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<document id="8E9BCCA4C33AE8AFEB26507F45A95C3C" ID-DOI="10.11646/zootaxa.1217.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5064796" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625283125096" checkinUser="felipe" docAuthor="Fuller, E. &amp; Platia, G." docDate="2006" docId="0F7C9B43FF84E92FFE8DFE5D9531FBFB" docLanguage="en" docName="zt01217p076.pdf" docOrigin="Zootaxa 1217 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1217.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Ctenoplus girardianus Fuller &amp; Platia 2006, n. sp." docType="treatment" docVersion="4" lastPageNumber="67" masterDocId="F345E33BFFC4E96DFF85FFFB9739FFFD" masterDocTitle="A revision of the click beetle genus Ctenoplus Candèze, 1863 (Coleoptera: Elateridae, Synaptina)" masterLastPageNumber="76" masterPageNumber="1" pageNumber="65" updateTime="1698986177451" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="C7BD5E7BF9F7046C97E18E860CA7BED0">A revision of the click beetle genus Ctenoplus Candèze, 1863 (Coleoptera: Elateridae, Synaptina)</mods:title>
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<mods:namePart id="62BA58D18DD5F3E98C1437E440DEAF75">Fuller, E.</mods:namePart>
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<mods:namePart id="7BFA89CC9CCD34194EE0EEB3CB838CBD">Platia, G.</mods:namePart>
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<treatment id="0F7C9B43FF84E92FFE8DFE5D9531FBFB" ID-DOI="http://doi.org/10.5281/zenodo.5066813" ID-Zenodo-Dep="5066813" LSID="urn:lsid:plazi:treatment:0F7C9B43FF84E92FFE8DFE5D9531FBFB" httpUri="http://treatment.plazi.org/id/0F7C9B43FF84E92FFE8DFE5D9531FBFB" lastPageId="66" lastPageNumber="67" pageId="64" pageNumber="65">
<subSubSection id="CFCF79DEFF84E92DFE8DFE5D9558FE3D" box="[264,609,422,448]" pageId="64" pageNumber="65" type="nomenclature">
<paragraph id="876A2A55FF84E92DFE8DFE5D9558FE3D" blockId="64.[264,609,422,483]" box="[264,609,422,448]" pageId="64" pageNumber="65">
<heading id="DC229D39FF84E92DFE8DFE5D9558FE3D" bold="true" box="[264,609,422,448]" fontSize="11" level="1" pageId="64" pageNumber="65" reason="1">
<emphasis id="B5A1F647FF84E92DFE8DFE5D9558FE3D" bold="true" box="[264,609,422,448]" pageId="64" pageNumber="65">
<taxonomicName id="40D551D6FF84E92DFE8DFE5D9521FE3D" authority="Fuller &amp; Platia, 2006" authorityName="Fuller &amp; Platia" authorityYear="2006" box="[264,536,422,448]" class="Insecta" family="Elateridae" genus="Ctenoplus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="girardianus" status="sp. nov.">
<emphasis id="B5A1F647FF84E92DFE8DFE5D9521FE3D" bold="true" box="[264,536,422,448]" italics="true" pageId="64" pageNumber="65">Ctenoplus girardianus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AE924B3CFF84E92DFD9AFE5D9558FE3D" box="[543,609,422,448]" pageId="64" pageNumber="65" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="CFCF79DEFF84E92FFE8DFE329531FBFB" lastPageId="66" lastPageNumber="67" pageId="64" pageNumber="65" type="description">
<paragraph id="876A2A55FF84E92DFE8DFE329568FE1E" blockId="64.[264,609,422,483]" box="[264,593,457,483]" pageId="64" pageNumber="65">
<figureCitation id="1FEE36D0FF84E92DFE8DFE3296BEFE1E" box="[264,391,457,483]" captionStart="FIGURES 4446" captionStartId="15.[264,382,1135,1159]" captionTargetBox="[267,1299,334,1105]" captionTargetId="figure-270@15.[264,1323,326,1125]" captionTargetPageId="15" captionText="FIGURES 4446. posterior part of hypomeron of Ctenoplus species. 44, C. alutaceus n. sp.; 45, C. semialutaceus n. sp.; 46, C. girardianus n. sp. 47, prosternum of C. nitidipennis Schwarz, ventral. 48, venter of mesothorax of C. dorsalis n. sp., ventral. 49, tarsus of C. castaneus (Fleut.), lateral. Abbreviations: ac, alutaceous cuticle; ag, anterior groove of mesepisternum; as, anterior articulating surface of mesosternum; c, mesocoxa; d, lateral depression of mesosternum; gr, groove on prosternum; l, lobe of tarsomere 4; lp, anterolateral projection of mesepimeron; mf, mesosternal cavity; pr, projection of mesal border of hypomeron. Scale bar = 4446, 48, 49: 0.25 mm; 48: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5064814" httpUri="https://zenodo.org/record/5064814/files/figure.png" pageId="64" pageNumber="65">Figures 46</figureCitation>
, 65, 82, 103, 112
</paragraph>
<paragraph id="876A2A55FF84E92DFE8DFDF49646FDD5" blockId="64.[264,1323,527,793]" box="[264,383,527,552]" pageId="64" pageNumber="65">
<emphasis id="B5A1F647FF84E92DFE8DFDF49646FDD5" box="[264,383,527,552]" italics="true" pageId="64" pageNumber="65">Diagnosis</emphasis>
</paragraph>
<paragraph id="876A2A55FF84E92DFEBDFDCC9331FCE4" blockId="64.[264,1323,527,793]" pageId="64" pageNumber="65">
Medium­sized, uniformly brown
<taxonomicName id="40D551D6FF84E92DFD4CFDCC9406FDAD" box="[713,831,567,592]" class="Insecta" family="Elateridae" genus="Ctenoplus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="B5A1F647FF84E92DFD4CFDCC9406FDAD" box="[713,831,567,592]" italics="true" pageId="64" pageNumber="65">Ctenoplus</emphasis>
</taxonomicName>
with a relatively short, broad prosternal spine and relatively rounded elytral apices.
<taxonomicName id="40D551D6FF84E92DFC81FDA49496FD85" authorityName="Fuller &amp; Platia" authorityYear="2006" box="[772,943,607,632]" class="Insecta" family="Elateridae" genus="Ctenoplus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="girardianus">
<emphasis id="B5A1F647FF84E92DFC81FDA49496FD85" box="[772,943,607,632]" italics="true" pageId="64" pageNumber="65">C. girardianus</emphasis>
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is similar to
<taxonomicName id="40D551D6FF84E92DFBCDFDA493A6FD85" authority="Fuller &amp; Platia, 2006" authorityName="Fuller &amp; Platia" authorityYear="2006" box="[1096,1183,607,632]" class="Insecta" family="Elateridae" genus="Ctenoplus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="balli" status="sp. nov.">
<emphasis id="B5A1F647FF84E92DFBCDFDA493A6FD85" box="[1096,1183,607,632]" italics="true" pageId="64" pageNumber="65">C. balli</emphasis>
</taxonomicName>
<emphasis id="B5A1F647FF84E92DFB23FDA593D1FD85" bold="true" box="[1190,1256,606,632]" pageId="64" pageNumber="65">
<taxonomicNameLabel id="AE924B3CFF84E92DFB23FDA593D1FD85" box="[1190,1256,606,632]" pageId="64" pageNumber="65" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, with which it shares the short, broad prosternal spine (c.f.,
<figureCitation id="1FEE36D0FF84E92DFC06FD7C94E3FD5C" box="[899,986,647,673]" captionStart="FIGURES 5057" captionStartId="19.[264,382,1487,1511]" captionTargetBox="[311,1265,365,1457]" captionTargetId="figure-138@19.[264,1323,330,1493]" captionTargetPageId="19" captionText="FIGURES 5057. Prosternal spine of Ctenoplus and Silesis species. 5057, lateral; 58, 59, ventral. 50, C. sanguinolentus (Cand.); 51, C. sanguinolentoides n. sp.; 52a, b, C. coomani Fleut.; 53, C. balli n. sp.; 54, C. gigas n. sp.; 55, C. divergens n. sp.; 56, C. alutaceus n. sp.; 57, C. dorsalis n. sp.; 58, C. brunneus Fleut.; 59, S. fulvus Fleut.; Abbreviations: da, dorsal apex; va, ventral apex. Scale bar = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5064816" httpUri="https://zenodo.org/record/5064816/files/figure.png" pageId="64" pageNumber="65">Fig. 53</figureCitation>
); however,
<taxonomicName id="40D551D6FF84E92DFBE6FD7C9236FD5D" authorityName="Fuller &amp; Platia" authorityYear="2006" box="[1123,1295,647,672]" class="Insecta" family="Elateridae" genus="Ctenoplus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="girardianus">
<emphasis id="B5A1F647FF84E92DFBE6FD7C9236FD5D" box="[1123,1295,647,672]" italics="true" pageId="64" pageNumber="65">C. girardianus</emphasis>
</taxonomicName>
is smaller (8.08.5
<emphasis id="B5A1F647FF84E92DFE4BFD5496D7FD35" box="[462,494,687,712]" italics="true" pageId="64" pageNumber="65">vs.</emphasis>
<quantity id="402D87B0FF84E92DFE70FD549566FD34" box="[501,607,687,713]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="64" pageNumber="65" unit="mm" value="10.5">10.5 mm</quantity>
) and the apex of the elytron is less attenuated than in
<taxonomicName id="40D551D6FF84E92DFB56FD549213FD35" authorityName="Fuller &amp; Platia" authorityYear="2006" box="[1235,1322,687,712]" class="Insecta" family="Elateridae" genus="Ctenoplus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="balli">
<emphasis id="B5A1F647FF84E92DFB56FD549213FD35" box="[1235,1322,687,712]" italics="true" pageId="64" pageNumber="65">C. balli</emphasis>
</taxonomicName>
(c.f., Figs. 64, 65). In addition,
<taxonomicName id="40D551D6FF84E92DFDF0FD2C9419FD0D" authorityName="Fuller &amp; Platia" authorityYear="2006" box="[629,800,727,752]" class="Insecta" family="Elateridae" genus="Ctenoplus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="girardianus">
<emphasis id="B5A1F647FF84E92DFDF0FD2C9419FD0D" box="[629,800,727,752]" italics="true" pageId="64" pageNumber="65">C. girardianus</emphasis>
</taxonomicName>
is known from the
<collectingCountry id="FFC26AC5FF84E92DFB81FD2C93B1FD0C" box="[1028,1160,727,753]" name="Philippines" pageId="64" pageNumber="65">Philippines</collectingCountry>
and
<collectingCountry id="FFC26AC5FF84E92DFB45FD2C921FFD0C" box="[1216,1318,727,753]" name="Vietnam" pageId="64" pageNumber="65">Vietnam</collectingCountry>
, whereas
<taxonomicName id="40D551D6FF84E92DFEEAFD0496F1FCE5" authorityName="Fuller &amp; Platia" authorityYear="2006" box="[367,456,767,792]" class="Insecta" family="Elateridae" genus="Ctenoplus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="balli">
<emphasis id="B5A1F647FF84E92DFEEAFD0496F1FCE5" box="[367,456,767,792]" italics="true" pageId="64" pageNumber="65">C. balli</emphasis>
</taxonomicName>
is known only from northern
<collectingCountry id="FFC26AC5FF84E92DFCA2FD0494B6FCE4" box="[807,911,767,793]" name="Thailand" pageId="64" pageNumber="65">Thailand</collectingCountry>
and
<collectingCountry id="FFC26AC5FF84E92DFC4DFD04933CFCE4" box="[968,1029,767,793]" name="Laos" pageId="64" pageNumber="65">Laos</collectingCountry>
.
</paragraph>
<paragraph id="876A2A55FF84E92DFE8DFCB496ABFC95" blockId="64.[264,1324,847,1873]" box="[264,402,847,872]" pageId="64" pageNumber="65">
<emphasis id="B5A1F647FF84E92DFE8DFCB496ABFC95" box="[264,402,847,872]" italics="true" pageId="64" pageNumber="65">Description</emphasis>
</paragraph>
<paragraph id="876A2A55FF84E92DFEBDFC8C9604FC44" blockId="64.[264,1324,847,1873]" pageId="64" pageNumber="65">
Medium brown; setae whitish­yellow throughout. Length: 8.0
<quantity id="402D87B0FF84E92DFBB9FC8C93A4FC6C" box="[1084,1181,887,913]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="64" pageNumber="65" unit="mm" value="8.5">8.5 mm</quantity>
; width: 2.0 mm.
</paragraph>
<paragraph id="876A2A55FF84E92DFEBDFC3C939BFA3C" blockId="64.[264,1324,847,1873]" pageId="64" pageNumber="65">Head. Punctures moderate, almost contiguous throughout, setae slightly longer than antennomere 4. Antenna reaching or almost reaching apex of pronotal hind angles; antennomere 1 with punctures 1/2 size of frontal punctures, separated by about own diameters, setae varying from 2/3 to subequal in length to frontal setae; antennomere 2 about 1/3 longer than wide, cylindrical; antennomere 3 2X as long as wide, somewhat flattened, cuticle as in antennomere 2, anterior and posterior margins subparallel; antennomere 4 2X as long as greatest width, flattened, anterior and posterior margins weakly divergent distally, anterodistal angle rounded, distal end yellowish, cuticle microrugose, subshiny; antennomeres 510 like antennomere 4 but subtriangular, antennomeres 810 narrower than antennomeres 57; antennomere 11 slightly longer and narrower than antennomere 10; cuticle microrugose. Cuticle between eye and supraantennal carina microrugose, dull. Gena microrugose, subshiny; punctures minute, separated by about 2X own diameter; setae shorter and finer than frontal setae.</paragraph>
<paragraph id="876A2A55FF84E92CFEBDFA3493DEFCC3" blockId="64.[264,1324,847,1873]" lastBlockId="65.[264,1324,284,1870]" lastPageId="65" lastPageNumber="66" pageId="64" pageNumber="65">
Prothorax. Pronotum slightly wider than long, lateral margins subparallel; punctures moderate sized, relatively shallow, simple, separated by about own diameter on disk, becoming almost contiguous anteriorly and laterally; setae same length as frontal setae, erect, directed posteriorly; posterior margin as in
<taxonomicName id="40D551D6FF84E92DFCDCF9BC94DBF99D" box="[857,994,1607,1632]" class="Insecta" family="Elateridae" genus="Ctenoplus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="coomani">
<emphasis id="B5A1F647FF84E92DFCDCF9BC94DBF99D" box="[857,994,1607,1632]" italics="true" pageId="64" pageNumber="65">C. coomani</emphasis>
</taxonomicName>
(c.f.
<figureCitation id="1FEE36D0FF84E92DFBA1F9BC9344F99C" box="[1060,1149,1607,1633]" captionStart="FIGURES 3143" captionStartId="14.[264,382,1697,1721]" captionTargetBox="[369,1208,554,1659]" captionTargetId="figure-170@14.[359,1241,544,1699]" captionTargetPageId="14" captionText="FIGURES 3143. Pronotal outline of Ctenoplus species, dorsal. 31, C. javanensis Cand.; 32, C. sumatrensis n. sp.; 33, C. sanguinolentoides n. sp.; 34, C. indicus n. sp.; 35, C. rufoantennatus n. sp.; 36, C. castaneus (Fleut.); 37, C. coomani Fleut.; 38, C. divergens n. sp., a: male, b: female; 39, C. alutaceus n. sp.; 40, C. semialutaceus n. sp.; 41, C. gigas n. sp.; 42, C. cateianus n. sp.; 43, C. dorsalis n. sp. Abbreviation: bf, sublateral longitudinal incision. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5064810" httpUri="https://zenodo.org/record/5064810/files/figure.png" pageId="64" pageNumber="65">Fig. 37</figureCitation>
); apex of hind angles acute. Hypomeron: posterior 1/4 glabrous (
<figureCitation id="1FEE36D0FF84E92DFCF1F99494E8F974" box="[884,977,1647,1673]" captionStart="FIGURES 4446" captionStartId="15.[264,382,1135,1159]" captionTargetBox="[267,1299,334,1105]" captionTargetId="figure-270@15.[264,1323,326,1125]" captionTargetPageId="15" captionText="FIGURES 4446. posterior part of hypomeron of Ctenoplus species. 44, C. alutaceus n. sp.; 45, C. semialutaceus n. sp.; 46, C. girardianus n. sp. 47, prosternum of C. nitidipennis Schwarz, ventral. 48, venter of mesothorax of C. dorsalis n. sp., ventral. 49, tarsus of C. castaneus (Fleut.), lateral. Abbreviations: ac, alutaceous cuticle; ag, anterior groove of mesepisternum; as, anterior articulating surface of mesosternum; c, mesocoxa; d, lateral depression of mesosternum; gr, groove on prosternum; l, lobe of tarsomere 4; lp, anterolateral projection of mesepimeron; mf, mesosternal cavity; pr, projection of mesal border of hypomeron. Scale bar = 4446, 48, 49: 0.25 mm; 48: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5064814" httpUri="https://zenodo.org/record/5064814/files/figure.png" pageId="64" pageNumber="65">Fig. 46</figureCitation>
); anterior 3/4 with shallow, simple punctures, punctures subequal in size to pronotal punctures, separated by about own diameters; setae same length as pronotal setae; mesal smooth border bearing triangular projection at level of procoxa (
<figureCitation id="1FEE36D0FF84E92DFD6CF91C9479F8FC" box="[745,832,1767,1793]" captionStart="FIGURES 4446" captionStartId="15.[264,382,1135,1159]" captionTargetBox="[267,1299,334,1105]" captionTargetId="figure-270@15.[264,1323,326,1125]" captionTargetPageId="15" captionText="FIGURES 4446. posterior part of hypomeron of Ctenoplus species. 44, C. alutaceus n. sp.; 45, C. semialutaceus n. sp.; 46, C. girardianus n. sp. 47, prosternum of C. nitidipennis Schwarz, ventral. 48, venter of mesothorax of C. dorsalis n. sp., ventral. 49, tarsus of C. castaneus (Fleut.), lateral. Abbreviations: ac, alutaceous cuticle; ag, anterior groove of mesepisternum; as, anterior articulating surface of mesosternum; c, mesocoxa; d, lateral depression of mesosternum; gr, groove on prosternum; l, lobe of tarsomere 4; lp, anterolateral projection of mesepimeron; mf, mesosternal cavity; pr, projection of mesal border of hypomeron. Scale bar = 4446, 48, 49: 0.25 mm; 48: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5064814" httpUri="https://zenodo.org/record/5064814/files/figure.png" pageId="64" pageNumber="65">Fig. 46</figureCitation>
), projection as long as basal width; mesal smooth border with single row of setae, setae about 3/4 length of other hypomeral setae, punctures minute; posterolateral angle projecting posterad posterior margin (
<figureCitation id="1FEE36D0FF84E92DFB39F8CC9222F8AC" box="[1212,1307,1847,1873]" captionStart="FIGURES 4446" captionStartId="15.[264,382,1135,1159]" captionTargetBox="[267,1299,334,1105]" captionTargetId="figure-270@15.[264,1323,326,1125]" captionTargetPageId="15" captionText="FIGURES 4446. posterior part of hypomeron of Ctenoplus species. 44, C. alutaceus n. sp.; 45, C. semialutaceus n. sp.; 46, C. girardianus n. sp. 47, prosternum of C. nitidipennis Schwarz, ventral. 48, venter of mesothorax of C. dorsalis n. sp., ventral. 49, tarsus of C. castaneus (Fleut.), lateral. Abbreviations: ac, alutaceous cuticle; ag, anterior groove of mesepisternum; as, anterior articulating surface of mesosternum; c, mesocoxa; d, lateral depression of mesosternum; gr, groove on prosternum; l, lobe of tarsomere 4; lp, anterolateral projection of mesepimeron; mf, mesosternal cavity; pr, projection of mesal border of hypomeron. Scale bar = 4446, 48, 49: 0.25 mm; 48: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5064814" httpUri="https://zenodo.org/record/5064814/files/figure.png" pageId="64" pageNumber="65">Fig. 46</figureCitation>
), projection about as long as basal width in ventrolateral aspect. Prosternum: anterior lobe with shallow inconspicuous punctures and row of marginal setae anteriorly; punctures posterad lobe subequal in size to hypomeral punctures, shallow, simple, separated by about own diameter anteriorly, grading to 1/2 own diameter separation posteriorly including prosternal spine; setae same length as on hypomeron, directed posteriorly; prosternal spine: in lateral aspect, relatively short: length to posterior concavity about 1.5X width immediately posterad procoxae, slightly curved dorsally, ventral apex slightly produced, as long as wide, acute, triangular, slightly recurved, dorsal apex triangular, about 3X as long as basal width, with low lateral carina, posterior margin between apices vertical, straight; ventral apex forming ventral surface of U­shaped ridge around posterior vertical margin, U­shaped ridge accentuating medial ridge on posterior surface, ventrolateral carinae fused in apical 1/5 but not darkly pigmented, extending slightly anterad anterior margin of procoxal cavity; lateral margin sparsely punctured, punctures minute, posterodorsal angle glabrous, impunctate, setae as on hypomeron, directed ventrally.
</paragraph>
<paragraph id="876A2A55FF85E92CFEBDFCB793BDFABB" blockId="65.[264,1324,284,1870]" pageId="65" pageNumber="66">Mesothorax. Scutellum flat, slightly longer than anterior width; anterior margin slightly produced, with transverse microstriae; setae as on pronotum, directed posteriorly, semi­erect; punctures minute, separated by 23X own diameters. Mesosternum in ventral aspect with posterior margin truncate; in lateral aspect mesosternum between mesocoxae raised, horizontal; margin of mesosternal cavity anterad mesocoxa sloping almost vertically to level of mesepisternum; punctures minute, separated by 13X own diameters. Mesepisternum: anterior margin bearing circular pit adjacent to mesosternum; posteromesal 1/2 glabrous. Mesepimeron: punctures shallow, simple, almost contiguous. Elytra (Fig. 65) about 4X as long as basal width, lateral margins convergent apically; anterior margin projecting; apex with very small acute tooth at suture; striae impressed in basal 1/4, grading into rows of punctures in apical 3/4, margins of impressed striae sharp; punctures circular, relatively deep, steep­sided, separated by about 1/2 own diameters, without setae; interval punctures minute, separated by 23X own diameters.</paragraph>
<paragraph id="876A2A55FF85E92CFEBDFAAF93C8FA43" blockId="65.[264,1324,284,1870]" pageId="65" pageNumber="66">Metathorax. Metasternum: carina posterad mesocoxa very short, present only as small rounded bump on marginal ridge; medial punctures minute, separated by 12X own diameters, grading into larger, shallow, simple, almost contiguous punctures laterally.</paragraph>
<paragraph id="876A2A55FF85E92CFEBDFA37940AF9CB" blockId="65.[264,1324,284,1870]" pageId="65" pageNumber="66">Legs. Tarsomere 4 with small ventrodistal lobe; claws with 56 subapical tines, base of claws relatively thick, subapical tines 1/32/3 length of apical tine, middle pair of subapical tines longest, innermost tine shortest.</paragraph>
<paragraph id="876A2A55FF85E92FFEBDF9BF9309FE7B" blockId="65.[264,1324,284,1870]" lastBlockId="66.[264,1322,284,390]" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66">
Abdomen. Terga lightly sclerotized. Dorsal submarginal carina of ventrite 5 broadly U­shaped. Setae, punctures and cuticle as on metasternum except on ventrite 5 where punctures closer throughout. Male genitalia,
<figureCitation id="1FEE36D0FF85E92CFCCAF96F9489F953" box="[847,944,1684,1710]" captionStart="FIGURES 6685" captionStartId="22.[264,382,1424,1448]" captionTargetBox="[293,1279,326,1378]" captionTargetId="figure-207@22.[264,1323,326,1438]" captionTargetPageId="22" captionText="FIGURES 6685. Male genitalia of Ctenoplus species, dorsal aspect unless otherwise specified. 66, C. javanensis Cand.; 67, C. sumatrensis n. sp.; 68, C. sanguinolentoides n. sp.; 69, C. nitidipennis Schwarz; 70, C. indicus n. sp.; 71, C. collaris Cand., a, dorsal, b, ventral; 72, C. pseudocollaris n. sp.; 73, C. neosiamensis n. sp.; 74, C. siamensis n. sp.; 75, C. rufoantennatus n. sp.; 76, C. castaneus (Fleut.), a, dorsal, b, lateral; 77, C. coomani Fleut.; 78, C. divergens n. sp.; 79, C. alutaceus n. sp., a, dorsal, b, lateral; 80, C. semialutaceus n. sp.; 81, C. balli n. sp.; 82, C. girardianus n. sp.; 83, C. gigas n. sp.; 84, C. cateianus n. sp.; 85, C. deceptus n. sp. Abbreviations: st, median struts; pa, parameres; ph, phallobase. Scale bar = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5064822" httpUri="https://zenodo.org/record/5064822/files/figure.png" pageId="65" pageNumber="66">Fig. 82</figureCitation>
; parameres in ventral aspect contiguous at anterior end; median lobe with apical portion curved ventrally, dorsal surface with sharp but only slightly raised V­shaped carina, ventral surface with elongate elliptical sclerite tapered to a point at each end. Female genitalia,
<figureCitation id="1FEE36D0FF85E92CFBA5F8F793BEF8DB" box="[1056,1159,1804,1830]" captionStart="FIGURES 98106" captionStartId="30.[264,382,1757,1781]" captionTargetBox="[341,1253,875,1748]" captionTargetId="figure-246@30.[319,1281,865,1761]" captionTargetPageId="30" captionText="FIGURES 98106. Female genitalia of Ctenoplus species. 98, C. coomani Fleut.; 99, C. brunneus Fleut.; 100, C. divergens n. sp.; 101, C. alutaceus n. sp.; 102, C. balli n. sp.; 103, C. girardianus n. sp.; 104, C. gigas n. sp.; 105, C. cateianus n. sp.; 106, C. deceptus n. sp. Abbreviations: sd, tubular extension. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5064828" httpUri="https://zenodo.org/record/5064828/files/figure.png" pageId="65" pageNumber="66">Fig. 103</figureCitation>
; bursal plates crescentic, symmetrical; tubular extension: arising dorsolaterally on bursa; with 2 bands of spines of approximately equal width; spines grading from subequal in appearance to bursal spines basally to more darkly pigmented spines with larger bases in mid­region to spines subequal in appearance to bursal spines distally; spermatheca absent.
</paragraph>
<paragraph id="876A2A55FF86E92FFE8DFE47955CFE28" blockId="66.[264,1325,444,710]" box="[264,613,444,469]" pageId="66" pageNumber="67">
<emphasis id="B5A1F647FF86E92FFE8DFE47955CFE28" box="[264,613,444,469]" italics="true" pageId="66" pageNumber="67">Material examined and range</emphasis>
</paragraph>
<paragraph id="876A2A55FF86E92FFEBDFE1F93F0FD3B" blockId="66.[264,1325,444,710]" pageId="66" pageNumber="67">
<materialsCitation id="37BD2008FF86E92FFEBDFE1F9210FDDB" collectionCode="MNHN" collectorName="E. Fleutiaux &amp; Coll. E. Fleutiaux" country="Philippines" location="Cuernos Mts." pageId="66" pageNumber="67" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="91D3E1DCFF86E92FFEBDFE1F9663FE0A" box="[312,346,483,510]" pageId="66" pageNumber="67" type="male">1♂</specimenCount>
,
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.
<emphasis id="B5A1F647FF86E92FFE35FE189547FE00" bold="true" box="[432,638,483,509]" pageId="66" pageNumber="67">
[
<collectingCountry id="FFC26AC5FF86E92FFE32FE18954EFE00" box="[439,631,483,509]" name="Philippines" pageId="66" pageNumber="67">PHILIPPINES</collectingCountry>
]
</emphasis>
<typeStatus id="586E94F7FF86E92FFD14FE1F95C7FE03" box="[657,766,484,510]" pageId="66" pageNumber="67" type="holotype">Holotype</typeStatus>
female: “
<location id="820A7C8EFF86E92FFC03FE1F930CFE03" LSID="urn:lsid:plazi:treatment:0F7C9B43FF84E92FFE8DFE5D9531FBFB:820A7C8EFF86E92FFC03FE1F930CFE03" box="[902,1077,484,510]" country="Philippines" name="Cuernos Mts." pageId="66" pageNumber="67">Cuernos Mts.</location>
; Negros I Baker”/ “Collection;
<collectorName id="2A204F83FF86E92FFE29FDF79572FDDB" box="[428,587,524,550]" pageId="66" pageNumber="67">E. Fleutiaux</collectorName>
”/ “Museum Paris;
<collectorName id="2A204F83FF86E92FFCD6FDF7937BFDDB" box="[851,1090,524,550]" pageId="66" pageNumber="67">Coll. E. Fleutiaux</collectorName>
; 1919&quot; [
<collectionCode id="E1C4B290FF86E92FFB3AFDF7921BFDDB" box="[1215,1314,524,550]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="66" pageNumber="67">MNHN</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="37BD2008FF86E92FFE8DFDCF93FCFD3B" collectingDate="1999-05-28" collectionCode="UV, MV, ROME" collectorName="B. Hubley." country="Vietnam" county="This" elevation="120" latitude="11.434055" location="Philippines" longLatPrecision="1" longitude="107.42933" municipality="Negros Island" pageId="66" pageNumber="67" specimenCode="ROM993041" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Dong Nai" typeStatus="paratype">
<typeStatus id="586E94F7FF86E92FFE8DFDCF9647FDB3" box="[264,382,564,590]" pageId="66" pageNumber="67" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="91D3E1DCFF86E92FFE09FDCF9696FDBA" box="[396,431,563,590]" pageId="66" pageNumber="67" type="male">1♂</specimenCount>
,
<specimenCount id="91D3E1DCFF86E92FFE45FDCF96DAFDBA" box="[448,483,563,590]" pageId="66" pageNumber="67" type="female">1♀</specimenCount>
,
<emphasis id="B5A1F647FF86E92FFE71FDC895B8FDB0" bold="true" box="[500,641,563,589]" pageId="66" pageNumber="67">
<collectingCountry id="FFC26AC5FF86E92FFE71FDC895B8FDB0" box="[500,641,563,589]" name="Vietnam" pageId="66" pageNumber="67">VIETNAM</collectingCountry>
</emphasis>
. “
<collectingRegion id="4511E4B7FF86E92FFD1BFDCF9423FDB3" box="[670,794,564,590]" country="Vietnam" name="Dong Nai" pageId="66" pageNumber="67">Dong Nai</collectingRegion>
: Cat Tien Natl. Pk., ca.
<quantity id="402D87B0FF86E92FFBC7FDCF939CFDB3" box="[1090,1189,564,590]" metricMagnitude="3" metricUnit="m" metricValue="1.23" pageId="66" pageNumber="67" unit="km" value="1.23">1.23 km</quantity>
on road N from park HQ,
<date id="F36B0C95FF86E92FFE3EFDA79561FD8B" box="[443,600,604,630]" pageId="66" pageNumber="67" value="1999-05-28">
<collectingDate id="E32FF57DFF86E92FFE3EFDA79561FD8B" box="[443,600,604,630]" pageId="66" pageNumber="67" value="1999-05-28">28 May 1999</collectingDate>
</date>
, DC Currie, DC Darling,
<collectorName id="2A204F83FF86E92FFC06FDA79338FD8B" box="[899,1025,604,630]" pageId="66" pageNumber="67">B. Hubley.</collectorName>
<specimenCode id="D773822EFF86E92FFB82FDA7939DFD8B" box="[1031,1188,604,630]" collectionCode="ROM" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="66" pageNumber="67" type="Museum">ROM993041</specimenCode>
” / “tropical lowland forest;
<collectionCode id="E1C4B290FF86E92FFE54FD7F96C2FD63" box="[465,507,644,670]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="66" pageNumber="67">UV</collectionCode>
/
<collectionCode id="E1C4B290FF86E92FFD81FD7F950AFD63" box="[516,563,644,670]" country="0" name="University of Montana Museum" pageId="66" pageNumber="67">MV</collectionCode>
light
<quantity id="402D87B0FF86E92FFD0DFD7F95E7FD63" box="[648,734,644,670]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="66" pageNumber="67" unit="m" value="120.0">
<elevation id="0CF8CD66FF86E92FFD0DFD7F95E7FD63" box="[648,734,644,670]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="66" pageNumber="67" unit="m" value="120.0">120 m</elevation>
</quantity>
;
<geoCoordinate id="E2E14C92FF86E92FFD77FD7F94A1FD63" box="[754,920,642,670]" degrees="11" direction="north" minutes="26" orientation="latitude" pageId="66" pageNumber="67" precision="1" seconds="02.6" value="11.434055">11°2602.6&quot;N</geoCoordinate>
<geoCoordinate id="E2E14C92FF86E92FFC2DFD7F9365FD63" box="[936,1116,642,670]" degrees="107" direction="east" minutes="25" orientation="longitude" pageId="66" pageNumber="67" precision="1" seconds="45.6" value="107.42933">107°2545.6&quot;E</geoCoordinate>
” [
<collectionCode id="E1C4B290FF86E92FFBFBFD7F93E5FD63" box="[1150,1244,644,670]" country="Canada" httpUri="http://grbio.org/cool/2yxb-9en0" name="Royal Ontario Museum - Entomology" pageId="66" pageNumber="67" type="Museum">ROME</collectionCode>
].
<collectingCounty id="6E0B52D9FF86E92FFB73FD7F9213FD63" box="[1270,1322,644,670]" pageId="66" pageNumber="67">This</collectingCounty>
species is known from
<collectingMunicipality id="670EB02FFF86E92FFD93FD579585FD3B" box="[534,700,684,710]" pageId="66" pageNumber="67">Negros Island</collectingMunicipality>
,
<location id="820A7C8EFF86E92FFD4CFD579474FD3B" LSID="urn:lsid:plazi:treatment:0F7C9B43FF84E92FFE8DFE5D9531FBFB:820A7C8EFF86E92FFD4CFD579474FD3B" box="[713,845,684,710]" country="Vietnam" county="This" latitude="11.434055" longLatPrecision="1" longitude="107.42933" municipality="Negros Island" name="Philippines" pageId="66" pageNumber="67" stateProvince="Dong Nai">Philippines</location>
and
<location id="820A7C8EFF86E92FFC03FD57937FFD3B" LSID="urn:lsid:plazi:treatment:0F7C9B43FF84E92FFE8DFE5D9531FBFB:820A7C8EFF86E92FFC03FD57937FFD3B" box="[902,1094,684,710]" country="Vietnam" county="This" latitude="11.434055" longLatPrecision="1" longitude="107.42933" municipality="Negros Island" name="southern Vietam" pageId="66" pageNumber="67" stateProvince="Dong Nai">southern Vietam</location>
(
<figureCitation id="1FEE36D0FF86E92FFBD3FD579385FD3B" box="[1110,1212,684,710]" captionStart="FIGURE 112" captionStartId="63.[264,367,1371,1395]" captionTargetBox="[330,1251,632,1350]" captionTargetId="figure-186@63.[330,1251,632,1396]" captionTargetPageId="63" captionText="FIGURE 112. Distribution of Ctenoplus species." figureDoi="http://doi.org/10.5281/zenodo.5064842" httpUri="https://zenodo.org/record/5064842/files/figure.png" pageId="66" pageNumber="67">Fig. 112</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="876A2A55FF86E92FFE8DFD0796BCFCE8" blockId="66.[264,1322,764,870]" box="[264,389,764,789]" pageId="66" pageNumber="67">
<emphasis id="B5A1F647FF86E92FFE8DFD0796BCFCE8" box="[264,389,764,789]" italics="true" pageId="66" pageNumber="67">Etymology</emphasis>
</paragraph>
<paragraph id="876A2A55FF86E92FFEBDFCDF9673FC9B" blockId="66.[264,1322,764,870]" pageId="66" pageNumber="67">
The species is named in honour of Dr. Claude Girard, curator of
<taxonomicName id="40D551D6FF86E92FFBABFCDF939DFCC3" baseAuthorityName="Hayek" baseAuthorityYear="1990" box="[1070,1188,804,830]" class="Insecta" family="Elateridae" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="family">Elateridae</taxonomicName>
at MNHN, Paris.
</paragraph>
<paragraph id="876A2A55FF86E92FFE8DFC679682FC48" blockId="66.[264,1323,924,1030]" box="[264,443,924,949]" pageId="66" pageNumber="67">
<emphasis id="B5A1F647FF86E92FFE8DFC679682FC48" box="[264,443,924,949]" italics="true" pageId="66" pageNumber="67">Natural history</emphasis>
</paragraph>
<paragraph id="876A2A55FF86E92FFEBDFC3F9531FBFB" blockId="66.[264,1323,924,1030]" pageId="66" pageNumber="67">
Specimens with data were collected at lights in tropical lowland forest in late May at an elevation of
<specimenCount id="91D3E1DCFF86E92FFE38FC179531FBFB" box="[445,520,1004,1030]" pageId="66" pageNumber="67" type="generic">
<quantity id="402D87B0FF86E92FFE38FC17953DFBFB" box="[445,516,1004,1030]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="66" pageNumber="67" unit="m" value="120.0">120m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
</treatment>
</document>