216 lines
23 KiB
XML
216 lines
23 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1067.72369" ID-GBIF-Dataset="1165ef89-9b15-4c8a-a7a3-5a80ffae5903" ID-PMC="PMC8571248" ID-Pensoft-Pub="1313-2970-1067-19" ID-Pensoft-UUID="34D3B870A92C50819DE85FF5DDE19A1D" ID-PubMed="34759718" ID-ZooBank="AEAF44EE2CCD4E319CBCA4C5ACB0E2EB" ModsDocID="1313-2970-1067-19" checkinTime="1635566834375" checkinUser="pensoft" docAuthor="Srisonchai, Ruttapon, Sutcharit, Chirasak & Likhitrakarn, Natdanai" docDate="2021" docId="2680FE1494DB5F0E8A812CC2DA652065" docLanguage="en" docName="ZooKeys 1067: 19-56" docOrigin="ZooKeys 1067" docPubDate="2021-10-29" docSource="http://dx.doi.org/10.3897/zookeys.1067.72369" docTitle="Zephronia phrain Likhitrakarn & Golovatch 2021" docType="treatment" docVersion="4" id="34D3B870A92C50819DE85FF5DDE19A1D" lastPageNumber="19" masterDocId="34D3B870A92C50819DE85FF5DDE19A1D" masterDocTitle="The giant pill-millipede genus Zephronia Gray, 1832 from Thailand, with a redescription of Z. siamensis Hirst, 1907 and descriptions of three new species (Diplopoda, Sphaerotheriida, Zephroniidae)" masterLastPageNumber="56" masterPageNumber="19" pageNumber="19" updateTime="1668151042475" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The giant pill-millipede genus Zephronia Gray, 1832 from Thailand, with a redescription of Z. siamensis Hirst, 1907 and descriptions of three new species (Diplopoda, Sphaerotheriida, Zephroniidae)</mods:title>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Srisonchai, Ruttapon</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7142-0999</mods:nameIdentifier>
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<mods:affiliation>Department of Biology, Faculty of Science, Khon Kaen University, Khon Kaen, 40002, Thailand</mods:affiliation>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Likhitrakarn, Natdanai</mods:namePart>
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<mods:affiliation>Division of Plant Protection, Faculty of Agricultural Production, Maejo University, Chiang Mai, 50290, Thailand & Biodiversity and Utilization Research Center of Maejo University, Maejo University, Chiang Mai, 50290, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">kongerrrr@hotmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-10-29</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1067</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>19</mods:start>
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<mods:end>56</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1067.72369</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1067.72369</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1067-19</mods:identifier>
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<mods:identifier type="ZooBank">AEAF44EE2CCD4E319CBCA4C5ACB0E2EB</mods:identifier>
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<treatment ID-GBIF-Taxon="189635865" LSID="urn:lsid:plazi:treatment:2680FE1494DB5F0E8A812CC2DA652065" httpUri="http://treatment.plazi.org/id/2680FE1494DB5F0E8A812CC2DA652065" lastPageNumber="19" pageId="0" pageNumber="19">
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<subSubSection pageId="0" pageNumber="19" type="nomenclature">
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<paragraph pageId="0" pageNumber="19">
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<taxonomicName LSID="2680FE14-94DB-5F0E-8A81-2CC2DA652065" authority="Likhitrakarn & Golovatch, 2021" authorityName="Likhitrakarn & Golovatch" authorityYear="2021" class="Diplopoda" family="Zephroniidae" genus="Zephronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zephronia phrain" order="Sphaerotheriida" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="phrain">Zephronia phrain Likhitrakarn & Golovatch, 2021</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="description">
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<paragraph pageId="0" pageNumber="19">
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Photographs of living Zephronia spp. A-D Zephronia siamensis E, F Zephronia phrain. Photographs not to scale." figureDoi="10.3897/zookeys.1067.72369.figure1" httpUri="https://binary.pensoft.net/fig/604301" pageId="0" pageNumber="19">Figures 1E, F</figureCitation>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Zephronia phrain A-D male specimen from Phawor Shrine, Tak Province E, F female A right antenna, ventral view B the ninth left leg, posterior view C first coxae with stigmatic plates, posterior view D coxae of second legs with gonopores, posterior view E coxae and prefemur of second legs with vulvae, posterior view F subanal plate, ventral view. Abbreviations: cx = coxa, o = operculum, pre = prefemur, syn-cx = syncoxite, st-pl = stigmatic plate." figureDoi="10.3897/zookeys.1067.72369.figure5" httpUri="https://binary.pensoft.net/fig/604305" pageId="0" pageNumber="19">; 5</figureCitation>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Zephronia phrain, male specimen from Phawor Shrine, Tak Province A telopods, anterior view B anterior telopod, anterior view C posterior telopod, posterior view D right anterior and posterior telopods, ventral view. Abbreviations: cr-T = crenulated teeth, cx = coxa, ML = membranous lobe, syn-cx = syncoxite." figureDoi="10.3897/zookeys.1067.72369.figure6" httpUri="https://binary.pensoft.net/fig/604306" pageId="0" pageNumber="19">; 6</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="reference_group">
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<paragraph pageId="0" pageNumber="19">
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<taxonomicName authorityName="Likhitrakarn & Golovatch" authorityYear="2021" class="Diplopoda" family="Zephroniidae" genus="Zephronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zephronia phrain" order="Sphaerotheriida" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="phrain">Zephronia phrain</taxonomicName>
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Likhitrakarn & Golovatch, 2021 in
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<bibRefCitation author="Likhitrakarn, N" journalOrPublisher="Tropical Natural History" pageId="0" pageNumber="19" pagination="12 - 26" publicationUrl="https://li01.tci-thaijo.org/index.php/tnh/article/view/247730" refId="B15" refString="Likhitrakarn, N, Golovatch, SI, Srisonchai, R, Sutcharit, C, 2021. Two new species of the giant pill-millipede genus Zephronia Gray, 1832 from Thailand (Diplopoda: Sphaerotheriida: Zephroniidae). Tropical Natural History 21 (1): 12 - 26, https://li01.tci-thaijo.org/index.php/tnh/article/view/247730" title="Two new species of the giant pill-millipede genus Zephronia Gray, 1832 from Thailand (Diplopoda: Sphaerotheriida: Zephroniidae)." url="https://li01.tci-thaijo.org/index.php/tnh/article/view/247730" volume="21" year="2021">Likhitrakarn et al. 2021</bibRefCitation>
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: 19.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="materials_examined">
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<paragraph pageId="0" pageNumber="19">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<materialsCitation collectingDate="2016-10" collectorName="S. Panha, R. Srisonchai, C. Sutcharit, W. Siriwut" country="Thailand" elevation="694" latitude="16.771334" location="Phawor Shrine" longLatPrecision="14" longitude="98.68694" municipality="Mae Sot District" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Tak Province">
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<collectingCountry name="Thailand">Thailand</collectingCountry>
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-
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<collectingRegion country="Thailand" name="Tak">Tak Province</collectingRegion>
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•
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<specimenCount type="male">2 ♂♂</specimenCount>
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<specimenCount type="female">2 ♀♀</specimenCount>
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;
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<collectingMunicipality>Mae Sot District</collectingMunicipality>
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,
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<location LSID="urn:lsid:plazi:treatment:2680FE1494DB5F0E8A812CC2DA652065:A2B5F21F94B3C509A3E9E8F677263156" country="Thailand" latitude="16.771334" longLatPrecision="14" longitude="98.68694" municipality="Mae Sot District" name="Phawor Shrine" stateProvince="Tak Province">Phawor Shrine</location>
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;
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<geoCoordinate degrees="16" direction="north" minutes="46" orientation="latitude" precision="1" seconds="16.8" value="16.771334">16°46'16.8"N</geoCoordinate>
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,
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<geoCoordinate degrees="98" direction="east" minutes="41" orientation="longitude" precision="15" seconds="13" value="98.68694">98°41'13"E</geoCoordinate>
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;
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<elevation metricMagnitude="2" metricUnit="m" metricValue="6.94" unit="m" value="694.0">
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<quantity metricMagnitude="2" metricUnit="m" metricValue="6.94" unit="m" value="694.0">694 m</quantity>
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a.s.l.
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</elevation>
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;
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<collectingDate value="2016-10">October 2016</collectingDate>
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;
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<collectorName>S. Panha</collectorName>
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,
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<collectorName>R. Srisonchai</collectorName>
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,
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<collectorName>C. Sutcharit</collectorName>
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,
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<collectorName>W. Siriwut</collectorName>
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leg.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="description">
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<paragraph pageId="0" pageNumber="19">Description of some characters for a population in Tak Province.</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="19">Body length</emphasis>
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: Length in male 29.0-31.0 mm (holotype 33.5 mm), female 30.0-33.0 mm; head 5.5 mm; thoracic shield 5.5-6.0 mm; anal shield 9.5-10.5 mm.
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="19">Body width</emphasis>
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: Width in male 16.5 mm (holotype 18.2 mm), female 16.5-17.0 mm; head 8.0-9.0 mm; thoracic shield 15.0-16.0 mm; anal shield 14.0-15.5 mm.
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="19">Body height</emphasis>
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: Height in male 10.0 mm (holotype 11.2 mm), female 10.0-11.0 mm; thoracic shield 9.0-10.5 mm; tergite 9.5-11.0 mm.
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="19">Color</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Photographs of living Zephronia spp. A-D Zephronia siamensis E, F Zephronia phrain. Photographs not to scale." figureDoi="10.3897/zookeys.1067.72369.figure1" httpUri="https://binary.pensoft.net/fig/604301" pageId="0" pageNumber="19">1E, F</figureCitation>
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): Specimens in life with brown or dark brown; head, antennae and collum, thoracic shield, paratergites, anal shield and legs brown or dark brown; anterior margins of thoracic shield, of tergites and of anal shield dark brown contrasting with the posterior brown ones; setose part of thoracic shield with golden sheen. Color in alcohol after six years not changed.
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="19">Tergites</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Photographs of living Zephronia spp. A-D Zephronia siamensis E, F Zephronia phrain. Photographs not to scale." figureDoi="10.3897/zookeys.1067.72369.figure1" httpUri="https://binary.pensoft.net/fig/604301" pageId="0" pageNumber="19">1E, F</figureCitation>
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): Quite shiny; surface glabrous, with sparse, tiny, inconspicuous pits; tip of paratergite of midbody tergites curved, directed posteroventrad; anterior half of lateral margin covered with long and conspicuous setae.
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="19">Legs</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Zephronia phrain A-D male specimen from Phawor Shrine, Tak Province E, F female A right antenna, ventral view B the ninth left leg, posterior view C first coxae with stigmatic plates, posterior view D coxae of second legs with gonopores, posterior view E coxae and prefemur of second legs with vulvae, posterior view F subanal plate, ventral view. Abbreviations: cx = coxa, o = operculum, pre = prefemur, syn-cx = syncoxite, st-pl = stigmatic plate." figureDoi="10.3897/zookeys.1067.72369.figure5" httpUri="https://binary.pensoft.net/fig/604305" pageId="0" pageNumber="19">5B</figureCitation>
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): Leg-pairs 1 and 2 without apical spine. Leg-pair 1 with four ventral spines, leg-pair 2 with four or five ventral spines. Leg-pair 3 with seven or eight ventral spines and one or two apical spines. Leg-pair 4 with nine or ten ventral spines and two or three apical spines. Leg-pairs 5-19 with 9-11 ventral spines and 1-3 apical spines. Last two leg-pairs with eight or nine ventral spines, and one or two apical spines. In leg 9, femur ca. 1.7
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<normalizedToken originalValue="×">x</normalizedToken>
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, tarsus ca. 3.4
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<normalizedToken originalValue="×">x</normalizedToken>
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longer than wide. Length of tarsus> femur> prefemur> coxa> tibia ≥ postfemur. All podomeres densely setose. Coxa large, with dentate ridge marginally (coxal process). Coxal process absent in leg-pairs 1 and 2. Prefemur without teeth. Femur slightly extended mesally; mesal margin with very small, tiny, inconspicuous teeth.
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="19">Subanal plate</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Zephronia phrain A-D male specimen from Phawor Shrine, Tak Province E, F female A right antenna, ventral view B the ninth left leg, posterior view C first coxae with stigmatic plates, posterior view D coxae of second legs with gonopores, posterior view E coxae and prefemur of second legs with vulvae, posterior view F subanal plate, ventral view. Abbreviations: cx = coxa, o = operculum, pre = prefemur, syn-cx = syncoxite, st-pl = stigmatic plate." figureDoi="10.3897/zookeys.1067.72369.figure5" httpUri="https://binary.pensoft.net/fig/604305" pageId="0" pageNumber="19">5F</figureCitation>
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): Trapeziform, undivided; central margin (tip) truncate, narrow; lateral margin straight. Densely setose.
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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Head, antenna,
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<normalizedToken originalValue="Tömösváry’s">Toemoesvary's</normalizedToken>
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organ, gnathochilarium, stigmatic plates, laterotergites, collum, thoracic shield, endotergum, anal shield, male sexual characters, anterior telopods, posterior telopods, and female sexual characters: Same as the original description in
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<bibRefCitation author="Likhitrakarn, N" journalOrPublisher="Tropical Natural History" pageId="0" pageNumber="19" pagination="12 - 26" publicationUrl="https://li01.tci-thaijo.org/index.php/tnh/article/view/247730" refId="B15" refString="Likhitrakarn, N, Golovatch, SI, Srisonchai, R, Sutcharit, C, 2021. Two new species of the giant pill-millipede genus Zephronia Gray, 1832 from Thailand (Diplopoda: Sphaerotheriida: Zephroniidae). Tropical Natural History 21 (1): 12 - 26, https://li01.tci-thaijo.org/index.php/tnh/article/view/247730" title="Two new species of the giant pill-millipede genus Zephronia Gray, 1832 from Thailand (Diplopoda: Sphaerotheriida: Zephroniidae)." url="https://li01.tci-thaijo.org/index.php/tnh/article/view/247730" volume="21" year="2021">Likhitrakarn et al. 2021</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="distribution">
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<paragraph pageId="0" pageNumber="19">Distribution and habitats</paragraph>
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<paragraph pageId="0" pageNumber="19">
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(Figs
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<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Limestone habitats of some Zephronia spp. A type locality of Zephronia siamensis (Koh Srichang, Chonburi Province) B, C type locality of Zephronia enghoffi sp. nov. (Tham Phaya Nakharat, Khon Kaen Province) D Habitat of Zephronia phrain at Phawor shrine, Tak Province." figureDoi="10.3897/zookeys.1067.72369.figure15" httpUri="https://binary.pensoft.net/fig/604315" pageId="0" pageNumber="19">15D</figureCitation>
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,
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<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Known distribution of Zephronia spp. in Thailand. Red triangle = Zephronia lannaensis; green circle = Zephronia phrain; blue triangle = Zephronia enghoffi sp. nov.; sky blue square = Zephronia golovatchi sp. nov.; purple circle = Zephronia panhai sp. nov.; yellow diamond = Zephronia siamensis Hirst, 1907; orange square = Zephronia viridisoma." figureDoi="10.3897/zookeys.1067.72369.figure16" httpUri="https://binary.pensoft.net/fig/604316" pageId="0" pageNumber="19">16</figureCitation>
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). Currently known to occur in northern Thailand (Chiang Mai and Tak provinces) in dry dipterocarp forest and from limestone areas. Observations made at Phawor Shrine found that most specimens were seen creeping on rocks, with some hiding in leaf litter. Notably, the specimens were found in syntopy with the dragon millipede (
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<taxonomicName authorityName="Srisonchai, Enghoff & Panha" authorityYear="2018" class="Diplopoda" family="Paradoxosomatidae" genus="Nagaxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nagaxytes spatula" order="Polydesmida" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="spatula">
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<emphasis italics="true" pageId="0" pageNumber="19">Nagaxytes spatula</emphasis>
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</taxonomicName>
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Srisonchai, Enghoff & Panha, 2018) at the same site (
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<bibRefCitation DOI="https://doi.org/10.5852/ejt.2018.462" author="Srisonchai, R" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="19" pagination="1 - 44" refId="B24" refString="Srisonchai, R, Enghoff, H, Likhitrakarn, N, Panha, S, 2018b. A revision of dragon millipedes II: The new genus Nagaxytes, with the description of three new species (Diplopoda, Polydesmida, Paradoxosomatidae). European Journal of Taxonomy 462: 1 - 44, DOI: https://doi.org/10.5852/ejt.2018.462" title="A revision of dragon millipedes II: The new genus Nagaxytes, with the description of three new species (Diplopoda, Polydesmida, Paradoxosomatidae)." url="https://doi.org/10.5852/ejt.2018.462" volume="462" year="2018 b">Srisonchai et al. 2018b</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="remarks">
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<paragraph pageId="0" pageNumber="19">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="19">
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Based on observations of live specimens in the field, two color patterns were found, dark green in type specimens and brown/dark brown in the others from Tak Province (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Photographs of living Zephronia spp. A, B Zephronia enghoffi sp. nov., paratypes (CUMZ-Zeph 0006) C, D Zephronia golovatchi sp. nov., paratypes (CUMZ-Zeph 0008) E, F Zephronia panhai sp. nov., paratypes (CUMZ-Zeph 0010). Photographs not to scale." figureDoi="10.3897/zookeys.1067.72369.figure2" httpUri="https://binary.pensoft.net/fig/604302" pageId="0" pageNumber="19">2E, F</figureCitation>
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; fig. 1C, D in
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<bibRefCitation author="Likhitrakarn, N" journalOrPublisher="Tropical Natural History" pageId="0" pageNumber="19" pagination="12 - 26" publicationUrl="https://li01.tci-thaijo.org/index.php/tnh/article/view/247730" refId="B15" refString="Likhitrakarn, N, Golovatch, SI, Srisonchai, R, Sutcharit, C, 2021. Two new species of the giant pill-millipede genus Zephronia Gray, 1832 from Thailand (Diplopoda: Sphaerotheriida: Zephroniidae). Tropical Natural History 21 (1): 12 - 26, https://li01.tci-thaijo.org/index.php/tnh/article/view/247730" title="Two new species of the giant pill-millipede genus Zephronia Gray, 1832 from Thailand (Diplopoda: Sphaerotheriida: Zephroniidae)." url="https://li01.tci-thaijo.org/index.php/tnh/article/view/247730" volume="21" year="2021">Likhitrakarn et al. 2021</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="19">
|
||
A species described by
|
||
<bibRefCitation author="Pocock, RI" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" pageId="0" pageNumber="19" pagination="384 - 395" refId="B20" refString="Pocock, RI, 1890. On the Myriopoda of Burma. Pt. 1. Report on the Oniscomorpha collected by Sig. L. Fea, by Mr. E.W. Oates and by the late Sig. G.B. Comotto. Annali del Museo Civico di Storia Naturale di Genova 30: 384 - 395" title="On the Myriopoda of Burma. Pt. 1. Report on the Oniscomorpha collected by Sig. L. Fea, by Mr. E. W. Oates and by the late Sig. G. B. Comotto." volume="30" year="1890">Pocock (1890)</bibRefCitation>
|
||
from Myanmar (
|
||
<normalizedToken originalValue="Thagatà">Thagata</normalizedToken>
|
||
, Mount Mooleyit, Kayah State),
|
||
<taxonomicName lsidName="Z. gestri" pageId="0" pageNumber="19" rank="species" species="gestri">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">Z. gestri</emphasis>
|
||
</taxonomicName>
|
||
Pocock, 1890 occurs close to the type locality of this widespread species, but
|
||
<taxonomicName lsidName="Z. phrain" pageId="0" pageNumber="19" rank="species" species="phrain">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">Z. phrain</emphasis>
|
||
</taxonomicName>
|
||
clearly differs from
|
||
<taxonomicName lsidName="Z. gestri" pageId="0" pageNumber="19" rank="species" species="gestri">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">Z. gestri</emphasis>
|
||
</taxonomicName>
|
||
by being longer in body length (vs. shorter, ca. 14 mm), having a longer immovable finger or longer process of telopoditomere 2 (vs. shorter) and having a truncate/round central margin of subanal plate (vs. convex).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="19">
|
||
Based on material described by
|
||
<bibRefCitation author="Likhitrakarn, N" journalOrPublisher="Tropical Natural History" pageId="0" pageNumber="19" pagination="12 - 26" publicationUrl="https://li01.tci-thaijo.org/index.php/tnh/article/view/247730" refId="B15" refString="Likhitrakarn, N, Golovatch, SI, Srisonchai, R, Sutcharit, C, 2021. Two new species of the giant pill-millipede genus Zephronia Gray, 1832 from Thailand (Diplopoda: Sphaerotheriida: Zephroniidae). Tropical Natural History 21 (1): 12 - 26, https://li01.tci-thaijo.org/index.php/tnh/article/view/247730" title="Two new species of the giant pill-millipede genus Zephronia Gray, 1832 from Thailand (Diplopoda: Sphaerotheriida: Zephroniidae)." url="https://li01.tci-thaijo.org/index.php/tnh/article/view/247730" volume="21" year="2021">Likhitrakarn et al. 2021</bibRefCitation>
|
||
, deposited in the CUMZ (holotype CUMZ-Zeph0003, paratypes CUMZ-Zeph0004).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |