329 lines
46 KiB
XML
329 lines
46 KiB
XML
<document id="0B5A12CBBC6E21CE0AA65593AF4021D2" ID-CLB-Dataset="27716" ID-DOI="10.5852/ejt.2019.510" ID-GBIF-Dataset="08fe4f6b-5422-45af-bc2e-29ad35fed42f" ID-Zenodo-Dep="2616378" ID-ZooBank="BF58E108-2C6C-4A80-BD7D-47FEF5805B06" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1553845906407" checkinUser="plazi" docAuthor="Fujimoto, Shinta & Hansen, Jesper Guldberg" docDate="2019" docId="03D3878DFF9258131C7F6562DA9BFBCF" docLanguage="en" docName="ejt-510_fujimoto_hansen.pdf.imf" docOrigin="European Journal of Taxonomy 510" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Angursa abyssalis Renaud-Mornant 1981" docType="treatment" docVersion="9" lastPageNumber="7" masterDocId="FFEAFFF5FF9658151E5F6411DB46FFE9" masterDocTitle="Revision of Angursa (Arthrotardigrada: Styraconyxidae) with the description of a new species from Japan" masterLastPageNumber="19" masterPageNumber="1" pageNumber="5" updateTime="1702002503705" updateUser="plazi">
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<mods:mods id="212494F80E9EC17A7E4F44F7F2FC1235" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="6516FC3D872C3E61DC7D3D68F90FC0A7">
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<mods:title id="2AF6F36CBD365F566A1AEA03E0401D65">Revision of Angursa (Arthrotardigrada: Styraconyxidae) with the description of a new species from Japan</mods:title>
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<mods:namePart id="579D1DE03B4845B6C3711317533D6AF8">Fujimoto, Shinta</mods:namePart>
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<mods:roleTerm id="D863AC92804DB279519ECA2B353F9A37">Author</mods:roleTerm>
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<mods:namePart id="B4817B22ADD7D0AC109F117FAD18B269">Hansen, Jesper Guldberg</mods:namePart>
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<mods:typeOfResource id="D9FEDD29A66BD4A688FF11DFF39A7884">text</mods:typeOfResource>
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<mods:title id="711840CDD96F2C59D4F965CA2BE713D8">European Journal of Taxonomy</mods:title>
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</mods:titleInfo>
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<mods:part id="A207286FD21A696C788875B2C52E4591">
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<mods:date id="A45C4694778C6678E793A8AE9A240FD0">2019</mods:date>
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<mods:number id="B91BBA8E0179C907D2CC7EFA27157582">2019-03-28</mods:number>
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<mods:detail id="8A785C06F05E12624C41DDC8CB4F49BC" type="volume">
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<mods:number id="D6867FBC30DBF70B60A198207807C02B">510</mods:number>
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<mods:classification id="A26AFA421C7833AE4FA28F06CDB113FF">journal article</mods:classification>
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<mods:identifier id="9E5F90E5DCFF2D502E46B3EC90DFCBD6" type="CLB-Dataset">27716</mods:identifier>
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<mods:identifier id="4A600716532010CA5395D2BA7B09336A" type="DOI">10.5852/ejt.2019.510</mods:identifier>
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<mods:identifier id="E847752F9F2875F7E2F577B121DA31F6" type="GBIF-Dataset">08fe4f6b-5422-45af-bc2e-29ad35fed42f</mods:identifier>
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<mods:identifier id="1C166DE3F805438A93A778CF1BFAFCA6" type="Zenodo-Dep">2616378</mods:identifier>
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<mods:identifier id="12665FE2667DC3E0D31E06C208A15A9F" type="ZooBank">BF58E108-2C6C-4A80-BD7D-47FEF5805B06</mods:identifier>
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<treatment id="03D3878DFF9258131C7F6562DA9BFBCF" ID-DOI="http://doi.org/10.5281/zenodo.5941829" ID-GBIF-Taxon="154522284" ID-Zenodo-Dep="5941829" LSID="urn:lsid:plazi:treatment:03D3878DFF9258131C7F6562DA9BFBCF" httpUri="http://treatment.plazi.org/id/03D3878DFF9258131C7F6562DA9BFBCF" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5">
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<subSubSection id="C3606510FF9258111C7F6562DF52FE64" box="[544,1044,371,398]" pageId="4" pageNumber="5" type="nomenclature">
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||
<paragraph id="8BC5369BFF9258111C7F6562DF52FE64" blockId="4.[544,1044,371,432]" box="[544,1044,371,398]" pageId="4" pageNumber="5">
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||
<heading id="D08D81F7FF9258111C7F6562DF52FE64" box="[544,1044,371,398]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="5" reason="2">
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<taxonomicName id="4C7A4D18FF9258111C7F6562DF52FE64" ID-CoL="85d4a8a4-7dd8-4e00-9e0c-3faf8b02d2ad" authority="Renaud-Mornant, 1981" authorityName="Renaud-Mornant" authorityYear="1981" box="[544,1044,371,398]" class="Heterotardigrada" family="Styraconyxidae" genus="Angursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrotardigrada" pageId="4" pageNumber="5" phylum="Tardigrada" rank="species" species="abyssalis">
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<emphasis id="B90EEA89FF9258111C7F6562D9BDFE67" bold="true" box="[544,763,371,398]" italics="true" pageId="4" pageNumber="5">Angursa abyssalis</emphasis>
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<bibRefCitation id="EFEB4B6AFF9258111D5D6562DF52FE64" author="Renaud-Mornant J." box="[770,1044,371,397]" pageId="4" pageNumber="5" pagination="161 - 164" refId="ref9565" refString="Renaud-Mornant J. 1981. Deux nouveaux Angursa Pollock, 1979, du domaine abyssal (Tardigrada, Arthrotardigrada). Tethys 10: 161 - 164." type="journal article" year="1981">Renaud-Mornant, 1981</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3606510FF9258111CB86587D80BFE59" box="[743,845,406,432]" pageId="4" pageNumber="5" type="description">
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<paragraph id="8BC5369BFF9258111CB86587D80BFE59" blockId="4.[544,1044,371,432]" box="[743,845,406,432]" pageId="4" pageNumber="5">
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<figureCitation id="13412A1EFF9258111CB86587D80BFE59" box="[743,845,406,432]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="4.[189,232,1965,1991]" captionStartId-1="5.[189,232,1824,1850]" captionTargetBox-0="[610,982,1583,1909]" captionTargetBox-1="[464,1075,865,1782]" captionTargetId-0="figure@4.[556,1010,1562,1934]" captionTargetId-1="figure@5.[464,1075,865,1782]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="Fig. 2. Drawing of Angursa abyssalis Renaud-Mornant, 1981, cephalic region (ventral view) (USNM W 50900)." captionText-1="Fig. 3. Micrographs of specimens observed by Renaud-Mornant (1981) to describe Angursa abyssalis Renaud-Mornant, 1981 (A–B, E) and of Angursa sp. (C–D, F) by the present authors. A. Cephalic region (dorsal view) (MNHN AP345/560Ma). B. Cephalic region (ventral view) (MNHN AP345/560Ma). C. Cephalic region (ventral view) (MNHN AE120/78Ma). D. Cephalic region with paired globular bodies (black arrowhead) (MNHN AE120/78Ma). E. Leg IV sensory organs (MNHN AF05/77Ma). F. Leg IV sensory organ (MNHN AE120/78Ma)." httpUri-0="https://zenodo.org/record/2616382/files/figure.png" httpUri-1="https://zenodo.org/record/2616384/files/figure.png" pageId="4" pageNumber="5">Figs 2–3</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3606510FF9258111EE265C9D9D6FD44" pageId="4" pageNumber="5" type="diagnosis">
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<paragraph id="8BC5369BFF9258111EE265C9DAE9FE1B" blockId="4.[189,431,472,498]" box="[189,431,472,498]" pageId="4" pageNumber="5">
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<heading id="D08D81F7FF9258111EE265C9DAE9FE1B" bold="true" box="[189,431,472,498]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
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<emphasis id="B90EEA89FF9258111EE265C9DAE9FE1B" bold="true" box="[189,431,472,498]" pageId="4" pageNumber="5">Emended diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC5369BFF9258111EE26616D9D6FD44" blockId="4.[189,1399,518,685]" pageId="4" pageNumber="5">
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<taxonomicName id="4C7A4D18FF9258111EE26616DA66FDC8" authorityName="Pollock" authorityYear="1979" box="[189,288,519,545]" class="Heterotardigrada" family="Styraconyxidae" genus="Angursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrotardigrada" pageId="4" pageNumber="5" phylum="Tardigrada" rank="genus">
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<emphasis id="B90EEA89FF9258111EE26616DA66FDC8" box="[189,288,519,545]" italics="true" pageId="4" pageNumber="5">Angursa</emphasis>
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</taxonomicName>
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with club-shaped primary clavae longer than lateral cirri; pedestals of primary clavae and lateral cirri absent; flat, distorted-oval secondary clavae; flat, oval tertiary clavae slightly indented near external cirri; tapering cirri E with basal scapi; leg I sensory organs present; legs II and III sensory organs absent; leg IV sensory organs each as papilla enveloped in hemispherical cuticular sheath with short, apical spine; anal papillae absent.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3606510FF9258111EE266C5DA23FBB9" pageId="4" pageNumber="5" type="materials_examined">
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||
<paragraph id="8BC5369BFF9258111EE266C5DAECFD07" blockId="4.[189,426,724,750]" box="[189,426,724,750]" pageId="4" pageNumber="5">
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||
<heading id="D08D81F7FF9258111EE266C5DAECFD07" bold="true" box="[189,426,724,750]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
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<emphasis id="B90EEA89FF9258111EE266C5DAECFD07" bold="true" box="[189,426,724,750]" pageId="4" pageNumber="5">Material examined</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="8BC5369BFF9258111EB56712D87BFC8D" blockId="4.[189,1399,771,868]" pageId="4" pageNumber="5">
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<materialsCitation id="3B123CC6FF9258111EB56712D87BFC8D" ID-GBIF-Occurrence="2147483711" collectingDate="2063" collectionCode="ATLANTIC, OCEAN, MNHN" collectorName="Dinet & Ma." country="Angola" elevation="2063" location="Southeast Atlantic" pageId="4" pageNumber="5" specimenCode="AF05" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
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<emphasis id="B90EEA89FF9258111EB56712DA1AFCF4" bold="true" box="[234,348,771,797]" pageId="4" pageNumber="5">
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||
<typeStatus id="54C18839FF9258111EB56712DA1AFCF4" box="[234,348,771,797]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
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||
</emphasis>
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<collectionCode id="ED6BAE5EFF9258111EE26737DA0AFCA9" box="[189,332,806,832]" pageId="4" pageNumber="5">ATLANTIC</collectionCode>
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||
<collectionCode id="ED6BAE5EFF9258111F066737DAF8FCA9" box="[345,446,806,832]" pageId="4" pageNumber="5">OCEAN</collectionCode>
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• adult
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<specimenCount id="9D7CFD12FF9258111C766736D906FCA8" box="[553,576,807,833]" pageId="4" pageNumber="5" type="female">♀</specimenCount>
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||
;
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||
<location id="8EA56040FF9258111C0B6737D875FCA9" LSID="urn:lsid:plazi:treatment:03D3878DFF9258131C7F6562DA9BFBCF:8EA56040FF9258111C0B6737D875FCA9" box="[596,819,806,832]" country="Angola" name="Southeast Atlantic" pageId="4" pageNumber="5">Southeast Atlantic</location>
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||
, collected from off
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<collectingCountry id="F36D760BFF9258111A736737DFC0FCA9" box="[1068,1158,806,832]" name="Angola" pageId="4" pageNumber="5">Angola</collectingCountry>
|
||
;
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||
<elevation id="0057D1A8FF9258111AC86737DFB3FCA8" box="[1175,1269,806,833]" metricMagnitude="3" metricUnit="m" metricValue="2.063" pageId="4" pageNumber="5" unit="m" value="2063.0">
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||
<quantity id="4C829B7EFF9258111AC86737DFB3FCA8" box="[1175,1269,806,833]" metricMagnitude="3" metricUnit="m" metricValue="2.063" pageId="4" pageNumber="5" unit="m" value="2063.0">
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||
<date id="FFC4105BFF9258111AC86737DF94FCA9" box="[1175,1234,806,832]" pageId="4" pageNumber="5" value="2063">2063</date>
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||
m
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||
</quantity>
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||
</elevation>
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||
deep site;
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<geoCoordinate id="EE4E505CFF9258111EE26759DA4DFC8D" box="[189,267,840,868]" pageId="4" pageNumber="5">12°03′</geoCoordinate>
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3
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<collectionCode id="ED6BAE5EFF9258111F7F675BDA74FC8D" box="[288,306,842,868]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="4" pageNumber="5" type="Herbarium">S</collectionCode>
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||
,
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<geoCoordinate id="EE4E505CFF9258111F606759DACBFC8D" box="[319,397,840,868]" pageId="4" pageNumber="5">12°20′</geoCoordinate>
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5
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<collectionCode id="ED6BAE5EFF9258111FFD675BDAF1FC8D" box="[418,439,842,868]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="4" pageNumber="5" type="Herbarium">E</collectionCode>
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||
;
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<collectorName id="268F534DFF9258111F9C6758D942FC8A" box="[451,516,841,867]" pageId="4" pageNumber="5">Dinet</collectorName>
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leg.;
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<collectionCode id="ED6BAE5EFF9258111C19675BD9E6FC8D" box="[582,672,842,868]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="5">MNHN</collectionCode>
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||
<specimenCode id="DBDC9EE0FF9258111CFA6758D98AFC8D" box="[677,716,841,868]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="5">AF</specimenCode>
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||
05/77
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<collectorName id="268F534DFF9258111D4F6758D87BFC8D" box="[784,829,841,868]" pageId="4" pageNumber="5">Ma.</collectorName>
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="8BC5369BFF9258111EB5679ADA34FC05" blockId="4.[189,1399,907,1004]" pageId="4" pageNumber="5">
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<emphasis id="B90EEA89FF9258111EB5679ADAB0FC4C" bold="true" box="[234,502,907,933]" pageId="4" pageNumber="5">Additional specimens</emphasis>
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||
<collectionCode id="ED6BAE5EFF9258111EE267BFDA0AFC21" box="[189,332,942,968]" pageId="4" pageNumber="5">ATLANTIC</collectionCode>
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||
OCEAN •
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||
<materialsCitation id="3B123CC6FF9258111FB467BFDA2BFC05" ID-GBIF-Occurrence="2147483707" collectingDate="2205" collectionCode="MNHN" collectorName="Dinet & Vivier" elevation="2205" location="North Atlantic" pageId="4" pageNumber="5" specimenCode="AP345" specimenCount="2" specimenCount-adult="1" specimenCount-female="1">
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||
<specimenCount id="9D7CFD12FF9258111FB467BFD902FC21" box="[491,580,942,968]" pageId="4" pageNumber="5" type="adult">1 adult</specimenCount>
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||
<specimenCount id="9D7CFD12FF9258111C0C67BED92CFC20" box="[595,618,943,969]" pageId="4" pageNumber="5" type="female">♀</specimenCount>
|
||
;
|
||
<location id="8EA56040FF9258111CDE67BFD870FC21" LSID="urn:lsid:plazi:treatment:03D3878DFF9258131C7F6562DA9BFBCF:8EA56040FF9258111CDE67BFD870FC21" box="[641,822,942,968]" name="North Atlantic" pageId="4" pageNumber="5">North Atlantic</location>
|
||
;
|
||
<elevation id="0057D1A8FF9258111D1567BFD8EDFC20" box="[842,939,942,969]" metricMagnitude="3" metricUnit="m" metricValue="2.205" pageId="4" pageNumber="5" unit="m" value="2205.0">
|
||
<quantity id="4C829B7EFF9258111D1567BFD8EDFC20" box="[842,939,942,969]" metricMagnitude="3" metricUnit="m" metricValue="2.205" pageId="4" pageNumber="5" unit="m" value="2205.0">
|
||
<date id="FFC4105BFF9258111D1567BFD8C3FC20" box="[842,901,942,969]" pageId="4" pageNumber="5" value="2205">2205</date>
|
||
m
|
||
</quantity>
|
||
</elevation>
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||
deep;
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||
<collectorName id="268F534DFF9258111A5567BFDF0DFC21" box="[1034,1099,942,968]" pageId="4" pageNumber="5">Dinet</collectorName>
|
||
&
|
||
<collectorName id="268F534DFF9258111ADE67BFDF8CFC21" box="[1153,1226,942,968]" pageId="4" pageNumber="5">Vivier</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED6BAE5EFF9258111B4267BEDE31FC20" box="[1309,1399,943,969]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="5">MNHN</collectionCode>
|
||
<specimenCode id="DBDC9EE0FF9258111EE267C0DBA2FC05" box="[189,228,977,1004]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="5">AP</specimenCode>
|
||
345/560Ma
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC5369BFF9258111EE26002DA23FBB9" blockId="4.[189,1399,1043,1104]" pageId="4" pageNumber="5">
|
||
MEDITERRANEAN SEA •
|
||
<materialsCitation id="3B123CC6FF9258111C4D6002DA27FBB9" ID-GBIF-Occurrence="2147483703" collectionCode="MNHN" collectorName="Cassidaigne Canyon & Vitiello & Vivier" elevation="460" pageId="4" pageNumber="5" specimenCode="AE120" specimenCount="1" specimenCount-adult="1">
|
||
<specimenCount id="9D7CFD12FF9258111C4D6002D921FBC4" box="[530,615,1043,1069]" pageId="4" pageNumber="5" type="adult">1 adult</specimenCount>
|
||
;
|
||
<collectorName id="268F534DFF9258111C2A6002D82CFBC4" box="[629,874,1043,1069]" pageId="4" pageNumber="5">Cassidaigne Canyon</collectorName>
|
||
;
|
||
<elevation id="0057D1A8FF9258111D296002D884FBC4" box="[886,962,1043,1069]" metricMagnitude="2" metricUnit="m" metricValue="4.6" pageId="4" pageNumber="5" unit="m" value="460.0">
|
||
<quantity id="4C829B7EFF9258111D296002D884FBC4" box="[886,962,1043,1069]" metricMagnitude="2" metricUnit="m" metricValue="4.6" pageId="4" pageNumber="5" unit="m" value="460.0">460 m</quantity>
|
||
</elevation>
|
||
deep;
|
||
<collectorName id="268F534DFF9258111A4E6002DF2FFBC4" box="[1041,1129,1043,1069]" pageId="4" pageNumber="5">Vitiello</collectorName>
|
||
&
|
||
<collectorName id="268F534DFF9258111ACF6002DF9FFBC4" box="[1168,1241,1043,1069]" pageId="4" pageNumber="5">Vivier</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED6BAE5EFF9258111B426002DE31FBC4" box="[1309,1399,1043,1069]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="5">MNHN</collectionCode>
|
||
<specimenCode id="DBDC9EE0FF9258111EE26027DA57FBB9" box="[189,273,1078,1104]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="5">AE120</specimenCode>
|
||
/78Ma
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3606510FF9258131EE26069DA9BFBCF" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5" type="discussion">
|
||
<paragraph id="8BC5369BFF9258111EE26069DA69FB7B" blockId="4.[189,303,1144,1170]" box="[189,303,1144,1170]" pageId="4" pageNumber="5">
|
||
<heading id="D08D81F7FF9258111EE26069DA69FB7B" bold="true" box="[189,303,1144,1170]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
|
||
<emphasis id="B90EEA89FF9258111EE26069DA69FB7B" bold="true" box="[189,303,1144,1170]" pageId="4" pageNumber="5">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC5369BFF9258111EE260B7D9BCFAA4" blockId="4.[189,1399,1190,1357]" pageId="4" pageNumber="5">
|
||
There were two primary purposes for the re-examination of this taxon. One purpose was to identify the secondary and tertiary clavae, which were not reported in the original description (
|
||
<bibRefCitation id="EFEB4B6AFF9258111AF360D8DBBAFAEF" author="Renaud-Mornant J." pageId="4" pageNumber="5" pagination="161 - 164" refId="ref9565" refString="Renaud-Mornant J. 1981. Deux nouveaux Angursa Pollock, 1979, du domaine abyssal (Tardigrada, Arthrotardigrada). Tethys 10: 161 - 164." type="journal article" year="1981">Renaud-Mornant 1981</bibRefCitation>
|
||
). The second purpose was to accurately understand the morphology of the leg IV sensory organs, which was described as hemispherical papillae in the text, but drawn as elongate papillae in the figure of the original description (
|
||
<bibRefCitation id="EFEB4B6AFF9258111F836122D9ABFAA4" author="Renaud-Mornant J." box="[476,749,1331,1357]" pageId="4" pageNumber="5" pagination="161 - 164" refId="ref9565" refString="Renaud-Mornant J. 1981. Deux nouveaux Angursa Pollock, 1979, du domaine abyssal (Tardigrada, Arthrotardigrada). Tethys 10: 161 - 164." type="journal article" year="1981">Renaud-Mornant 1981</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC5369BFF9258101EE26165D996FE15" blockId="4.[189,1399,1396,1493]" lastBlockId="5.[189,1399,267,508]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
|
||
Unfortunately, the secondary and tertiary clavae were not recognised in the
|
||
<typeStatus id="54C18839FF9258111AC16165DE4EFA67" box="[1182,1288,1396,1422]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
|
||
, MNHN AF05/77Ma. However, they were evident in the additional specimen, MNHN AP345/560Ma. In this specimen, the paired secondary clavae were positioned in the dorsal area between the median cirrus and the primary clavae and each was in the form of a flat, distorted-oval lobe (the complete contour was not recognisable) (
|
||
<figureCitation id="13412A1EFF9358101F34653FDAFEFEA0" box="[363,440,302,329]" captionStart="Fig" captionStartId="4.[189,232,1965,1991]" captionTargetBox="[610,982,1583,1909]" captionTargetId="figure@4.[556,1010,1562,1934]" captionTargetPageId="4" captionText="Fig. 2. Drawing of Angursa abyssalis Renaud-Mornant, 1981, cephalic region (ventral view) (USNM W 50900)." httpUri="https://zenodo.org/record/2616382/files/figure.png" pageId="5" pageNumber="6">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation id="13412A1EFF9358101F9A653EDAAAFEA0" box="[453,492,303,329]" captionStart="Fig" captionStartId="5.[189,232,1824,1850]" captionTargetBox="[464,1075,865,1782]" captionTargetId="figure@5.[464,1075,865,1782]" captionTargetPageId="5" captionText="Fig. 3. Micrographs of specimens observed by Renaud-Mornant (1981) to describe Angursa abyssalis Renaud-Mornant, 1981 (A–B, E) and of Angursa sp. (C–D, F) by the present authors. A. Cephalic region (dorsal view) (MNHN AP345/560Ma). B. Cephalic region (ventral view) (MNHN AP345/560Ma). C. Cephalic region (ventral view) (MNHN AE120/78Ma). D. Cephalic region with paired globular bodies (black arrowhead) (MNHN AE120/78Ma). E. Leg IV sensory organs (MNHN AF05/77Ma). F. Leg IV sensory organ (MNHN AE120/78Ma)." httpUri="https://zenodo.org/record/2616384/files/figure.png" pageId="5" pageNumber="6">3A</figureCitation>
|
||
). The paired tertiary clavae were position ventrally, immediately posterior to the external cirri and each was in the form of a flat, transversally-elongated lobe with a slight indentation where the external cirrus arises (
|
||
<figureCitation id="13412A1EFF9358101C6B6564D939FE66" box="[564,639,373,399]" captionStart="Fig" captionStartId="4.[189,232,1965,1991]" captionTargetBox="[610,982,1583,1909]" captionTargetId="figure@4.[556,1010,1562,1934]" captionTargetPageId="4" captionText="Fig. 2. Drawing of Angursa abyssalis Renaud-Mornant, 1981, cephalic region (ventral view) (USNM W 50900)." httpUri="https://zenodo.org/record/2616382/files/figure.png" pageId="5" pageNumber="6">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation id="13412A1EFF9358101CD56564D9E9FE66" box="[650,687,373,399]" captionStart="Fig" captionStartId="5.[189,232,1824,1850]" captionTargetBox="[464,1075,865,1782]" captionTargetId="figure@5.[464,1075,865,1782]" captionTargetPageId="5" captionText="Fig. 3. Micrographs of specimens observed by Renaud-Mornant (1981) to describe Angursa abyssalis Renaud-Mornant, 1981 (A–B, E) and of Angursa sp. (C–D, F) by the present authors. A. Cephalic region (dorsal view) (MNHN AP345/560Ma). B. Cephalic region (ventral view) (MNHN AP345/560Ma). C. Cephalic region (ventral view) (MNHN AE120/78Ma). D. Cephalic region with paired globular bodies (black arrowhead) (MNHN AE120/78Ma). E. Leg IV sensory organs (MNHN AF05/77Ma). F. Leg IV sensory organ (MNHN AE120/78Ma)." httpUri="https://zenodo.org/record/2616384/files/figure.png" pageId="5" pageNumber="6">3B</figureCitation>
|
||
). In the other additional specimen, MNHN AE120/78Ma, the secondary clavae were not recognised, but a paired structure indicative of clavae (tertiary clavae?) was observed in the latero-ventral position of the cephalic region (
|
||
<figureCitation id="13412A1EFF9358101DC965AAD8B5FE3C" box="[918,1011,443,469]" captionStart="Fig" captionStartId="5.[189,232,1824,1850]" captionTargetBox="[464,1075,865,1782]" captionTargetId="figure@5.[464,1075,865,1782]" captionTargetPageId="5" captionText="Fig. 3. Micrographs of specimens observed by Renaud-Mornant (1981) to describe Angursa abyssalis Renaud-Mornant, 1981 (A–B, E) and of Angursa sp. (C–D, F) by the present authors. A. Cephalic region (dorsal view) (MNHN AP345/560Ma). B. Cephalic region (ventral view) (MNHN AP345/560Ma). C. Cephalic region (ventral view) (MNHN AE120/78Ma). D. Cephalic region with paired globular bodies (black arrowhead) (MNHN AE120/78Ma). E. Leg IV sensory organs (MNHN AF05/77Ma). F. Leg IV sensory organ (MNHN AE120/78Ma)." httpUri="https://zenodo.org/record/2616384/files/figure.png" pageId="5" pageNumber="6">Fig. 3C</figureCitation>
|
||
). Further, a paired globular body fringing the mouth was recognised (
|
||
<figureCitation id="13412A1EFF9358101C3B65F3D985FE15" box="[612,707,482,508]" captionStart="Fig" captionStartId="5.[189,232,1824,1850]" captionTargetBox="[464,1075,865,1782]" captionTargetId="figure@5.[464,1075,865,1782]" captionTargetPageId="5" captionText="Fig. 3. Micrographs of specimens observed by Renaud-Mornant (1981) to describe Angursa abyssalis Renaud-Mornant, 1981 (A–B, E) and of Angursa sp. (C–D, F) by the present authors. A. Cephalic region (dorsal view) (MNHN AP345/560Ma). B. Cephalic region (ventral view) (MNHN AP345/560Ma). C. Cephalic region (ventral view) (MNHN AE120/78Ma). D. Cephalic region with paired globular bodies (black arrowhead) (MNHN AE120/78Ma). E. Leg IV sensory organs (MNHN AF05/77Ma). F. Leg IV sensory organ (MNHN AE120/78Ma)." httpUri="https://zenodo.org/record/2616384/files/figure.png" pageId="5" pageNumber="6">Fig. 3D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF056613FF9258111EE263BCDA7CF802" httpUri="https://zenodo.org/record/2616382/files/figure.png" pageId="4" pageNumber="5" startId="4.[189,232,1965,1991]" targetBox="[610,982,1583,1909]" targetPageId="4">
|
||
<paragraph id="8BC5369BFF9258111EE263BCDA7CF802" blockId="4.[189,1399,1965,2027]" pageId="4" pageNumber="5">
|
||
<emphasis id="B90EEA89FF9258111EE263BCDA4AF82E" bold="true" box="[189,268,1965,1991]" pageId="4" pageNumber="5">Fig. 2.</emphasis>
|
||
Drawing of
|
||
<taxonomicName id="4C7A4D18FF9258111FF963BFD8DAF82E" authority="Renaud-Mornant, 1981" authorityName="Renaud-Mornant" authorityYear="1981" box="[422,924,1965,1992]" class="Heterotardigrada" family="Styraconyxidae" genus="Angursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrotardigrada" pageId="4" pageNumber="5" phylum="Tardigrada" rank="species" species="abyssalis">
|
||
<emphasis id="B90EEA89FF9258111FF963BFD938F82E" box="[422,638,1965,1992]" italics="true" pageId="4" pageNumber="5">Angursa abyssalis</emphasis>
|
||
Renaud-Mornant, 1981
|
||
</taxonomicName>
|
||
, cephalic region (ventral view) (USNM
|
||
<accessionNumber id="9429AB78FF9258111EE263C0DA68F802" box="[189,302,2000,2027]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/W50900" pageId="4" pageNumber="5" type="EnaNcbi">W 50900</accessionNumber>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC5369BFF9358101EE26632D91EFCF9" blockId="5.[189,1399,547,785]" pageId="5" pageNumber="6">
|
||
The leg IV sensory organs were recognised in all three specimens.The
|
||
<typeStatus id="54C18839FF9358101D896632DF7BFDD4" box="[982,1085,547,573]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
|
||
and MNHN AP345/560Ma had leg IV sensory organs consisting of papillae enveloped in hemispherical cuticular sheaths with short, apical spines (
|
||
<figureCitation id="13412A1EFF9358101F3B6678DAFAFD6D" box="[356,444,617,644]" captionStart="Fig" captionStartId="5.[189,232,1824,1850]" captionTargetBox="[464,1075,865,1782]" captionTargetId="figure@5.[464,1075,865,1782]" captionTargetPageId="5" captionText="Fig. 3. Micrographs of specimens observed by Renaud-Mornant (1981) to describe Angursa abyssalis Renaud-Mornant, 1981 (A–B, E) and of Angursa sp. (C–D, F) by the present authors. A. Cephalic region (dorsal view) (MNHN AP345/560Ma). B. Cephalic region (ventral view) (MNHN AP345/560Ma). C. Cephalic region (ventral view) (MNHN AE120/78Ma). D. Cephalic region with paired globular bodies (black arrowhead) (MNHN AE120/78Ma). E. Leg IV sensory organs (MNHN AF05/77Ma). F. Leg IV sensory organ (MNHN AE120/78Ma)." httpUri="https://zenodo.org/record/2616384/files/figure.png" pageId="5" pageNumber="6">Fig. 3E</figureCitation>
|
||
) similar to that described in the text of the original description (
|
||
<bibRefCitation id="EFEB4B6AFF9358101AF36678DBBAFD4E" author="Renaud-Mornant J." pageId="5" pageNumber="6" pagination="161 - 164" refId="ref9565" refString="Renaud-Mornant J. 1981. Deux nouveaux Angursa Pollock, 1979, du domaine abyssal (Tardigrada, Arthrotardigrada). Tethys 10: 161 - 164." type="journal article" year="1981">Renaud-Mornant 1981</bibRefCitation>
|
||
). On the other hand, the leg IV sensory organs of MNHN AE120/78Ma lacked the enveloping cuticular sheaths and the papillae appeared slightly elongate and also constricted at the base (
|
||
<figureCitation id="13412A1EFF9358101AA766A1DE08FD23" box="[1272,1358,688,714]" captionStart="Fig" captionStartId="5.[189,232,1824,1850]" captionTargetBox="[464,1075,865,1782]" captionTargetId="figure@5.[464,1075,865,1782]" captionTargetPageId="5" captionText="Fig. 3. Micrographs of specimens observed by Renaud-Mornant (1981) to describe Angursa abyssalis Renaud-Mornant, 1981 (A–B, E) and of Angursa sp. (C–D, F) by the present authors. A. Cephalic region (dorsal view) (MNHN AP345/560Ma). B. Cephalic region (ventral view) (MNHN AP345/560Ma). C. Cephalic region (ventral view) (MNHN AE120/78Ma). D. Cephalic region with paired globular bodies (black arrowhead) (MNHN AE120/78Ma). E. Leg IV sensory organs (MNHN AF05/77Ma). F. Leg IV sensory organ (MNHN AE120/78Ma)." httpUri="https://zenodo.org/record/2616384/files/figure.png" pageId="5" pageNumber="6">Fig. 3F</figureCitation>
|
||
) as in fig. 1A of
|
||
<bibRefCitation id="EFEB4B6AFF9358101F1066C2D936FD04" author="Renaud-Mornant J." box="[335,624,723,749]" pageId="5" pageNumber="6" pagination="161 - 164" refId="ref9565" refString="Renaud-Mornant J. 1981. Deux nouveaux Angursa Pollock, 1979, du domaine abyssal (Tardigrada, Arthrotardigrada). Tethys 10: 161 - 164." type="journal article" year="1981">Renaud-Mornant (1981)</bibRefCitation>
|
||
. Further, it is clear that fig. 1C of
|
||
<bibRefCitation id="EFEB4B6AFF9358101DA966C2DE55FD04" author="Renaud-Mornant J." box="[1014,1299,723,749]" pageId="5" pageNumber="6" pagination="161 - 164" refId="ref9565" refString="Renaud-Mornant J. 1981. Deux nouveaux Angursa Pollock, 1979, du domaine abyssal (Tardigrada, Arthrotardigrada). Tethys 10: 161 - 164." type="journal article" year="1981">Renaud-Mornant (1981)</bibRefCitation>
|
||
is based on MNHN AE120/78Ma (
|
||
<figureCitation id="13412A1EFF9358101FAE66E7D90CFCF9" box="[497,586,758,784]" captionStart="Fig" captionStartId="5.[189,232,1824,1850]" captionTargetBox="[464,1075,865,1782]" captionTargetId="figure@5.[464,1075,865,1782]" captionTargetPageId="5" captionText="Fig. 3. Micrographs of specimens observed by Renaud-Mornant (1981) to describe Angursa abyssalis Renaud-Mornant, 1981 (A–B, E) and of Angursa sp. (C–D, F) by the present authors. A. Cephalic region (dorsal view) (MNHN AP345/560Ma). B. Cephalic region (ventral view) (MNHN AP345/560Ma). C. Cephalic region (ventral view) (MNHN AE120/78Ma). D. Cephalic region with paired globular bodies (black arrowhead) (MNHN AE120/78Ma). E. Leg IV sensory organs (MNHN AF05/77Ma). F. Leg IV sensory organ (MNHN AE120/78Ma)." httpUri="https://zenodo.org/record/2616384/files/figure.png" pageId="5" pageNumber="6">Fig. 3F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF056613FF9358101EE26331D9BBF803" httpUri="https://zenodo.org/record/2616384/files/figure.png" pageId="5" pageNumber="6" startId="5.[189,232,1824,1850]" targetBox="[464,1075,865,1782]" targetPageId="5">
|
||
<paragraph id="8BC5369BFF9358101EE26331D9BBF803" blockId="5.[189,1399,1824,2026]" pageId="5" pageNumber="6">
|
||
<emphasis id="B90EEA89FF9358101EE26331DA4AF8D3" bold="true" box="[189,268,1824,1850]" pageId="5" pageNumber="6">Fig. 3.</emphasis>
|
||
Micrographs of specimens observed by Renaud-Mornant (1981) to describe
|
||
<taxonomicName id="4C7A4D18FF9358101AC06331DA88F8B4" authority="Renaud-Mornant, 1981" authorityName="Renaud-Mornant" authorityYear="1981" class="Heterotardigrada" family="Styraconyxidae" genus="Angursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrotardigrada" pageId="5" pageNumber="6" phylum="Tardigrada" rank="species" species="abyssalis">
|
||
<emphasis id="B90EEA89FF9358101AC06331DE31F8D3" box="[1183,1399,1824,1850]" italics="true" pageId="5" pageNumber="6">Angursa abyssalis</emphasis>
|
||
Renaud-Mornant, 1981
|
||
</taxonomicName>
|
||
(A–B, E) and of
|
||
<taxonomicName id="4C7A4D18FF9358101CCE6352D9B2F8B4" authorityName="Pollock" authorityYear="1979" box="[657,756,1859,1885]" class="Heterotardigrada" family="Styraconyxidae" genus="Angursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrotardigrada" pageId="5" pageNumber="6" phylum="Tardigrada" rank="genus">
|
||
<emphasis id="B90EEA89FF9358101CCE6352D9B2F8B4" box="[657,756,1859,1885]" italics="true" pageId="5" pageNumber="6">Angursa</emphasis>
|
||
</taxonomicName>
|
||
sp. (C–D, F) by the present authors.
|
||
<emphasis id="B90EEA89FF9358101AC46352DFF6F8B4" bold="true" box="[1179,1200,1859,1885]" pageId="5" pageNumber="6">A</emphasis>
|
||
. Cephalic region (dorsal view) (MNHN AP345/560Ma).
|
||
<emphasis id="B90EEA89FF9358101CFD6377D9F0F869" bold="true" box="[674,694,1894,1920]" pageId="5" pageNumber="6">B</emphasis>
|
||
. Cephalic region (ventral view) (MNHN AP345/560Ma).
|
||
<emphasis id="B90EEA89FF9358101EE26398DB94F84A" bold="true" box="[189,210,1929,1955]" pageId="5" pageNumber="6">C</emphasis>
|
||
. Cephalic region (ventral view) (MNHN AE120/78Ma).
|
||
<emphasis id="B90EEA89FF9358101DD1639BD8E5F84D" bold="true" box="[910,931,1930,1956]" pageId="5" pageNumber="6">D</emphasis>
|
||
. Cephalic region with paired globular bodies (black arrowhead) (MNHN AE120/78Ma).
|
||
<emphasis id="B90EEA89FF9358101D7C63BCD871F82E" bold="true" box="[803,823,1965,1991]" pageId="5" pageNumber="6">E</emphasis>
|
||
. Leg IV sensory organs (MNHN AF05/77Ma).
|
||
<emphasis id="B90EEA89FF9358101EE263C1DB89F803" bold="true" box="[189,207,2000,2026]" pageId="5" pageNumber="6">F</emphasis>
|
||
. Leg IV sensory organ (MNHN AE120/78Ma).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC5369BFF9058131EE2651ADE09FE7B" blockId="6.[189,1398,267,403]" pageId="6" pageNumber="7">
|
||
Other morphological characters of the three specimens confirm the original description (
|
||
<bibRefCitation id="EFEB4B6AFF9058131AF3651ADBBFFEA0" author="Renaud-Mornant J." pageId="6" pageNumber="7" pagination="161 - 164" refId="ref9565" refString="Renaud-Mornant J. 1981. Deux nouveaux Angursa Pollock, 1979, du domaine abyssal (Tardigrada, Arthrotardigrada). Tethys 10: 161 - 164." type="journal article" year="1981">Renaud-Mornant 1981</bibRefCitation>
|
||
) except that a scapus was observed in one of the external cirri of the MNHN AP345/560Ma (
|
||
<figureCitation id="13412A1EFF9058131B7D653FDE2EFEA0" box="[1314,1384,302,329]" captionStart="Fig" captionStartId="4.[189,232,1965,1991]" captionTargetBox="[610,982,1583,1909]" captionTargetId="figure@4.[556,1010,1562,1934]" captionTargetPageId="4" captionText="Fig. 2. Drawing of Angursa abyssalis Renaud-Mornant, 1981, cephalic region (ventral view) (USNM W 50900)." httpUri="https://zenodo.org/record/2616382/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
|
||
). Here we provide measurements of the
|
||
<typeStatus id="54C18839FF9058131C246543D9A4FE85" box="[635,738,338,364]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
|
||
missing in the original description: internal cirrus, 3 μm; external cirrus, 6 μm; primary clava, 10 μm; leg IV sensory organ, 5 μm long and 3 μm in diameter.
|
||
</paragraph>
|
||
<paragraph id="8BC5369BFF9058131EE265ABDF88FD4E" blockId="6.[189,1399,441,679]" pageId="6" pageNumber="7">
|
||
Our re-examination reveals that the term “balloon-shaped” used by
|
||
<bibRefCitation id="EFEB4B6AFF9058131D8465ABDFC3FE3D" author="Bussau C." box="[987,1157,442,468]" pageId="6" pageNumber="7" pagination="79 - 91" refId="ref8261" refString="Bussau C. 1992. New deep-sea Tardigrada (Arthrotardigrada, Halechiniscidae) from manganese nodule area of the eastern South Pacific. Zoologica Scripta 21: 79 - 91. https: // doi. org / 10.1111 / j. 1463 - 6409.1992. tb 00311. x" type="journal article" year="1992">Bussau (1992)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFEB4B6AFF9058131AE165A8DA4AFE1E" author="Villora-Moreno S." pageId="6" pageNumber="7" pagination="336 - 341" refId="ref9804" refString="Villora-Moreno S. 1998. Deep-sea Tardigrada from South Shetland Islands (Antarctica) with description of Angursa antarctica sp. nov. (Arthrotardigrada, Halechiniscidae). Polar Biology 19: 336 - 341. https: // doi. org / 10.1007 / s 003000050255" type="journal article" year="1998">Villora-Moreno (1998)</bibRefCitation>
|
||
(probably based on the figure in the original description of
|
||
<bibRefCitation id="EFEB4B6AFF9058131D8865CCDFA2FE1E" author="Renaud-Mornant J." box="[983,1252,477,503]" pageId="6" pageNumber="7" pagination="161 - 164" refId="ref9565" refString="Renaud-Mornant J. 1981. Deux nouveaux Angursa Pollock, 1979, du domaine abyssal (Tardigrada, Arthrotardigrada). Tethys 10: 161 - 164." type="journal article" year="1981">Renaud-Mornant 1981</bibRefCitation>
|
||
) to describe the sensory organs of leg IV of
|
||
<taxonomicName id="4C7A4D18FF9058131C146611D99AFDF3" authorityName="Renaud-Mornant" authorityYear="1981" box="[587,732,512,538]" class="Heterotardigrada" family="Styraconyxidae" genus="Angursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="7" phylum="Tardigrada" rank="species" species="abyssalis">
|
||
<emphasis id="B90EEA89FF9058131C146611D99AFDF3" box="[587,732,512,538]" italics="true" pageId="6" pageNumber="7">A. abyssalis</emphasis>
|
||
</taxonomicName>
|
||
is inadequate since those of the
|
||
<typeStatus id="54C18839FF9058131A2A6611DF9AFDF3" box="[1141,1244,512,538]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
|
||
and MNHN AP345/560Ma had a hemispherical appearance. Further, we consider MNHN AE120/78Ma with “balloon-shaped” leg IV sensory organs (in the sense of
|
||
<bibRefCitation id="EFEB4B6AFF9058131D046656D8B5FD89" author="Bussau C." box="[859,1011,582,609]" pageId="6" pageNumber="7" pagination="79 - 91" refId="ref8261" refString="Bussau C. 1992. New deep-sea Tardigrada (Arthrotardigrada, Halechiniscidae) from manganese nodule area of the eastern South Pacific. Zoologica Scripta 21: 79 - 91. https: // doi. org / 10.1111 / j. 1463 - 6409.1992. tb 00311. x" type="journal article" year="1992">Bussau 1992</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFEB4B6AFF9058131A726657DE6FFD89" author="Villora-Moreno S." box="[1069,1321,582,608]" pageId="6" pageNumber="7" pagination="336 - 341" refId="ref9804" refString="Villora-Moreno S. 1998. Deep-sea Tardigrada from South Shetland Islands (Antarctica) with description of Angursa antarctica sp. nov. (Arthrotardigrada, Halechiniscidae). Polar Biology 19: 336 - 341. https: // doi. org / 10.1007 / s 003000050255" type="journal article" year="1998">Villora-Moreno 1998</bibRefCitation>
|
||
) as an undescribed species. The difference in the morphology of the tertiary clavae also supports this view. However, MNHN AE120/78Ma is not in an acceptable condition for species description.
|
||
</paragraph>
|
||
<paragraph id="8BC5369BFF9058131EE266DFDA9BFBCF" blockId="6.[189,1399,718,1062]" pageId="6" pageNumber="7">
|
||
Based on the observation of the
|
||
<typeStatus id="54C18839FF9058131C0F66DFD9F1FD01" box="[592,695,718,744]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
|
||
and MNHN AP345/560Ma,
|
||
<taxonomicName id="4C7A4D18FF9058131A4466DEDFB0FD01" authorityName="Renaud-Mornant" authorityYear="1981" box="[1051,1270,718,745]" class="Heterotardigrada" family="Styraconyxidae" genus="Angursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="7" phylum="Tardigrada" rank="species" species="abyssalis">
|
||
<emphasis id="B90EEA89FF9058131A4466DEDFB0FD01" box="[1051,1270,718,745]" italics="true" pageId="6" pageNumber="7">Angursa abyssalis</emphasis>
|
||
</taxonomicName>
|
||
resembles
|
||
<taxonomicName id="4C7A4D18FF9058131EE266E3DA01FCE2" authorityName="Noda" authorityYear="1985" box="[189,327,753,780]" class="Heterotardigrada" family="Styraconyxidae" genus="Angursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="7" phylum="Tardigrada" rank="species" species="clavifera">
|
||
<emphasis id="B90EEA89FF9058131EE266E3DA01FCE2" box="[189,327,753,780]" italics="true" pageId="6" pageNumber="7">A. clavifera</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7A4D18FF9058131FDB66E3D947FCE2" authorityName="Bussau" authorityYear="1992" box="[388,513,753,780]" class="Heterotardigrada" family="Styraconyxidae" genus="Angursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="7" phylum="Tardigrada" rank="species" species="capsula">
|
||
<emphasis id="B90EEA89FF9058131FDB66E3D947FCE2" box="[388,513,753,780]" italics="true" pageId="6" pageNumber="7">A. capsula</emphasis>
|
||
</taxonomicName>
|
||
by the club-shaped primary clavae longer than the lateral cirri, the leg IV sensory organs in hemispherical shape, and the lack of the anal papillae (
|
||
<bibRefCitation id="EFEB4B6AFF9058131A6C6704DFFBFCC6" author="Noda H." box="[1075,1213,788,815]" pageId="6" pageNumber="7" pagination="269 - 276" refId="ref9298" refString="Noda H. 1985. Description of a new subspecies of Angursa bicuspis Pollock (Heterotardigrada, Halechiniscidae) from Tanabe Bay, Japan. Publications of the Seto Marine Biological Laboratory 30: 269 - 276. Available from https: // hdl. handle. net / 2433 / 176110 [accessed 4 Feb. 2019]." type="journal article" year="1985">Noda 1985</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEB4B6AFF9058131A936704DE2CFCC6" author="Bussau C." box="[1228,1386,789,815]" pageId="6" pageNumber="7" pagination="79 - 91" refId="ref8261" refString="Bussau C. 1992. New deep-sea Tardigrada (Arthrotardigrada, Halechiniscidae) from manganese nodule area of the eastern South Pacific. Zoologica Scripta 21: 79 - 91. https: // doi. org / 10.1111 / j. 1463 - 6409.1992. tb 00311. x" type="journal article" year="1992">Bussau 1992</bibRefCitation>
|
||
). However, it differs from
|
||
<taxonomicName id="4C7A4D18FF9058131F816729D920FCBB" authorityName="Noda" authorityYear="1985" box="[478,614,824,850]" class="Heterotardigrada" family="Styraconyxidae" genus="Angursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="7" phylum="Tardigrada" rank="species" species="clavifera">
|
||
<emphasis id="B90EEA89FF9058131F816729D920FCBB" box="[478,614,824,850]" italics="true" pageId="6" pageNumber="7">A. clavifera</emphasis>
|
||
</taxonomicName>
|
||
by the larger secondary clavae, the tertiary clavae slightly indented around the external cirri, and the absence of sensory organs on legs II and III (
|
||
<bibRefCitation id="EFEB4B6AFF9058131A30674ADFBFFC9F" author="Noda H." box="[1135,1273,859,886]" pageId="6" pageNumber="7" pagination="269 - 276" refId="ref9298" refString="Noda H. 1985. Description of a new subspecies of Angursa bicuspis Pollock (Heterotardigrada, Halechiniscidae) from Tanabe Bay, Japan. Publications of the Seto Marine Biological Laboratory 30: 269 - 276. Available from https: // hdl. handle. net / 2433 / 176110 [accessed 4 Feb. 2019]." type="journal article" year="1985">Noda 1985</bibRefCitation>
|
||
). It differs from
|
||
<taxonomicName id="4C7A4D18FF9058131F5D676EDAC4FC71" authorityName="Bussau" authorityYear="1992" box="[258,386,894,921]" class="Heterotardigrada" family="Styraconyxidae" genus="Angursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="7" phylum="Tardigrada" rank="species" species="capsula">
|
||
<emphasis id="B90EEA89FF9058131F5D676EDAC4FC71" box="[258,386,894,921]" italics="true" pageId="6" pageNumber="7">A. capsula</emphasis>
|
||
</taxonomicName>
|
||
by the larger and closer situated pair of secondary clavae (3 μm apart in MNHN AP345/560Ma and 9 μm apart in
|
||
<taxonomicName id="4C7A4D18FF9058131C0D67B3D996FC52" authorityName="Bussau" authorityYear="1992" box="[594,720,929,956]" class="Heterotardigrada" family="Styraconyxidae" genus="Angursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="7" phylum="Tardigrada" rank="species" species="capsula">
|
||
<emphasis id="B90EEA89FF9058131C0D67B3D996FC52" box="[594,720,929,956]" italics="true" pageId="6" pageNumber="7">A. capsula</emphasis>
|
||
</taxonomicName>
|
||
measured from the drawing of a paratypic female in the original description see
|
||
<bibRefCitation id="EFEB4B6AFF9058131FBE67D4D9C6FC36" author="Bussau C." box="[481,640,965,991]" pageId="6" pageNumber="7" pagination="79 - 91" refId="ref8261" refString="Bussau C. 1992. New deep-sea Tardigrada (Arthrotardigrada, Halechiniscidae) from manganese nodule area of the eastern South Pacific. Zoologica Scripta 21: 79 - 91. https: // doi. org / 10.1111 / j. 1463 - 6409.1992. tb 00311. x" type="journal article" year="1992">Bussau 1992</bibRefCitation>
|
||
), the tertiary clavae slightly indented around the external cirri (
|
||
<bibRefCitation id="EFEB4B6AFF9058131E9A67F9DA19FBEB" author="Bussau C." box="[197,351,1000,1026]" pageId="6" pageNumber="7" pagination="79 - 91" refId="ref8261" refString="Bussau C. 1992. New deep-sea Tardigrada (Arthrotardigrada, Halechiniscidae) from manganese nodule area of the eastern South Pacific. Zoologica Scripta 21: 79 - 91. https: // doi. org / 10.1111 / j. 1463 - 6409.1992. tb 00311. x" type="journal article" year="1992">Bussau 1992</bibRefCitation>
|
||
) and a much shorter cirrus E (10 μm (body length, 141 μm) in
|
||
<taxonomicName id="4C7A4D18FF9058131A1A67F9DF94FBEB" authorityName="Renaud-Mornant" authorityYear="1981" box="[1093,1234,1000,1026]" class="Heterotardigrada" family="Styraconyxidae" genus="Angursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="7" phylum="Tardigrada" rank="species" species="abyssalis">
|
||
<emphasis id="B90EEA89FF9058131A1A67F9DF94FBEB" box="[1093,1234,1000,1026]" italics="true" pageId="6" pageNumber="7">A. abyssalis</emphasis>
|
||
</taxonomicName>
|
||
versus 19 μm (170 μm) in
|
||
<taxonomicName id="4C7A4D18FF9058131F0F601ADA8DFBCC" authorityName="Bussau" authorityYear="1992" box="[336,459,1035,1061]" class="Heterotardigrada" family="Styraconyxidae" genus="Angursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="7" phylum="Tardigrada" rank="species" species="capsula">
|
||
<emphasis id="B90EEA89FF9058131F0F601ADA8DFBCC" box="[336,459,1035,1061]" italics="true" pageId="6" pageNumber="7">A. capsula</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |