172,1385,1406]" box="[685,1172,1385,1406]" pageId="41" pageNumber="42">
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<bibRefCitation id="EFAFDC9FFFD9BC6DFD77FACFFCFACB48" author="Hatch" box="[685,839,1385,1406]" pageId="41" pageNumber="42" refId="ref28862" refString="Hatch, M. H. 1957. The Beetles of the Pacific Northwest. Part II: Staphyliniformia. University of Washington Publications in Biology, Seattle, WA." type="book" year="1957">Hatch 1957: 461</bibRefCitation>
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[misidentification as
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<taxonomicName id="4C3EDAEDFFD9BC6DFBDDFACFFB3ACB48" authorityName="Mullen & Campbell & Sikes" authorityYear="2018" baseAuthorityName="Casey" baseAuthorityYear="1885" box="[1031,1159,1385,1406]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="longipennis">
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||
<emphasis id="B94A7D7CFFD9BC6DFBDDFACFFB3ACB48" box="[1031,1159,1385,1406]" italics="true" pageId="41" pageNumber="42">V. longipennis</emphasis>
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</taxonomicName>
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].
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</paragraph>
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</subSubSection>
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<subSubSection id="C324F2E5FFD9BC6DFD1FFA05FC64CBE5" pageId="41" pageNumber="42" type="materials_examined">
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<paragraph id="8B81A16EFFD9BC6DFD1FFA05FC64CBE5" blockId="41.[685,1173,1443,1730]" pageId="41" pageNumber="42">
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<materialsCitation id="3B56AB33FFD9BC6DFD1FFA05FC6BCBE5" country="United States of America" location="Berkeley Park" municipality="Mt. Rainer National Park" pageId="41" pageNumber="42" specimenCount="1" stateProvince="Washington" typeStatus="holotype">
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<emphasis id="B94A7D7CFFD9BC6DFD1FFA05FCE5CB8E" bold="true" box="[709,856,1443,1464]" pageId="41" pageNumber="42">
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<typeStatus id="54851FCCFFD9BC6DFD1FFA05FD4ACB8E" box="[709,759,1443,1464]" pageId="41" pageNumber="42">Type</typeStatus>
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Locality.
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||
</emphasis>
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||
<location id="8EE1F7B5FFD9BC6DFCB8FA05FC52CB8E" LSID="urn:lsid:plazi:treatment:03971078FFD9BC68FD06FB17FE28C8F4:8EE1F7B5FFD9BC6DFCB8FA05FC52CB8E" box="[866,1007,1443,1464]" country="United States of America" municipality="Mt. Rainer National Park" name="Berkeley Park" pageId="41" pageNumber="42" stateProvince="Washington">Berkeley Park</location>
|
||
,
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||
<collectingMunicipality id="6BE53B14FFD9BC6DFC26FA05FCA4CBE5" pageId="41" pageNumber="42">Mt. Rainer National Park</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49FA6F8CFFD9BC6DFCFEFA18FC25CBE5" box="[804,920,1470,1491]" country="United States of America" name="Washington" pageId="41" pageNumber="42">Washington</collectingRegion>
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||
,
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||
<collectingCountry id="F329E1FEFFD9BC6DFC79FA18FC6BCBE5" box="[931,982,1470,1491]" name="United States of America" pageId="41" pageNumber="42">USA</collectingCountry>
|
||
</materialsCitation>
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||
.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C324F2E5FFD9BC6FFD1FFA7EFE30C83E" lastPageId="43" lastPageNumber="44" pageId="41" pageNumber="42" type="description">
|
||
<paragraph id="8B81A16EFFD9BC6FFD1FFA7EFE30C83E" blockId="41.[685,1173,1443,1730]" lastBlockId="43.[151,639,1417,1730]" lastPageId="43" lastPageNumber="44" pageId="41" pageNumber="42">
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||
<emphasis id="B94A7D7CFFD9BC6DFD1FFA7EFC14CBDB" bold="true" box="[709,937,1496,1517]" pageId="41" pageNumber="42">Description. Habitus:</emphasis>
|
||
Length
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||
<quantity id="4CC60C8BFFD9BC6DFBCAFA7EFB2CCBDB" box="[1040,1169,1496,1518]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" metricValueMax="3.9" metricValueMin="2.9" pageId="41" pageNumber="42" unit="mm" value="3.4" valueMax="3.9" valueMin="2.9">2.9–3.9 mm</quantity>
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||
. Light brown to dark brown, legs lighter, antennae sometimes lighter (
|
||
<figureCitation id="1305BDEBFFD9BC6DFCB0F9A8FC04C815" box="[874,953,1550,1571]" captionStart="Fig" captionStartId="13.[175,206,1742,1761]" captionTargetBox="[157,1166,196,1725]" captionTargetId="figure-1@13.[155,1168,194,1727]" captionTargetPageId="13" captionText="Fig. 4. Phlaeopterus species, habitus. A) P. loganensis, female (UAMObs:Ento:233072), B) P. longipennis, male" figureDoi="http://doi.org/10.5281/zenodo.5102849" httpUri="https://zenodo.org/record/5102849/files/figure.png" pageId="41" pageNumber="42">Fig. 4C</figureCitation>
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).
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<emphasis id="B94A7D7CFFD9BC6DFC17F9A8FBB6C815" bold="true" box="[973,1035,1550,1571]" pageId="41" pageNumber="42">Head:</emphasis>
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Width across eyes to head length slightly wider than long (
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||
<figureCitation id="1305BDEBFFD9BC6DFBB7F98EFD4DC86E" captionStart="Fig" captionStartId="41.[175,206,1236,1255]" captionTargetBox="[154,634,195,1220]" captionTargetId="figure-493@41.[153,635,194,1221]" captionTargetPageId="41" captionText="Fig. 34. Phlaeopterus species, head, dorsal and dorsal oblique views. A–B) P. lagrandeuri, C–D) P. houkae, E–F) P. obsoletus." figureDoi="http://doi.org/10.5281/zenodo.5102897" httpUri="https://zenodo.org/record/5102897/files/figure.png" pageId="41" pageNumber="42">Fig. 34E–F</figureCitation>
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). Interantennal groove absent. Anteocellar foveae moderately large, deeply impressed. Eyes moderately pubescent on ventral half, with more than 10 setae. Antennomeres 3–10 each 1.3–1.9 times longer than wide, antennomeres 4–11 each with many sensory pits with groups of pore-like openings (
|
||
<figureCitation id="1305BDEBFFDABC6EFECBFAF2FEC3CB5F" box="[273,382,1363,1385]" captionStart="Fig" captionStartId="43.[175,206,1294,1313]" captionTargetBox="[156,1167,196,1276]" captionTargetId="figure-165@43.[154,1169,194,1278]" captionTargetPageId="43" captionText="Fig. 37. Sensory structures on antenna of A) Lesteva pallipes, B–C) Lesteva pubescens, D–E) Unamis sp. undescribed, F–G) Phlaeopterus houkae, H–I) Phlaeopterus obsoletus, J–K) Phlaeopterus elongatus. In B, D, F, H, and J, white boxes indicate inset area that is magnified in the image to the right." figureDoi="http://doi.org/10.5281/zenodo.5102903" httpUri="https://zenodo.org/record/5102903/files/figure.png" pageId="42" pageNumber="43">Fig. 37H–I</figureCitation>
|
||
). Ocelli absent. Nuchal constriction vague. Mandibular molar area with Lshaped row of setae. Epipharynx as in Fig. 33G. Maxillary palpus as in
|
||
<figureCitation id="1305BDEBFFDABC6EFEA5FA05FE4CCB8E" box="[383,497,1443,1465]" captionStart="Fig" captionStartId="44.[731,762,1213,1232]" captionTargetBox="[711,1189,196,1195]" captionTargetId="figure-182@44.[709,1191,194,1197]" captionTargetPageId="44" captionText="Fig. 39. Phlaeopterus species, maxillae, dorsal and ventral views. A–B) P. houkae, C–D) P. obsoletus, E–F) P. elongatus." figureDoi="http://doi.org/10.5281/zenodo.5102905" httpUri="https://zenodo.org/record/5102905/files/figure.png" pageId="42" pageNumber="43">Fig. 39C–D</figureCitation>
|
||
. Hypopharynx as in
|
||
<figureCitation id="1305BDEBFFDABC6EFF16FA18FE97CBE5" box="[204,298,1470,1491]" captionStart="Fig" captionStartId="45.[709,740,1185,1204]" captionTargetBox="[154,1169,195,1168]" captionTargetPageId="45" captionText="Fig. 41. Phlaeopterus species, hypopharynges, dorsal view, of A) P. obsoletus, B) P. kavanaughi, C) P. elongatus. D) Phlaeopterus lagrandeuri, metatrochanter with apical tooth. Heads, ventral view, of E) Lesteva pallipes, F) Lesteva pubescens." figureDoi="http://doi.org/10.5281/zenodo.5102909" httpUri="https://zenodo.org/record/5102909/files/figure.png" pageId="42" pageNumber="43">Fig. 41A</figureCitation>
|
||
. Labrum with sensory pores along anterior margin only (
|
||
<figureCitation id="1305BDEBFFDABC6EFE5BFA7EFE61CBDB" box="[385,476,1496,1517]" captionStart="Fig" captionStartId="38.[197,228,1598,1617]" captionTargetBox="[177,1188,196,1581]" captionTargetId="figure-0@38.[175,1190,194,1583]" captionTargetPageId="38" captionText="Fig. 31. Labra of A) Lesteva pallipes, B) Lesteva pubescens, C) Unamis sp. undescribed, D) Unamis sp., E) Phlaeopterus lagrandeuri, F) Phlaeopterus houkae, G) Phlaeopterus obsoletus, H) Phlaeopterus filicornis. White arrows indicate labrum." figureDoi="http://doi.org/10.5281/zenodo.5102893" httpUri="https://zenodo.org/record/5102893/files/figure.png" pageId="42" pageNumber="43">Fig. 31G</figureCitation>
|
||
). Labial palpi with 3
|
||
<superScript id="7C4B0C26FFDABC6EFF63FA48FF75CBCA" attach="left" box="[185,200,1518,1532]" fontSize="6" pageId="42" pageNumber="43">rd</superScript>
|
||
palpomere 1.4–1.8 times longer than palpomere 2.
|
||
<emphasis id="B94A7D7CFFDABC6EFF1FF9A8FEA5C815" bold="true" box="[197,280,1550,1571]" pageId="42" pageNumber="43">Thorax:</emphasis>
|
||
Pronotum narrow (
|
||
<figureCitation id="1305BDEBFFDABC6EFE0EF9A8FD90C815" box="[468,557,1550,1571]" captionStart="Fig" captionStartId="21.[175,206,1672,1691]" captionTargetBox="[157,1166,199,1650]" captionTargetId="figure-1@21.[154,1169,194,1656]" captionTargetPageId="21" captionText="Fig. 15. Pronota, dorsal view, of A) Lesteva pallipes LeConte, 1863, B) Lesteva longoelytrata (Goeze, 1777), C) Unamis sp. undescribed, D) Unamis sp., E) Phlaeopterus lagrandeuri, F) Phlaeopterus houkae, G) Phlaeopterus longipennis, H) Phlaeopterus obsoletus." figureDoi="http://doi.org/10.5281/zenodo.5102865" httpUri="https://zenodo.org/record/5102865/files/figure.png" pageId="42" pageNumber="43">Fig. 15H</figureCitation>
|
||
), length to width ratio = 0.79–0.88; ratio of pronotal width to head width = 1.25–1.33; maximum width slightly narrower than elytra at base; punctures separated by average distance subequal to or slightly greater than diameter of a puncture; lateral margins not explanate anterad or posterad lateral fovea; lateral foveae moderately deeply impressed, narrowly separated from lateral margins. Elytra with humeral angles convex; epipleural carina not projecting; 2.0–2.3 times longer than pronotum; apical margins convex. Wings developed or brachypterous. Mesosternum with projecting tooth; longitudinal carina along midline of mesosternum vaguely present, becoming obscured on posterior half (
|
||
<figureCitation id="1305BDEBFFDABC6EFC03F98EFB89C80B" box="[985,1076,1576,1597]" captionStart="Fig" captionStartId="25.[709,740,1185,1204]" captionTargetBox="[689,1169,195,1168]" captionTargetId="figure-245@25.[688,1170,194,1169]" captionTargetPageId="25" captionText="Fig. 20. Phlaeopterus species, mesosterna, ventral oblique view. A) P. longipennis, B) P. obsoletus, C) P. olympicus, D) P. frosti, E) P. occidentalis, F) P. elongatus." figureDoi="http://doi.org/10.5281/zenodo.5102871" httpUri="https://zenodo.org/record/5102871/files/figure.png" pageId="42" pageNumber="43">Fig. 20B</figureCitation>
|
||
).
|
||
<emphasis id="B94A7D7CFFDABC6EFB93F98EFB3DC80B" bold="true" box="[1097,1152,1576,1597]" pageId="42" pageNumber="43">Legs:</emphasis>
|
||
All tibiae evenly pubescent to apex. Metatrochanter without tooth on apical margin.
|
||
<emphasis id="B94A7D7CFFDABC6EFC34F9FBFBE5C844" bold="true" box="[1006,1112,1629,1650]" pageId="42" pageNumber="43">Abdomen:</emphasis>
|
||
Tergites IV and V with wing-folding spicules very broad, nearly contiguous or contiguous in shape of single transverse band. Apical palisade fringe on tergite VII absent.
|
||
<emphasis id="B94A7D7CFFDBBC6FFEC3FA2FFE3FCBA8" bold="true" box="[281,386,1417,1438]" pageId="43" pageNumber="44">Aedeagus:</emphasis>
|
||
Length
|
||
<quantity id="4CC60C8BFFDBBC6FFE3EFA2FFDC6CBA8" box="[484,635,1417,1438]" metricMagnitude="-4" metricUnit="m" metricValue="5.75" metricValueMax="6.3" metricValueMin="5.2" pageId="43" pageNumber="44" unit="mm" value="0.575" valueMax="0.63" valueMin="0.52">0.52–0.63 mm</quantity>
|
||
. Median lobe short, oblong, sides subparallel, apex broadly convex. Parameres diverging in basal half (Fig. 21D). Internal sac large; with two mediolateral patches of microspinules.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF41F1E6FFDABC6EFF1FFB0AFE27CB31" ID-DOI="http://doi.org/10.5281/zenodo.5102899" ID-Zenodo-Dep="5102899" httpUri="https://zenodo.org/record/5102899/files/figure.png" pageId="42" pageNumber="43" startId="42.[197,228,1196,1215]" targetBox="[176,656,195,1179]" targetPageId="42">
|
||
<paragraph id="8B81A16EFFDABC6EFF1FFB0AFE27CB31" blockId="42.[173,659,1196,1287]" pageId="42" pageNumber="43">
|
||
<emphasis id="B94A7D7CFFDABC6EFF1FFB0AFEB4CA89" bold="true" box="[197,265,1196,1215]" pageId="42" pageNumber="43">Fig. 35.</emphasis>
|
||
Heads, dorsal and dorsal oblique views, of A–B)
|
||
<taxonomicName id="4C3EDAEDFFDABC6EFF37FB62FE6CCAE1" authorityName="Mullen and Campbell" authorityYear="2018" box="[237,465,1220,1239]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="kavanaughi">
|
||
<emphasis id="B94A7D7CFFDABC6EFF37FB62FE6CCAE1" box="[237,465,1220,1239]" italics="true" pageId="42" pageNumber="43">Phlaeopterus kavanaughi</emphasis>
|
||
</taxonomicName>
|
||
, C–D
|
||
<taxonomicName id="4C3EDAEDFFDABC6EFDF9FB62FF5FCAD9" authorityName="Mullen and Campbell" authorityYear="2018" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="hatchi">
|
||
<emphasis id="B94A7D7CFFDABC6EFDF9FB62FF5FCAD9" italics="true" pageId="42" pageNumber="43">Phlaeopterus hatchi</emphasis>
|
||
</taxonomicName>
|
||
. Eyes, dorsal view, of E)
|
||
<taxonomicName id="4C3EDAEDFFDABC6EFE09FB7AFD30CAD9" baseAuthorityName="Goeze" baseAuthorityYear="1777" box="[467,653,1244,1263]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="longoelytrata">
|
||
<emphasis id="B94A7D7CFFDABC6EFE09FB7AFD30CAD9" box="[467,653,1244,1263]" italics="true" pageId="42" pageNumber="43">Lesteva longoelytrata</emphasis>
|
||
</taxonomicName>
|
||
, F)
|
||
<taxonomicName id="4C3EDAEDFFDABC6EFF1CFB52FE95CB31" box="[198,296,1268,1287]" class="Insecta" family="Staphylinidae" genus="Unamis" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B94A7D7CFFDABC6EFF1CFB52FEB5CB31" box="[198,264,1268,1287]" italics="true" pageId="42" pageNumber="43">Unamis</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
undescribed.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF41F1E6FFDABC6EFD01FB5EFBA7CB0D" ID-DOI="http://doi.org/10.5281/zenodo.5102901" ID-Zenodo-Dep="5102901" httpUri="https://zenodo.org/record/5102901/files/figure.png" pageId="42" pageNumber="43" startId="42.[731,762,1272,1291]" targetBox="[711,1189,196,1255]" targetPageId="42">
|
||
<paragraph id="8B81A16EFFDABC6EFD01FB5EFBA7CB0D" blockId="42.[707,1194,1272,1339]" pageId="42" pageNumber="43">
|
||
<emphasis id="B94A7D7CFFDABC6EFD01FB5EFCA3CB3D" bold="true" box="[731,798,1272,1291]" pageId="42" pageNumber="43">Fig. 36.</emphasis>
|
||
<taxonomicName id="4C3EDAEDFFDABC6EFCEEFB5EFC57CB3D" box="[820,1002,1272,1291]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B94A7D7CFFDABC6EFCEEFB5EFC19CB3D" box="[820,932,1272,1291]" italics="true" pageId="42" pageNumber="43">Phlaeopterus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, eyes, dorsal view. A)
|
||
<taxonomicName id="4C3EDAEDFFDABC6EFD19FAB6FCFCCB15" authorityName="Hatch" authorityYear="1957" box="[707,833,1296,1315]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="lagrandeuri">
|
||
<emphasis id="B94A7D7CFFDABC6EFD19FAB6FCFCCB15" box="[707,833,1296,1315]" italics="true" pageId="42" pageNumber="43">P. lagrandeuri</emphasis>
|
||
</taxonomicName>
|
||
, B)
|
||
<taxonomicName id="4C3EDAEDFFDABC6EFCB7FAB6FC7ECB15" authorityName="Hatch" authorityYear="1957" box="[877,963,1296,1315]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="houkae">
|
||
<emphasis id="B94A7D7CFFDABC6EFCB7FAB6FC7ECB15" box="[877,963,1296,1315]" italics="true" pageId="42" pageNumber="43">P. houkae</emphasis>
|
||
</taxonomicName>
|
||
, C)
|
||
<taxonomicName id="4C3EDAEDFFDABC6EFC2AFAB6FBD6CB15" authorityName="Mullen & Campbell & Sikes" authorityYear="2018" baseAuthorityName="Casey" baseAuthorityYear="1885" box="[1008,1131,1296,1315]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="longipennis">
|
||
<emphasis id="B94A7D7CFFDABC6EFC2AFAB6FBD6CB15" box="[1008,1131,1296,1315]" italics="true" pageId="42" pageNumber="43">P. longipennis</emphasis>
|
||
</taxonomicName>
|
||
, D)
|
||
<taxonomicName id="4C3EDAEDFFDABC6EFB43FAB6FD52CB0D" authorityName="Hatch" authorityYear="1957" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="frosti">
|
||
<emphasis id="B94A7D7CFFDABC6EFB43FAB6FD52CB0D" italics="true" pageId="42" pageNumber="43">P. frosti</emphasis>
|
||
</taxonomicName>
|
||
, E)
|
||
<taxonomicName id="4C3EDAEDFFDABC6EFCCFFA8EFC39CB0D" authorityName="Mullen & Campbell & Sikes" authorityYear="2018" box="[789,900,1320,1339]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="bakerensis">
|
||
<emphasis id="B94A7D7CFFDABC6EFCCFFA8EFC39CB0D" box="[789,900,1320,1339]" italics="true" pageId="42" pageNumber="43">P. bakerensis</emphasis>
|
||
</taxonomicName>
|
||
, F)
|
||
<taxonomicName id="4C3EDAEDFFDABC6EFC70FA8EFBA8CB0D" authorityName="Mullen and Campbell" authorityYear="2018" box="[938,1045,1320,1339]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="olympicus">
|
||
<emphasis id="B94A7D7CFFDABC6EFC70FA8EFBA8CB0D" box="[938,1045,1320,1339]" italics="true" pageId="42" pageNumber="43">P. olympicus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF41F1E6FFDBBC6FFF75FAA8FCF0CB67" ID-DOI="http://doi.org/10.5281/zenodo.5102903" ID-Zenodo-Dep="5102903" httpUri="https://zenodo.org/record/5102903/files/figure.png" pageId="43" pageNumber="44" startId="43.[175,206,1294,1313]" targetBox="[156,1167,196,1276]" targetPageId="43">
|
||
<paragraph id="8B81A16EFFDBBC6FFF75FAA8FCF0CB67" blockId="43.[151,1173,1294,1361]" pageId="43" pageNumber="44">
|
||
<emphasis id="B94A7D7CFFDBBC6FFF75FAA8FF45CB17" bold="true" box="[175,248,1294,1313]" pageId="43" pageNumber="44">Fig. 37.</emphasis>
|
||
Sensory structures on antenna of A)
|
||
<taxonomicName id="4C3EDAEDFFDBBC6FFDB2FAA8FD4BCB17" authorityName="LeConte" authorityYear="1863" box="[616,758,1294,1313]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="pallipes">
|
||
<emphasis id="B94A7D7CFFDBBC6FFDB2FAA8FD4BCB17" box="[616,758,1294,1313]" italics="true" pageId="43" pageNumber="44">Lesteva pallipes</emphasis>
|
||
</taxonomicName>
|
||
, B–C)
|
||
<taxonomicName id="4C3EDAEDFFDBBC6FFC9BFAA8FC59CB17" authorityName="Mannerheim" authorityYear="1830" box="[833,996,1294,1313]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="pubescens">
|
||
<emphasis id="B94A7D7CFFDBBC6FFC9BFAA8FC59CB17" box="[833,996,1294,1313]" italics="true" pageId="43" pageNumber="44">Lesteva pubescens</emphasis>
|
||
</taxonomicName>
|
||
, D–E)
|
||
<taxonomicName id="4C3EDAEDFFDBBC6FFBF4FAA8FB28CB17" box="[1070,1173,1294,1313]" class="Insecta" family="Staphylinidae" genus="Unamis" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B94A7D7CFFDBBC6FFBF4FAA8FBCDCB17" box="[1070,1136,1294,1313]" italics="true" pageId="43" pageNumber="44">Unamis</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
undescribed, F–G)
|
||
<taxonomicName id="4C3EDAEDFFDBBC6FFE9AFA80FE4BCB0F" authorityName="Hatch" authorityYear="1957" box="[320,502,1318,1337]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="houkae">
|
||
<emphasis id="B94A7D7CFFDBBC6FFE9AFA80FE4BCB0F" box="[320,502,1318,1337]" italics="true" pageId="43" pageNumber="44">Phlaeopterus houkae</emphasis>
|
||
</taxonomicName>
|
||
, H–I)
|
||
<taxonomicName id="4C3EDAEDFFDBBC6FFDEEFA80FD46CB0F" authorityName="Mullen and Campbell" authorityYear="2018" box="[564,763,1318,1337]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="obsoletus">
|
||
<emphasis id="B94A7D7CFFDBBC6FFDEEFA80FD46CB0F" box="[564,763,1318,1337]" italics="true" pageId="43" pageNumber="44">Phlaeopterus obsoletus</emphasis>
|
||
</taxonomicName>
|
||
, J–K)
|
||
<taxonomicName id="4C3EDAEDFFDBBC6FFCE1FA80FBBBCB0F" authorityName="Mullen and Campbell" authorityYear="2018" box="[827,1030,1318,1337]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="elongatus">
|
||
<emphasis id="B94A7D7CFFDBBC6FFCE1FA80FBBBCB0F" box="[827,1030,1318,1337]" italics="true" pageId="43" pageNumber="44">Phlaeopterus elongatus</emphasis>
|
||
</taxonomicName>
|
||
. In B, D, F, H, and J, white boxes indicate inset area that is magnified in the image to the right.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C324F2E5FFDBBC6FFF75F9A8FCD3CBDB" pageId="43" pageNumber="44" type="materials_examined">
|
||
<paragraph id="8B81A16EFFDBBC6FFF75F9A8FCD3CBDB" blockId="43.[151,639,1417,1730]" lastBlockId="43.[685,1172,1417,1730]" pageId="43" pageNumber="44">
|
||
<emphasis id="B94A7D7CFFDBBC6FFF75F9A8FEDDC815" bold="true" box="[175,352,1550,1571]" pageId="43" pageNumber="44">Type Specimens.</emphasis>
|
||
<materialsCitation id="3B56AB33FFDBBC6FFEB4F9A8FE5CC8BB" collectingDate="1975-08-08" collectorName="Mt. Rainer N. P. & W. Sunrise & J. M. & B. A. Campbell" location="Mt. Rainer" municipality="Berkeley Park" pageId="43" pageNumber="44" specimenCount="2" typeStatus="holotype">
|
||
<typeStatus id="54851FCCFFDBBC6FFEB4F9A8FE7BC815" box="[366,454,1550,1571]" pageId="43" pageNumber="44" type="holotype">Holotype</typeStatus>
|
||
male (UAMObs: Ento:235217) and
|
||
<typeStatus id="54851FCCFFDBBC6FFEBAF98EFE12C80B" box="[352,431,1576,1597]" pageId="43" pageNumber="44" type="allotype">allotype</typeStatus>
|
||
female (UAMObs: Ento:235837) labeled as follows: WASH,
|
||
<collectorName id="26CBC4B8FFDBBC6FFD81F9E5FEBBC844" pageId="43" pageNumber="44">
|
||
<location id="8EE1F7B5FFDBBC6FFD81F9E5FF65C844" LSID="urn:lsid:plazi:treatment:03971078FFD9BC68FD06FB17FE28C8F4:8EE1F7B5FFDBBC6FFD81F9E5FF65C844" municipality="Berkeley Park" name="Mt. Rainer" pageId="43" pageNumber="44">Mt. Rainer</location>
|
||
N.P.
|
||
</collectorName>
|
||
,
|
||
<collectingMunicipality id="6BE53B14FFDBBC6FFED7F9FBFE32C844" box="[269,399,1629,1650]" pageId="43" pageNumber="44">Berkeley Park</collectingMunicipality>
|
||
,
|
||
<quantity id="4CC60C8BFFDBBC6FFE4CF9FBFE68C844" box="[406,469,1629,1650]" metricMagnitude="3" metricUnit="m" metricValue="5.632704" pageId="43" pageNumber="44" unit="mi" value="3.5">3.5 mi.</quantity>
|
||
<collectorName id="26CBC4B8FFDBBC6FFE02F9FBFD86C844" box="[472,571,1629,1650]" pageId="43" pageNumber="44">W.Sunrise</collectorName>
|
||
,6700’,
|
||
<date id="FF8087AEFFDBBC6FFF4DF9DEFEB8C8BB" box="[151,261,1656,1677]" pageId="43" pageNumber="44" value="1975-08-08">
|
||
<collectingDate id="EFC47E46FFDBBC6FFF4DF9DEFEB8C8BB" box="[151,261,1656,1677]" pageId="43" pageNumber="44" value="1975-08-08">8.VIII.1975</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26CBC4B8FFDBBC6FFED5F9DEFE5CC8BB" box="[271,481,1656,1677]" pageId="43" pageNumber="44">J.M. & B.A. Campbell</collectorName>
|
||
</materialsCitation>
|
||
/
|
||
<typeStatus id="54851FCCFFDBBC6FFE34F9DEFDD5C8BB" box="[494,616,1656,1677]" pageId="43" pageNumber="44" type="holotype">HOLOTYPE</typeStatus>
|
||
³ (or
|
||
<typeStatus id="54851FCCFFDBBC6FFF61F934FE8AC891" box="[187,311,1682,1703]" pageId="43" pageNumber="44" type="allotype">ALLOTYPE</typeStatus>
|
||
<specimenCount id="9D386AE7FFDBBC6FFEE5F933FEF1C891" box="[319,332,1685,1703]" pageId="43" pageNumber="44" type="female">♀</specimenCount>
|
||
)
|
||
<taxonomicName id="4C3EDAEDFFDBBC6FFE86F934FD80C891" authorityName="Mullen and Campbell" authorityYear="2018" box="[348,573,1682,1703]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="obsoletus">Phlaeopterus obsoletus</taxonomicName>
|
||
desig. L.J. Mullen and J.M. Campbell, CNC No. 18470 (red label). Both specimens are deposited in the CNC.
|
||
<typeStatus id="54851FCCFFDBBC6FFD2AFA05FCF3CB8E" box="[752,846,1443,1464]" pageId="43" pageNumber="44" type="paratype">Paratypes</typeStatus>
|
||
(
|
||
<emphasis id="B94A7D7CFFDBBC6FFCBAFA05FCD1CB8E" box="[864,876,1443,1464]" italics="true" pageId="43" pageNumber="44">n</emphasis>
|
||
= 878) are deposited in the AMNH, BMNH, CAS, CNC, CSCA, FMNH, MCZ, OSAC, and USNM.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C324F2E5FFDBBC6FFD1FFA55FCDCC8F4" pageId="43" pageNumber="44" type="distribution">
|
||
<paragraph id="8B81A16EFFDBBC6FFD1FFA55FB80C86E" blockId="43.[685,1172,1417,1730]" pageId="43" pageNumber="44">
|
||
<emphasis id="B94A7D7CFFDBBC6FFD1FFA55FCFBC83E" bold="true" box="[709,838,1523,1544]" pageId="43" pageNumber="44">Distribution.</emphasis>
|
||
<taxonomicName id="4C3EDAEDFFDBBC6FFC8AFA55FB8CC83E" authorityName="Mullen and Campbell" authorityYear="2018" box="[848,1073,1523,1544]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="obsoletus">
|
||
<emphasis id="B94A7D7CFFDBBC6FFC8AFA55FB8CC83E" box="[848,1073,1523,1544]" italics="true" pageId="43" pageNumber="44">Phlaeopterus obsoletus</emphasis>
|
||
</taxonomicName>
|
||
is known from the Cascade Range in
|
||
<collectingRegion id="49FA6F8CFFDBBC6FFC3BF9A8FB2DC815" box="[993,1168,1550,1571]" country="Canada" name="British Columbia" pageId="43" pageNumber="44">British Columbia</collectingRegion>
|
||
,
|
||
<collectingRegion id="49FA6F8CFFDBBC6FFD77F98EFD4AC80B" box="[685,759,1576,1597]" country="United States of America" name="Oregon" pageId="43" pageNumber="44">Oregon</collectingRegion>
|
||
,
|
||
<collectingRegion id="49FA6F8CFFDBBC6FFCD3F98EFCC0C80B" box="[777,893,1576,1597]" country="United States of America" name="Washington" pageId="43" pageNumber="44">Washington</collectingRegion>
|
||
, and
|
||
<collectingRegion id="49FA6F8CFFDBBC6FFC1AF98EFB9FC80B" box="[960,1058,1576,1597]" country="United States of America" name="California" pageId="43" pageNumber="44">California</collectingRegion>
|
||
, and from Jasper National Park,
|
||
<collectingRegion id="49FA6F8CFFDBBC6FFC58F9E5FC74C86E" box="[898,969,1603,1624]" country="Canada" name="Alberta" pageId="43" pageNumber="44">Alberta</collectingRegion>
|
||
(
|
||
<figureCitation id="1305BDEBFFDBBC6FFC0DF9E5FB8CC86E" box="[983,1073,1603,1624]" captionStart="Fig" captionStartId="19.[175,206,1027,1046]" captionText="Fig. 13. Distributions of Phlaeopterus species based on all georeferenced specimen records. A) P. obsoletus, B) P. occidentalis. Grey dots represent localities where at least one Phlaeopterus species, but not the species of interest, was collected. Red dots represent localities where the species of interest was collected." pageId="43" pageNumber="44">Fig. 13A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B81A16EFFDBBC6FFD1FF9FBFCDCC8F4" blockId="43.[685,1172,1417,1730]" pageId="43" pageNumber="44">
|
||
<emphasis id="B94A7D7CFFDBBC6FFD1FF9FBFC89C844" bold="true" box="[709,820,1629,1650]" pageId="43" pageNumber="44">Bionomics.</emphasis>
|
||
Adults have been collected at
|
||
<quantity id="4CC60C8BFFDBBC6FFD77F9DEFCFFC8BB" box="[685,834,1656,1677]" metricMagnitude="3" metricUnit="m" metricValue="1.565" metricValueMax="2.13" metricValueMin="1.0" pageId="43" pageNumber="44" unit="m" value="1565.0" valueMax="2130.0" valueMin="1000.0">1,000–2,130 m</quantity>
|
||
elevation during May–August in wet moss at the edges of streams or in the splash zone of waterfalls.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF41F1E6FFDCBC68FD01FB1BFC8DCAC9" ID-DOI="http://doi.org/10.5281/zenodo.5102905" ID-Zenodo-Dep="5102905" httpUri="https://zenodo.org/record/5102905/files/figure.png" pageId="44" pageNumber="45" startId="44.[731,762,1213,1232]" targetBox="[711,1189,196,1195]" targetPageId="44">
|
||
<paragraph id="8B81A16EFFDCBC68FD01FB1BFC8DCAC9" blockId="44.[707,1193,1213,1279]" pageId="44" pageNumber="45">
|
||
<emphasis id="B94A7D7CFFDCBC68FD01FB1BFC9DCAE6" bold="true" box="[731,800,1213,1232]" pageId="44" pageNumber="45">Fig. 39.</emphasis>
|
||
<taxonomicName id="4C3EDAEDFFDCBC68FCEFFB1BFC50CAE6" box="[821,1005,1213,1232]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B94A7D7CFFDCBC68FCEFFB1BFC18CAE6" box="[821,933,1213,1232]" italics="true" pageId="44" pageNumber="45">Phlaeopterus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, maxillae, dorsal and ventral views. A–B)
|
||
<taxonomicName id="4C3EDAEDFFDCBC68FCADFB73FC76CADE" authorityName="Hatch" authorityYear="1957" box="[887,971,1237,1256]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="houkae">
|
||
<emphasis id="B94A7D7CFFDCBC68FCADFB73FC76CADE" box="[887,971,1237,1256]" italics="true" pageId="44" pageNumber="45">P. houkae</emphasis>
|
||
</taxonomicName>
|
||
, C–D)
|
||
<taxonomicName id="4C3EDAEDFFDCBC68FBD7FB73FBCECADE" authorityName="Mullen and Campbell" authorityYear="2018" box="[1037,1139,1237,1256]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="obsoletus">
|
||
<emphasis id="B94A7D7CFFDCBC68FBD7FB73FBCECADE" box="[1037,1139,1237,1256]" italics="true" pageId="44" pageNumber="45">P. obsoletus</emphasis>
|
||
</taxonomicName>
|
||
, E–F)
|
||
<taxonomicName id="4C3EDAEDFFDCBC68FD19FB4AFC96CAC9" authorityName="Mullen and Campbell" authorityYear="2018" box="[707,811,1260,1279]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="elongatus">
|
||
<emphasis id="B94A7D7CFFDCBC68FD19FB4AFC96CAC9" box="[707,811,1260,1279]" italics="true" pageId="44" pageNumber="45">P. elongatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF41F1E6FFDCBC68FF1FFAB9FE13CB54" ID-DOI="http://doi.org/10.5281/zenodo.5102907" ID-Zenodo-Dep="5102907" httpUri="https://zenodo.org/record/5102907/files/figure.png" pageId="44" pageNumber="45" startId="44.[197,228,1311,1330]" targetBox="[177,655,196,1293]" targetPageId="44">
|
||
<paragraph id="8B81A16EFFDCBC68FF1FFAB9FE13CB54" blockId="44.[173,660,1311,1378]" pageId="44" pageNumber="45">
|
||
<emphasis id="B94A7D7CFFDCBC68FF1FFAB9FEADCB04" bold="true" box="[197,272,1311,1330]" pageId="44" pageNumber="45">Fig. 38.</emphasis>
|
||
Maxillae, dorsal and ventral views, of A–B)
|
||
<taxonomicName id="4C3EDAEDFFDCBC68FF38FA91FE33CB7C" baseAuthorityName="Goeze" baseAuthorityYear="1777" box="[226,398,1335,1354]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="longoelytrata">
|
||
<emphasis id="B94A7D7CFFDCBC68FF38FA91FE33CB7C" box="[226,398,1335,1354]" italics="true" pageId="44" pageNumber="45">Lesteva longoelytrata</emphasis>
|
||
</taxonomicName>
|
||
, C–D)
|
||
<taxonomicName id="4C3EDAEDFFDCBC68FE17FA91FD97CB7C" box="[461,554,1335,1354]" class="Insecta" family="Staphylinidae" genus="Unamis" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B94A7D7CFFDCBC68FE17FA91FDB6CB7C" box="[461,523,1335,1354]" italics="true" pageId="44" pageNumber="45">Unamis</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
undescribed, E–F)
|
||
<taxonomicName id="4C3EDAEDFFDCBC68FF06FAE9FE14CB54" authorityName="Hatch" authorityYear="1957" box="[220,425,1359,1378]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="lagrandeuri">
|
||
<emphasis id="B94A7D7CFFDCBC68FF06FAE9FE14CB54" box="[220,425,1359,1378]" italics="true" pageId="44" pageNumber="45">Phlaeopterus lagrandeuri</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C324F2E5FFDCBC68FF1FFA2FFF4EC844" pageId="44" pageNumber="45" type="discussion">
|
||
<paragraph id="8B81A16EFFDCBC68FF1FFA2FFF4EC844" blockId="44.[173,660,1417,1730]" pageId="44" pageNumber="45">
|
||
<emphasis id="B94A7D7CFFDCBC68FF1FFA2FFE9BCBA8" bold="true" box="[197,294,1417,1438]" pageId="44" pageNumber="45">Remarks.</emphasis>
|
||
<taxonomicName id="4C3EDAEDFFDCBC68FEE8FA2FFDABCBA8" authorityName="Mullen and Campbell" authorityYear="2018" box="[306,534,1417,1438]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="obsoletus">
|
||
<emphasis id="B94A7D7CFFDCBC68FEE8FA2FFDABCBA8" box="[306,534,1417,1438]" italics="true" pageId="44" pageNumber="45">Phlaeopterus obsoletus</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from all other
|
||
<taxonomicName id="4C3EDAEDFFDCBC68FE1BFA05FD2ECB8E" box="[449,659,1443,1464]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B94A7D7CFFDCBC68FE1BFA05FD82CB8E" box="[449,575,1443,1464]" italics="true" pageId="44" pageNumber="45">Phlaeopterus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
except
|
||
<taxonomicName id="4C3EDAEDFFDCBC68FF2DFA18FE38CBE5" authorityName="Mullen & Campbell & Sikes" authorityYear="2018" baseAuthorityName="Casey" baseAuthorityYear="1885" box="[247,389,1470,1491]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="longipennis">
|
||
<emphasis id="B94A7D7CFFDCBC68FF2DFA18FE38CBE5" box="[247,389,1470,1491]" italics="true" pageId="44" pageNumber="45">P. longipennis</emphasis>
|
||
</taxonomicName>
|
||
by the absence of ocelli.
|
||
<taxonomicName id="4C3EDAEDFFDCBC68FF77FA7EFE36CBDB" authorityName="Mullen and Campbell" authorityYear="2018" box="[173,395,1496,1517]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="obsoletus">
|
||
<emphasis id="B94A7D7CFFDCBC68FF77FA7EFE36CBDB" box="[173,395,1496,1517]" italics="true" pageId="44" pageNumber="45">Phlaeopterus obsoletus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3EDAEDFFDCBC68FE60FA7EFDFDCBDB" authorityName="Mullen & Campbell & Sikes" authorityYear="2018" baseAuthorityName="Casey" baseAuthorityYear="1885" box="[442,576,1496,1517]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="longipennis">
|
||
<emphasis id="B94A7D7CFFDCBC68FE60FA7EFDFDCBDB" box="[442,576,1496,1517]" italics="true" pageId="44" pageNumber="45">P. longipennis</emphasis>
|
||
</taxonomicName>
|
||
are very similar but can be differentiated by characters discussed in the Remarks section of the latter species. This species has been misidentified as
|
||
<taxonomicName id="4C3EDAEDFFDCBC68FD99F98EFF4CC86E" authorityName="Mullen & Campbell & Sikes" authorityYear="2018" baseAuthorityName="Casey" baseAuthorityYear="1885" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="longipennis">
|
||
<emphasis id="B94A7D7CFFDCBC68FD99F98EFF4CC86E" italics="true" pageId="44" pageNumber="45">P. longipennis</emphasis>
|
||
</taxonomicName>
|
||
in most collections as well as by
|
||
<bibRefCitation id="EFAFDC9FFFDCBC68FD80F9E5FF52C844" author="Hatch" pageId="44" pageNumber="45" refId="ref28862" refString="Hatch, M. H. 1957. The Beetles of the Pacific Northwest. Part II: Staphyliniformia. University of Washington Publications in Biology, Seattle, WA." type="book" year="1957">Hatch (1957)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C324F2E5FFDCBC68FF1FF9DEFE28C8F4" pageId="44" pageNumber="45" type="etymology">
|
||
<paragraph id="8B81A16EFFDCBC68FF1FF9DEFE28C8F4" blockId="44.[173,660,1417,1730]" pageId="44" pageNumber="45">
|
||
<emphasis id="B94A7D7CFFDCBC68FF1FF9DEFE8AC8BB" bold="true" box="[197,311,1656,1677]" pageId="44" pageNumber="45">Etymology.</emphasis>
|
||
This species name is derived from the Latin adjective for effaced or indistinct, referring to the absence of ocelli.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |