244 lines
37 KiB
XML
244 lines
37 KiB
XML
<document id="561FCE767F9BCFB9F748F16D19A24303" ID-DOI="10.11646/zootaxa.1217.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5064796" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625283125096" checkinUser="felipe" docAuthor="Fuller, E. & Platia, G." docDate="2006" docId="0F7C9B43FFFDE956FE8DFC2F969BFA93" docLanguage="en" docName="zt01217p076.pdf" docOrigin="Zootaxa 1217 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1217.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Ctenoplus alutaceus Fuller & Platia 2006, n. sp." docType="treatment" docVersion="4" lastPageNumber="60" masterDocId="F345E33BFFC4E96DFF85FFFB9739FFFD" masterDocTitle="A revision of the click beetle genus Ctenoplus Candèze, 1863 (Coleoptera: Elateridae, Synaptina)" masterLastPageNumber="76" masterPageNumber="1" pageNumber="58" updateTime="1698986177451" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="4D257857F1DABD66FE658F1D5C1CEA61">A revision of the click beetle genus Ctenoplus Candèze, 1863 (Coleoptera: Elateridae, Synaptina)</mods:title>
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<mods:namePart id="6434CB046FB488541C91178ECCD6FCA0">Fuller, E.</mods:namePart>
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<mods:namePart id="0014FC19FBE6D8ED31963840537D46F6">Platia, G.</mods:namePart>
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<mods:title id="1CE90D7561AFCDFFA52EF7C8A0C704EF">Zootaxa</mods:title>
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<mods:part id="EC9C00E51D73C8114BB0C9A9F76C3E0D">
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<mods:date id="1817CB55DB71DD919845A7ED2F2C0E40">2006</mods:date>
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<mods:number id="AA87AE89C96AD547A85D78E16FB324C7">2006-05-26</mods:number>
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<mods:number id="4FF56FF8B36A89CC2E5D76863D3F7A63">1217</mods:number>
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<mods:url id="04FB24655EEB5DABBEACACF86E77921C">https://biotaxa.org/Zootaxa/article/view/zootaxa.1217.1.1</mods:url>
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<mods:classification id="C1A8B8D13C38BC8EC49FA28375764C63">journal article</mods:classification>
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<mods:identifier id="C7ACEC95380BD6EB20DAE17DB60E25FF" type="DOI">10.11646/zootaxa.1217.1.1</mods:identifier>
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<treatment id="0F7C9B43FFFDE956FE8DFC2F969BFA93" ID-DOI="http://doi.org/10.5281/zenodo.5066809" ID-Zenodo-Dep="5066809" LSID="urn:lsid:plazi:treatment:0F7C9B43FFFDE956FE8DFC2F969BFA93" httpUri="http://treatment.plazi.org/id/0F7C9B43FFFDE956FE8DFC2F969BFA93" lastPageId="59" lastPageNumber="60" pageId="57" pageNumber="58">
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<subSubSection id="CFCF79DEFFFDE954FE8DFC2F957FFC13" box="[264,582,980,1006]" pageId="57" pageNumber="58" type="nomenclature">
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<paragraph id="876A2A55FFFDE954FE8DFC2F957FFC13" blockId="57.[264,679,980,1041]" box="[264,582,980,1006]" pageId="57" pageNumber="58">
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<heading id="DC229D39FFFDE954FE8DFC2F957FFC13" bold="true" box="[264,582,980,1006]" fontSize="11" level="1" pageId="57" pageNumber="58" reason="1">
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<emphasis id="B5A1F647FFFDE954FE8DFC2F957FFC13" bold="true" box="[264,582,980,1006]" pageId="57" pageNumber="58">
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<taxonomicName id="40D551D6FFFDE954FE8DFC2F96C4FC13" authority="Fuller & Platia, 2006" authorityName="Fuller & Platia" authorityYear="2006" box="[264,509,980,1006]" class="Insecta" family="Elateridae" genus="Ctenoplus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="alutaceus" status="sp. nov.">
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<emphasis id="B5A1F647FFFDE954FE8DFC2F96C4FC13" bold="true" box="[264,509,980,1006]" italics="true" pageId="57" pageNumber="58">Ctenoplus alutaceus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="AE924B3CFFFDE954FD81FC2F957FFC13" box="[516,582,980,1006]" pageId="57" pageNumber="58" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="CFCF79DEFFFDE956FE8DFC0C969BFA93" lastPageId="59" lastPageNumber="60" pageId="57" pageNumber="58" type="description">
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<paragraph id="876A2A55FFFDE954FE8DFC0C959EFBEC" blockId="57.[264,679,980,1041]" box="[264,679,1015,1041]" pageId="57" pageNumber="58">
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<figureCitation id="1FEE36D0FFFDE954FE8DFC0C96BEFBEC" box="[264,391,1015,1041]" captionStart="FIGURES 15–30" captionStartId="11.[264,382,1146,1170]" captionTargetBox="[297,1282,330,1113]" captionTargetId="figure-245@11.[293,1583,326,1140]" captionTargetPageId="11" captionText="FIGURES 15–30. Antenna of Ctenoplus species. 15, C. javanensis Cand.; 16, C. sumatrensis n. sp.; 17, C. sanguinolentus (Cand.); 18, C. sanguinolentoides n. sp.; 19, C. nitidipennis Schwarz; 20, C. indicus n. sp.; 21, C. neosiamensis n. sp.; 22, C. siamensis n. sp.; 23, C. rufoantennatus n. sp.; 24, C. castaneus (Fleut.); 25, C. coomani Fleut.; 26, C. divergens n. sp.; 27, C. alutaceus n. sp.; 28, C. semialutaceus n. sp.; 29, C. cateianus n. sp.; 30, C. dorsalis n. sp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5064806" httpUri="https://zenodo.org/record/5064806/files/figure.png" pageId="57" pageNumber="58">Figures 27</figureCitation>
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,
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<figureCitation id="1FEE36D0FFFDE954FE16FC0C968AFBEC" box="[403,435,1015,1041]" captionStart="FIGURES 31–43" captionStartId="14.[264,382,1697,1721]" captionTargetBox="[369,1208,554,1659]" captionTargetId="figure-170@14.[359,1241,544,1699]" captionTargetPageId="14" captionText="FIGURES 31–43. Pronotal outline of Ctenoplus species, dorsal. 31, C. javanensis Cand.; 32, C. sumatrensis n. sp.; 33, C. sanguinolentoides n. sp.; 34, C. indicus n. sp.; 35, C. rufoantennatus n. sp.; 36, C. castaneus (Fleut.); 37, C. coomani Fleut.; 38, C. divergens n. sp., a: male, b: female; 39, C. alutaceus n. sp.; 40, C. semialutaceus n. sp.; 41, C. gigas n. sp.; 42, C. cateianus n. sp.; 43, C. dorsalis n. sp. Abbreviation: bf, sublateral longitudinal incision. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5064810" httpUri="https://zenodo.org/record/5064810/files/figure.png" pageId="57" pageNumber="58">39</figureCitation>
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,
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<figureCitation id="1FEE36D0FFFDE954FE3AFC0C96E6FBEC" box="[447,479,1015,1041]" captionStart="FIGURES 44–46" captionStartId="15.[264,382,1135,1159]" captionTargetBox="[267,1299,334,1105]" captionTargetId="figure-270@15.[264,1323,326,1125]" captionTargetPageId="15" captionText="FIGURES 44–46. posterior part of hypomeron of Ctenoplus species. 44, C. alutaceus n. sp.; 45, C. semialutaceus n. sp.; 46, C. girardianus n. sp. 47, prosternum of C. nitidipennis Schwarz, ventral. 48, venter of mesothorax of C. dorsalis n. sp., ventral. 49, tarsus of C. castaneus (Fleut.), lateral. Abbreviations: ac, alutaceous cuticle; ag, anterior groove of mesepisternum; as, anterior articulating surface of mesosternum; c, mesocoxa; d, lateral depression of mesosternum; gr, groove on prosternum; l, lobe of tarsomere 4; lp, anterolateral projection of mesepimeron; mf, mesosternal cavity; pr, projection of mesal border of hypomeron. Scale bar = 44–46, 48, 49: 0.25 mm; 48: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5064814" httpUri="https://zenodo.org/record/5064814/files/figure.png" pageId="57" pageNumber="58">44</figureCitation>
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,
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<figureCitation id="1FEE36D0FFFDE954FE6EFC0C9532FBEC" box="[491,523,1015,1041]" captionStart="FIGURES 50–57" captionStartId="19.[264,382,1487,1511]" captionTargetBox="[311,1265,365,1457]" captionTargetId="figure-138@19.[264,1323,330,1493]" captionTargetPageId="19" captionText="FIGURES 50–57. Prosternal spine of Ctenoplus and Silesis species. 50–57, lateral; 58, 59, ventral. 50, C. sanguinolentus (Cand.); 51, C. sanguinolentoides n. sp.; 52a, b, C. coomani Fleut.; 53, C. balli n. sp.; 54, C. gigas n. sp.; 55, C. divergens n. sp.; 56, C. alutaceus n. sp.; 57, C. dorsalis n. sp.; 58, C. brunneus Fleut.; 59, S. fulvus Fleut.; Abbreviations: da, dorsal apex; va, ventral apex. Scale bar = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5064816" httpUri="https://zenodo.org/record/5064816/files/figure.png" pageId="57" pageNumber="58">56</figureCitation>
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,
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<figureCitation id="1FEE36D0FFFDE954FD93FC0C950FFBEC" box="[534,566,1015,1041]" captionStart="FIGURES 66–85" captionStartId="22.[264,382,1424,1448]" captionTargetBox="[293,1279,326,1378]" captionTargetId="figure-207@22.[264,1323,326,1438]" captionTargetPageId="22" captionText="FIGURES 66–85. Male genitalia of Ctenoplus species, dorsal aspect unless otherwise specified. 66, C. javanensis Cand.; 67, C. sumatrensis n. sp.; 68, C. sanguinolentoides n. sp.; 69, C. nitidipennis Schwarz; 70, C. indicus n. sp.; 71, C. collaris Cand., a, dorsal, b, ventral; 72, C. pseudocollaris n. sp.; 73, C. neosiamensis n. sp.; 74, C. siamensis n. sp.; 75, C. rufoantennatus n. sp.; 76, C. castaneus (Fleut.), a, dorsal, b, lateral; 77, C. coomani Fleut.; 78, C. divergens n. sp.; 79, C. alutaceus n. sp., a, dorsal, b, lateral; 80, C. semialutaceus n. sp.; 81, C. balli n. sp.; 82, C. girardianus n. sp.; 83, C. gigas n. sp.; 84, C. cateianus n. sp.; 85, C. deceptus n. sp. Abbreviations: st, median struts; pa, parameres; ph, phallobase. Scale bar = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5064822" httpUri="https://zenodo.org/record/5064822/files/figure.png" pageId="57" pageNumber="58">79</figureCitation>
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,
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<figureCitation id="1FEE36D0FFFDE954FDC6FC0C9548FBEC" box="[579,625,1015,1041]" captionStart="FIGURES 98–106" captionStartId="30.[264,382,1757,1781]" captionTargetBox="[341,1253,875,1748]" captionTargetId="figure-246@30.[319,1281,865,1761]" captionTargetPageId="30" captionText="FIGURES 98–106. Female genitalia of Ctenoplus species. 98, C. coomani Fleut.; 99, C. brunneus Fleut.; 100, C. divergens n. sp.; 101, C. alutaceus n. sp.; 102, C. balli n. sp.; 103, C. girardianus n. sp.; 104, C. gigas n. sp.; 105, C. cateianus n. sp.; 106, C. deceptus n. sp. Abbreviations: sd, tubular extension. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5064828" httpUri="https://zenodo.org/record/5064828/files/figure.png" pageId="57" pageNumber="58">101</figureCitation>
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,
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<figureCitation id="1FEE36D0FFFDE954FDF8FC0C959EFBEC" box="[637,679,1015,1041]" captionStart="FIGURE 111" captionStartId="58.[264,367,1149,1173]" captionTargetBox="[340,1260,398,1129]" captionTargetId="figure-221@58.[340,1260,398,1162]" captionTargetPageId="58" captionText="FIGURE 111. Distribution of Ctenoplus species." figureDoi="http://doi.org/10.5281/zenodo.5064840" httpUri="https://zenodo.org/record/5064840/files/figure.png" pageId="57" pageNumber="58">111</figureCitation>
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</paragraph>
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<paragraph id="876A2A55FFFDE954FE8DFBB49646FB95" blockId="57.[264,1323,1103,1289]" box="[264,383,1103,1128]" pageId="57" pageNumber="58">
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<emphasis id="B5A1F647FFFDE954FE8DFBB49646FB95" box="[264,383,1103,1128]" italics="true" pageId="57" pageNumber="58">Diagnosis</emphasis>
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</paragraph>
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<paragraph id="876A2A55FFFDE954FEBDFB8C9584FAF4" blockId="57.[264,1323,1103,1289]" pageId="57" pageNumber="58">
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Small, uniformly yellowishbrown
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<taxonomicName id="40D551D6FFFDE954FD52FB8C9477FB6D" box="[727,846,1143,1168]" class="Insecta" family="Elateridae" genus="Ctenoplus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="genus">
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<emphasis id="B5A1F647FFFDE954FD52FB8C9477FB6D" box="[727,846,1143,1168]" italics="true" pageId="57" pageNumber="58">Ctenoplus</emphasis>
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</taxonomicName>
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with a broad patch of alutaceous cuticle on the posterolateral margin of the hypomeron (
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<figureCitation id="1FEE36D0FFFDE954FCCCFB64949BFB44" box="[841,930,1183,1209]" captionStart="FIGURES 44–46" captionStartId="15.[264,382,1135,1159]" captionTargetBox="[267,1299,334,1105]" captionTargetId="figure-270@15.[264,1323,326,1125]" captionTargetPageId="15" captionText="FIGURES 44–46. posterior part of hypomeron of Ctenoplus species. 44, C. alutaceus n. sp.; 45, C. semialutaceus n. sp.; 46, C. girardianus n. sp. 47, prosternum of C. nitidipennis Schwarz, ventral. 48, venter of mesothorax of C. dorsalis n. sp., ventral. 49, tarsus of C. castaneus (Fleut.), lateral. Abbreviations: ac, alutaceous cuticle; ag, anterior groove of mesepisternum; as, anterior articulating surface of mesosternum; c, mesocoxa; d, lateral depression of mesosternum; gr, groove on prosternum; l, lobe of tarsomere 4; lp, anterolateral projection of mesepimeron; mf, mesosternal cavity; pr, projection of mesal border of hypomeron. Scale bar = 44–46, 48, 49: 0.25 mm; 48: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5064814" httpUri="https://zenodo.org/record/5064814/files/figure.png" pageId="57" pageNumber="58">Fig. 44</figureCitation>
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) and with the pronotum broader than long (
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<figureCitation id="1FEE36D0FFFDE954FE08FB3C96D0FB1C" box="[397,489,1223,1249]" captionStart="FIGURES 31–43" captionStartId="14.[264,382,1697,1721]" captionTargetBox="[369,1208,554,1659]" captionTargetId="figure-170@14.[359,1241,544,1699]" captionTargetPageId="14" captionText="FIGURES 31–43. Pronotal outline of Ctenoplus species, dorsal. 31, C. javanensis Cand.; 32, C. sumatrensis n. sp.; 33, C. sanguinolentoides n. sp.; 34, C. indicus n. sp.; 35, C. rufoantennatus n. sp.; 36, C. castaneus (Fleut.); 37, C. coomani Fleut.; 38, C. divergens n. sp., a: male, b: female; 39, C. alutaceus n. sp.; 40, C. semialutaceus n. sp.; 41, C. gigas n. sp.; 42, C. cateianus n. sp.; 43, C. dorsalis n. sp. Abbreviation: bf, sublateral longitudinal incision. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5064810" httpUri="https://zenodo.org/record/5064810/files/figure.png" pageId="57" pageNumber="58">Fig. 39</figureCitation>
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).
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<taxonomicName id="40D551D6FFFDE954FD84FB3C95AEFB1D" authorityName="Fuller & Platia" authorityYear="2006" box="[513,663,1223,1248]" class="Insecta" family="Elateridae" genus="Ctenoplus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="alutaceus">
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<emphasis id="B5A1F647FFFDE954FD84FB3C95AEFB1D" box="[513,663,1223,1248]" italics="true" pageId="57" pageNumber="58">C. alutaceus</emphasis>
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</taxonomicName>
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is similar to
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<taxonomicName id="40D551D6FFFDE954FCBAFB3C94EFFB1D" authorityName="Fuller & Platia" authorityYear="2006" box="[831,982,1223,1248]" class="Insecta" family="Elateridae" genus="Ctenoplus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="divergens">
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<emphasis id="B5A1F647FFFDE954FCBAFB3C94EFFB1D" box="[831,982,1223,1248]" italics="true" pageId="57" pageNumber="58">C. divergens</emphasis>
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</taxonomicName>
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, but the latter species lacks alutaceous cuticle on the hypomeron.
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</paragraph>
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<paragraph id="876A2A55FFFDE954FE8DFAC496ABFAA5" blockId="57.[264,1324,1343,1889]" box="[264,402,1343,1368]" pageId="57" pageNumber="58">
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<emphasis id="B5A1F647FFFDE954FE8DFAC496ABFAA5" box="[264,402,1343,1368]" italics="true" pageId="57" pageNumber="58">Description</emphasis>
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</paragraph>
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<paragraph id="876A2A55FFFDE954FEBDFA9C941BFA7C" blockId="57.[264,1324,1343,1889]" box="[312,802,1383,1409]" pageId="57" pageNumber="58">
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As in
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<taxonomicName id="40D551D6FFFDE954FEF8FA9C9528FA7D" authorityName="Fuller & Platia" authorityYear="2006" box="[381,529,1383,1408]" class="Insecta" family="Elateridae" genus="Ctenoplus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="divergens">
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<emphasis id="B5A1F647FFFDE954FEF8FA9C9528FA7D" box="[381,529,1383,1408]" italics="true" pageId="57" pageNumber="58">C. divergens</emphasis>
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</taxonomicName>
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except as noted below.
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</paragraph>
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<paragraph id="876A2A55FFFDE954FEBDFA7494E4FA2C" blockId="57.[264,1324,1343,1889]" pageId="57" pageNumber="58">
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Body light yellowishbrown; antennae, legs and palpi light brownishyellow; setae opaque light yellow. Length: 6.0–
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<quantity id="402D87B0FFFDE954FD10FA4C95C8FA2C" box="[661,753,1463,1489]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="57" pageNumber="58" unit="mm" value="6.5">6.5 mm</quantity>
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; width 1.0–2.0 mm.
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</paragraph>
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<paragraph id="876A2A55FFFDE954FEBDFA2495EBF9DC" blockId="57.[264,1324,1343,1889]" pageId="57" pageNumber="58">
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Head. Punctures shallow, almost contiguous throughout; antenna (
|
||
<figureCitation id="1FEE36D0FFFDE954FBCBFA24939CFA04" box="[1102,1189,1503,1529]" captionStart="FIGURES 15–30" captionStartId="11.[264,382,1146,1170]" captionTargetBox="[297,1282,330,1113]" captionTargetId="figure-245@11.[293,1583,326,1140]" captionTargetPageId="11" captionText="FIGURES 15–30. Antenna of Ctenoplus species. 15, C. javanensis Cand.; 16, C. sumatrensis n. sp.; 17, C. sanguinolentus (Cand.); 18, C. sanguinolentoides n. sp.; 19, C. nitidipennis Schwarz; 20, C. indicus n. sp.; 21, C. neosiamensis n. sp.; 22, C. siamensis n. sp.; 23, C. rufoantennatus n. sp.; 24, C. castaneus (Fleut.); 25, C. coomani Fleut.; 26, C. divergens n. sp.; 27, C. alutaceus n. sp.; 28, C. semialutaceus n. sp.; 29, C. cateianus n. sp.; 30, C. dorsalis n. sp. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5064806" httpUri="https://zenodo.org/record/5064806/files/figure.png" pageId="57" pageNumber="58">Fig. 27</figureCitation>
|
||
) extending almost to apex of pronotal hind angles.
|
||
</paragraph>
|
||
<paragraph id="876A2A55FFFDE957FEBDF9D49428FEA3" blockId="57.[264,1324,1343,1889]" lastBlockId="58.[264,1323,284,350]" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58">
|
||
Prothorax. Pronotum (
|
||
<figureCitation id="1FEE36D0FFFDE954FDC1F9D49598F9B4" box="[580,673,1583,1609]" captionStart="FIGURES 31–43" captionStartId="14.[264,382,1697,1721]" captionTargetBox="[369,1208,554,1659]" captionTargetId="figure-170@14.[359,1241,544,1699]" captionTargetPageId="14" captionText="FIGURES 31–43. Pronotal outline of Ctenoplus species, dorsal. 31, C. javanensis Cand.; 32, C. sumatrensis n. sp.; 33, C. sanguinolentoides n. sp.; 34, C. indicus n. sp.; 35, C. rufoantennatus n. sp.; 36, C. castaneus (Fleut.); 37, C. coomani Fleut.; 38, C. divergens n. sp., a: male, b: female; 39, C. alutaceus n. sp.; 40, C. semialutaceus n. sp.; 41, C. gigas n. sp.; 42, C. cateianus n. sp.; 43, C. dorsalis n. sp. Abbreviation: bf, sublateral longitudinal incision. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5064810" httpUri="https://zenodo.org/record/5064810/files/figure.png" pageId="57" pageNumber="58">Fig. 39</figureCitation>
|
||
): punctures grading from separated by about 1/2 own diameters anteriorly to separated by about own diameters posteriorly, punctures shallow, simple; carina of hind angles slightly less than 1/2 length of pronotum in lateral aspect. Hypomeron: mesal margin bearing linear row of five punctures and with small rounded projection adjacent to procoxa; posterolateral margin with broad band of alutaceous cuticle (
|
||
<figureCitation id="1FEE36D0FFFDE954FE94F90C9653F8EC" box="[273,362,1783,1809]" captionStart="FIGURES 44–46" captionStartId="15.[264,382,1135,1159]" captionTargetBox="[267,1299,334,1105]" captionTargetId="figure-270@15.[264,1323,326,1125]" captionTargetPageId="15" captionText="FIGURES 44–46. posterior part of hypomeron of Ctenoplus species. 44, C. alutaceus n. sp.; 45, C. semialutaceus n. sp.; 46, C. girardianus n. sp. 47, prosternum of C. nitidipennis Schwarz, ventral. 48, venter of mesothorax of C. dorsalis n. sp., ventral. 49, tarsus of C. castaneus (Fleut.), lateral. Abbreviations: ac, alutaceous cuticle; ag, anterior groove of mesepisternum; as, anterior articulating surface of mesosternum; c, mesocoxa; d, lateral depression of mesosternum; gr, groove on prosternum; l, lobe of tarsomere 4; lp, anterolateral projection of mesepimeron; mf, mesosternal cavity; pr, projection of mesal border of hypomeron. Scale bar = 44–46, 48, 49: 0.25 mm; 48: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5064814" httpUri="https://zenodo.org/record/5064814/files/figure.png" pageId="57" pageNumber="58">Fig. 44</figureCitation>
|
||
), band separated from lateral margin by single row of punctures, band about 1/3 width of hypomeron at level of procoxa. Prosternum: punctures grading from almost contiguous anteriorly to separated by about own diameters around procoxa; prosternal spine (
|
||
<figureCitation id="1FEE36D0FFFEE957FEDEFEE79680FECB" box="[347,441,284,310]" captionStart="FIGURES 50–57" captionStartId="19.[264,382,1487,1511]" captionTargetBox="[311,1265,365,1457]" captionTargetId="figure-138@19.[264,1323,330,1493]" captionTargetPageId="19" captionText="FIGURES 50–57. Prosternal spine of Ctenoplus and Silesis species. 50–57, lateral; 58, 59, ventral. 50, C. sanguinolentus (Cand.); 51, C. sanguinolentoides n. sp.; 52a, b, C. coomani Fleut.; 53, C. balli n. sp.; 54, C. gigas n. sp.; 55, C. divergens n. sp.; 56, C. alutaceus n. sp.; 57, C. dorsalis n. sp.; 58, C. brunneus Fleut.; 59, S. fulvus Fleut.; Abbreviations: da, dorsal apex; va, ventral apex. Scale bar = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5064816" httpUri="https://zenodo.org/record/5064816/files/figure.png" pageId="58" pageNumber="59">Fig. 56</figureCitation>
|
||
): ventral margin concave in lateral aspect; ventrolateral carinae joining about 2/3 length of spine posterad procoxae.
|
||
</paragraph>
|
||
<caption id="D3AA7ADDFFFEE957FE8DFB86942CFB69" ID-DOI="http://doi.org/10.5281/zenodo.5064840" ID-Zenodo-Dep="5064840" box="[264,789,1149,1173]" httpUri="https://zenodo.org/record/5064840/files/figure.png" pageId="58" pageNumber="59" startId="58.[264,367,1149,1173]" targetBox="[340,1260,398,1129]" targetPageId="58">
|
||
<paragraph id="876A2A55FFFEE957FE8DFB86942CFB69" blockId="58.[264,789,1149,1173]" box="[264,789,1149,1173]" pageId="58" pageNumber="59">
|
||
<emphasis id="B5A1F647FFFEE957FE8DFB86969BFB68" bold="true" box="[264,418,1149,1173]" pageId="58" pageNumber="59">FIGURE 111.</emphasis>
|
||
Distribution of
|
||
<taxonomicName id="40D551D6FFFEE957FDC8FB86942CFB69" box="[589,789,1149,1172]" class="Insecta" family="Elateridae" genus="Ctenoplus" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B5A1F647FFFEE957FDC8FB869580FB69" box="[589,697,1149,1172]" italics="true" pageId="58" pageNumber="59">Ctenoplus</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="876A2A55FFFEE957FEBDFB3396BBFA07" blockId="58.[264,1323,1224,1890]" pageId="58" pageNumber="59">
|
||
Mesothorax. Scutellum flat; anterior margin convex; scutellum depressed below level of adjacent elytra anteriorly. Mesosternum moderately raised between mesocoxae, margin convex in lateral aspect, declivous beaded margin angled at
|
||
<emphasis id="B5A1F647FFFEE957FC59FAE394C7FACC" box="[988,1022,1304,1329]" italics="true" pageId="58" pageNumber="59">ca.</emphasis>
|
||
45°; posterior margin of mesosternum slightly concave; posterolateral angle glabrous. Mesepisternum glabrous on posterior 1/2. Elytra not attenuated posteriorly, broadly rounded at apex; apices subtruncate, without sutural teeth; striae shallowly impressed; intervals moderately densely punctured with minute punctures, punctures separated by about 2X own diameters.
|
||
</paragraph>
|
||
<paragraph id="876A2A55FFFEE957FEBDF9F39440F9B7" blockId="58.[264,1323,1224,1890]" pageId="58" pageNumber="59">Metathorax. Metasternum: carina posterad mesocoxa 3–5X as long as basal width; punctures separated by own diameters along midline.</paragraph>
|
||
<paragraph id="876A2A55FFFEE957FEBDF9A395BAF967" blockId="58.[264,1323,1224,1890]" pageId="58" pageNumber="59">Legs. Claws with 4 subapical tines, apical tine slightly longer than subapical tines; tarsomere 4 with indistinct lobe.</paragraph>
|
||
<paragraph id="876A2A55FFFEE956FEBDF9539590FDDB" blockId="58.[264,1323,1224,1890]" lastBlockId="59.[264,1323,284,550]" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59">
|
||
Abdomen. Terga not heavily sclerotized; spiracles marginal on terga 2–6 and appearing narrowly enclosed in cuticle laterally; dorsal carina of ventrite 5 broadly Ushaped. Male genitalia,
|
||
<figureCitation id="1FEE36D0FFFEE957FDA7F9039540F8EF" box="[546,633,1784,1810]" captionStart="FIGURES 66–85" captionStartId="22.[264,382,1424,1448]" captionTargetBox="[293,1279,326,1378]" captionTargetId="figure-207@22.[264,1323,326,1438]" captionTargetPageId="22" captionText="FIGURES 66–85. Male genitalia of Ctenoplus species, dorsal aspect unless otherwise specified. 66, C. javanensis Cand.; 67, C. sumatrensis n. sp.; 68, C. sanguinolentoides n. sp.; 69, C. nitidipennis Schwarz; 70, C. indicus n. sp.; 71, C. collaris Cand., a, dorsal, b, ventral; 72, C. pseudocollaris n. sp.; 73, C. neosiamensis n. sp.; 74, C. siamensis n. sp.; 75, C. rufoantennatus n. sp.; 76, C. castaneus (Fleut.), a, dorsal, b, lateral; 77, C. coomani Fleut.; 78, C. divergens n. sp.; 79, C. alutaceus n. sp., a, dorsal, b, lateral; 80, C. semialutaceus n. sp.; 81, C. balli n. sp.; 82, C. girardianus n. sp.; 83, C. gigas n. sp.; 84, C. cateianus n. sp.; 85, C. deceptus n. sp. Abbreviations: st, median struts; pa, parameres; ph, phallobase. Scale bar = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5064822" httpUri="https://zenodo.org/record/5064822/files/figure.png" pageId="58" pageNumber="59">Fig. 79</figureCitation>
|
||
; parameres not enclosing median lobe ventrally, separated by slightly less than width of median lobe across basal struts; median lobe evenly tapered from basal struts to apex, curved ventrally. Female genitalia,
|
||
<figureCitation id="1FEE36D0FFFEE957FB97F8B39344F89F" box="[1042,1149,1864,1890]" captionStart="FIGURES 98–106" captionStartId="30.[264,382,1757,1781]" captionTargetBox="[341,1253,875,1748]" captionTargetId="figure-246@30.[319,1281,865,1761]" captionTargetPageId="30" captionText="FIGURES 98–106. Female genitalia of Ctenoplus species. 98, C. coomani Fleut.; 99, C. brunneus Fleut.; 100, C. divergens n. sp.; 101, C. alutaceus n. sp.; 102, C. balli n. sp.; 103, C. girardianus n. sp.; 104, C. gigas n. sp.; 105, C. cateianus n. sp.; 106, C. deceptus n. sp. Abbreviations: sd, tubular extension. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5064828" httpUri="https://zenodo.org/record/5064828/files/figure.png" pageId="58" pageNumber="59">Fig. 101</figureCitation>
|
||
; bursal plates subcrescentic, symmetrical; band of bursal spines extending almost completely around bursa anterad base of tubular extension; tubular extension: base very wide, length of base about 1/2 length of bursa; base almost completely covered with pale spines, spines smaller and finer than bursal spines, grading to size of bursal spines, band of pale spines continuous around extension proximally, interrupted distally; distal bulbous spineless section of extension arising laterally on base; spermatheca absent; spermathecal gland tubular, with subreniform reservoir.
|
||
</paragraph>
|
||
<paragraph id="876A2A55FFFFE956FE8DFDA7955CFD88" blockId="59.[264,1323,604,790]" box="[264,613,604,629]" pageId="59" pageNumber="60">
|
||
<emphasis id="B5A1F647FFFFE956FE8DFDA7955CFD88" box="[264,613,604,629]" italics="true" pageId="59" pageNumber="60">Material examined and range</emphasis>
|
||
</paragraph>
|
||
<paragraph id="876A2A55FFFFE956FEBDFD7F96BBFCEB" blockId="59.[264,1323,604,790]" pageId="59" pageNumber="60">
|
||
<materialsCitation id="37BD2008FFFFE956FEBDFD7F9571FD3B" collectingDate="1995-05-21" collectingDateMax="1995-06-11" collectingDateMin="1995-05-21" collectionCode="SRGI, CRG" country="Thailand" location="THAILAND" pageId="59" pageNumber="60" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="holotype">
|
||
<specimenCount id="91D3E1DCFFFFE956FEBDFD7F9663FD6A" box="[312,346,643,670]" pageId="59" pageNumber="60" type="male">1♂</specimenCount>
|
||
,
|
||
<specimenCount id="91D3E1DCFFFFE956FEEDFD7F96B2FD6A" box="[360,395,643,670]" pageId="59" pageNumber="60" type="female">2♀</specimenCount>
|
||
.
|
||
<emphasis id="B5A1F647FFFFE956FE1DFD7F9506FD63" bold="true" box="[408,575,644,670]" pageId="59" pageNumber="60">
|
||
<collectingCountry id="FFC26AC5FFFFE956FE1DFD7F9503FD63" box="[408,570,644,670]" name="Thailand" pageId="59" pageNumber="60">THAILAND</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<typeStatus id="586E94F7FFFFE956FDC2FD7F958DFD63" box="[583,692,644,670]" pageId="59" pageNumber="60" type="holotype">Holotype</typeStatus>
|
||
female, “Thailand, Chiang Dao,
|
||
<date id="F36B0C95FFFFE956FBB3FD7F923CFD63" box="[1078,1285,644,670]" pageId="59" pageNumber="60" value="1995-05-21" valueMax="1995-06-11" valueMin="1995-05-21">
|
||
<collectingDate id="E32FF57DFFFFE956FBB3FD7F923CFD63" box="[1078,1285,644,670]" pageId="59" pageNumber="60" value="1995-05-21" valueMax="1995-06-11" valueMin="1995-05-21">21.v –11.vi. 1995</collectingDate>
|
||
</date>
|
||
, M Snížek lgt.”/
|
||
<collectionCode id="E1C4B290FFFFE956FE2DFD5796D2FD3B" box="[424,491,684,710]" pageId="59" pageNumber="60">SRGI</collectionCode>
|
||
[
|
||
<collectionCode id="E1C4B290FFFFE956FE79FD579578FD3B" box="[508,577,684,710]" pageId="59" pageNumber="60">CRG</collectionCode>
|
||
]
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="37BD2008FFFFE956FDD3FD579647FCEB" collectingDate="1995-06-04" collectionCode="CHIANG, CPG" collectorName="M. Snizek & This" country="Thailand" location="northern Thailand" pageId="59" pageNumber="60" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
|
||
<typeStatus id="586E94F7FFFFE956FDD3FD5795F5FD3B" box="[598,716,684,710]" pageId="59" pageNumber="60" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<specimenCount id="91D3E1DCFFFFE956FD5FFD5795C5FD42" box="[730,764,683,710]" pageId="59" pageNumber="60" type="male">1♂</specimenCount>
|
||
,
|
||
<specimenCount id="91D3E1DCFFFFE956FC88FD579409FD42" box="[781,816,683,710]" pageId="59" pageNumber="60" type="female">1♀</specimenCount>
|
||
, “THAILAND bor.,
|
||
<collectionCode id="E1C4B290FFFFE956FBABFD579399FD3B" box="[1070,1184,684,710]" pageId="59" pageNumber="60">CHIANG</collectionCode>
|
||
DAO env., 21.5–
|
||
<date id="F36B0C95FFFFE956FECFFD2F968AFD13" box="[330,435,724,750]" pageId="59" pageNumber="60" value="1995-06-04">
|
||
<collectingDate id="E32FF57DFFFFE956FECFFD2F968AFD13" box="[330,435,724,750]" pageId="59" pageNumber="60" value="1995-06-04">4.6.1995</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="2A204F83FFFFE956FE41FD2F9579FD13" box="[452,576,723,750]" pageId="59" pageNumber="60">M. Snížek</collectorName>
|
||
leg.” [
|
||
<collectionCode id="E1C4B290FFFFE956FD1CFD2F95E2FD13" box="[665,731,724,750]" pageId="59" pageNumber="60">CPG</collectionCode>
|
||
].
|
||
<collectorName id="2A204F83FFFFE956FD74FD2F941CFD13" box="[753,805,724,750]" pageId="59" pageNumber="60">This</collectorName>
|
||
species is known from
|
||
<location id="820A7C8EFFFFE956FBD6FD2F9212FD13" LSID="urn:lsid:plazi:treatment:0F7C9B43FFFDE956FE8DFC2F969BFA93:820A7C8EFFFFE956FBD6FD2F9212FD13" box="[1107,1323,724,750]" country="Thailand" name="northern Thailand" pageId="59" pageNumber="60">northern Thailand</location>
|
||
(
|
||
<figureCitation id="1FEE36D0FFFFE956FE94FD07964CFCEB" box="[273,373,764,790]" captionStart="FIGURE 111" captionStartId="58.[264,367,1149,1173]" captionTargetBox="[340,1260,398,1129]" captionTargetId="figure-221@58.[340,1260,398,1162]" captionTargetPageId="58" captionText="FIGURE 111. Distribution of Ctenoplus species." figureDoi="http://doi.org/10.5281/zenodo.5064840" httpUri="https://zenodo.org/record/5064840/files/figure.png" pageId="59" pageNumber="60">Fig. 111</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="876A2A55FFFFE956FE8DFCB7935DFC73" blockId="59.[264,1124,844,910]" box="[264,1124,844,910]" pageId="59" pageNumber="60">
|
||
<emphasis id="B5A1F647FFFFE956FE8DFCB796BCFC98" box="[264,389,844,869]" italics="true" pageId="59" pageNumber="60">Etymology</emphasis>
|
||
The species epithet refers to the alutaceous cuticle of the hypomeron.
|
||
</paragraph>
|
||
<paragraph id="876A2A55FFFFE956FE8DFC3F94E0FBFB" blockId="59.[264,985,964,1030]" pageId="59" pageNumber="60">
|
||
<emphasis id="B5A1F647FFFFE956FE8DFC3F9682FC20" box="[264,443,964,989]" italics="true" pageId="59" pageNumber="60">Natural history</emphasis>
|
||
Adults have been collected between 21 May and 11 June.
|
||
</paragraph>
|
||
<paragraph id="876A2A55FFFFE956FE8DFBC796B1FBA8" blockId="59.[264,1323,1084,1390]" box="[264,392,1084,1109]" pageId="59" pageNumber="60">
|
||
<emphasis id="B5A1F647FFFFE956FE8DFBC796B1FBA8" box="[264,392,1084,1109]" italics="true" pageId="59" pageNumber="60">Discussion</emphasis>
|
||
</paragraph>
|
||
<paragraph id="876A2A55FFFFE956FEBDFB9F969BFA93" blockId="59.[264,1323,1084,1390]" pageId="59" pageNumber="60">
|
||
Female genitalia are similar to those of
|
||
<taxonomicName id="40D551D6FFFFE956FC96FB9F9491FB80" authorityName="Fuller & Platia" authorityYear="2006" box="[787,936,1124,1149]" class="Insecta" family="Elateridae" genus="Ctenoplus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="divergens">
|
||
<emphasis id="B5A1F647FFFFE956FC96FB9F9491FB80" box="[787,936,1124,1149]" italics="true" pageId="59" pageNumber="60">C. divergens</emphasis>
|
||
</taxonomicName>
|
||
, suggesting a close relationship, but can be distinguished by the bursal plates, and the length and shape of the tubular extension (c.f.,
|
||
<figureCitation id="1FEE36D0FFFFE956FE40FB4F9556FB33" box="[453,623,1204,1230]" captionStart="FIGURES 98–106" captionStartId="30.[264,382,1757,1781]" captionTargetBox="[341,1253,875,1748]" captionTargetId="figure-246@30.[319,1281,865,1761]" captionTargetPageId="30" captionText="FIGURES 98–106. Female genitalia of Ctenoplus species. 98, C. coomani Fleut.; 99, C. brunneus Fleut.; 100, C. divergens n. sp.; 101, C. alutaceus n. sp.; 102, C. balli n. sp.; 103, C. girardianus n. sp.; 104, C. gigas n. sp.; 105, C. cateianus n. sp.; 106, C. deceptus n. sp. Abbreviations: sd, tubular extension. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5064828" httpUri="https://zenodo.org/record/5064828/files/figure.png" pageId="59" pageNumber="60">Figs 100, 101</figureCitation>
|
||
). The bursal plates of
|
||
<taxonomicName id="40D551D6FFFFE956FC00FB4F9322FB30" authorityName="Fuller & Platia" authorityYear="2006" box="[901,1051,1204,1229]" class="Insecta" family="Elateridae" genus="Ctenoplus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="alutaceus">
|
||
<emphasis id="B5A1F647FFFFE956FC00FB4F9322FB30" box="[901,1051,1204,1229]" italics="true" pageId="59" pageNumber="60">C. alutaceus</emphasis>
|
||
</taxonomicName>
|
||
are narrower and less heavily spined than those of
|
||
<taxonomicName id="40D551D6FFFFE956FDDFFB2795D4FB08" authorityName="Fuller & Platia" authorityYear="2006" box="[602,749,1244,1269]" class="Insecta" family="Elateridae" genus="Ctenoplus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="divergens">
|
||
<emphasis id="B5A1F647FFFFE956FDDFFB2795D4FB08" box="[602,749,1244,1269]" italics="true" pageId="59" pageNumber="60">C. divergens</emphasis>
|
||
</taxonomicName>
|
||
. The tubular extension of
|
||
<taxonomicName id="40D551D6FFFFE956FBA1FB27938FFB08" authorityName="Fuller & Platia" authorityYear="2006" box="[1060,1206,1244,1269]" class="Insecta" family="Elateridae" genus="Ctenoplus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="alutaceus">
|
||
<emphasis id="B5A1F647FFFFE956FBA1FB27938FFB08" box="[1060,1206,1244,1269]" italics="true" pageId="59" pageNumber="60">C. alutaceus</emphasis>
|
||
</taxonomicName>
|
||
is shorter (about half the length of the bursa
|
||
<emphasis id="B5A1F647FFFFE956FD2DFAFF95F1FAE0" box="[680,712,1284,1309]" italics="true" pageId="59" pageNumber="60">vs.</emphasis>
|
||
subequal in length to the bursa), and the extension appears to branch laterally from the wide base, rather than being continuous distally as in
|
||
<taxonomicName id="40D551D6FFFFE956FE8DFAAF96A2FA90" authorityName="Fuller & Platia" authorityYear="2006" box="[264,411,1364,1389]" class="Insecta" family="Elateridae" genus="Ctenoplus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="divergens">
|
||
<emphasis id="B5A1F647FFFFE956FE8DFAAF96A2FA90" box="[264,411,1364,1389]" italics="true" pageId="59" pageNumber="60">C. divergens</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |