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<document id="F50108EC4C3513C61138A78645450D2C" ID-DOI="10.11646/zootaxa.4040.1.1" ID-GBIF-Dataset="3b927167-5686-4666-861f-75b65e82fcdb" ID-ISSN="1175-5326" ID-Zenodo-Dep="289973" ID-ZooBank="51219593-CC2D-4077-B9FA-5E2F3255FC24" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461333548483" checkinUser="plazi" docAuthor="Cardoso, Giovanna Monticelli, Taiti, Stefano &amp; Sfenthourakis, Spyros" docDate="2015" docId="03B287A20E31403AFF06F8F81F93FE45" docLanguage="en" docName="zt04040p016.pdf" docOrigin="Zootaxa 4040 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Levantoniscus wahrmani Strouhal 1968" docType="treatment" docVersion="6" lastPageNumber="15" masterDocId="FF8BFFDA0E394034FF91FFD51D0EFFB1" masterDocTitle="A new genus of Trachelipodidae Strouhal, 1953 (Crustacea, Isopoda, Oniscidea) from the eastern Mediterranean" masterLastPageNumber="16" masterPageNumber="1" pageNumber="9" updateTime="1698618859390" updateUser="plazi">
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<mods:title id="B2B04634CB6D6E8D7087F7566A810C2B">A new genus of Trachelipodidae Strouhal, 1953 (Crustacea, Isopoda, Oniscidea) from the eastern Mediterranean</mods:title>
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<emphasis id="B96FEAA60E31403CFF06F8F81CC6F8F6" bold="true" box="[151,456,1837,1863]" italics="true" pageId="8" pageNumber="9">Levantoniscus wahrmani</emphasis>
(
<bibRefCitation id="EF8A4B450E31403CFE46F8FB1F81F8F9" author="Strouhal" box="[471,655,1838,1864]" pageId="8" pageNumber="9" refString="Strouhal, H. (1968) Eine neue, vorderasiatische Bathytropa - Art (Oniscoidea, Oniscidae, Bathytropinae). Crustaceana, 15, 67 - 78. http: // dx. doi. org / 10.1163 / 156854068 X 00764" type="journal article" year="1968">Strouhal, 1968</bibRefCitation>
)
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<figureCitation id="13202A310E31403CFF06F8851C0CF8D8" box="[151,258,1872,1897]" captionStart="FIGURE 9" captionStartId="11.[151,249,1725,1747]" captionTargetBox="[161,1419,202,1687]" captionTargetId="figure@11.[151,1436,193,1704]" captionTargetPageId="11" captionText="FIGURE 9. Levantoniscus wahrmani. ♀: A, adult specimen, dorsal view; B, lateral view; C, dorsal scale-seta; D, cephalon, dorsal view; E, cephalon, posterior view; F, cephalon, frontal view, G, pleonite 5, telson and uropods, dorsal view. ♂: H, antennula; I, antenna." httpUri="https://zenodo.org/record/289982/files/figure.png" pageId="8" pageNumber="9">Figures 9</figureCitation>
,
<figureCitation id="13202A310E31403CFE9FF8841C25F8D8" box="[270,299,1873,1897]" captionStart="FIGURE 10" captionStartId="12.[151,249,1947,1969]" captionTargetBox="[151,1417,207,1909]" captionTargetId="figure@12.[151,1436,193,1925]" captionTargetPageId="12" captionText="FIGURE 10. Levantoniscus wahrmani. ♂: A, uropod; B, pereopod 1; C, pereopod 7; D, pleopod 1, dorsal view; E, pleopod 2, dorsal view; F, pleopod 2 exopodite, ventral view; G, pleopod 3 exopodite, ventral view; H, pleopod 4 exopodite, ventral view; I, pleopod 5 exopodite, ventral view." httpUri="https://zenodo.org/record/289983/files/figure.png" pageId="8" pageNumber="9">10</figureCitation>
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<emphasis id="B96FEAA60E31403CFF06F8461C78F818" box="[151,374,1939,1961]" italics="true" pageId="8" pageNumber="9">Bathytropa wahrmani</emphasis>
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<bibRefCitation id="EF8A4B450E31403CFEEDF8461F1EF818" author="Strouhal" box="[380,528,1939,1961]" pageId="8" pageNumber="9" refString="Strouhal, H. (1968) Eine neue, vorderasiatische Bathytropa - Art (Oniscoidea, Oniscidae, Bathytropinae). Crustaceana, 15, 67 - 78. http: // dx. doi. org / 10.1163 / 156854068 X 00764" type="journal article" year="1968">Strouhal 1968</bibRefCitation>
: 69
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,
<figureCitation id="13202A310E31403CFDD1F8461F72F818" box="[576,636,1939,1961]" captionStart="FIGURE 1" captionStartId="3.[151,249,1821,1843]" captionTargetBox="[173,1402,366,1772]" captionTargetId="figure@3.[160,1426,363,1787]" captionTargetPageId="3" captionText="FIGURE 1. Levantoniscus bicostulatus n. sp. Paratype ♀: A, adult specimen, dorsal view; B, adult specimen, lateral view; C, dorsal scale-seta; D, cephalon, frontal view. Paratype ♂: E, antennula; F, antenna." httpUri="https://zenodo.org/record/289974/files/figure.png" pageId="8" pageNumber="9">Figs 1</figureCitation>
25;
<treatmentCitation id="0ABA10A50E31403CFD22F8461E51F818" author="Caruso" box="[691,863,1939,1961]" page="96" pageId="8" pageNumber="9" year="1973">
<bibRefCitation id="EF8A4B450E31403CFD22F8461E37F818" author="Caruso" box="[691,825,1939,1961]" pageId="8" pageNumber="9" refString="Caruso, D. (1973) Una nuova specie di isopodo terrestre di Sicilia (Arthropoda, Crustacea). Bollettino delle Sedute dell'Accademia Gioenia di Scienze Naturali in Catania (IV), 11, 95 - 103." type="journal article" year="1973">Caruso 1973</bibRefCitation>
: 96
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,
<figureCitation id="13202A310E31403CFCF8F8461EABF818" box="[873,933,1939,1961]" captionStart="FIGURE 1" captionStartId="3.[151,249,1821,1843]" captionTargetBox="[173,1402,366,1772]" captionTargetId="figure@3.[160,1426,363,1787]" captionTargetPageId="3" captionText="FIGURE 1. Levantoniscus bicostulatus n. sp. Paratype ♀: A, adult specimen, dorsal view; B, adult specimen, lateral view; C, dorsal scale-seta; D, cephalon, frontal view. Paratype ♂: E, antennula; F, antenna." httpUri="https://zenodo.org/record/289974/files/figure.png" pageId="8" pageNumber="9">Fig. 1</figureCitation>
;
<treatmentCitation id="0ABA10A50E31403CFC22F84619BAF818" author="Warburg" box="[947,1204,1939,1961]" page="159" pageId="8" pageNumber="9" year="1978">
<bibRefCitation id="EF8A4B450E31403CFC22F846198CF818" author="Warburg" box="[947,1154,1939,1961]" pageId="8" pageNumber="9" refString="Warburg, M. R., Rankevich, D. &amp; Chasanmus, K. (1978) Isopod species diversity and community structure in mesic and xeric habitats of the Mediterranean region. Journal of arid Environments, 1, 157 - 163." type="journal article" year="1978">
Warburg
<emphasis id="B96FEAA60E31403CFB83F8411931F818" box="[1042,1087,1939,1961]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
. 1978
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: 159
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, 159, 162;
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<bibRefCitation id="EF8A4B450E31403CFAB6F8411C55F879" author="Hornung" pageId="8" pageNumber="9" refString="Hornung, E. &amp; Warburg, M. (1996) Intra-habitat distribution of terrestrial isopods. European Journal of Soil Biology, 32, 179 - 185." type="journal article" year="1996">Hornung &amp; Warburg 1996</bibRefCitation>
: 181
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;
<treatmentCitation id="0ABA10A50E31403CFE0AF8671F58F879" author="Warburg" box="[411,598,1970,1992]" page="82" pageId="8" pageNumber="9" year="1993">
<bibRefCitation id="EF8A4B450E31403CFE0AF8671F21F879" author="Warburg" box="[411,559,1970,1992]" pageId="8" pageNumber="9" refString="Warburg, M. R. (1993) Evolutionary Biology of Land Isopods. Springer-Verlag, Berlin, 159 pp." type="book" year="1993">Warburg 1993</bibRefCitation>
: 82
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,
<figureCitation id="13202A310E31403CFDF0F8671FAFF879" box="[609,673,1970,1992]" captionStart="FIGURE 1" captionStartId="3.[151,249,1821,1843]" captionTargetBox="[173,1402,366,1772]" captionTargetId="figure@3.[160,1426,363,1787]" captionTargetPageId="3" captionText="FIGURE 1. Levantoniscus bicostulatus n. sp. Paratype ♀: A, adult specimen, dorsal view; B, adult specimen, lateral view; C, dorsal scale-seta; D, cephalon, frontal view. Paratype ♂: E, antennula; F, antenna." httpUri="https://zenodo.org/record/289974/files/figure.png" pageId="8" pageNumber="9">Figs 1</figureCitation>
, 2.1, 2.2, 2.8, 9.7, 9.8, 9.12;
<treatmentCitation id="0ABA10A50E31403CFC5BF8671812F879" author="Warburg" box="[970,1308,1970,1992]" page="1473" pageId="8" pageNumber="9" year="1999">
<bibRefCitation id="EF8A4B450E31403CFC5BF86719D5F879" author="Warburg" box="[970,1243,1970,1992]" pageId="8" pageNumber="9" refString="Warburg, M. R. &amp; Hornung, E. (1999) Diversity of terrestrial isopod species along a transect through northern Israel. Biodiversity and Conservation, 8, 1469 - 1478." type="journal article" year="1999">Warburg &amp; Hornung 1999</bibRefCitation>
: 1473
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;
<treatmentCitation id="0ABA10A50E31403CFAB6F8671C2DF856" author="Schmalfuss" page="61" pageId="8" pageNumber="9" year="2003">
<bibRefCitation id="EF8A4B450E31403CFAB6F8671DF3F856" author="Schmalfuss" pageId="8" pageNumber="9" refString="Schmalfuss, H. (2003) World catalog of terrestrial isopods (Isopoda: Oniscidea). Stuttgarter Beitrage zur Naturkunde (A), 654, 1 - 341." type="journal article" year="2003">Schmalfuss 2003</bibRefCitation>
: 61
</treatmentCitation>
;
<treatmentCitation id="0ABA10A50E31403CFEBCF8041F0EF856" author="Warburg" box="[301,512,2001,2023]" page="1228" pageId="8" pageNumber="9" year="2007">
<bibRefCitation id="EF8A4B450E31403CFEBCF8041CCFF856" author="Warburg" box="[301,449,2001,2023]" pageId="8" pageNumber="9" refString="Warburg, M. R. (2007) Distribution, reproduction, and relative abundance of oniscids: a long-term study on isopods (Isopoda: Oniscidea) in Israel. Crustaceana, 80 (10), 1223 - 1252. http: // dx. doi. org / 10.1163 / 156854007782321218" type="journal article" year="2007">Warburg 2007</bibRefCitation>
: 1228
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, 1230, 1233, 1235, 1248,
<figureCitation id="13202A310E31403CFC99F8041E4BF856" box="[776,837,2001,2023]" captionStart="FIGURE 5" captionStartId="7.[151,249,1944,1966]" captionTargetBox="[218,1369,214,1868]" captionTargetId="figure@7.[199,1386,193,1888]" captionTargetPageId="7" captionText="FIGURE 5. Levantoniscus bicostulatus n. sp. Paratype ♀: A, adult specimen, dorsal view; B, cephalon and pereonite 1, dorsal view; C, pereonites 6 and 7, pleon, telson and uropods, dorsal view; D, pereonites 2 and 3, details of the cuticular structure; E, tubercle on pereonite 7, medial surface." httpUri="https://zenodo.org/record/289978/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
.
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<paragraph id="8BA436B40E30403DFF06F8C41955F8F4" blockId="9.[151,1435,1809,1861]" pageId="9" pageNumber="10">
<emphasis id="B96FEAA60E30403DFF06F8C41C1AF897" bold="true" box="[151,276,1809,1831]" pageId="9" pageNumber="10">FIGURE 7.</emphasis>
<taxonomicName id="4C1B4D370E30403DFE8DF8C71F0FF896" box="[284,513,1809,1831]" class="Malacostraca" family="Trachelipodidae" genus="Levantoniscus" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="makrisi" status="sp. nov.">
<emphasis id="B96FEAA60E30403DFE8DF8C71F0FF896" box="[284,513,1809,1831]" italics="true" pageId="9" pageNumber="10">Levantoniscus makrisi</emphasis>
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<emphasis id="B96FEAA60E30403DFD98F8C41F4AF897" bold="true" box="[521,580,1809,1830]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="A25C57DD0E30403DFD98F8C41F4AF897" box="[521,580,1809,1830]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
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Holotype: A, specimen, dorsal view; B, lateral view; C, cephalon, dorsal view; D, cephalon, frontal view, E, pleonite 5, telson and uropods, dorsal view; E, antennula; F, antenna.
</paragraph>
</caption>
<caption id="DF64663C0E33403EFF06F96C1E7FF8BD" httpUri="https://zenodo.org/record/289981/files/figure.png" pageId="10" pageNumber="11" targetBox="[179,1426,198,1684]" targetPageId="10">
<paragraph id="8BA436B40E33403EFF06F96C1E7FF8BD" blockId="10.[151,1435,1721,1804]" pageId="10" pageNumber="11">
<emphasis id="B96FEAA60E33403EFF06F96C1C1DF97F" bold="true" box="[151,275,1721,1743]" pageId="10" pageNumber="11">FIGURE 8.</emphasis>
<taxonomicName id="4C1B4D370E33403EFE8DF96F1F0FF97E" box="[284,513,1721,1743]" class="Malacostraca" family="Trachelipodidae" genus="Levantoniscus" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="makrisi" status="sp. nov.">
<emphasis id="B96FEAA60E33403EFE8DF96F1F0FF97E" box="[284,513,1721,1743]" italics="true" pageId="10" pageNumber="11">Levantoniscus makrisi</emphasis>
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<taxonomicNameLabel id="A25C57DD0E33403EFD98F96C1F4AF97F" box="[521,580,1721,1742]" pageId="10" pageNumber="11" rank="species">n. sp.</taxonomicNameLabel>
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Holotype: A, pereopod 1; B, pereopod 7; C, pleopod 1 exopodite, dorsal view; D, pleopod 1 endopodite; E, pleopod 2 exopodite, ventral view; F, pleopod 2 endopodite; G, pleopod 3 exopodite, ventral view; H, pleopod 4 exopodite, ventral view; I, pleopod 5 exopodite, ventral view.
</paragraph>
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<paragraph id="8BA436B40E32403FFF06F9681C7CF8A0" blockId="11.[151,1436,1725,1809]" pageId="11" pageNumber="12">
<emphasis id="B96FEAA60E32403FFF06F9681C1DF962" bold="true" box="[151,275,1725,1747]" pageId="11" pageNumber="12">FIGURE 9.</emphasis>
<emphasis id="B96FEAA60E32403FFE8DF96A1F2DF962" box="[284,547,1725,1748]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="4C1B4D370E32403FFE8DF96A1F11F962" box="[284,543,1725,1748]" class="Malacostraca" family="Trachelipodidae" genus="Levantoniscus" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="wahrmani">Levantoniscus wahrmani</taxonomicName>
.
</emphasis>
♀: A, adult specimen, dorsal view; B, lateral view; C, dorsal scale-seta; D, cephalon, dorsal view; E, cephalon, posterior view; F, cephalon, frontal view, G, pleonite 5, telson and uropods, dorsal view. ♂: H, antennula; I, antenna.
</paragraph>
</caption>
<caption id="DF64663C0E354038FF06F84E1F05F85F" httpUri="https://zenodo.org/record/289983/files/figure.png" pageId="12" pageNumber="13" targetBox="[151,1417,207,1909]" targetPageId="12">
<paragraph id="8BA436B40E354038FF06F84E1F05F85F" blockId="12.[151,1436,1947,2030]" pageId="12" pageNumber="13">
<emphasis id="B96FEAA60E354038FF06F84E1C11F801" bold="true" box="[151,287,1947,1969]" pageId="12" pageNumber="13">FIGURE 10.</emphasis>
<emphasis id="B96FEAA60E354038FEB7F8491F25F800" box="[294,555,1947,1969]" italics="true" pageId="12" pageNumber="13">
<taxonomicName id="4C1B4D370E354038FEB7F8491F29F800" box="[294,551,1947,1969]" class="Malacostraca" family="Trachelipodidae" genus="Levantoniscus" kingdom="Animalia" order="Isopoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="wahrmani">Levantoniscus wahrmani</taxonomicName>
.
</emphasis>
♂: A, uropod; B, pereopod 1; C, pereopod 7; D, pleopod 1, dorsal view; E, pleopod 2, dorsal view; F, pleopod 2 exopodite, ventral view; G, pleopod 3 exopodite, ventral view; H, pleopod 4 exopodite, ventral view; I, pleopod 5 exopodite, ventral view.
</paragraph>
</caption>
<subSubSection id="C301653F0E344039FF06FF42196AFE1D" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="8BA436B40E344039FF06FF421DD8FEAD" blockId="13.[151,1436,151,717]" pageId="13" pageNumber="14">
<emphasis id="B96FEAA60E344039FF06FF421C8DFF01" bold="true" box="[151,387,151,176]" pageId="13" pageNumber="14">Material examined.</emphasis>
<collectingCountry id="F30C76240E344039FE1CFF421CDEFF01" box="[397,464,151,176]" name="Israel" pageId="13" pageNumber="14">Israel</collectingCountry>
:
<specimenCount id="9D1DFD3D0E344039FE4FFF4D1F03FF1E" box="[478,525,152,176]" pageId="13" pageNumber="14" type="generic">1 ♂</specimenCount>
, 1 ♀ (
<collectingCountry id="F30C76240E344039FDCAFF4D1F9AFF01" box="[603,660,152,176]" name="China" pageId="13" pageNumber="14">
<collectionCode id="ED0AAE710E344039FDCAFF4D1F9AFF01" box="[603,660,152,176]" httpUri="http://grbio.org/cool/ze71-842p" name="Tel-Aviv University" pageId="13" pageNumber="14">TAU</collectionCode>
</collectingCountry>
2406), Jerusalem,
<date id="FFA510740E344039FCE3FF4D1EE9FF01" box="[882,999,152,176]" pageId="13" pageNumber="14" value="1943-05-24">24.V.1943</date>
, det. H. Schmalfuss 1987; 2 ♀ (
<collectingCountry id="F30C76240E344039FAF2FF4D1892FF01" box="[1379,1436,152,176]" name="China" pageId="13" pageNumber="14">
<collectionCode id="ED0AAE710E344039FAF2FF4D1892FF01" box="[1379,1436,152,176]" httpUri="http://grbio.org/cool/ze71-842p" name="Tel-Aviv University" pageId="13" pageNumber="14">TAU</collectionCode>
</collectingCountry>
1985), no locality, date and collector, det. H. Schmalfuss 1987; 1 ♀ (
<collectingCountry id="F30C76240E344039FC4DFF69191BFF65" box="[988,1045,188,212]" name="China" pageId="13" pageNumber="14">
<collectionCode id="ED0AAE710E344039FC4DFF69191BFF65" box="[988,1045,188,212]" httpUri="http://grbio.org/cool/ze71-842p" name="Tel-Aviv University" pageId="13" pageNumber="14">TAU</collectionCode>
</collectingCountry>
3118), no locality and collector,
<date id="FFA510740E344039FF06FF351C18FF49" box="[151,278,224,248]" pageId="13" pageNumber="14" value="1974-04-21">21.IV.1974</date>
, det. H. Schmalfuss 1987; 1 ♀ (
<collectingCountry id="F30C76240E344039FDEEFF351FB6FF49" box="[639,696,224,248]" name="China" pageId="13" pageNumber="14">
<collectionCode id="ED0AAE710E344039FDEEFF351FB6FF49" box="[639,696,224,248]" httpUri="http://grbio.org/cool/ze71-842p" name="Tel-Aviv University" pageId="13" pageNumber="14">TAU</collectionCode>
</collectingCountry>
1924), no locality and collector,
<date id="FFA510740E344039FBA1FF3519B2FF49" box="[1072,1212,224,248]" pageId="13" pageNumber="14" value="1973-12-20">20.XII.1973</date>
, det. H. Schmalfuss 1987.
</paragraph>
<paragraph id="8BA436B40E344039FF56FEF2196AFE1D" blockId="13.[151,1436,151,717]" pageId="13" pageNumber="14">
<emphasis id="B96FEAA60E344039FF56FEF21C90FEF1" bold="true" box="[199,414,295,320]" pageId="13" pageNumber="14">Previous records.</emphasis>
<collectingCountry id="F30C76240E344039FE3CFEF21F0FFEF1" box="[429,513,295,320]" name="Turkey" pageId="13" pageNumber="14">Turkey</collectingCountry>
: Vilayet
<collectingCountry id="F30C76240E344039FDE4FEFD1FB6FEF1" box="[629,696,296,320]" name="Turkey" pageId="13" pageNumber="14">Hatay</collectingCountry>
(Amanus, Bitiyas cave);
<collectingCountry id="F30C76240E344039FC67FEF21937FEF1" box="[1014,1081,295,320]" name="Israel" pageId="13" pageNumber="14">Israel</collectingCountry>
: Geiah stream, south of Yad Mordechai; Lachish stream, south of Ashdod; Juda Mt.; Aqua Bella (
<bibRefCitation id="EF8A4B450E344039FC38FE991943FED4" author="Strouhal" box="[937,1101,332,357]" pageId="13" pageNumber="14" refString="Strouhal, H. (1968) Eine neue, vorderasiatische Bathytropa - Art (Oniscoidea, Oniscidae, Bathytropinae). Crustaceana, 15, 67 - 78. http: // dx. doi. org / 10.1163 / 156854068 X 00764" type="journal article" year="1968">Strouhal 1968</bibRefCitation>
); Mt. Carmel (
<bibRefCitation id="EF8A4B450E344039FB65FE991899FED4" author="Strouhal" box="[1268,1431,332,357]" pageId="13" pageNumber="14" refString="Strouhal, H. (1968) Eine neue, vorderasiatische Bathytropa - Art (Oniscoidea, Oniscidae, Bathytropinae). Crustaceana, 15, 67 - 78. http: // dx. doi. org / 10.1163 / 156854068 X 00764" type="journal article" year="1968">Strouhal 1968</bibRefCitation>
;
<bibRefCitation id="EF8A4B450E344039FF06FEBA1CCAFE39" author="Warburg" box="[151,452,367,392]" pageId="13" pageNumber="14" refString="Warburg, M. R. &amp; Hornung, E. (1999) Diversity of terrestrial isopod species along a transect through northern Israel. Biodiversity and Conservation, 8, 1469 - 1478." type="journal article" year="1999">Warburg &amp; Hornung 1999</bibRefCitation>
;
<bibRefCitation id="EF8A4B450E344039FE40FEBA1F78FE39" author="Warburg" box="[465,630,367,392]" pageId="13" pageNumber="14" refString="Warburg, M. R. (2007) Distribution, reproduction, and relative abundance of oniscids: a long-term study on isopods (Isopoda: Oniscidea) in Israel. Crustaceana, 80 (10), 1223 - 1252. http: // dx. doi. org / 10.1163 / 156854007782321218" type="journal article" year="2007">Warburg 2007</bibRefCitation>
); Bet Oren; Muhraqa (
<bibRefCitation id="EF8A4B450E344039FCE6FEBA1951FE39" author="Warburg" box="[887,1119,367,392]" pageId="13" pageNumber="14" refString="Warburg, M. R., Rankevich, D. &amp; Chasanmus, K. (1978) Isopod species diversity and community structure in mesic and xeric habitats of the Mediterranean region. Journal of arid Environments, 1, 157 - 163." type="journal article" year="1978">
Warburg
<emphasis id="B96FEAA60E344039FC73FEA4191DFE39" box="[994,1043,367,392]" italics="true" pageId="13" pageNumber="14">et al</emphasis>
. 1978
</bibRefCitation>
); Segev, Lower Galilee Mts (
<bibRefCitation id="EF8A4B450E344039FF0FFE411CC5FE1C" author="Hornung" box="[158,459,404,429]" pageId="13" pageNumber="14" refString="Hornung, E. &amp; Warburg, M. (1996) Intra-habitat distribution of terrestrial isopods. European Journal of Soil Biology, 32, 179 - 185." type="journal article" year="1996">Hornung &amp; Warburg 1996</bibRefCitation>
); Galilee Mts; Golan Heights; Judean Mts (
<bibRefCitation id="EF8A4B450E344039FC20FE411956FE1D" author="Warburg" box="[945,1112,404,429]" pageId="13" pageNumber="14" refString="Warburg, M. R. (2007) Distribution, reproduction, and relative abundance of oniscids: a long-term study on isopods (Isopoda: Oniscidea) in Israel. Crustaceana, 80 (10), 1223 - 1252. http: // dx. doi. org / 10.1163 / 156854007782321218" type="journal article" year="2007">Warburg 2007</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C301653F0E344039FF56FE621E62FE61" box="[199,876,439,464]" pageId="13" pageNumber="14" type="distribution">
<paragraph id="8BA436B40E344039FF56FE621E62FE61" blockId="13.[151,1436,151,717]" box="[199,876,439,464]" pageId="13" pageNumber="14">
<emphasis id="B96FEAA60E344039FF56FE621C6EFE61" bold="true" box="[199,352,439,464]" pageId="13" pageNumber="14">Distribution.</emphasis>
Southern
<collectingCountry id="F30C76240E344039FE42FE621F2DFE61" box="[467,547,439,464]" name="Turkey" pageId="13" pageNumber="14">Turkey</collectingCountry>
and northern
<collectingCountry id="F30C76240E344039FD2EFE621FF3FE61" box="[703,765,439,464]" name="Israel" pageId="13" pageNumber="14">Israel</collectingCountry>
(
<figureCitation id="13202A310E344039FC9CFE621E51FE61" box="[781,863,439,464]" captionStart="FIGURE 11" captionStartId="13.[151,250,1947,1969]" captionTargetBox="[169,1421,785,1906]" captionTargetId="figure@13.[151,1436,769,1925]" captionTargetPageId="13" captionText="FIGURE 11. Distribution of Levantoniscus species." httpUri="https://zenodo.org/record/289984/files/figure.png" pageId="13" pageNumber="14">Fig. 11</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C301653F0E34403AFF56FE091F93FE45" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="discussion">
<paragraph id="8BA436B40E344039FF56FE091C5AFD7C" blockId="13.[151,1436,151,717]" pageId="13" pageNumber="14">
<emphasis id="B96FEAA60E344039FF56FE091C35FE44" bold="true" box="[199,315,476,501]" pageId="13" pageNumber="14">Remarks.</emphasis>
<bibRefCitation id="EF8A4B450E344039FED5FE091CF8FE44" author="Strouhal" box="[324,502,476,501]" pageId="13" pageNumber="14" refString="Strouhal, H. (1968) Eine neue, vorderasiatische Bathytropa - Art (Oniscoidea, Oniscidae, Bathytropinae). Crustaceana, 15, 67 - 78. http: // dx. doi. org / 10.1163 / 156854068 X 00764" type="journal article" year="1968">Strouhal (1968)</bibRefCitation>
had described this species as
<taxonomicName id="4C1B4D370E344039FCC2FE091944FE44" box="[851,1098,476,501]" class="Malacostraca" family="Bathytropidae" genus="Bathytropa" kingdom="Animalia" order="Isopoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="wahrmani">
<emphasis id="B96FEAA60E344039FCC2FE091944FE44" box="[851,1098,476,501]" italics="true" pageId="13" pageNumber="14">Bathytropa wahrmani</emphasis>
</taxonomicName>
and included it in the family
<taxonomicName id="4C1B4D370E344039FF06FE2A1C37FDA9" box="[151,313,511,536]" class="Malacostraca" family="Bathytropidae" kingdom="Animalia" order="Isopoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="family">Bathytropidae</taxonomicName>
, which is definitely not correct because a diagnostic feature of this family is the absence of lungs from all pleopods (see
<bibRefCitation id="EF8A4B450E344039FE06FDF11F36FD8D" author="Schmidt" box="[407,568,548,573]" pageId="13" pageNumber="14" refString="Schmidt, C. (2003) Contribution to the phylogenetic system of the Crinocheta (Crustacea, Isopoda). Part 2. (Oniscoidea to Armadillidiidae). Mitteilungen aus dem Museum fur Naturkunde in Berlin, Zoologische Reihe, 79, 3 - 179. http: // dx. doi. org / 10.1002 / mmnz. 20030790102" type="journal article" year="2003">Schmidt 2003</bibRefCitation>
), while this species has distinct uncovered lungs on pleopod exopodites. Also the hour-glass-shaped telson is a distinguishing character from all the other species of
<taxonomicName id="4C1B4D370E344039FBFAFD9219E6FDD1" box="[1131,1256,583,608]" class="Malacostraca" family="Bathytropidae" genus="Bathytropa" kingdom="Animalia" order="Isopoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEAA60E344039FBFAFD9219E6FDD1" box="[1131,1256,583,608]" italics="true" pageId="13" pageNumber="14">Bathytropa</emphasis>
</taxonomicName>
. This species is similar to the two new species from
<collectingCountry id="F30C76240E344039FDD1FDB91F9FFD35" box="[576,657,620,644]" name="Cyprus" pageId="13" pageNumber="14">Cyprus</collectingCountry>
described above, especially in the interlocking pleopods and in the same structure of the pleopodal lungs. It is therefore transferred to the new genus
<taxonomicName id="4C1B4D370E344039FBE3FD44181CFD19" box="[1138,1298,657,680]" class="Malacostraca" family="Trachelipodidae" genus="Levantoniscus" kingdom="Animalia" order="Isopoda" pageId="13" pageNumber="2" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96FEAA60E344039FBE3FD44181CFD19" box="[1138,1298,657,680]" italics="true" pageId="13" pageNumber="14">Levantoniscus</emphasis>
</taxonomicName>
and family
<taxonomicName id="4C1B4D370E344039FF06FD611C5EFD7C" box="[151,336,692,717]" class="Malacostraca" family="Trachelipodidae" kingdom="Animalia" order="Isopoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="family">Trachelipodidae</taxonomicName>
.
</paragraph>
<caption id="DF64663C0E344039FF06F84E1FA6F800" box="[151,680,1947,1969]" httpUri="https://zenodo.org/record/289984/files/figure.png" pageId="13" pageNumber="14" targetBox="[169,1421,785,1906]" targetPageId="13">
<paragraph id="8BA436B40E344039FF06F84E1FA6F800" blockId="13.[151,680,1947,1969]" box="[151,680,1947,1969]" pageId="13" pageNumber="14">
<emphasis id="B96FEAA60E344039FF06F84E1C16F801" bold="true" box="[151,280,1947,1969]" pageId="13" pageNumber="14">FIGURE 11</emphasis>
. Distribution of
<taxonomicName id="4C1B4D370E344039FE51F8491F5FF800" box="[448,593,1948,1969]" class="Malacostraca" family="Trachelipodidae" genus="Levantoniscus" kingdom="Animalia" order="Isopoda" pageId="13" pageNumber="2" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96FEAA60E344039FE51F8491F5FF800" box="[448,593,1948,1969]" italics="true" pageId="13" pageNumber="14">Levantoniscus</emphasis>
</taxonomicName>
species.
</paragraph>
</caption>
<paragraph id="8BA436B40E37403AFF56FF42199EFE39" blockId="14.[151,1437,151,501]" pageId="14" pageNumber="15">
The species is here fully illustrated (
<figureCitation id="13202A310E37403AFDC8FF421FAFFF01" box="[601,673,151,176]" captionStart="FIGURE 9" captionStartId="11.[151,249,1725,1747]" captionTargetBox="[161,1419,202,1687]" captionTargetId="figure@11.[151,1436,193,1704]" captionTargetPageId="11" captionText="FIGURE 9. Levantoniscus wahrmani. ♀: A, adult specimen, dorsal view; B, lateral view; C, dorsal scale-seta; D, cephalon, dorsal view; E, cephalon, posterior view; F, cephalon, frontal view, G, pleonite 5, telson and uropods, dorsal view. ♂: H, antennula; I, antenna." httpUri="https://zenodo.org/record/289982/files/figure.png" pageId="14" pageNumber="15">Figs 9</figureCitation>
,
<figureCitation id="13202A310E37403AFD3CFF4D1FC4FF01" box="[685,714,152,176]" captionStart="FIGURE 10" captionStartId="12.[151,249,1947,1969]" captionTargetBox="[151,1417,207,1909]" captionTargetId="figure@12.[151,1436,193,1925]" captionTargetPageId="12" captionText="FIGURE 10. Levantoniscus wahrmani. ♂: A, uropod; B, pereopod 1; C, pereopod 7; D, pleopod 1, dorsal view; E, pleopod 2, dorsal view; F, pleopod 2 exopodite, ventral view; G, pleopod 3 exopodite, ventral view; H, pleopod 4 exopodite, ventral view; I, pleopod 5 exopodite, ventral view." httpUri="https://zenodo.org/record/289983/files/figure.png" pageId="14" pageNumber="15">10</figureCitation>
) for comparison with
<taxonomicName id="4C1B4D370E37403AFC50FF4C19E0FF01" box="[961,1262,151,176]" class="Malacostraca" family="Trachelipodidae" genus="Levantoniscus" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="bicostulatus">
<emphasis id="B96FEAA60E37403AFC50FF4C19E0FF01" box="[961,1262,151,176]" italics="true" pageId="14" pageNumber="15">Levantoniscus bicostulatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1B4D370E37403AFAB5FF4C189BFF01" box="[1316,1429,151,176]" class="Malacostraca" family="Trachelipodidae" genus="Levantoniscus" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="makrisi">
<emphasis id="B96FEAA60E37403AFAB5FF4C189BFF01" box="[1316,1429,151,176]" italics="true" pageId="14" pageNumber="15">L. makrisi</emphasis>
</taxonomicName>
, from which it is readily distinguished by the pattern of dorsal ornamentation (
<figureCitation id="13202A310E37403AFC69FF691934FF65" box="[1016,1082,188,213]" captionStart="FIGURE 9" captionStartId="11.[151,249,1725,1747]" captionTargetBox="[161,1419,202,1687]" captionTargetId="figure@11.[151,1436,193,1704]" captionTargetPageId="11" captionText="FIGURE 9. Levantoniscus wahrmani. ♀: A, adult specimen, dorsal view; B, lateral view; C, dorsal scale-seta; D, cephalon, dorsal view; E, cephalon, posterior view; F, cephalon, frontal view, G, pleonite 5, telson and uropods, dorsal view. ♂: H, antennula; I, antenna." httpUri="https://zenodo.org/record/289982/files/figure.png" pageId="14" pageNumber="15">Fig. 9</figureCitation>
A, B). In particular, it has a row of four pointed tubercles at the back of the head instead of two in the new species, a couple of forward-pointed tubercles on the front of the head which are lacking in the new species, plus a narrower and acuter middle frontal lobe on the cephalon (
<figureCitation id="13202A310E37403AFE1CFEF21CC1FEF1" box="[397,463,295,320]" captionStart="FIGURE 9" captionStartId="11.[151,249,1725,1747]" captionTargetBox="[161,1419,202,1687]" captionTargetId="figure@11.[151,1436,193,1704]" captionTargetPageId="11" captionText="FIGURE 9. Levantoniscus wahrmani. ♀: A, adult specimen, dorsal view; B, lateral view; C, dorsal scale-seta; D, cephalon, dorsal view; E, cephalon, posterior view; F, cephalon, frontal view, G, pleonite 5, telson and uropods, dorsal view. ♂: H, antennula; I, antenna." httpUri="https://zenodo.org/record/289982/files/figure.png" pageId="14" pageNumber="15">Fig. 9</figureCitation>
DF). Seven pointed tubercles are present on the pereonite 1, six on pereonites 24 and four on pereonites 57, while the two new species only have two lamellar tubercles on all pereonites. Finally, it has two tubercles on pleonites 15, while the new species have two tubercles on pleonites 14.
</paragraph>
<paragraph id="8BA436B40E37403AFF56FE401F93FE45" blockId="14.[151,1437,151,501]" pageId="14" pageNumber="15">
In the detailed description provided by
<bibRefCitation id="EF8A4B450E37403AFD08FE411E40FE1C" author="Strouhal" box="[665,846,404,429]" pageId="14" pageNumber="15" refString="Strouhal, H. (1968) Eine neue, vorderasiatische Bathytropa - Art (Oniscoidea, Oniscidae, Bathytropinae). Crustaceana, 15, 67 - 78. http: // dx. doi. org / 10.1163 / 156854068 X 00764" type="journal article" year="1968">Strouhal (1968)</bibRefCitation>
there is no mention on the position of the noduli laterales. They are located on the tip of the external tubercle on the pereonites 14 and on the postero-lateral corners of the pereonites 57 (see
<figureCitation id="13202A310E37403AFD86FE091F56FE45" box="[535,600,476,501]" captionStart="FIGURE 9" captionStartId="11.[151,249,1725,1747]" captionTargetBox="[161,1419,202,1687]" captionTargetId="figure@11.[151,1436,193,1704]" captionTargetPageId="11" captionText="FIGURE 9. Levantoniscus wahrmani. ♀: A, adult specimen, dorsal view; B, lateral view; C, dorsal scale-seta; D, cephalon, dorsal view; E, cephalon, posterior view; F, cephalon, frontal view, G, pleonite 5, telson and uropods, dorsal view. ♂: H, antennula; I, antenna." httpUri="https://zenodo.org/record/289982/files/figure.png" pageId="14" pageNumber="15">Fig. 9</figureCitation>
A, B).
</paragraph>
</subSubSection>
</treatment>
</document>