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<document id="04CD5E54697524210DE3CD2E6EEBBA94" ID-CLB-Dataset="3175" ID-DOI="10.11646/zootaxa.5072.3.1" ID-GBIF-Dataset="a5454a8b-04cd-4cc6-ae0b-f13f31a7af34" ID-ISSN="1175-5326" ID-Zenodo-Dep="5744549" ID-ZooBank="CAB2C8C4-E0A4-4805-9DEB-94769C2F90EB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638283643950" checkinUser="plazi" docAuthor="Mahecha-J, Oscar, Florczyk, Klaudia, Willmott, Keith, Cerdeña, José, Zubek, Anna, Boyer, Pierre, Farfán, Jackie, Lachowska-Cierlik, Dorota &amp; Pyrcz, Tomasz W." docDate="2021" docId="864387EE113F7245F8A10DC884A22041" docLanguage="en" docName="zootaxa.5072.3.1.pdf" docOrigin="Zootaxa 5072 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Manerebia granatus Willmott, Radford &amp; Pyrcz 2021, n. sp." docType="treatment" docVersion="8" lastPageNumber="223" masterDocId="7A7AFF9611307253F836080285142354" masterDocTitle="Solving the cryptic diversity of the genus Manerebia Staudinger in northern Peru description of new species and considerations on the biogeographical role of the Huancabamba Deflection (Nymphalidae: Satyrinae: Pronophilina)" masterLastPageNumber="237" masterPageNumber="201" pageNumber="216" updateTime="1699471567071" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4E2C2D536D26C0C6DB0B49950320D702">Solving the cryptic diversity of the genus Manerebia Staudinger in northern Peru description of new species and considerations on the biogeographical role of the Huancabamba Deflection (Nymphalidae: Satyrinae: Pronophilina)</mods:title>
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<mods:namePart id="D864A382A90C7B19D363A4173008E4FA">Florczyk, Klaudia</mods:namePart>
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<mods:affiliation id="ED963F4BA8B0242E69C97E6020FBB83E">Universidad Nacional de San Agustín de Arequipa, Museo de Historia Natural, Av. Alcides Carrión s / n, Escuela de Biología UNSA, Arequipa, Perú. &amp; jjackie 4 u @ gmail. com; https: // orcid. org / 0000 - 0001 - 5913 - 0756</mods:affiliation>
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<mods:affiliation id="DE7FE46893A5E50795F2509525D9042A">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland &amp; dorota. lachowska-cierlik @ uj. edu. pl; https: // orcid. org / 0000 - 0001 - 9072 - 1677</mods:affiliation>
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<mods:namePart id="EDF8B677A92E3B356CFCF422B09C390C">Pyrcz, Tomasz W.</mods:namePart>
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<mods:affiliation id="80EF5925A1C13DF3B06BB2B380A53336">Nature Education Centre, Jagiellonian University, Gronostajowa 5, 30 - 387 Kraków, Poland. &amp; Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland &amp; tomasz. pyrcz @ uj. edu. pl; https: // orcid. org / 0000 - 0003 - 4822 - 0670</mods:affiliation>
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<taxonomicName id="C9EA4D7B113F725CF8A10DC887FA26B1" authority="Willmott, Radford &amp; Pyrcz" authorityName="Willmott, Radford &amp; Pyrcz" authorityYear="2021" box="[151,750,1482,1509]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="granatus" status="sp. nov.">
<emphasis id="3C9EEAEA113F725CF8A10DC8849B26B1" bold="true" box="[151,399,1482,1509]" italics="true" pageId="15" pageNumber="216">Manerebia granatus</emphasis>
Willmott, Radford &amp; Pyrcz
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(
<figureCitation id="96D12A7D113F725CF8A90DED85FB255D" box="[159,239,1519,1545]" captionStart="FIGURE 4" captionStartId="10.[151,250,1864,1888]" captionTargetBox="[245,1350,210,1839]" captionTargetId="figure-23@10.[223,1364,181,1840]" captionTargetPageId="10" captionText="FIGURE 4. Adults (left—dorsum, right—venter) A. M. granatus n. sp. ♂ Holotype, Ecuador, Paquisha Alto B. M. granatus n. sp. ♂ Paratype, Peru, Alto Río Nieva C. M. granatus n. sp.♀, Ecuador, San Andrés D. M. placida n. sp. ♂ Holotype, Ecuador, Zamora - Arcoiris" figureDoi="http://doi.org/10.5281/zenodo.5744557" httpUri="https://zenodo.org/record/5744557/files/figure.png" pageId="15" pageNumber="216">Figs. 4</figureCitation>
,
<figureCitation id="96D12A7D113F725CF8CD0DED841E255E" box="[251,266,1519,1546]" captionStart="FIGURE 6" captionStartId="17.[151,250,1847,1871]" captionTargetBox="[237,1370,197,1811]" captionTargetId="figure-21@17.[201,1386,181,1823]" captionTargetPageId="17" captionText="FIGURE 6. Male genitalia (top—lateral view, middle—plan view, bottom—aedeagus extracted, lateral view) A. M. granatus n. sp. Peru, Rodríguez de Mendoza, prep. genit. KF-2736 B. M. placida n. sp. Ecuador, Zamora, prep. genit. KW-20-021 C. M. benigni tessmanni, Peru, Alto Río Nieva, prep. genit. KF-H_241 D. M. benigni tessmanni, Ecuador, Río Troya, prep. genit. KF-6282" figureDoi="http://doi.org/10.5281/zenodo.5744563" httpUri="https://zenodo.org/record/5744563/files/figure.png" pageId="15" pageNumber="216">6</figureCitation>
)
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locality.
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,
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,
<collectingMunicipality id="EE31AC82113F725CFB250E35811F2505" box="[787,1035,1591,1617]" pageId="15" pageNumber="216">Cordillera del Cóndor</collectingMunicipality>
,
<location id="0B356023113F725CFC200E35804C2505" LSID="urn:lsid:plazi:treatment:864387EE113F7245F8A10DC884A22041:0B356023113F725CFC200E35804C2505" box="[1046,1368,1591,1617]" country="Ecuador" county="Department" municipality="Cordillera del Condor" name="Destacamento Paquisha Alto" pageId="15" pageNumber="216" stateProvince="Zamora-Chinchipe">Destacamento Paquisha Alto</location>
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<emphasis id="3C9EEAEA113F725CF8F10E5884562520" bold="true" box="[199,322,1626,1652]" pageId="15" pageNumber="216">Diagnosis.</emphasis>
This species superficially resembles a number of congeners, including its sister-species
<taxonomicName id="C9EA4D7B113F725CFD150E5980882521" authority="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz, 2021" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[1315,1436,1627,1653]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="placida" status="sp. nov.">
<emphasis id="3C9EEAEA113F725CFD150E59802E2520" box="[1315,1338,1627,1652]" italics="true" pageId="15" pageNumber="216">M</emphasis>
.
<emphasis id="3C9EEAEA113F725CFD7E0E5980882521" box="[1352,1436,1627,1653]" italics="true" pageId="15" pageNumber="216">placida</emphasis>
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<taxonomicNameLabel id="27AD5791113F725CF8A10E7D85C225CD" box="[151,214,1663,1689]" pageId="15" pageNumber="216" rank="species">n. sp.</taxonomicNameLabel>
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, with which it may be broadly sympatric. It differs from
<taxonomicName id="C9EA4D7B113F725CFB6F0E7D86C625CD" authority="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz, 2021" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[857,978,1663,1689]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="placida" status="sp. nov.">
<emphasis id="3C9EEAEA113F725CFB6F0E7D866425CC" box="[857,880,1663,1688]" italics="true" pageId="15" pageNumber="216">M</emphasis>
.
<emphasis id="3C9EEAEA113F725CFB480E7D86C625CD" box="[894,978,1663,1689]" italics="true" pageId="15" pageNumber="216">placida</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA113F725CFBEC0E7D810D25CD" bold="true" box="[986,1049,1663,1689]" pageId="15" pageNumber="216">
<taxonomicNameLabel id="27AD5791113F725CFBEC0E7D810D25CD" box="[986,1049,1663,1689]" pageId="15" pageNumber="216" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
as follows (characters in the latter species in parentheses): the VHW is uniform reddish brown in the distal half, including distally of the submarginal ocelli and white dots (rather than just basally of these markings); the VHW ocellus in cell CuA2-CuA1 is very small, with no distinct ocellar ring (larger with distinct yellowish orange ocellar ring); the VHW ocellus in M2-M1 is not developed (developed, with black centre and ocellar ring); the VHW submarginal red-brown line is more undulate (less undulate); the VHW has virtually no trace of a pale postdiscal line (variable but distinct line present). The species is also similar to other
<taxonomicName id="C9EA4D7B113F725CF9CF0F5587672425" box="[505,627,1879,1905]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="genus">
<emphasis id="3C9EEAEA113F725CF9CF0F5587672425" box="[505,627,1879,1905]" italics="true" pageId="15" pageNumber="216">Manerebia</emphasis>
</taxonomicName>
that lack a VHW pale postdiscal line (e.g.,
<taxonomicName id="C9EA4D7B113F725CFC5E0F55845724C1" authority="Pyrcz &amp; Willmott, 2006" authorityName="Pyrcz &amp; Willmott" authorityYear="2006" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="lamasi" status="stat. nov.">
<emphasis id="3C9EEAEA113F725CFC5E0F5580222425" box="[1128,1334,1879,1905]" italics="true" pageId="15" pageNumber="216">Manerebia lamasi</emphasis>
<bibRefCitation id="6A7B4B09113F725CFD080F55845724C1" author="Pyrcz, T. W. &amp; Willmott, K. R. &amp; Hall, J. P. W. &amp; Viloria, A. L." pageId="15" pageNumber="216" pagination="37 - 79" refId="ref22357" refString="Pyrcz, T. W., Willmott, K. R., Hall, J. P. W. &amp; Viloria, A. L. (2006) A review of the genus Manerebia Staudinger (Lepidoptera: Nymphalidae: Satyrinae) in the northern Andes. Journal of Research on the Lepidoptera, 39, 37 - 79." type="journal article" year="2006">Pyrcz &amp; Willmott, 2006</bibRefCitation>
</taxonomicName>
<emphasis id="3C9EEAEA113F725CF97D0F79848924C1" bold="true" box="[331,413,1915,1941]" pageId="15" pageNumber="216">
<taxonomicNameLabel id="27AD5791113F725CF97D0F79848924C1" box="[331,413,1915,1941]" pageId="15" pageNumber="216">n. stat.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="C9EA4D7B113F725CF99A0F7987B624C0" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" baseAuthorityName="Pyrcz &amp; Willmott" baseAuthorityYear="2006" box="[428,674,1915,1941]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="pauperata" status="stat. nov.">
<emphasis id="3C9EEAEA113F725CF99A0F7987B624C0" box="[428,674,1915,1941]" italics="true" pageId="15" pageNumber="216">Manerebia pauperata</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA113F725CFA9C0F7987E824C1" bold="true" box="[682,764,1915,1941]" pageId="15" pageNumber="216">
<taxonomicNameLabel id="27AD5791113F725CFA9C0F7987E824C1" box="[682,764,1915,1941]" pageId="15" pageNumber="216">n. stat.</taxonomicNameLabel>
</emphasis>
), but may be distinguished from them by the very reduced VHW submarginal ocelli, especially that in cell CuA2-CuA1. Two males each of
<taxonomicName id="C9EA4D7B113F725CFC170F9D81BE24EC" authority="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz, 2021" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[1057,1194,1951,1977]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="granatus" status="sp. nov.">
<emphasis id="3C9EEAEA113F725CFC170F9D812C24EC" box="[1057,1080,1951,1976]" italics="true" pageId="15" pageNumber="216">M</emphasis>
.
<emphasis id="3C9EEAEA113F725CFC700F9D81BE24EC" box="[1094,1194,1951,1976]" italics="true" pageId="15" pageNumber="216">granatus</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA113F725CFC860F9D81FA24ED" bold="true" box="[1200,1262,1951,1977]" pageId="15" pageNumber="216">
<taxonomicNameLabel id="27AD5791113F725CFC860F9D81FA24ED" box="[1200,1262,1951,1977]" pageId="15" pageNumber="216" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="C9EA4D7B113F725CFD150F9D808824ED" authority="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz, 2021" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[1315,1436,1951,1977]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="placida" status="sp. nov.">
<emphasis id="3C9EEAEA113F725CFD150F9D802E24EC" box="[1315,1338,1951,1976]" italics="true" pageId="15" pageNumber="216">M</emphasis>
.
<emphasis id="3C9EEAEA113F725CFD7E0F9D808824ED" box="[1352,1436,1951,1977]" italics="true" pageId="15" pageNumber="216">placida</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA113F725CF8A10FC185CD2489" bold="true" box="[151,217,1987,2013]" pageId="15" pageNumber="216">
<taxonomicNameLabel id="27AD5791113F725CF8A10FC185CD2489" box="[151,217,1987,2013]" pageId="15" pageNumber="216" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
were dissected and they showed consistent differences between the two species, in particular the arms of gnathos parallel in dorsal view in
<taxonomicName id="C9EA4D7B11207243FA2B089587BC23E4" authority="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz, 2021" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[541,680,151,177]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="granatus" status="sp. nov.">
<emphasis id="3C9EEAEA11207243FA2B0895872023E4" box="[541,564,151,176]" italics="true" pageId="16" pageNumber="217">M</emphasis>
.
<emphasis id="3C9EEAEA11207243FA72089587BC23E4" box="[580,680,151,176]" italics="true" pageId="16" pageNumber="217">granatus</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA11207243FA86089587E523E5" bold="true" box="[688,753,151,177]" pageId="16" pageNumber="217">
<taxonomicNameLabel id="27AD579111207243FA86089587E523E5" box="[688,753,151,177]" pageId="16" pageNumber="217" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
but directed inwards in
<taxonomicName id="C9EA4D7B11207243FC3F0895819023E5" authority="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz, 2021" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[1033,1156,151,177]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="placida" status="sp. nov.">
<emphasis id="3C9EEAEA11207243FC3F0895813423E4" box="[1033,1056,151,176]" italics="true" pageId="16" pageNumber="217">M</emphasis>
.
<emphasis id="3C9EEAEA11207243FC060895819023E5" box="[1072,1156,151,177]" italics="true" pageId="16" pageNumber="217">placida</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA11207243FCBB089581D923E5" bold="true" box="[1165,1229,151,177]" pageId="16" pageNumber="217">
<taxonomicNameLabel id="27AD579111207243FCBB089581D923E5" box="[1165,1229,151,177]" pageId="16" pageNumber="217" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, and the spines at distal tip of valvae extending further basally along the inner edge in
<taxonomicName id="C9EA4D7B11207243FBBD08B981002380" authority="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz, 2021" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[907,1044,187,213]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="granatus" status="sp. nov.">
<emphasis id="3C9EEAEA11207243FBBD08B986B62380" box="[907,930,187,212]" italics="true" pageId="16" pageNumber="217">M</emphasis>
.
<emphasis id="3C9EEAEA11207243FB8608B981002380" box="[944,1044,187,212]" italics="true" pageId="16" pageNumber="217">granatus</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA11207243FC2C08B981432381" bold="true" box="[1050,1111,187,213]" pageId="16" pageNumber="217">
<taxonomicNameLabel id="27AD579111207243FC2C08B981432381" box="[1050,1111,187,213]" pageId="16" pageNumber="217" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, but confined to the distal tip in
<taxonomicName id="C9EA4D7B11207243F88208DD843923AD" authority="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz, 2021" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[180,301,223,249]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="placida" status="sp. nov.">
<emphasis id="3C9EEAEA11207243F88208DD85DF23AC" box="[180,203,223,248]" italics="true" pageId="16" pageNumber="217">M</emphasis>
.
<emphasis id="3C9EEAEA11207243F8EF08DD843923AD" box="[217,301,223,249]" italics="true" pageId="16" pageNumber="217">placida</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA11207243F90208DD846723AD" bold="true" box="[308,371,223,249]" pageId="16" pageNumber="217">
<taxonomicNameLabel id="27AD579111207243F90208DD846723AD" box="[308,371,223,249]" pageId="16" pageNumber="217" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
The male genitalia of both species may be distinguished from the otherwise similar species
<taxonomicName id="C9EA4D7B11207243FD4808DD846C2249" authority="Pyrcz, 2004" authorityName="Pyrcz" authorityYear="2004" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="benigni">
<emphasis id="3C9EEAEA11207243FD4808DD808123AC" box="[1406,1429,223,248]" italics="true" pageId="16" pageNumber="217">M</emphasis>
.
<emphasis id="3C9EEAEA11207243F8A1090185FF2249" box="[151,235,259,285]" italics="true" pageId="16" pageNumber="217">benigni</emphasis>
<bibRefCitation id="6A7B4B0911207243F8C40901846C2249" author="Pyrcz, T. W." box="[242,376,259,285]" pageId="16" pageNumber="217" pagination="455 - 622" refId="ref22314" refString="Pyrcz, T. W. (2004) Pronophiline butterflies of the highlands of Chachapoyas in northern Peru: faunal survey, diversity and distribution patterns (Lepidoptera, Nymphalidae, Satyrinae). Genus, 15, 455 - 622." type="journal article" year="2004">Pyrcz, 2004</bibRefCitation>
</taxonomicName>
as described below under
<taxonomicName id="C9EA4D7B11207243FA940901860F2249" authority="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz, 2021" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[674,795,259,285]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="placida" status="sp. nov.">
<emphasis id="3C9EEAEA11207243FA94090187AD2248" box="[674,697,259,284]" italics="true" pageId="16" pageNumber="217">M</emphasis>
.
<emphasis id="3C9EEAEA11207243FAF10901860F2249" box="[711,795,259,285]" italics="true" pageId="16" pageNumber="217">placida</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA11207243FB14090186752249" bold="true" box="[802,865,259,285]" pageId="16" pageNumber="217">
<taxonomicNameLabel id="27AD579111207243FB14090186752249" box="[802,865,259,285]" pageId="16" pageNumber="217" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="46F0657311207243F8F1092484CF26B1" pageId="16" pageNumber="217" type="description">
<paragraph id="0E5536F811207243F8F1092481582601" blockId="16.[151,1437,151,2013]" pageId="16" pageNumber="217">
<emphasis id="3C9EEAEA11207243F8F10924844D2214" bold="true" box="[199,345,294,320]" pageId="16" pageNumber="217">Description.</emphasis>
MALE (
<figureCitation id="96D12A7D11207243F988092587542215" box="[446,576,295,321]" captionStart="FIGURE 4" captionStartId="10.[151,250,1864,1888]" captionTargetBox="[245,1350,210,1839]" captionTargetId="figure-23@10.[223,1364,181,1840]" captionTargetPageId="10" captionText="FIGURE 4. Adults (left—dorsum, right—venter) A. M. granatus n. sp. ♂ Holotype, Ecuador, Paquisha Alto B. M. granatus n. sp. ♂ Paratype, Peru, Alto Río Nieva C. M. granatus n. sp.♀, Ecuador, San Andrés D. M. placida n. sp. ♂ Holotype, Ecuador, Zamora - Arcoiris" figureDoi="http://doi.org/10.5281/zenodo.5744557" httpUri="https://zenodo.org/record/5744557/files/figure.png" pageId="16" pageNumber="217">Fig 4. A, B</figureCitation>
):
<emphasis id="3C9EEAEA11207243FA63092587862215" box="[597,658,295,321]" italics="true" pageId="16" pageNumber="217">Head</emphasis>
: eyes chocolate brown, naked, lustrous; labial palpi two times length of head, covered with black hair-like scales, longer ventrally; antennae slender, 2/5
<superScript id="F99F9BB011207243FC7609488159220C" attach="left" box="[1088,1101,330,344]" fontSize="6" pageId="16" pageNumber="217">th</superScript>
length of costa, 38 segments, mostly naked, dorsally brown, ventrally paler brown, club formed gradually of terminal 12 segments.
<emphasis id="3C9EEAEA11207243FCCC096D805F22DD" box="[1274,1355,367,393]" italics="true" pageId="16" pageNumber="217">Thorax</emphasis>
: black, with long black hair-like scales; legs dorsally dark brown, ventrally pale yellowish brown.
<emphasis id="3C9EEAEA11207243FCBE099181D822F8" box="[1160,1228,403,428]" italics="true" pageId="16" pageNumber="217">Wings</emphasis>
: FW length
<quantity id="C9129B1D11207243FD60099185D12285" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.1" metricValueMin="2.0" pageId="16" pageNumber="217" unit="mm" value="20.5" valueMax="21.0" valueMin="20.0">2021 mm</quantity>
, mean: 20.4 mm, n=8; FWD uniform chocolate brown, except for patch of dense, black, elongate rectangular androconial scales in basal half to third of cells 2A-M1 and extending into adjacent posterior half of discal cell. HWD with hair-like scales in median half, uniform chocolate brown, with scattered reddish brown scaling in tornus. FWV dark brown, darker blackish brown in areas with dorsal androconial scales, with a well-marked dark reddish brown, undulating and rather irregular submarginal line, and a narrow reddish brown marginal line; series of five white submarginal dots in cells Cu2-Cu1 to M1-R5, first of these in centre of a small black spot surrounded by a reddish brown ring. HWV dark blackish brown (similar to basal half of FWV) basal to an indistinct, straight, dark brown postdiscal line, distally of this line reddish brown (also extending along tornus) up to a dark reddish brown, undulate submarginal line, then dark brown, with dark reddish brown marginal line; series of seven white submarginal dots in cells 2A-CuA2 (two dots), CuA2-CuA1 to M1-Rs, first three of these in centre of small black spots.
<emphasis id="3C9EEAEA11207243FD1D0AF980812041" box="[1323,1429,763,789]" italics="true" pageId="16" pageNumber="217">Abdomen</emphasis>
: Covered with dense, dark brown scales dorsally and laterally, and slightly paler greyish brown scales ventrally, with long dark hair-like scales increasing in density anteriorly.
<emphasis id="3C9EEAEA11207243FB2A0B4186922009" box="[796,902,835,861]" italics="true" pageId="16" pageNumber="217">Genitalia</emphasis>
(
<figureCitation id="96D12A7D11207243FBA30B4186FB200A" box="[917,1007,835,862]" captionStart="FIGURE 6" captionStartId="17.[151,250,1847,1871]" captionTargetBox="[237,1370,197,1811]" captionTargetId="figure-21@17.[201,1386,181,1823]" captionTargetPageId="17" captionText="FIGURE 6. Male genitalia (top—lateral view, middle—plan view, bottom—aedeagus extracted, lateral view) A. M. granatus n. sp. Peru, Rodríguez de Mendoza, prep. genit. KF-2736 B. M. placida n. sp. Ecuador, Zamora, prep. genit. KW-20-021 C. M. benigni tessmanni, Peru, Alto Río Nieva, prep. genit. KF-H_241 D. M. benigni tessmanni, Ecuador, Río Troya, prep. genit. KF-6282" figureDoi="http://doi.org/10.5281/zenodo.5744563" httpUri="https://zenodo.org/record/5744563/files/figure.png" pageId="16" pageNumber="217">Fig. 6A</figureCitation>
): Uncus slightly curving and 1.5 times longer than tegumen shoulder, gnathos half-length of uncus, slightly curving upwards and pointed; pedunculus with a massive base and apex curved downwards; saccus short, bulbous; valva with a broad basal half ending in squared-off mid-dorsal process, and a narrower apical half with a distal series of 8 or so squat teeth oriented subhorizontally; aedeagus curving evenly upwards, tapering anteriorly, shorter than valva, smooth, no visible cornuti. FEMALE (
<figureCitation id="96D12A7D11207243F92F0BF584612745" box="[281,373,1015,1041]" captionStart="FIGURE 4" captionStartId="10.[151,250,1864,1888]" captionTargetBox="[245,1350,210,1839]" captionTargetId="figure-23@10.[223,1364,181,1840]" captionTargetPageId="10" captionText="FIGURE 4. Adults (left—dorsum, right—venter) A. M. granatus n. sp. ♂ Holotype, Ecuador, Paquisha Alto B. M. granatus n. sp. ♂ Paratype, Peru, Alto Río Nieva C. M. granatus n. sp.♀, Ecuador, San Andrés D. M. placida n. sp. ♂ Holotype, Ecuador, Zamora - Arcoiris" figureDoi="http://doi.org/10.5281/zenodo.5744557" httpUri="https://zenodo.org/record/5744557/files/figure.png" pageId="16" pageNumber="217">Fig. 4C</figureCitation>
): Similar to male except: slightly larger (FW length
<quantity id="C9129B1D11207243FBE40BF5813C2745" box="[978,1064,1015,1041]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="16" pageNumber="217" unit="mm" value="22.0">22 mm</quantity>
); wings slightly paler brown, no FWD androconial patch; reddish brown scales in distal part of HWV scattered, less dense than in male; HWV with narrow, even, straight white postdiscal band; submarginal ocelli larger in cells CuA2-CuA1 of FWV and HWV, and submarginal white spot in HWV cell surrounded by a small black spot.
<emphasis id="3C9EEAEA11207243FBE40C6181282729" box="[978,1084,1123,1149]" italics="true" pageId="16" pageNumber="217">Genitalia</emphasis>
(not illustrated): Anal papillae prominent, covered with setae of varying length, with a strongly sclerotized, amorphous basal plate; membrane below papillae a lightly sclerotized, grooved plate terminating in a point posteriorly, bordered anteriorly by a band of fine spine-like protrusions; postvaginal lamella sclerotized and forming a broad curved plate, produced into two prominent lateral, folded flaps with smooth edges; antevaginal lamella slightly sclerotized, pocket-like and folded; antrum sclerotized; ductus bursae wide and short, opening into a large, oval corpus bursae, with two narrow bands of dorsal signa that converge slightly anteriorly and extend over half of bursa length.
</paragraph>
<paragraph id="0E5536F811207243F8F10D5D84CF26B1" blockId="16.[151,1437,151,2013]" pageId="16" pageNumber="217">
<emphasis id="3C9EEAEA11207243F8F10D5D849C262D" bold="true" box="[199,392,1375,1401]" pageId="16" pageNumber="217">Molecular data.</emphasis>
BI (
<figureCitation id="96D12A7D11207243F9F40D5D870C262D" box="[450,536,1375,1401]" captionStart="FIGURE 12" captionStartId="25.[151,250,1943,1967]" captionTargetBox="[572,1012,184,1914]" captionTargetId="figure-21@25.[572,1014,181,1919]" captionTargetPageId="25" captionText="FIGURE 12. Bayesian Inference tree (BI) reconstructed from a fragment of COI mitochondrial gene. Posterior probability values indicated on nodes." figureDoi="http://doi.org/10.5281/zenodo.5744583" httpUri="https://zenodo.org/record/5744583/files/figure.png" pageId="16" pageNumber="217">Fig. 12</figureCitation>
), ML (
<figureCitation id="96D12A7D11207243FA580D5D87D6262D" box="[622,706,1375,1401]" captionStart="FIGURE 13" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[575,1017,184,1919]" captionTargetId="figure-23@26.[570,1017,181,1919]" captionTargetPageId="26" captionText="FIGURE 13. Maximum likelihood tree (ML) reconstructed from a fragment of COI mitochondrial gene. Ultrafast bootstrap support values indicated on nodes." figureDoi="http://doi.org/10.5281/zenodo.5744585" httpUri="https://zenodo.org/record/5744585/files/figure.png" pageId="16" pageNumber="217">Fig. 13</figureCitation>
) trees, species-delimitation analyses (Figs. 14,15), and genetic distances (Supplementary material 2) indicate that
<taxonomicName id="C9EA4D7B11207243FAD20D81866426C8" authority="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz, 2021" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[740,880,1411,1437]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="granatus" status="sp. nov.">
<emphasis id="3C9EEAEA11207243FAD20D8187EF26C8" box="[740,763,1411,1436]" italics="true" pageId="16" pageNumber="217">M</emphasis>
.
<emphasis id="3C9EEAEA11207243FB3A0D81866426C8" box="[780,880,1411,1436]" italics="true" pageId="16" pageNumber="217">granatus</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA11207243FB4C0D8186A826C9" bold="true" box="[890,956,1411,1437]" pageId="16" pageNumber="217">
<taxonomicNameLabel id="27AD579111207243FB4C0D8186A826C9" box="[890,956,1411,1437]" pageId="16" pageNumber="217" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is a distinct species, and on both trees it clusters with
<taxonomicName id="C9EA4D7B11207243F91D0DA584B02695" authority="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz, 2021" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[299,420,1447,1473]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="placida" status="sp. nov.">
<emphasis id="3C9EEAEA11207243F91D0DA584B02695" box="[299,420,1447,1473]" italics="true" pageId="16" pageNumber="217">M. placida</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA11207243F99C0DA584FC2695" bold="true" box="[426,488,1447,1473]" pageId="16" pageNumber="217">
<taxonomicNameLabel id="27AD579111207243F99C0DA584FC2695" box="[426,488,1447,1473]" pageId="16" pageNumber="217" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and a new, undescribed species from Central
<collectingCountry id="76FD766811207243FBD20DA5810E2695" box="[996,1050,1447,1473]" name="Peru" pageId="16" pageNumber="217">Peru</collectingCountry>
, AZ-650
<taxonomicName id="C9EA4D7B11207243FCB50DA580302695" box="[1155,1316,1447,1473]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="3C9EEAEA11207243FCB50DA581E92695" box="[1155,1277,1447,1473]" italics="true" pageId="16" pageNumber="217">Manerebia</emphasis>
sp.
</taxonomicName>
, the sisterspecies to
<taxonomicName id="C9EA4D7B11207243F93A0DC9848126B0" authority="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz, 2021" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[268,405,1483,1509]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="granatus" status="sp. nov.">
<emphasis id="3C9EEAEA11207243F93A0DC9843726B0" box="[268,291,1483,1508]" italics="true" pageId="16" pageNumber="217">M</emphasis>
.
<emphasis id="3C9EEAEA11207243F9070DC9848126B0" box="[305,405,1483,1508]" italics="true" pageId="16" pageNumber="217">granatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="27AD579111207243F9AA0DC984CF26B1" box="[412,475,1483,1509]" pageId="16" pageNumber="217" rank="species">
<emphasis id="3C9EEAEA11207243F9AA0DC984C726B1" bold="true" box="[412,467,1483,1509]" pageId="16" pageNumber="217">n. sp</emphasis>
.
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="46F0657311207246F8F10DED870125D1" lastPageId="21" lastPageNumber="222" pageId="16" pageNumber="217" type="materials_examined">
<paragraph id="0E5536F811207243F8F10DED87B22489" blockId="16.[151,1437,151,2013]" pageId="16" pageNumber="217">
<emphasis id="3C9EEAEA11207243F8F10DED8461255C" bold="true" box="[199,373,1518,1545]" pageId="16" pageNumber="217">Type material:</emphasis>
<materialsCitation id="BE823CA511207243F94F0DED808D2579" ID-GBIF-Occurrence="3413048325" collectingDate="2010-08-31" collectionCode="FLMNH, INABIO" collectorName="Ecuador &amp; K. Buckland" elevation="2010" latitude="-3.91" location="Zamora-Chinchipe" longLatPrecision="21" longitude="-78.48611" municipality="Destacamento Paquisha Alto" pageId="16" pageNumber="217" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectorName id="A31F532E11207243F94F0DED84CF255D" box="[377,475,1519,1545]" pageId="16" pageNumber="217">Ecuador</collectorName>
:
<typeStatus id="D151885A11207243F9D40DED875E255D" box="[482,586,1519,1545]" pageId="16" pageNumber="217" type="holotype">Holotype</typeStatus>
<specimenCount id="18ECFD7111207243FA790DED8773255D" box="[591,615,1519,1545]" pageId="16" pageNumber="217" type="male"></specimenCount>
:
<location id="0B35602311207243FA460DED8653255D" LSID="urn:lsid:plazi:treatment:864387EE113F7245F8A10DC884A22041:0B35602311207243FA460DED8653255D" box="[624,839,1519,1545]" latitude="-3.91" longLatPrecision="21" longitude="-78.48611" municipality="Destacamento Paquisha Alto" name="Zamora-Chinchipe" pageId="16" pageNumber="217">Zamora-Chinchipe</location>
,
<collectingMunicipality id="EE31AC8211207243FB790DED819B255D" box="[847,1167,1519,1545]" pageId="16" pageNumber="217">Destacamento Paquisha Alto</collectingMunicipality>
,
<geoCoordinate id="6BDE503F11207243FCA10DEC8017255D" box="[1175,1283,1518,1545]" degrees="3" direction="south" minutes="54" orientation="latitude" pageId="16" pageNumber="217" precision="15" seconds="36" value="-3.91">3°54'36''S</geoCoordinate>
/
<geoCoordinate id="6BDE503F11207243FD3B0DEC808C255D" box="[1293,1432,1518,1545]" degrees="78" direction="west" minutes="29" orientation="longitude" pageId="16" pageNumber="217" precision="15" seconds="10" value="-78.48611">78°29'10''W</geoCoordinate>
,
<quantity id="C9129B1D11207243F8A10E1185FB2579" box="[151,239,1555,1581]" metricMagnitude="3" metricUnit="m" metricValue="2.01" pageId="16" pageNumber="217" unit="m" value="2010.0">
<elevation id="85C7D1CB11207243F8A10E1185FB2579" box="[151,239,1555,1581]" metricMagnitude="3" metricUnit="m" metricValue="2.01" pageId="16" pageNumber="217" unit="m" value="2010.0">2010 m</elevation>
</quantity>
,
<date id="7A54103811207243F8CA0E1184922579" box="[252,390,1555,1581]" pageId="16" pageNumber="217" value="2010-08-31">
<collectingDate id="6A10E9D011207243F8CA0E1184922579" box="[252,390,1555,1581]" pageId="16" pageNumber="217" value="2010-08-31">31.viii.2010</collectingDate>
</date>
,
<collectorName id="A31F532E11207243F9A40E1187342579" box="[402,544,1555,1581]" pageId="16" pageNumber="217">K. Buckland</collectorName>
leg., [PAN11; dissection, KW-20-018],
<collectionCode id="68FBAE3D11207243FBD00E11815C2579" box="[998,1096,1555,1581]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="16" pageNumber="217" type="Museum">FLMNH</collectionCode>
(to be deposited in
<collectionCode id="68FBAE3D11207243FD1C0E1180852579" box="[1322,1425,1555,1581]" pageId="16" pageNumber="217">INABIO</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="BE823CA511207243F8A10E3587222521" ID-GBIF-Occurrence="3413048307" collectingDate="2010-09-01" collectionCode="FLMNH" collectorName="J. Radford" elevation="2100" latitude="-3.9077778" location="Zamora-Chinchipe" longLatPrecision="21" longitude="-78.48472" municipality="Destacamento Paquisha Alto" pageId="16" pageNumber="217" specimenCount="8" specimenCount-male="8" typeStatus="paratype">
<typeStatus id="D151885A11207243F8A10E3584102505" box="[151,260,1591,1617]" pageId="16" pageNumber="217" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="18ECFD7111207243F9210E35845C2505" box="[279,328,1591,1617]" pageId="16" pageNumber="217" type="male">7 ♂</specimenCount>
):
<location id="0B35602311207243F9560E3587232505" LSID="urn:lsid:plazi:treatment:864387EE113F7245F8A10DC884A22041:0B35602311207243F9560E3587232505" box="[352,567,1591,1617]" latitude="-3.9077778" longLatPrecision="21" longitude="-78.48472" municipality="Destacamento Paquisha Alto" name="Zamora-Chinchipe" pageId="16" pageNumber="217">Zamora-Chinchipe</location>
,
<collectingMunicipality id="EE31AC8211207243FA720E3586842505" box="[580,912,1591,1617]" pageId="16" pageNumber="217">Destacamento Paquisha Alto</collectingMunicipality>
,
<geoCoordinate id="6BDE503F11207243FBA80E34811F2505" box="[926,1035,1590,1617]" degrees="3" direction="south" minutes="54" orientation="latitude" pageId="16" pageNumber="217" precision="15" seconds="28" value="-3.9077778">3°54'28''S</geoCoordinate>
/
<geoCoordinate id="6BDE503F11207243FC220E3481842505" box="[1044,1168,1590,1617]" degrees="78" direction="west" minutes="29" orientation="longitude" pageId="16" pageNumber="217" precision="15" seconds="5" value="-78.48472">78°29'5''W</geoCoordinate>
,
<quantity id="C9129B1D11207243FCA90E3581ED2505" box="[1183,1273,1591,1617]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="16" pageNumber="217" unit="m" value="2100.0">
<elevation id="85C7D1CB11207243FCA90E3581ED2505" box="[1183,1273,1591,1617]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="16" pageNumber="217" unit="m" value="2100.0">2100 m</elevation>
</quantity>
,
<date id="7A54103811207243FD3E0E35806F2505" box="[1288,1403,1591,1617]" pageId="16" pageNumber="217" value="2010-09-01">
<collectingDate id="6A10E9D011207243FD3E0E35806F2505" box="[1288,1403,1591,1617]" pageId="16" pageNumber="217" value="2010-09-01">1.IX.2010</collectingDate>
</date>
,
<collectorName id="A31F532E11207243FDBC0E3585E72521" pageId="16" pageNumber="217">J. Radford</collectorName>
leg.,
<specimenCount id="18ECFD7111207243F9060E59844D2521" box="[304,345,1627,1653]" pageId="16" pageNumber="217" type="male">1 ♂</specimenCount>
[PAN52],
<collectionCode id="68FBAE3D11207243F9E60E5987222521" box="[464,566,1627,1653]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="16" pageNumber="217" type="Museum">FLMNH</collectionCode>
</materialsCitation>
;
<materialsCitation id="BE823CA511207243FA760E59875725CD" ID-GBIF-Occurrence="3413048345" collectingDate="2010-08-31" collectionCode="FLMNH" collectorName="K. Buckland" elevation="2088" latitude="-3.9077778" location="Destacamento Paquisha Alto" longLatPrecision="21" longitude="-78.48499" pageId="16" pageNumber="217" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<location id="0B35602311207243FA760E5986902521" LSID="urn:lsid:plazi:treatment:864387EE113F7245F8A10DC884A22041:0B35602311207243FA760E5986902521" box="[576,900,1627,1653]" latitude="-3.9077778" longLatPrecision="21" longitude="-78.48499" name="Destacamento Paquisha Alto" pageId="16" pageNumber="217">Destacamento Paquisha Alto</location>
,
<geoCoordinate id="6BDE503F11207243FBB80E5886EE2521" box="[910,1018,1626,1653]" degrees="3" direction="south" minutes="54" orientation="latitude" pageId="16" pageNumber="217" precision="15" seconds="28" value="-3.9077778">3°54'28''S</geoCoordinate>
/
<geoCoordinate id="6BDE503F11207243FC320E5881942521" box="[1028,1152,1626,1653]" degrees="78" direction="west" minutes="29" orientation="longitude" pageId="16" pageNumber="217" precision="15" seconds="6" value="-78.48499">78°29'6''W</geoCoordinate>
,
<quantity id="C9129B1D11207243FCBD0E5981F52521" box="[1163,1249,1627,1653]" metricMagnitude="3" metricUnit="m" metricValue="2.088" pageId="16" pageNumber="217" unit="m" value="2088.0">
<elevation id="85C7D1CB11207243FCBD0E5981F52521" box="[1163,1249,1627,1653]" metricMagnitude="3" metricUnit="m" metricValue="2.088" pageId="16" pageNumber="217" unit="m" value="2088.0">2088 m</elevation>
</quantity>
,
<date id="7A54103811207243FCDA0E5980632521" box="[1260,1399,1627,1653]" pageId="16" pageNumber="217" value="2010-08-31">
<collectingDate id="6A10E9D011207243FCDA0E5980632521" box="[1260,1399,1627,1653]" pageId="16" pageNumber="217" value="2010-08-31">31.viii.2010</collectingDate>
</date>
,
<collectorName id="A31F532E11207243FDB70E59841625CD" pageId="16" pageNumber="217">K. Buckland</collectorName>
leg.,
<specimenCount id="18ECFD7111207243F9080E7D847325CD" box="[318,359,1663,1689]" pageId="16" pageNumber="217" type="male">1 ♂</specimenCount>
[PAN13],
<collectionCode id="68FBAE3D11207243F9EB0E7D875725CD" box="[477,579,1663,1689]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="16" pageNumber="217" type="Museum">FLMNH</collectionCode>
</materialsCitation>
;
<materialsCitation id="BE823CA511207243FA7A0E7D875225E9" ID-GBIF-Occurrence="3413048362" collectingDate="2010-09-03" collectionCode="FLMNH" collectorName="K. Buckland" elevation="2010" latitude="-3.91" location="Destacamento Paquisha Alto" longLatPrecision="21" longitude="-78.48611" pageId="16" pageNumber="217" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<location id="0B35602311207243FA7A0E7D869B25CD" LSID="urn:lsid:plazi:treatment:864387EE113F7245F8A10DC884A22041:0B35602311207243FA7A0E7D869B25CD" box="[588,911,1663,1689]" latitude="-3.91" longLatPrecision="21" longitude="-78.48611" name="Destacamento Paquisha Alto" pageId="16" pageNumber="217">Destacamento Paquisha Alto</location>
,
<geoCoordinate id="6BDE503F11207243FBAE0E7C811025CD" box="[920,1028,1662,1689]" degrees="3" direction="south" minutes="54" orientation="latitude" pageId="16" pageNumber="217" precision="15" seconds="36" value="-3.91">3°54'36''S</geoCoordinate>
/
<geoCoordinate id="6BDE503F11207243FC380E7C818D25CD" box="[1038,1177,1662,1689]" degrees="78" direction="west" minutes="29" orientation="longitude" pageId="16" pageNumber="217" precision="15" seconds="10" value="-78.48611">78°29'10''W</geoCoordinate>
,
<quantity id="C9129B1D11207243FC950E7D81EC25CD" box="[1187,1272,1663,1689]" metricMagnitude="3" metricUnit="m" metricValue="2.01" pageId="16" pageNumber="217" unit="m" value="2010.0">
<elevation id="85C7D1CB11207243FC950E7D81EC25CD" box="[1187,1272,1663,1689]" metricMagnitude="3" metricUnit="m" metricValue="2.01" pageId="16" pageNumber="217" unit="m" value="2010.0">2010 m</elevation>
</quantity>
,
<date id="7A54103811207243FD350E7D806325CD" box="[1283,1399,1663,1689]" pageId="16" pageNumber="217" value="2010-09-03">
<collectingDate id="6A10E9D011207243FD350E7D806325CD" box="[1283,1399,1663,1689]" pageId="16" pageNumber="217" value="2010-09-03">3.IX.2010</collectingDate>
</date>
,
<collectorName id="A31F532E11207243FDB70E7D841625E9" pageId="16" pageNumber="217">K. Buckland</collectorName>
leg.,
<specimenCount id="18ECFD7111207243F9090EA1847D25E9" box="[319,361,1699,1725]" pageId="16" pageNumber="217" type="male">1 ♂</specimenCount>
[PAN69],
<collectionCode id="68FBAE3D11207243F9D70EA1875225E9" box="[481,582,1699,1725]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="16" pageNumber="217" type="Museum">FLMNH</collectionCode>
</materialsCitation>
,
<materialsCitation id="BE823CA511207243FA660EA1808325E9" ID-GBIF-Occurrence="3413048348" box="[592,1431,1698,1725]" collectingDate="2010-08-31" collectionCode="FLMNH" collectorName="Hartley" pageId="16" pageNumber="217" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<date id="7A54103811207243FA660EA187F125E9" box="[592,741,1699,1725]" pageId="16" pageNumber="217" value="2010-08-31">
<collectingDate id="6A10E9D011207243FA660EA187F125E9" box="[592,741,1699,1725]" pageId="16" pageNumber="217" value="2010-08-31">31.VIII.2010</collectingDate>
</date>
,
<specimenCount id="18ECFD7111207243FAC60EA1860D25E9" box="[752,793,1699,1725]" pageId="16" pageNumber="217" type="male">1 ♂</specimenCount>
[PAN12; dissection KW-20-023],
<collectionCode id="68FBAE3D11207243FCAD0EA1801425E9" box="[1179,1280,1699,1725]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="16" pageNumber="217" type="Museum">FLMNH</collectionCode>
, (
<collectorName id="A31F532E11207243FD240EA1807D25E9" box="[1298,1385,1699,1725]" pageId="16" pageNumber="217">Hartley</collectorName>
, E.)
</materialsCitation>
,
<materialsCitation id="BE823CA511207243F8A10EC5870925B5" ID-GBIF-Occurrence="3413048361" box="[151,541,1734,1761]" collectingDate="2010-09-03" collectionCode="FLMNH" pageId="16" pageNumber="217" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<date id="7A54103811207243F8A10EC5841725B5" box="[151,259,1735,1761]" pageId="16" pageNumber="217" value="2010-09-03">
<collectingDate id="6A10E9D011207243F8A10EC5841725B5" box="[151,259,1735,1761]" pageId="16" pageNumber="217" value="2010-09-03">3.ix.2010</collectingDate>
</date>
,
<specimenCount id="18ECFD7111207243F9260EC5842825B5" box="[272,316,1735,1761]" pageId="16" pageNumber="217" type="male">1 ♂</specimenCount>
[PAN70],
<collectionCode id="68FBAE3D11207243F98E0EC5870925B5" box="[440,541,1735,1761]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="16" pageNumber="217" type="Museum">FLMNH</collectionCode>
</materialsCitation>
,
<materialsCitation id="BE823CA511207243FA1F0EC5819025B5" ID-GBIF-Occurrence="3413048313" box="[553,1156,1734,1761]" collectingDate="2010-08-31" collectionCode="FLMNH" collectorName="J. Radford" pageId="16" pageNumber="217" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<date id="7A54103811207243FA1F0EC587A725B5" box="[553,691,1735,1761]" pageId="16" pageNumber="217" value="2010-08-31">
<collectingDate id="6A10E9D011207243FA1F0EC587A725B5" box="[553,691,1735,1761]" pageId="16" pageNumber="217" value="2010-08-31">31.viii.2010</collectingDate>
</date>
,
<collectorName id="A31F532E11207243FAF60EC5862225B5" box="[704,822,1735,1761]" pageId="16" pageNumber="217">J. Radford</collectorName>
leg.,
<specimenCount id="18ECFD7111207243FB410EC586B625B5" box="[887,930,1735,1761]" pageId="16" pageNumber="217" type="male">1 ♂</specimenCount>
[PAN10],
<collectionCode id="68FBAE3D11207243FC280EC5819025B5" box="[1054,1156,1735,1761]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="16" pageNumber="217" type="Museum">FLMNH</collectionCode>
</materialsCitation>
;
<materialsCitation id="BE823CA511207243FCA60EC581622451" ID-GBIF-Occurrence="3413048373" collectingDate="2010-09-08" collectionCode="FLMNH" collectorName="E. Hartley" elevation="2299" latitude="-3.901389" location="Destacamento Paquisha Alto" longLatPrecision="21" longitude="-78.48167" pageId="16" pageNumber="217" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<location id="0B35602311207243FCA60EC585D82451" LSID="urn:lsid:plazi:treatment:864387EE113F7245F8A10DC884A22041:0B35602311207243FCA60EC585D82451" latitude="-3.901389" longLatPrecision="21" longitude="-78.48167" name="Destacamento Paquisha Alto" pageId="16" pageNumber="217">Destacamento Paquisha Alto</location>
,
<geoCoordinate id="6BDE503F11207243F8ED0EE884232451" box="[219,311,1770,1797]" degrees="3" direction="south" minutes="54" orientation="latitude" pageId="16" pageNumber="217" precision="15" seconds="5" value="-3.901389">3°54'5''S</geoCoordinate>
/
<geoCoordinate id="6BDE503F11207243F9760EE884D92451" box="[320,461,1770,1797]" degrees="78" direction="west" minutes="28" orientation="longitude" pageId="16" pageNumber="217" precision="15" seconds="54" value="-78.48167">78°28'54''W</geoCoordinate>
,
<quantity id="C9129B1D11207243F9EB0EE9872C2451" box="[477,568,1771,1797]" metricMagnitude="3" metricUnit="m" metricValue="2.299" pageId="16" pageNumber="217" unit="m" value="2299.0">
<elevation id="85C7D1CB11207243F9EB0EE9872C2451" box="[477,568,1771,1797]" metricMagnitude="3" metricUnit="m" metricValue="2.299" pageId="16" pageNumber="217" unit="m" value="2299.0">2299 m</elevation>
</quantity>
,
<date id="7A54103811207243FA7E0EE987A82451" box="[584,700,1771,1797]" pageId="16" pageNumber="217" value="2010-09-08">
<collectingDate id="6A10E9D011207243FA7E0EE987A82451" box="[584,700,1771,1797]" pageId="16" pageNumber="217" value="2010-09-08">8.IX.2010</collectingDate>
</date>
,
<collectorName id="A31F532E11207243FAFA0EE986502451" box="[716,836,1771,1797]" pageId="16" pageNumber="217">E. Hartley</collectorName>
,
<specimenCount id="18ECFD7111207243FB650EE986952451" box="[851,897,1771,1797]" pageId="16" pageNumber="217" type="male">1 ♂</specimenCount>
[PAN197],
<collectionCode id="68FBAE3D11207243FC260EE981622451" box="[1040,1142,1771,1797]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="16" pageNumber="217" type="Museum">FLMNH</collectionCode>
</materialsCitation>
;
<materialsCitation id="BE823CA511207243FCB30EE986642419" ID-GBIF-Occurrence="3413048341" collectingDate="2002-06" collectingDateMax="2020-06-25" collectingDateMin="2002-06" collectionCode="T" collectorName="B. Calderon &amp; K. Florczyk" country="Peru" elevation="2350" location="Amazonas" municipality="Rio Nieva" pageId="16" pageNumber="217" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<collectingCountry id="76FD766811207243FCB30EE981C52451" box="[1157,1233,1771,1797]" name="Peru" pageId="16" pageNumber="217">PERU</collectingCountry>
:
<specimenCount id="18ECFD7111207243FCD60EE980052451" box="[1248,1297,1771,1797]" pageId="16" pageNumber="217" type="male">1 ♂</specimenCount>
:
<location id="0B35602311207243FD290EE9808C2451" LSID="urn:lsid:plazi:treatment:864387EE113F7245F8A10DC884A22041:0B35602311207243FD290EE9808C2451" box="[1311,1432,1771,1797]" country="Peru" municipality="Rio Nieva" name="Amazonas" pageId="16" pageNumber="217">Amazonas</location>
, Alto
<collectingMunicipality id="EE31AC8211207243F8E40F0D845E247D" box="[210,330,1807,1833]" pageId="16" pageNumber="217">Río Nieva</collectingMunicipality>
,
<quantity id="C9129B1D11207243F96E0F0C84E3247D" box="[344,503,1806,1833]" metricMagnitude="3" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="16" pageNumber="217" unit="m" value="2350.0" valueMax="2500.0" valueMin="2200.0">
<elevation id="85C7D1CB11207243F96E0F0C84E3247D" box="[344,503,1806,1833]" metricMagnitude="3" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="16" pageNumber="217" unit="m" value="2350.0" valueMax="2500.0" valueMin="2200.0">22002500 m</elevation>
</quantity>
,
<date id="7A54103811207243FA300F0D8749247D" box="[518,605,1807,1833]" pageId="16" pageNumber="217" value="2002-06">
<collectingDate id="6A10E9D011207243FA300F0D8749247D" box="[518,605,1807,1833]" pageId="16" pageNumber="217" value="2002-06">vi.2002</collectingDate>
</date>
,
<collectorName id="A31F532E11207243FA5D0F0D87E1247D" box="[619,757,1807,1833]" pageId="16" pageNumber="217">B. Calderón</collectorName>
leg., prep. genit. 2743,
<date id="7A54103811207243FC3D0F0C8199247D" box="[1035,1165,1806,1833]" pageId="16" pageNumber="217" value="2020-06-25">
<collectingDate id="6A10E9D011207243FC3D0F0C8199247D" box="[1035,1165,1806,1833]" pageId="16" pageNumber="217" value="2020-06-25">25.06.2020</collectingDate>
</date>
/
<collectorName id="A31F532E11207243FCA00F0C8007247D" box="[1174,1299,1806,1833]" pageId="16" pageNumber="217">K.Florczyk</collectorName>
, H 397, red label saying:
<taxonomicName id="C9EA4D7B11207243F9030F3187B32419" authority="Pyrcz, 2004" authorityName="Pyrcz" authorityYear="2004" box="[309,679,1843,1869]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="benigni">
<emphasis id="3C9EEAEA11207243F9030F31871A2419" box="[309,526,1843,1869]" italics="true" pageId="16" pageNumber="217">Manerebia benigni</emphasis>
<bibRefCitation id="6A7B4B0911207243FA2C0F3187B32419" author="Pyrcz, T. W." box="[538,679,1843,1869]" pageId="16" pageNumber="217" pagination="455 - 622" refId="ref22314" refString="Pyrcz, T. W. (2004) Pronophiline butterflies of the highlands of Chachapoyas in northern Peru: faunal survey, diversity and distribution patterns (Lepidoptera, Nymphalidae, Satyrinae). Genus, 15, 455 - 622." type="journal article" year="2004">Pyrcz, 2004</bibRefCitation>
</taxonomicName>
,
<typeStatus id="D151885A11207243FA800F31860F2419" box="[694,795,1843,1869]" pageId="16" pageNumber="217" type="paratype">Paratype</typeStatus>
, det.
<collectionCode id="68FBAE3D11207243FB680F3186642419" box="[862,880,1843,1869]" name="Tavera, Department of Geology and Geophysics" pageId="16" pageNumber="217">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="BE823CA511207243FBB60F3181A52419" ID-GBIF-Occurrence="3413048343" box="[896,1201,1843,1869]" collectionCode="R" collectorName="Pyrcz" country="Peru" location="Amazonas" pageId="16" pageNumber="217" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<collectorName id="A31F532E11207243FBB60F3186D72419" box="[896,963,1843,1869]" pageId="16" pageNumber="217">Pyrcz</collectorName>
;
<specimenCount id="18ECFD7111207243FBE50F3181102419" box="[979,1028,1843,1869]" pageId="16" pageNumber="217" type="male">1 ♂</specimenCount>
:
<location id="0B35602311207243FC250F3181982419" LSID="urn:lsid:plazi:treatment:864387EE113F7245F8A10DC884A22041:0B35602311207243FC250F3181982419" box="[1043,1164,1843,1869]" country="Peru" name="Amazonas" pageId="16" pageNumber="217">Amazonas</location>
,
<collectionCode id="68FBAE3D11207243FCAD0F3181A52419" box="[1179,1201,1843,1869]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="16" pageNumber="217">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="BE823CA511207243FCF60F31846724C1" ID-GBIF-Occurrence="3413048352" collectingDate="1998-10-02" collectingDateMax="2020-06-24" collectingDateMin="1998-10-02" collectionCode="CEPUJ" collectorName="B. Calderon &amp; K. Florczyk" country="Peru" elevation="1900" location="De Mendoza" municipality="Qda. Llanohuaico" pageId="16" pageNumber="217" specimenCode="CEPUJ 202006110" specimenCount="1" typeStatus="paratype">
<location id="0B35602311207243FCF60F3180432419" LSID="urn:lsid:plazi:treatment:864387EE113F7245F8A10DC884A22041:0B35602311207243FCF60F3180432419" box="[1216,1367,1843,1869]" country="Peru" municipality="Qda. Llanohuaico" name="De Mendoza" pageId="16" pageNumber="217">De Mendoza</location>
,
<collectingMunicipality id="EE31AC8211207243FD510F3184322425" pageId="16" pageNumber="217">Qda. Llanohuaico</collectingMunicipality>
,
<quantity id="C9129B1D11207243F9020F5584C02425" box="[308,468,1879,1905]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="16" pageNumber="217" unit="m" value="1900.0" valueMax="2000.0" valueMin="1800.0">
<elevation id="85C7D1CB11207243F9020F5584C02425" box="[308,468,1879,1905]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="16" pageNumber="217" unit="m" value="1900.0" valueMax="2000.0" valueMin="1800.0">18002000 m</elevation>
</quantity>
,
<date id="7A54103811207243F9D50F5587412425" box="[483,597,1879,1905]" pageId="16" pageNumber="217" value="1998-10-02">
<collectingDate id="6A10E9D011207243F9D50F5587412425" box="[483,597,1879,1905]" pageId="16" pageNumber="217" value="1998-10-02">02.x.1998</collectingDate>
</date>
,
<collectorName id="A31F532E11207243FA520F5587FB2425" box="[612,751,1879,1905]" pageId="16" pageNumber="217">B. Calderón</collectorName>
leg., prep. genit. 2736,
<date id="7A54103811207243FC310F55819C2425" box="[1031,1160,1879,1905]" pageId="16" pageNumber="217" value="2020-06-24">
<collectingDate id="6A10E9D011207243FC310F55819C2425" box="[1031,1160,1879,1905]" pageId="16" pageNumber="217" value="2020-06-24">24.06.2020</collectingDate>
</date>
/
<collectorName id="A31F532E11207243FCA70F55800C2425" box="[1169,1304,1879,1905]" pageId="16" pageNumber="217">K. Florczyk</collectorName>
, prep. mol.
<specimenCode id="5E4C9E8311207243F8A10F79846724C1" box="[151,371,1915,1941]" collectionCode="CEPUJ" pageId="16" pageNumber="217">CEPUJ 202006110</specimenCode>
</materialsCitation>
/
<materialsCitation id="BE823CA511207243F94A0F79805324C1" ID-GBIF-Occurrence="3413048329" box="[380,1351,1915,1942]" collectionCode="T" collectorName="Florczyk" country="Peru" pageId="16" pageNumber="217" specimenCount="1" typeStatus="paratype">
K.
<collectorName id="A31F532E11207243F9AB0F79871524C1" box="[413,513,1915,1941]" pageId="16" pageNumber="217">Florczyk</collectorName>
(lab), red label saying:
<taxonomicName id="C9EA4D7B11207243FB210F79819724C1" authority="Pyrcz, 2004" authorityName="Pyrcz" authorityYear="2004" box="[791,1155,1915,1941]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="benigni">
<emphasis id="3C9EEAEA11207243FB210F7986FA24C1" box="[791,1006,1915,1941]" italics="true" pageId="16" pageNumber="217">Manerebia benigni</emphasis>
<bibRefCitation id="6A7B4B0911207243FBCE0F79819724C1" author="Pyrcz, T. W." box="[1016,1155,1915,1941]" pageId="16" pageNumber="217" pagination="455 - 622" refId="ref22314" refString="Pyrcz, T. W. (2004) Pronophiline butterflies of the highlands of Chachapoyas in northern Peru: faunal survey, diversity and distribution patterns (Lepidoptera, Nymphalidae, Satyrinae). Genus, 15, 455 - 622." type="journal article" year="2004">Pyrcz, 2004</bibRefCitation>
</taxonomicName>
,
<typeStatus id="D151885A11207243FCA60F7981E124C1" box="[1168,1269,1915,1941]" pageId="16" pageNumber="217" type="paratype">Paratype</typeStatus>
, det.
<collectionCode id="68FBAE3D11207243FD030F79805324C1" box="[1333,1351,1915,1941]" name="Tavera, Department of Geology and Geophysics" pageId="16" pageNumber="217">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="BE823CA511207243FD630F7987F024ED" ID-GBIF-Occurrence="3413048360" collectionCode="VI" collectorName="Pyrcz" country="Peru" elevation="2350" location="Amazonas" municipality="Rio Nieva" pageId="16" pageNumber="217" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<collectorName id="A31F532E11207243FD630F79808C24C1" box="[1365,1432,1915,1941]" pageId="16" pageNumber="217">Pyrcz</collectorName>
;
<specimenCount id="18ECFD7111207243F8A10F9D85D224ED" box="[151,198,1951,1977]" pageId="16" pageNumber="217" type="male">1 ♂</specimenCount>
:
<location id="0B35602311207243F8E50F9D845824ED" LSID="urn:lsid:plazi:treatment:864387EE113F7245F8A10DC884A22041:0B35602311207243F8E50F9D845824ED" box="[211,332,1951,1977]" country="Peru" municipality="Rio Nieva" name="Amazonas" pageId="16" pageNumber="217">Amazonas</location>
, Alto
<collectingMunicipality id="EE31AC8211207243F9A50F9D871F24ED" box="[403,523,1951,1977]" pageId="16" pageNumber="217">Río Nieva</collectingMunicipality>
,
<quantity id="C9129B1D11207243FA2E0F9C87A324ED" box="[536,695,1950,1977]" metricMagnitude="3" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="16" pageNumber="217" unit="m" value="2350.0" valueMax="2500.0" valueMin="2200.0">
<elevation id="85C7D1CB11207243FA2E0F9C87A324ED" box="[536,695,1950,1977]" metricMagnitude="3" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="16" pageNumber="217" unit="m" value="2350.0" valueMax="2500.0" valueMin="2200.0">22002500 m</elevation>
</quantity>
,
<date id="7A54103811207243FAF30F9D87F024ED" box="[709,740,1951,1977]" pageId="16" pageNumber="217" value="2002-06">
<collectionCode id="68FBAE3D11207243FAF30F9D87F024ED" box="[709,740,1951,1977]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="16" pageNumber="217">VI</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="BE823CA511207243FAD20F9D87402489" ID-GBIF-Occurrence="3413048356" collectionCode="T" collectorName="B. Calderon" country="Peru" pageId="16" pageNumber="217" specimenCount="1" typeStatus="paratype">
.2002,
<collectorName id="A31F532E11207243FB040F9D86A824ED" box="[818,956,1951,1977]" pageId="16" pageNumber="217">B. Calderón</collectorName>
leg., red label saying:
<taxonomicName id="C9EA4D7B11207243FCF30F9D84832489" authority="Pyrcz, 2004" authorityName="Pyrcz" authorityYear="2004" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="subSpecies" species="benigni" subSpecies="tessmanni">
<emphasis id="3C9EEAEA11207243FCF30F9D84132488" italics="true" pageId="16" pageNumber="217">Manerebia benigni tessmanni</emphasis>
<bibRefCitation id="6A7B4B0911207243F9380FC184832489" author="Pyrcz, T. W." box="[270,407,1987,2013]" pageId="16" pageNumber="217" pagination="455 - 622" refId="ref22314" refString="Pyrcz, T. W. (2004) Pronophiline butterflies of the highlands of Chachapoyas in northern Peru: faunal survey, diversity and distribution patterns (Lepidoptera, Nymphalidae, Satyrinae). Genus, 15, 455 - 622." type="journal article" year="2004">Pyrcz, 2004</bibRefCitation>
</taxonomicName>
,
<typeStatus id="D151885A11207243F9940FC187132489" box="[418,519,1987,2013]" pageId="16" pageNumber="217" type="paratype">Paratype</typeStatus>
, det.
<collectionCode id="68FBAE3D11207243FA740FC187402489" box="[578,596,1987,2013]" name="Tavera, Department of Geology and Geophysics" pageId="16" pageNumber="217">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="BE823CA511207243FA690FC187B62489" ID-GBIF-Occurrence="3413048366" box="[607,674,1987,2013]" country="Peru" location="Pyrcz" pageId="16" pageNumber="217" specimenCount="1" typeStatus="paratype">
<location id="0B35602311207243FA690FC187B62489" LSID="urn:lsid:plazi:treatment:864387EE113F7245F8A10DC884A22041:0B35602311207243FA690FC187B62489" box="[607,674,1987,2013]" country="Peru" name="Pyrcz" pageId="16" pageNumber="217">Pyrcz</location>
</materialsCitation>
.
</paragraph>
<caption id="5A95667011217242F8A10F358641248B" ID-DOI="http://doi.org/10.5281/zenodo.5744563" ID-Zenodo-Dep="5744563" httpUri="https://zenodo.org/record/5744563/files/figure.png" pageId="17" pageNumber="218" startId="17.[151,250,1847,1871]" targetBox="[237,1370,197,1811]" targetPageId="17">
<paragraph id="0E5536F811217242F8A10F3581E4241B" blockId="17.[151,1264,1847,2016]" box="[151,1264,1847,1871]" pageId="17" pageNumber="218">
<emphasis id="3C9EEAEA11217242F8A10F358407241B" bold="true" box="[151,275,1847,1871]" pageId="17" pageNumber="218">FIGURE 6.</emphasis>
Male genitalia (top—lateral view, middle—plan view, bottom—aedeagus extracted, lateral view)
</paragraph>
<paragraph id="0E5536F811217242F8A10F5E86852427" blockId="17.[151,1264,1847,2016]" box="[151,913,1883,1908]" pageId="17" pageNumber="218">
A.
<taxonomicName id="C9EA4D7B11217242F8F10F5E84572427" authority="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz, 2021" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[199,323,1884,1907]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="granatus" status="sp. nov.">
<emphasis id="3C9EEAEA11217242F8F10F5E85C82427" box="[199,220,1884,1907]" italics="true" pageId="17" pageNumber="218">M</emphasis>
.
<emphasis id="3C9EEAEA11217242F8DF0F5E84572427" box="[233,323,1884,1907]" italics="true" pageId="17" pageNumber="218">granatus</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA11217242F97F0F5E84962427" bold="true" box="[329,386,1884,1908]" pageId="17" pageNumber="218">
<taxonomicNameLabel id="27AD579111217242F97F0F5E84962427" box="[329,386,1884,1908]" pageId="17" pageNumber="218" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
<collectingCountry id="76FD766811217242F9BF0F5E84AE2427" box="[393,442,1884,1907]" name="Peru" pageId="17" pageNumber="218">Peru</collectingCountry>
, Rodríguez de Mendoza, prep. genit. KF-2736
</paragraph>
<paragraph id="0E5536F811217242F8A10F82863C24CC" blockId="17.[151,1264,1847,2016]" box="[151,808,1919,1944]" pageId="17" pageNumber="218">
B.
<taxonomicName id="C9EA4D7B11217242F8F10F82842124C3" authority="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz, 2021" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[199,309,1919,1943]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="placida" status="sp. nov.">
<emphasis id="3C9EEAEA11217242F8F10F8285C824C3" box="[199,220,1920,1943]" italics="true" pageId="17" pageNumber="218">M</emphasis>
.
<emphasis id="3C9EEAEA11217242F8DF0F7D842124C3" box="[233,309,1919,1943]" italics="true" pageId="17" pageNumber="218">placida</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA11217242F90D0F82846024C3" bold="true" box="[315,372,1920,1944]" pageId="17" pageNumber="218">
<taxonomicNameLabel id="27AD579111217242F90D0F82846024C3" box="[315,372,1920,1944]" pageId="17" pageNumber="218" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
<collectingCountry id="76FD766811217242F94D0F7D84C524C3" box="[379,465,1919,1943]" name="Ecuador" pageId="17" pageNumber="218">Ecuador</collectingCountry>
, Zamora, prep. genit. KW-20-021
</paragraph>
<paragraph id="0E5536F811217242F8A10FA6866124E8" blockId="17.[151,1264,1847,2016]" box="[151,885,1955,1980]" pageId="17" pageNumber="218">
C.
<taxonomicName id="C9EA4D7B11217242F8F10FA684B424EF" authorityName="Pyrcz" authorityYear="2004" box="[199,416,1955,1979]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="subSpecies" species="benigni" subSpecies="tessmanni">
<emphasis id="3C9EEAEA11217242F8F10FA685C824EF" box="[199,220,1956,1979]" italics="true" pageId="17" pageNumber="218">M</emphasis>
.
<emphasis id="3C9EEAEA11217242F8DF0FA184B424EF" box="[233,416,1955,1979]" italics="true" pageId="17" pageNumber="218">benigni tessmanni</emphasis>
</taxonomicName>
,
<collectingCountry id="76FD766811217242F99B0FA684CA24EF" box="[429,478,1956,1979]" name="Peru" pageId="17" pageNumber="218">Peru</collectingCountry>
, Alto Río Nieva, prep. genit. KF-H_241
</paragraph>
<paragraph id="0E5536F811217242F8A10FCA8641248B" blockId="17.[151,1264,1847,2016]" box="[151,853,1991,2016]" pageId="17" pageNumber="218">
D.
<taxonomicName id="C9EA4D7B11217242F8F10FCA84B4248B" authorityName="Pyrcz" authorityYear="2004" box="[199,416,1991,2015]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="subSpecies" species="benigni" subSpecies="tessmanni">
<emphasis id="3C9EEAEA11217242F8F10FCA85C8248B" box="[199,220,1992,2015]" italics="true" pageId="17" pageNumber="218">M</emphasis>
.
<emphasis id="3C9EEAEA11217242F8DF0FC584B4248B" box="[233,416,1991,2015]" italics="true" pageId="17" pageNumber="218">benigni tessmanni</emphasis>
</taxonomicName>
,
<collectingCountry id="76FD766811217242F99B0FC58717248B" box="[429,515,1991,2015]" name="Ecuador" pageId="17" pageNumber="218">Ecuador</collectingCountry>
, Río Troya, prep. genit. KF-6282
</paragraph>
</caption>
<caption id="5A95667011227241F8A10F3C866624B3" ID-DOI="http://doi.org/10.5281/zenodo.5744565" ID-Zenodo-Dep="5744565" httpUri="https://zenodo.org/record/5744565/files/figure.png" pageId="18" pageNumber="219" startId="18.[151,250,1854,1878]" targetBox="[216,1381,183,1809]" targetPageId="18">
<paragraph id="0E5536F811227241F8A10F3C81E42403" blockId="18.[151,1264,1854,2023]" box="[151,1264,1854,1879]" pageId="18" pageNumber="219">
<emphasis id="3C9EEAEA11227241F8A10F3C84072403" bold="true" box="[151,275,1854,1879]" pageId="18" pageNumber="219">FIGURE 7.</emphasis>
Male genitalia (top—lateral view, middle—plan view, bottom—aedeagus extracted, lateral view)
</paragraph>
<paragraph id="0E5536F811227241F8A10F6187E0242E" blockId="18.[151,1264,1854,2023]" box="[151,756,1890,1915]" pageId="18" pageNumber="219">
<emphasis id="3C9EEAEA11227241F8A10F6185BA242E" box="[151,174,1891,1914]" italics="true" pageId="18" pageNumber="219">A.</emphasis>
<taxonomicName id="C9EA4D7B11227241F8F10F618431242F" authority="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz, 2021" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[199,293,1891,1915]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="ronda" status="sp. nov.">
<emphasis id="3C9EEAEA11227241F8F10F6185C8242E" box="[199,220,1891,1914]" italics="true" pageId="18" pageNumber="219">M</emphasis>
.
<emphasis id="3C9EEAEA11227241F8DF0F618431242F" box="[233,293,1891,1915]" italics="true" pageId="18" pageNumber="219">ronda</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA11227241F91D0F618470242E" bold="true" box="[299,356,1891,1915]" pageId="18" pageNumber="219">
<taxonomicNameLabel id="27AD579111227241F91D0F618470242E" box="[299,356,1891,1915]" pageId="18" pageNumber="219" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, Peru, Salhuindo, prep. genit. KF-1571
</paragraph>
<paragraph id="0E5536F811227241F8A10F85863C24CA" blockId="18.[151,1264,1854,2023]" box="[151,808,1926,1951]" pageId="18" pageNumber="219">
<emphasis id="3C9EEAEA11227241F8A10F85846524CB" box="[151,369,1927,1951]" italics="true" pageId="18" pageNumber="219">
B.
<taxonomicName id="C9EA4D7B11227241F8F10F85846524CB" authorityName="Pyrcz &amp; Boyer" authorityYear="2021" box="[199,369,1927,1951]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="subSpecies" species="ronda" subSpecies="amplia">M. ronda amplia</taxonomicName>
</emphasis>
<emphasis id="3C9EEAEA11227241F94E0F8584AE24CA" bold="true" box="[376,442,1927,1951]" pageId="18" pageNumber="219">n. ssp.</emphasis>
, Peru, Pucara, prep. genit. KF-1572
</paragraph>
<paragraph id="0E5536F811227241F8A10FA9862E2496" blockId="18.[151,1264,1854,2023]" box="[151,826,1962,1987]" pageId="18" pageNumber="219">
<emphasis id="3C9EEAEA11227241F8A10FA985BB2496" box="[151,175,1963,1986]" italics="true" pageId="18" pageNumber="219">C.</emphasis>
<taxonomicName id="C9EA4D7B11227241F8F10FA984332497" authority="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz, 2021" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[199,295,1963,1987]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="punku" status="sp. nov.">
<emphasis id="3C9EEAEA11227241F8F10FA985C82496" box="[199,220,1963,1986]" italics="true" pageId="18" pageNumber="219">M</emphasis>
.
<emphasis id="3C9EEAEA11227241F8DF0FA984332497" box="[233,295,1963,1987]" italics="true" pageId="18" pageNumber="219">punku</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA11227241F91B0FA984722496" bold="true" box="[301,358,1963,1987]" pageId="18" pageNumber="219">
<taxonomicNameLabel id="27AD579111227241F91B0FA984722496" box="[301,358,1963,1987]" pageId="18" pageNumber="219" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, Peru, Abra de Porculla, prep. genit. KF-1613
</paragraph>
<paragraph id="0E5536F811227241F8A10FCD866624B3" blockId="18.[151,1264,1854,2023]" box="[151,882,1999,2023]" pageId="18" pageNumber="219">
<emphasis id="3C9EEAEA11227241F8A10FCD85A424B2" box="[151,176,1999,2022]" italics="true" pageId="18" pageNumber="219">D.</emphasis>
<taxonomicName id="C9EA4D7B11227241F8F10FCD848124B3" authorityName="Pyrcz &amp; Boyer" authorityYear="2021" box="[199,405,1999,2023]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="subSpecies" species="prattorum" subSpecies="udima">
<emphasis id="3C9EEAEA11227241F8F10FCD85C824B2" box="[199,220,1999,2022]" italics="true" pageId="18" pageNumber="219">M</emphasis>
.
<emphasis id="3C9EEAEA11227241F8DF0FCD848124B3" box="[233,405,1999,2023]" italics="true" pageId="18" pageNumber="219">prattorum udima</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA11227241F9AA0FCD84CA24B2" bold="true" box="[412,478,1999,2023]" pageId="18" pageNumber="219">n. ssp.</emphasis>
,
<collectingCountry id="76FD766811227241F9DD0FCD870824B2" box="[491,540,1999,2022]" name="Peru" pageId="18" pageNumber="219">Peru</collectingCountry>
, La Florida, prep. genit. KF-1472
</paragraph>
</caption>
<caption id="5A95667011237240F8A10F52866624AC" ID-DOI="http://doi.org/10.5281/zenodo.5744569" ID-Zenodo-Dep="5744569" httpUri="https://zenodo.org/record/5744569/files/figure.png" pageId="19" pageNumber="220" startId="19.[151,250,1872,1896]" targetBox="[222,1362,240,1829]" targetPageId="19">
<paragraph id="0E5536F811237240F8A10F5287F224D8" blockId="19.[151,1437,1872,2040]" pageId="19" pageNumber="220">
<emphasis id="3C9EEAEA11237240F8A10F528406243C" bold="true" box="[151,274,1872,1896]" pageId="19" pageNumber="220">FIGURE 8.</emphasis>
Male genitalia (A, B) (top—lateral view, middle—plan view, bottom—aedeagus extracted, lateral view) C. Female genitalia (top—lateral view; bottom—ventral view, detail)
</paragraph>
<paragraph id="0E5536F811237240F8A10F9B86D424E4" blockId="19.[151,1437,1872,2040]" box="[151,960,1944,1969]" pageId="19" pageNumber="220">
<emphasis id="3C9EEAEA11237240F8A10F9B85BA24E4" box="[151,174,1945,1968]" italics="true" pageId="19" pageNumber="220">A.</emphasis>
<taxonomicName id="C9EA4D7B11237240F8F10F9B842124E4" authority="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz, 2021" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[199,309,1944,1968]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="220" phylum="Arthropoda" rank="species" species="inducta" status="sp. nov.">
<emphasis id="3C9EEAEA11237240F8F10F9B85C824E4" box="[199,220,1945,1968]" italics="true" pageId="19" pageNumber="220">M</emphasis>
.
<emphasis id="3C9EEAEA11237240F8DF0F9A842124E4" box="[233,309,1944,1968]" italics="true" pageId="19" pageNumber="220">inducta</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA11237240F90D0F9A846024E4" bold="true" box="[315,372,1944,1968]" pageId="19" pageNumber="220">
<taxonomicNameLabel id="27AD579111237240F90D0F9A846024E4" box="[315,372,1944,1968]" pageId="19" pageNumber="220" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, Peru, Lagunas Arrebiatadas, prep. genit. MUSA_015149
</paragraph>
<paragraph id="0E5536F811237240F8A10FBF864C2480" blockId="19.[151,1437,1872,2040]" box="[151,856,1980,2004]" pageId="19" pageNumber="220">
<emphasis id="3C9EEAEA11237240F8A10FBF85BA2480" box="[151,174,1981,2004]" italics="true" pageId="19" pageNumber="220">B.</emphasis>
<taxonomicName id="C9EA4D7B11237240F8F10FBF845D2480" authority="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz, 2021" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[199,329,1980,2004]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="220" phylum="Arthropoda" rank="species" species="huamanii" status="sp. nov.">
<emphasis id="3C9EEAEA11237240F8F10FBF85C82480" box="[199,220,1981,2004]" italics="true" pageId="19" pageNumber="220">M</emphasis>
.
<emphasis id="3C9EEAEA11237240F8DF0FBE845D2480" box="[233,329,1980,2004]" italics="true" pageId="19" pageNumber="220">huamanii</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA11237240F9790FBE849C2480" bold="true" box="[335,392,1980,2004]" pageId="19" pageNumber="220">
<taxonomicNameLabel id="27AD579111237240F9790FBE849C2480" box="[335,392,1980,2004]" pageId="19" pageNumber="220" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, Peru, Tres Ríos, prep. genit. MUSA_015150
</paragraph>
<paragraph id="0E5536F811237240F8A10FE3866624AC" blockId="19.[151,1437,1872,2040]" box="[151,882,2016,2040]" pageId="19" pageNumber="220">
<emphasis id="3C9EEAEA11237240F8A10FE385BB24AC" box="[151,175,2017,2040]" italics="true" pageId="19" pageNumber="220">C.</emphasis>
<taxonomicName id="C9EA4D7B11237240F8F10FE3848124AC" authorityName="Pyrcz &amp; Boyer" authorityYear="2021" box="[199,405,2016,2040]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="220" phylum="Arthropoda" rank="subSpecies" species="prattorum" subSpecies="udima">
<emphasis id="3C9EEAEA11237240F8F10FE385C824AC" box="[199,220,2017,2040]" italics="true" pageId="19" pageNumber="220">M</emphasis>
.
<emphasis id="3C9EEAEA11237240F8DF0FE3848124AC" box="[233,405,2016,2040]" italics="true" pageId="19" pageNumber="220">prattorum udima</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA11237240F9AA0FE284CA24AC" bold="true" box="[412,478,2016,2040]" pageId="19" pageNumber="220">n. ssp.</emphasis>
, Peru, La Florida, prep. genit. KF-1473
</paragraph>
</caption>
<caption id="5A95667011247247F8A10F9586B924A3" ID-DOI="http://doi.org/10.5281/zenodo.5744571" ID-Zenodo-Dep="5744571" httpUri="https://zenodo.org/record/5744571/files/figure.png" pageId="20" pageNumber="221" startId="20.[151,250,1943,1967]" targetBox="[202,1350,202,1901]" targetPageId="20">
<paragraph id="0E5536F811247247F8A10F9586AD24E4" blockId="20.[151,953,1943,2040]" box="[151,953,1943,1968]" pageId="20" pageNumber="221">
<emphasis id="3C9EEAEA11247247F8A10F95840724FB" bold="true" box="[151,275,1943,1967]" pageId="20" pageNumber="221">FIGURE 9.</emphasis>
Female genitalia (top—lateral view; bottom—ventral view, detail)
</paragraph>
<paragraph id="0E5536F811247247F8A10FBE866E2487" blockId="20.[151,953,1943,2040]" box="[151,890,1979,2004]" pageId="20" pageNumber="221">
A.
<taxonomicName id="C9EA4D7B11247247F8800FBE84002480" authority="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz, 2021" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[182,276,1980,2004]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="ronda" status="sp. nov.">
<emphasis id="3C9EEAEA11247247F8800FBE85DF2487" box="[182,203,1980,2003]" italics="true" pageId="20" pageNumber="221">M</emphasis>
.
<emphasis id="3C9EEAEA11247247F8EE0FBE84002480" box="[216,276,1980,2004]" italics="true" pageId="20" pageNumber="221">ronda</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA11247247F92C0FBE84472487" bold="true" box="[282,339,1980,2004]" pageId="20" pageNumber="221">
<taxonomicNameLabel id="27AD579111247247F92C0FBE84472487" box="[282,339,1980,2004]" pageId="20" pageNumber="221" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, Peru, Laguna Salahuido, prep. genit. MUSA_015151
</paragraph>
<paragraph id="0E5536F811247247F8A10FE286B924A3" blockId="20.[151,953,1943,2040]" box="[151,941,2015,2040]" pageId="20" pageNumber="221">
B.
<taxonomicName id="C9EA4D7B11247247F8830FE2843724AC" authority="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz, 2021" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[181,291,2016,2040]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="inducta" status="sp. nov.">
<emphasis id="3C9EEAEA11247247F8830FE285DE24A3" box="[181,202,2016,2039]" italics="true" pageId="20" pageNumber="221">M</emphasis>
.
<emphasis id="3C9EEAEA11247247F8E10FE2843724AC" box="[215,291,2016,2040]" italics="true" pageId="20" pageNumber="221">inducta</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA11247247F91F0FE2847624A3" bold="true" box="[297,354,2016,2040]" pageId="20" pageNumber="221">
<taxonomicNameLabel id="27AD579111247247F91F0FE2847624A3" box="[297,354,2016,2040]" pageId="20" pageNumber="221" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, Peru, Lagunas Arrebiatadas, prep. genit. MUSA_015152
</paragraph>
</caption>
<caption id="5A95667011257246F8A10D06861526A1" ID-DOI="http://doi.org/10.5281/zenodo.5744577" ID-Zenodo-Dep="5744577" httpUri="https://zenodo.org/record/5744577/files/figure.png" pageId="21" pageNumber="222" startId="21.[151,250,1284,1308]" targetBox="[171,1415,185,1248]" targetPageId="21">
<paragraph id="0E5536F811257246F8A10D0687CF2615" blockId="21.[151,1436,1284,1525]" pageId="21" pageNumber="222">
<emphasis id="3C9EEAEA11257246F8A10D0684342649" bold="true" box="[151,288,1284,1309]" pageId="21" pageNumber="222">FIGURE 10.</emphasis>
Male genitalia x adults comparison indicating morphological similarities and genetic affinities (outer—genitalia in lateral view, aedeagus omitted, middle—wings venter)
</paragraph>
<paragraph id="0E5536F811257246F8A10D4F863D2631" blockId="21.[151,1436,1284,1525]" box="[151,809,1357,1381]" pageId="21" pageNumber="222">
<emphasis id="3C9EEAEA11257246F8A10D4F85BA2630" box="[151,174,1357,1380]" italics="true" pageId="21" pageNumber="222">A.</emphasis>
<taxonomicName id="C9EA4D7B11257246F8F10D4F84312631" authority="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz, 2021" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[199,293,1357,1381]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="222" phylum="Arthropoda" rank="species" species="ronda" status="sp. nov.">
<emphasis id="3C9EEAEA11257246F8F10D4F85C82630" box="[199,220,1357,1380]" italics="true" pageId="21" pageNumber="222">M</emphasis>
.
<emphasis id="3C9EEAEA11257246F8DF0D4F84312631" box="[233,293,1357,1381]" italics="true" pageId="21" pageNumber="222">ronda</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA11257246F91D0D4F84702631" bold="true" box="[299,356,1357,1381]" pageId="21" pageNumber="222">
<taxonomicNameLabel id="27AD579111257246F91D0D4F84702631" box="[299,356,1357,1381]" pageId="21" pageNumber="222" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, Peru, Laguna Salahuindo, prep. genit. 1571
</paragraph>
<paragraph id="0E5536F811257246F8A10D73860E26DD" blockId="21.[151,1436,1284,1525]" box="[151,794,1393,1417]" pageId="21" pageNumber="222">
<emphasis id="3C9EEAEA11257246F8A10D7385BA26DC" box="[151,174,1393,1416]" italics="true" pageId="21" pageNumber="222">B.</emphasis>
<taxonomicName id="C9EA4D7B11257246F8F10D73849926DD" authorityName="Pyrcz &amp; Willmott" authorityYear="2006" box="[199,397,1393,1417]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="222" phylum="Arthropoda" rank="subSpecies" species="inderena" subSpecies="similis">
<emphasis id="3C9EEAEA11257246F8F10D7385C826DC" box="[199,220,1393,1416]" italics="true" pageId="21" pageNumber="222">M</emphasis>
.
<emphasis id="3C9EEAEA11257246F8DF0D73849926DD" box="[233,397,1393,1417]" italics="true" pageId="21" pageNumber="222">inderena similis</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA11257246F9BB0D73848526DD" box="[397,401,1393,1417]" italics="true" pageId="21" pageNumber="222">,</emphasis>
Peru, Balzapamba, prep. genit. H_161
</paragraph>
<paragraph id="0E5536F811257246F8A10D97867126F9" blockId="21.[151,1436,1284,1525]" box="[151,869,1429,1453]" pageId="21" pageNumber="222">
<emphasis id="3C9EEAEA11257246F8A10D9785BB26F8" box="[151,175,1429,1452]" italics="true" pageId="21" pageNumber="222">C.</emphasis>
<taxonomicName id="C9EA4D7B11257246F8F10D97846526F9" authorityName="Pyrcz &amp; Boyer" authorityYear="2021" box="[199,369,1429,1453]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="222" phylum="Arthropoda" rank="subSpecies" species="ronda" subSpecies="amplia">
<emphasis id="3C9EEAEA11257246F8F10D9785C826F8" box="[199,220,1429,1452]" italics="true" pageId="21" pageNumber="222">M</emphasis>
.
<emphasis id="3C9EEAEA11257246F8DF0D97846526F9" box="[233,369,1429,1453]" italics="true" pageId="21" pageNumber="222">ronda amplia</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA11257246F94E0D9784AE26F8" bold="true" box="[376,442,1429,1453]" pageId="21" pageNumber="222">n. ssp.</emphasis>
, Peru, Abra de Porculla, prep. genit. 1572
</paragraph>
<paragraph id="0E5536F811257246F8A10DBB87EB2685" blockId="21.[151,1436,1284,1525]" box="[151,767,1465,1489]" pageId="21" pageNumber="222">
<emphasis id="3C9EEAEA11257246F8A10DBB85A42684" box="[151,176,1465,1488]" italics="true" pageId="21" pageNumber="222">D.</emphasis>
<taxonomicName id="C9EA4D7B11257246F8F10DBB848F2685" box="[199,411,1465,1489]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="222" phylum="Arthropoda" rank="subSpecies" species="undulata" subSpecies="milaena">
<emphasis id="3C9EEAEA11257246F8F10DBB85C82684" box="[199,220,1465,1488]" italics="true" pageId="21" pageNumber="222">M</emphasis>
.
<emphasis id="3C9EEAEA11257246F8DF0DBB848F2685" box="[233,411,1465,1489]" italics="true" pageId="21" pageNumber="222">undulata milaena</emphasis>
</taxonomicName>
, Peru, Jimbura, prep. genit. H_175
</paragraph>
<paragraph id="0E5536F811257246F8A10DDF861526A1" blockId="21.[151,1436,1284,1525]" box="[151,769,1501,1525]" pageId="21" pageNumber="222">
<emphasis id="3C9EEAEA11257246F8A10DDF85BA26A0" box="[151,174,1501,1524]" italics="true" pageId="21" pageNumber="222">E.</emphasis>
<taxonomicName id="C9EA4D7B11257246F8F10DDF844526A0" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" baseAuthorityName="Pyrcz &amp; Willmott" baseAuthorityYear="2006" box="[199,337,1501,1524]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="222" phylum="Arthropoda" rank="species" species="pauperata" status="stat. nov.">
<emphasis id="3C9EEAEA11257246F8F10DDF85C826A0" box="[199,220,1501,1524]" italics="true" pageId="21" pageNumber="222">M</emphasis>
.
<emphasis id="3C9EEAEA11257246F8DF0DDF844526A0" box="[233,337,1501,1524]" italics="true" pageId="21" pageNumber="222">pauperata</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA11257246F9610DDF84B426A1" bold="true" box="[343,416,1501,1525]" pageId="21" pageNumber="222">
<taxonomicNameLabel id="27AD579111257246F9610DDF84B426A1" box="[343,416,1501,1525]" pageId="21" pageNumber="222">n. stat.</taxonomicNameLabel>
</emphasis>
, Peru, Zamora, prep. genit. H_246
</paragraph>
</caption>
<paragraph id="0E5536F811257246F8F10E21870125D1" blockId="21.[151,1437,1571,2029]" pageId="21" pageNumber="222">
<materialsCitation id="BE823CA511257246F8F10E21870125D1" ID-GBIF-Occurrence="3413048302" collectingDate="2010-10-13" collectionCode="R" collectorName="Willmott" country="Ecuador" elevation="2020" latitude="-4.799722" location="San Andres-Jimbura rd." longLatPrecision="21" longitude="-79.305" pageId="21" pageNumber="222" specimenCount="1" specimenCount-female="1" stateProvince="Zamora-Chinchipe" typeStatus="paratype">
<emphasis id="3C9EEAEA11257246F8F10E21871B2569" bold="true" box="[199,527,1571,1597]" pageId="21" pageNumber="222">Other specimens examined</emphasis>
(not considered
<typeStatus id="D151885A11257246FAE20E2186512569" box="[724,837,1571,1597]" pageId="21" pageNumber="222" type="paratype">paratypes</typeStatus>
):
<collectingCountry id="76FD766811257246FB6C0E2186F22569" box="[858,998,1571,1597]" name="Ecuador" pageId="21" pageNumber="222">ECUADOR</collectingCountry>
:
<specimenCount id="18ECFD7111257246FBC50E2181372569" box="[1011,1059,1571,1597]" pageId="21" pageNumber="222" type="female">1 ♀</specimenCount>
:
<collectingRegion id="CC2EF81A11257246FC050E21801D2569" box="[1075,1289,1571,1597]" country="Ecuador" name="Zamora-Chinchipe" pageId="21" pageNumber="222">Zamora-Chinchipe</collectingRegion>
, km 4.3
<location id="0B35602311257246FD440E2184612535" LSID="urn:lsid:plazi:treatment:864387EE113F7245F8A10DC884A22041:0B35602311257246FD440E2184612535" country="Ecuador" latitude="-4.799722" longLatPrecision="21" longitude="-79.305" name="San Andres-Jimbura rd." pageId="21" pageNumber="222" stateProvince="Zamora-Chinchipe">San Andrés-Jimbura rd.</location>
, [
<geoCoordinate id="6BDE503F11257246F9BF0E4484E82535" box="[393,508,1606,1633]" degrees="4" direction="south" minutes="47" orientation="latitude" pageId="21" pageNumber="222" precision="15" seconds="59" value="-4.799722">4°47'59''S</geoCoordinate>
,
<geoCoordinate id="6BDE503F11257246FA370E44879B2535" box="[513,655,1606,1633]" degrees="79" direction="west" minutes="18" orientation="longitude" pageId="21" pageNumber="222" precision="15" seconds="18" value="-79.305">79°18'18''W</geoCoordinate>
],
<quantity id="C9129B1D11257246FA950E4587EF2535" box="[675,763,1607,1633]" metricMagnitude="3" metricUnit="m" metricValue="2.02" pageId="21" pageNumber="222" unit="m" value="2020.0">
<elevation id="85C7D1CB11257246FA950E4587EF2535" box="[675,763,1607,1633]" metricMagnitude="3" metricUnit="m" metricValue="2.02" pageId="21" pageNumber="222" unit="m" value="2020.0">2020 m</elevation>
</quantity>
,
<date id="7A54103811257246FB3E0E45866E2535" box="[776,890,1607,1633]" pageId="21" pageNumber="222" value="2010-10-13">
<collectingDate id="6A10E9D011257246FB3E0E45866E2535" box="[776,890,1607,1633]" pageId="21" pageNumber="222" value="2010-10-13">13.x.2010</collectingDate>
</date>
, K.
<collectionCode id="68FBAE3D11257246FB9D0E4586D52535" box="[939,961,1607,1633]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="21" pageNumber="222">R</collectionCode>
.
<collectorName id="A31F532E11257246FBFB0E4581252535" box="[973,1073,1607,1633]" pageId="21" pageNumber="222">Willmott</collectorName>
leg., [FLMNH-MGCL-145842; dissection KW-20-022],
<collectionCode id="68FBAE3D11257246F99D0E69870525D1" box="[427,529,1643,1669]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="21" pageNumber="222" type="Museum">FLMNH</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="46F0657311257246F8F10E8D864F2599" pageId="21" pageNumber="222" type="etymology">
<paragraph id="0E5536F811257246F8F10E8D864F2599" blockId="21.[151,1437,1571,2029]" pageId="21" pageNumber="222">
<emphasis id="3C9EEAEA11257246F8F10E8D845A25FD" bold="true" box="[199,334,1679,1705]" pageId="21" pageNumber="222">Etymology.</emphasis>
The species name is the Latin word for garnet, in reference to the deep reddish brown colours of this species, and it is treated as a masculine noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="46F0657311257245F8F10ED584A22041" lastPageId="22" lastPageNumber="223" pageId="21" pageNumber="222" type="discussion">
<paragraph id="0E5536F811257245F8F10ED580702169" blockId="21.[151,1437,1571,2029]" lastBlockId="22.[151,1437,151,789]" lastPageId="22" lastPageNumber="223" pageId="21" pageNumber="222">
<emphasis id="3C9EEAEA11257246F8F10ED5842F25A5" bold="true" box="[199,315,1751,1777]" pageId="21" pageNumber="222">Remarks.</emphasis>
This species is externally most similar to
<taxonomicName id="C9EA4D7B11257246FB150EDA868A25A5" authority="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz, 2021" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[803,926,1751,1777]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="222" phylum="Arthropoda" rank="species" species="placida" status="sp. nov.">
<emphasis id="3C9EEAEA11257246FB150EDA862E25A5" box="[803,826,1752,1777]" italics="true" pageId="21" pageNumber="222">M</emphasis>
.
<emphasis id="3C9EEAEA11257246FB7C0ED5868A25A5" box="[842,926,1751,1777]" italics="true" pageId="21" pageNumber="222">placida</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA11257246FB910ED586FC25A5" bold="true" box="[935,1000,1751,1777]" pageId="21" pageNumber="222">
<taxonomicNameLabel id="27AD579111257246FB910ED586FC25A5" box="[935,1000,1751,1777]" pageId="21" pageNumber="222" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, although there are a number of wing pattern differences as mentioned in the Diagnosis that are consistent in all examined specimens. Nevertheless, we initially considered that the two taxa might represent subspecies of a single species, given that no males have been collected in sympatry, and the lack of substantial differences in the dorsal androconial scales or male genitalia. However, the DNA barcode of a single female from the Jimbura-San Andrés road in southern
<collectingCountry id="76FD766811257246FC930F65801624D5" box="[1189,1282,1895,1921]" name="Ecuador" pageId="21" pageNumber="222">Ecuador</collectingCountry>
grouped with those of males of
<taxonomicName id="C9EA4D7B11257246F9690F8E84FD24F1" authority="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz, 2021" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[351,489,1931,1957]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="222" phylum="Arthropoda" rank="species" species="granatus" status="sp. nov.">
<emphasis id="3C9EEAEA11257246F9690F8E846224F1" box="[351,374,1932,1957]" italics="true" pageId="21" pageNumber="222">M</emphasis>
.
<emphasis id="3C9EEAEA11257246F9B30F8E84FD24F1" box="[389,489,1932,1957]" italics="true" pageId="21" pageNumber="222">granatus</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA11257246F9C60F89873A24F1" bold="true" box="[496,558,1931,1957]" pageId="21" pageNumber="222">
<taxonomicNameLabel id="27AD579111257246F9C60F89873A24F1" box="[496,558,1931,1957]" pageId="21" pageNumber="222" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
from the Cordillera del Cóndor, based on which it is tentatively considered as belonging to this species, although we exclude it from the type series. This female specimen also has reddish brown surrounding the HWV black ocellus in Cu2-Cu1, as in males of
<taxonomicName id="C9EA4D7B11257246FB5F0FD686E724B9" authority="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz, 2021" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[873,1011,2003,2029]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="222" phylum="Arthropoda" rank="species" species="granatus" status="sp. nov.">
<emphasis id="3C9EEAEA11257246FB5F0FD6869424B9" box="[873,896,2004,2029]" italics="true" pageId="21" pageNumber="222">M</emphasis>
.
<emphasis id="3C9EEAEA11257246FBB90FD686E724B9" box="[911,1011,2004,2029]" italics="true" pageId="21" pageNumber="222">granatus</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA11257246FBCC0FD1812D24B9" bold="true" box="[1018,1081,2003,2029]" pageId="21" pageNumber="222">
<taxonomicNameLabel id="27AD579111257246FBCC0FD1812D24B9" box="[1018,1081,2003,2029]" pageId="21" pageNumber="222" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
but in contrast to the yellowish brown ring around the ocellus in
<taxonomicName id="C9EA4D7B11267245FA3D0895879023E5" authority="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz, 2021" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[523,644,151,177]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="placida" status="sp. nov.">
<emphasis id="3C9EEAEA11267245FA3D0895873623E4" box="[523,546,151,176]" italics="true" pageId="22" pageNumber="223">M</emphasis>
.
<emphasis id="3C9EEAEA11267245FA060895879023E5" box="[560,644,151,177]" italics="true" pageId="22" pageNumber="223">placida</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA11267245FABD089587DE23E5" bold="true" box="[651,714,151,177]" pageId="22" pageNumber="223">
<taxonomicNameLabel id="27AD579111267245FABD089587DE23E5" box="[651,714,151,177]" pageId="22" pageNumber="223" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="C9EA4D7B11267245FB370895866E23E5" authorityName="Pyrcz" authorityYear="2004" box="[769,890,151,177]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="benigni">
<emphasis id="3C9EEAEA11267245FB370895860C23E4" box="[769,792,151,176]" italics="true" pageId="22" pageNumber="223">M</emphasis>
.
<emphasis id="3C9EEAEA11267245FB100895866E23E5" box="[806,890,151,177]" italics="true" pageId="22" pageNumber="223">benigni</emphasis>
</taxonomicName>
. Assuming this female is conspecific with males from the Cordillera del Cóndor,
<taxonomicName id="C9EA4D7B11267245FA3708B9879E2380" authority="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz, 2021" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[513,650,187,213]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="granatus" status="sp. nov.">
<emphasis id="3C9EEAEA11267245FA3708B9870C2380" box="[513,536,187,212]" italics="true" pageId="22" pageNumber="223">M</emphasis>
.
<emphasis id="3C9EEAEA11267245FA1008B9879E2380" box="[550,650,187,212]" italics="true" pageId="22" pageNumber="223">granatus</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA11267245FAA708B987C42381" bold="true" box="[657,720,187,213]" pageId="22" pageNumber="223">
<taxonomicNameLabel id="27AD579111267245FAA708B987C42381" box="[657,720,187,213]" pageId="22" pageNumber="223" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
may be relatively widespread in southern
<collectingCountry id="76FD766811267245FC9D08B9801E2381" box="[1195,1290,187,213]" name="Ecuador" pageId="22" pageNumber="223">Ecuador</collectingCountry>
, and perhaps even sympatric or locally elevationally parapatric with
<taxonomicName id="C9EA4D7B11267245FB3608DD866D23AD" authority="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz, 2021" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[768,889,223,249]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="placida" status="sp. nov.">
<emphasis id="3C9EEAEA11267245FB3608DD860323AC" box="[768,791,223,248]" italics="true" pageId="22" pageNumber="223">M</emphasis>
.
<emphasis id="3C9EEAEA11267245FB1308DD866D23AD" box="[805,889,223,249]" italics="true" pageId="22" pageNumber="223">placida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="27AD579111267245FBB608DD86AB23AD" box="[896,959,223,249]" pageId="22" pageNumber="223" rank="species">
<emphasis id="3C9EEAEA11267245FBB608DD86A323AD" bold="true" box="[896,951,223,249]" pageId="22" pageNumber="223">n. sp</emphasis>
.
</taxonomicNameLabel>
However, so far no males are known from the area where the putative female of
<taxonomicName id="C9EA4D7B11267245FA01090187AB2248" authority="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz, 2021" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[567,703,259,285]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="granatus" status="sp. nov.">
<emphasis id="3C9EEAEA11267245FA010901875A2248" box="[567,590,259,284]" italics="true" pageId="22" pageNumber="223">M</emphasis>
.
<emphasis id="3C9EEAEA11267245FA6D090187AB2248" box="[603,703,259,284]" italics="true" pageId="22" pageNumber="223">granatus</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA11267245FAF3090186162249" bold="true" box="[709,770,259,285]" pageId="22" pageNumber="223">
<taxonomicNameLabel id="27AD579111267245FAF3090186162249" box="[709,770,259,285]" pageId="22" pageNumber="223" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
was collected. In addition, aside from the rather remarkable differences in wing pattern exhibited between the pierid
<taxonomicName id="C9EA4D7B11267245FB120925808C2215" authority="Radford &amp; Willmott, 2013" authorityName="Radford &amp; Willmott" authorityYear="2013" box="[804,1432,295,321]" class="Insecta" family="Pieridae" genus="Catasticta" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="subSpecies" species="poujadei" subSpecies="condor">
<emphasis id="3C9EEAEA11267245FB120925814F2215" box="[804,1115,295,321]" italics="true" pageId="22" pageNumber="223">Catasticta poujadei condor</emphasis>
Radford &amp; Willmott, 2013
</taxonomicName>
, from the Cordillera del Cóndor, and the nominate subspecies in the adjacent Andes, there are few examples of butterfly species with different subspecies in these two regions.
<taxonomicName id="C9EA4D7B11267245FB44096D86E922DC" authority="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz, 2021" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[882,1021,367,393]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="granatus" status="sp. nov.">
<emphasis id="3C9EEAEA11267245FB44096D869D22DC" box="[882,905,367,392]" italics="true" pageId="22" pageNumber="223">M</emphasis>
.
<emphasis id="3C9EEAEA11267245FBAF096D86E922DC" box="[921,1021,367,392]" italics="true" pageId="22" pageNumber="223">granatus</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA11267245FC30096D815222DD" bold="true" box="[1030,1094,367,393]" pageId="22" pageNumber="223">
<taxonomicNameLabel id="27AD579111267245FC30096D815222DD" box="[1030,1094,367,393]" pageId="22" pageNumber="223" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
was also detected in northern
<collectingCountry id="76FD766811267245F8A1099185D922F9" box="[151,205,403,429]" name="Peru" pageId="22" pageNumber="223">Peru</collectingCountry>
, in the highlands of Chachapoyas, where it was originally mistaken for
<taxonomicName id="C9EA4D7B11267245FBD20991814822F9" authorityName="Pyrcz" authorityYear="2004" box="[996,1116,403,429]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="benigni">
<emphasis id="3C9EEAEA11267245FBD2099186EF22F8" box="[996,1019,403,428]" italics="true" pageId="22" pageNumber="223">M</emphasis>
.
<emphasis id="3C9EEAEA11267245FC3E0991814822F9" box="[1032,1116,403,429]" italics="true" pageId="22" pageNumber="223">benigni</emphasis>
</taxonomicName>
, and three among the known Peruvian specimens were actually included as
<typeStatus id="D151885A11267245FA8109B586362285" box="[695,802,439,465]" pageId="22" pageNumber="223" type="paratype">paratypes</typeStatus>
of that species (
<bibRefCitation id="6A7B4B0911267245FBD109B581642285" author="Pyrcz, T. W." box="[999,1136,439,465]" pageId="22" pageNumber="223" pagination="455 - 622" refId="ref22314" refString="Pyrcz, T. W. (2004) Pronophiline butterflies of the highlands of Chachapoyas in northern Peru: faunal survey, diversity and distribution patterns (Lepidoptera, Nymphalidae, Satyrinae). Genus, 15, 455 - 622." type="journal article" year="2004">Pyrcz 2004</bibRefCitation>
). However, their genitalia match the specimens from the type locality of
<taxonomicName id="C9EA4D7B11267245FA9709D9863F22A0" authority="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz, 2021" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[673,811,475,501]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="granatus" status="sp. nov.">
<emphasis id="3C9EEAEA11267245FA9709D987AC22A0" box="[673,696,475,500]" italics="true" pageId="22" pageNumber="223">M</emphasis>
.
<emphasis id="3C9EEAEA11267245FAF109D9863F22A0" box="[711,811,475,500]" italics="true" pageId="22" pageNumber="223">granatus</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA11267245FB0409D9866522A1" bold="true" box="[818,881,475,501]" pageId="22" pageNumber="223">
<taxonomicNameLabel id="27AD579111267245FB0409D9866522A1" box="[818,881,475,501]" pageId="22" pageNumber="223" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Finally, the divergence in DNA barcode between
<taxonomicName id="C9EA4D7B11267245F8A109FD8436214C" authority="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz, 2021" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[151,290,511,537]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="granatus" status="sp. nov.">
<emphasis id="3C9EEAEA11267245F8A109FD85BA214C" box="[151,174,511,536]" italics="true" pageId="22" pageNumber="223">M</emphasis>
.
<emphasis id="3C9EEAEA11267245F88809FD8436214C" box="[190,290,511,536]" italics="true" pageId="22" pageNumber="223">granatus</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA11267245F91F09FD847D214D" bold="true" box="[297,361,511,537]" pageId="22" pageNumber="223">
<taxonomicNameLabel id="27AD579111267245F91F09FD847D214D" box="[297,361,511,537]" pageId="22" pageNumber="223" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="C9EA4D7B11267245F99409FD8708214D" authority="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz, 2021" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[418,540,511,537]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="placida" status="sp. nov.">
<emphasis id="3C9EEAEA11267245F99409FD84AD214C" box="[418,441,511,536]" italics="true" pageId="22" pageNumber="223">M</emphasis>
.
<emphasis id="3C9EEAEA11267245F9FE09FD8708214D" box="[456,540,511,537]" italics="true" pageId="22" pageNumber="223">placida</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA11267245FA1209FD8770214D" bold="true" box="[548,612,511,537]" pageId="22" pageNumber="223">
<taxonomicNameLabel id="27AD579111267245FA1209FD8770214D" box="[548,612,511,537]" pageId="22" pageNumber="223" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is comparable or even higher than that between other related
<taxonomicName id="C9EA4D7B11267245FD1409FD85E42169" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="3C9EEAEA11267245FD1409FD8088214D" box="[1314,1436,511,537]" italics="true" pageId="22" pageNumber="223">Manerebia</emphasis>
species.
</taxonomicName>
In summary, we consider that
<taxonomicName id="C9EA4D7B11267245FA710A2187C52168" authority="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz, 2021" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[583,721,547,573]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="granatus" status="sp. nov.">
<emphasis id="3C9EEAEA11267245FA710A21874A2168" box="[583,606,547,572]" italics="true" pageId="22" pageNumber="223">M</emphasis>
.
<emphasis id="3C9EEAEA11267245FA5B0A2187C52168" box="[621,721,547,572]" italics="true" pageId="22" pageNumber="223">granatus</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA11267245FAE10A2186022169" bold="true" box="[727,790,547,573]" pageId="22" pageNumber="223">
<taxonomicNameLabel id="27AD579111267245FAE10A2186022169" box="[727,790,547,573]" pageId="22" pageNumber="223" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="C9EA4D7B11267245FB7B0A2186D22169" authority="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz, 2021" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[845,966,547,573]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="placida" status="sp. nov.">
<emphasis id="3C9EEAEA11267245FB7B0A2186702168" box="[845,868,547,572]" italics="true" pageId="22" pageNumber="223">M</emphasis>
.
<emphasis id="3C9EEAEA11267245FB440A2186D22169" box="[882,966,547,573]" italics="true" pageId="22" pageNumber="223">placida</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA11267245FBFB0A2181182169" bold="true" box="[973,1036,547,573]" pageId="22" pageNumber="223">
<taxonomicNameLabel id="27AD579111267245FBFB0A2181182169" box="[973,1036,547,573]" pageId="22" pageNumber="223" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
represent two distinct species.
</paragraph>
<paragraph id="0E5536F811267245F8F10A4584A22041" blockId="22.[151,1437,151,789]" pageId="22" pageNumber="223">
Most known individuals of
<taxonomicName id="C9EA4D7B11267245FA340A4587992134" authority="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz, 2021" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" box="[514,653,583,608]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="granatus" status="sp. nov.">
<emphasis id="3C9EEAEA11267245FA340A4587992134" box="[514,653,583,608]" italics="true" pageId="22" pageNumber="223">M. granatus</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA11267245FAA30A4587C12135" bold="true" box="[661,725,583,609]" pageId="22" pageNumber="223">
<taxonomicNameLabel id="27AD579111267245FAA30A4587C12135" box="[661,725,583,609]" pageId="22" pageNumber="223" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
were collected at the
<typeStatus id="D151885A11267245FBE40A4581162135" box="[978,1026,583,609]" pageId="22" pageNumber="223">type</typeStatus>
locality, a sandstone tepui in southeastern
<collectingCountry id="76FD766811267245F8D90A69845A21D1" box="[239,334,619,645]" name="Ecuador" pageId="22" pageNumber="223">Ecuador</collectingCountry>
, along a steeply climbing trail through cloud forest from
<quantity id="C9129B1D11267245FBD50A69819721D1" box="[995,1155,619,645]" metricMagnitude="3" metricUnit="m" metricValue="2.055" metricValueMax="2.1" metricValueMin="2.01" pageId="22" pageNumber="223" unit="m" value="2055.0" valueMax="2100.0" valueMin="2010.0">20102100 m</quantity>
. A single individual was collected on the top of the tepui in stunted elfin forest near
<quantity id="C9129B1D11267245FB740A8D868821FD" box="[834,924,655,681]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="22" pageNumber="223" unit="m" value="2300.0">2300 m</quantity>
. A single female that may also represent this species (discussed above) was collected flying
<quantity id="C9129B1D11267245FA990AB187C82199" box="[687,732,691,717]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="22" pageNumber="223" unit="m" value="1.0">1 m</quantity>
above the ground along the edge of a dirt road through cloud forest. Other congeners present at the
<typeStatus id="D151885A11267245FA7B0AD5876921A5" box="[589,637,727,753]" pageId="22" pageNumber="223">type</typeStatus>
locality included
<taxonomicName id="C9EA4D7B11267245FB7B0AD581C121A5" authority="Pyrcz, 2004" authorityName="Pyrcz" authorityYear="2004" box="[845,1237,727,753]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="subSpecies" species="benigni" subSpecies="tessmanni">
<emphasis id="3C9EEAEA11267245FB7B0AD5867021A4" box="[845,868,727,752]" italics="true" pageId="22" pageNumber="223">M</emphasis>
.
<emphasis id="3C9EEAEA11267245FB420AD5815521A4" box="[884,1089,727,753]" italics="true" pageId="22" pageNumber="223">benigni tessmanni</emphasis>
<bibRefCitation id="6A7B4B0911267245FC7C0AD581C121A5" author="Pyrcz, T. W." box="[1098,1237,727,753]" pageId="22" pageNumber="223" pagination="455 - 622" refId="ref22314" refString="Pyrcz, T. W. (2004) Pronophiline butterflies of the highlands of Chachapoyas in northern Peru: faunal survey, diversity and distribution patterns (Lepidoptera, Nymphalidae, Satyrinae). Genus, 15, 455 - 622." type="journal article" year="2004">Pyrcz, 2004</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="C9EA4D7B11267245FCD40AD5806821A4" authorityName="Mahecha-J &amp; Florczyk &amp; Willmott &amp; Cerdeña &amp; Zubek &amp; Boyer &amp; Farfán &amp; Lachowska-Cierlik &amp; Pyrcz" authorityYear="2021" baseAuthorityName="Pyrcz &amp; Willmott" baseAuthorityYear="2006" box="[1250,1404,727,753]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="pauperata" status="stat. nov.">
<emphasis id="3C9EEAEA11267245FCD40AD581ED21A4" box="[1250,1273,727,752]" italics="true" pageId="22" pageNumber="223">M</emphasis>
.
<emphasis id="3C9EEAEA11267245FD3F0AD5806821A4" box="[1289,1404,727,752]" italics="true" pageId="22" pageNumber="223">pauperata</emphasis>
</taxonomicName>
<emphasis id="3C9EEAEA11267245FDB00AD585DE2041" bold="true" pageId="22" pageNumber="223">
<taxonomicNameLabel id="27AD579111267245FDB00AD585DE2041" pageId="22" pageNumber="223">n. stat.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="C9EA4D7B11267245F9310AF984BB2041" baseAuthorityName="Hewitson" baseAuthorityYear="1870" box="[263,431,763,789]" class="Insecta" family="Nymphalidae" genus="Manerebia" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="trimaculata">
<emphasis id="3C9EEAEA11267245F9310AF9840A2040" box="[263,286,763,788]" italics="true" pageId="22" pageNumber="223">M</emphasis>
.
<emphasis id="3C9EEAEA11267245F91A0AF984BB2041" box="[300,431,763,789]" italics="true" pageId="22" pageNumber="223">trimaculata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>