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<document id="ACD1F5C04561338C2BAC832074D4C307" ID-DOI="https://doi.org/10.5852/ejt.2023.881.2175" ID-ISSN="2118-9773" ID-Zenodo-Dep="8178280" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1690201937279" checkinUser="felipe" docAuthor="Moonlight, Peter. W., Jara-Muñoz, Orlando A., Purvis, David A., Delves, Jay, Allen, Josh P. &amp; Reynel, Carlos" docDate="2023" docId="03B26B4BFF38FF66FD8BF9EFAC9DEE0E" docLanguage="en" docName="ejt-881_1-334.imf" docOrigin="European Journal of Taxonomy 881" docSource="http://dx.doi.org/10.5852/ejt.2023.881.2175" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Begonia weberbaueri Irmsch." docType="treatment" docVersion="7" lastPageNumber="160" masterDocId="FF8B1333FFA5FFF9FFD9FFBFAD27EA79" masterDocTitle="The genus Begonia (Begoniaceae) in Peru" masterLastPageNumber="334" masterPageNumber="1" pageNumber="158" updateTime="1707123162844" updateUser="tatiana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="DC15643431741BEDA39AA93B68F11DA7">The genus Begonia (Begoniaceae) in Peru</mods:title>
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<mods:namePart id="FC1E0F8E681A11499DDF3CE65D9325C3">Moonlight, Peter. W.</mods:namePart>
<mods:affiliation id="8ACE6AE00E54F938C46E70A97FB41DB3">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK. &amp; Botany Department, School of Natural Sciences, Trinity College Dublin, Dublin 2, Ireland.</mods:affiliation>
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<mods:namePart id="63EC8ECEDBE212790E7C8CFE789AC6A5">Jara-Muñoz, Orlando A.</mods:namePart>
<mods:affiliation id="1DDF599A57230B53B38F584ACCB6415B">Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá D. C., Colombia.</mods:affiliation>
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<mods:namePart id="0EB642C1AC973529578863C40A740D5D">Purvis, David A.</mods:namePart>
<mods:affiliation id="FF301E9A0BEBA24D3FC08D2A26722169">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
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<mods:namePart id="A1E67193C15B7AE8BBF9E4CA1A8DB282">Delves, Jay</mods:namePart>
<mods:affiliation id="F255826B073FA33B8A8CA3D0EE445BD9">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
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<mods:namePart id="41EB3DD2EA11AE0ED8310B649977BD9E">Allen, Josh P.</mods:namePart>
<mods:affiliation id="91F0F652AEB4F882358301B04590B56E">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
<mods:nameIdentifier id="34086599FFA8AEBADCF7CC47FC56EB11" type="email">myrmeciaman@gmail.com</mods:nameIdentifier>
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<mods:namePart id="E9B7F97AB5321D17B4B7B7D84FF1CD30">Reynel, Carlos</mods:namePart>
<mods:affiliation id="B65D7E867B4F1B099078066882BE6F7C">Facultad de Ciencias Forestales, Universidad Nacional Agraria-La Molina, Lima 12, Peru.</mods:affiliation>
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<mods:date id="1B4EF622C46401A90EA6DCC579F961DA">2023</mods:date>
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34.
<taxonomicName id="4C1BA1DEFF38FF64FDA7F9EFAEC6EC13" ID-CoL="87ZG5" ID-ENA="1684063" authority="Irmsch." authorityName="Irmsch." box="[638,993,1615,1642]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="157" pageNumber="158" phylum="Tracheophyta" rank="species" species="weberbaueri">
<emphasis id="B96F064FFF38FF64FDA7F9EFAEA6EC13" bold="true" box="[638,897,1615,1642]" italics="true" pageId="157" pageNumber="158">Begonia weberbaueri</emphasis>
Irmsch.
</taxonomicName>
</paragraph>
</subSubSection>
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<paragraph id="8BA4DA5DFF38FF64FD3FF9CCAE6AECF4" blockId="157.[594,993,1615,1677]" box="[742,845,1651,1677]" pageId="157" pageNumber="158">
<figureCitation id="1320C6D8FF38FF64FD3FF9CCAE6AECF4" box="[742,845,1651,1677]" captionStart="Fig" captionStartId="147.[189,232,1892,1918]" captionTargetBox="[194,1392,1228,1847]" captionTargetId="figure-12@147.[189,1398,1218,1853]" captionTargetPageId="147" captionText="Fig. 48. Distribution of Begonia sect. Eupetalum (Lindl.) A.DC. in Peru and surrounding countries. A. B. geraniifolia Hook. (red) and B. joshii Moonlight (blue). B. B. neoharlingii L.B.Sm. &amp; Wassh. (red) and B. weberbaueri Irmsch. (blue). Black lines indicate country borders, grey lines indicate major administrative divisions, blue lines indicate rivers, and shading indicates elevation." figureDoi="http://doi.org/10.5281/zenodo.8178403" httpUri="https://zenodo.org/record/8178403/files/figure.png" pageId="157" pageNumber="158">Fig. 48B</figureCitation>
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<bibCitation id="5E31D231FF38FF64FF64F90BAC13EC8A" journal="Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie" page="78" pageId="157" pageNumber="158" score="478.66328617067893" volume="76" year="1953">
<emphasis id="B96F064FFF38FF64FF64F90BA981ECB7" box="[189,1190,1716,1743]" italics="true" pageId="157" pageNumber="158">
Botanische Jahrbücher für
<bibCitation id="5E31D231FF38FF64FDD1F90BA981ECB7" author="Systematik" box="[520,1190,1716,1742]" journal="Pflanzengeschichte und Pflanzengeographie" page="78" pageId="157" pageNumber="158" score="109.65844492058366" volume="76" year="1953">Systematik, Pflanzengeschichte und Pflanzengeographie</bibCitation>
</emphasis>
76: 78 (
<bibRefCitation id="EF8AA7ACFF38FF64FAD5F90BAC0CEC8A" author="Irmscher" firstAuthor="Irmscher" pageId="157" pageNumber="158" pagination="1 - 102" refId="ref228521" refString="Irmscher E. 1953. Systematische Studien uber Begoniaceen des tropischen Sudamerikas, besonders Brasiliens. Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie 76 (1): 1 - 102." type="journal article" year="1953">Irmscher 1953</bibRefCitation>
)
</bibCitation>
.
</paragraph>
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<emphasis id="B96F064FFF38FF64FE9AF966AC75EC8B" box="[323,338,1753,1778]" italics="true" pageId="157" pageNumber="158">–</emphasis>
<emphasis id="B96F064FFF38FF64FE87F967ACBBEC8B" bold="true" box="[350,412,1752,1778]" pageId="157" pageNumber="158">Type</emphasis>
:
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<collectingCountry id="F30C9ACDFF38FF64FE68F967ACDBEC8B" box="[433,508,1752,1778]" name="Peru" pageId="157" pageNumber="158">PERU</collectingCountry>
–
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<collectingRegion id="49DF14BFFF38FF64FDFDF968AE3CEC88" box="[548,795,1751,1777]" country="Peru" name="Cajamarca" pageId="157" pageNumber="158">Cajamarca Region</collectingRegion>
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•
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; [
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a.s.l.
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<collectorName id="26EEBF8BFF38FF64FCC6F944AEF5ED6C" box="[799,978,1787,1813]" pageId="157" pageNumber="158">A. Weberbauer</collectorName>
3824
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;
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:
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[
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;
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:
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[
<specimenCode id="DBBD7226FF38FF64FD53F8A1AE0DED40" box="[650,810,1822,1849]" collectionCode="B" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="157" pageNumber="158" type="Herbarium">B100186581</specimenCode>
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[
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<bibRefCitation id="EF8AA7ACFF38FF64FF64F8FDAF00ED25" author="Leon &amp; Monsalve" box="[189,551,1857,1884]" firstAuthor="Leon" page="170" pageId="157" pageNumber="158" pagination="165 - 170" refId="ref229226" refString="Leon B. &amp; Monsalve C. 2006. Begoniaceae endemicas del Peru. Revista Peruana de Biologia 13 (2): 165 - 170. https: // doi. org / 10.15381 / rpb. v 13 i 2.1810" type="journal article" year="2006">
<collectingRegion id="49DF14BFFF38FF64FF64F8FDADDEED25" box="[189,249,1858,1884]" country="Nicaragua" name="Leon" pageId="157" pageNumber="158">León</collectingRegion>
&amp; Monsalve (2006: 170)
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.
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<paragraph id="8BA4DA5DFF38FF64FF64F8DAAE8BEDDB" blockId="157.[189,1400,1716,2025]" pageId="157" pageNumber="158">
<taxonomicName id="4C1BA1DEFF38FF64FF64F8DAAEB4ED06" authority="L. B. Sm. &amp; B. G. Schub., Publicaciones" authorityName="L. B. Sm. &amp; B. G. Schub., Publicaciones" box="[189,915,1893,1919]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="157" pageNumber="158" phylum="Tracheophyta" rank="species" species="lichenoides">
<emphasis id="B96F064FFF38FF64FF64F8DAAC96ED06" box="[189,433,1893,1919]" italics="true" pageId="157" pageNumber="158">Begonia lichenoides</emphasis>
L.B.Sm. &amp; B.G.Schub.,
<emphasis id="B96F064FFF38FF64FD32F8DAAEB4ED06" box="[747,915,1893,1919]" italics="true" pageId="157" pageNumber="158">Publicaciones</emphasis>
</taxonomicName>
<emphasis id="B96F064FFF38FF64FC79F8DAAF3EEDDB" italics="true" pageId="157" pageNumber="158">
del Museo de Historia Natural Javier Prado, Serie B,
<collectingMunicipality id="6BC04027FF38FF64FE77F837AF3EEDDB" box="[430,537,1928,1954]" pageId="157" pageNumber="158">Botánica</collectingMunicipality>
</emphasis>
17: 10 (
<bibRefCitation id="EF8AA7ACFF38FF64FDA7F837AEB8EDDB" author="Smith &amp; Schubert" box="[638,927,1928,1954]" firstAuthor="Smith" pageId="157" pageNumber="158" pagination="1 - 19" refId="ref231666" refString="Smith L. B. &amp; Schubert B. G. 1964. Nuevas especies de la familia Begoniaceae. Publicaciones del Museo de Historia Natural Javier Prado, Serie B, Botanica 17: 1 - 19." type="journal article" year="1964">Smith &amp; Schubert 1964</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF38FF64FC6EF836A929ED90" pageId="157" pageNumber="158" type="materials_examined">
<paragraph id="8BA4DA5DFF38FF64FC6EF836A929ED90" blockId="157.[189,1400,1716,2025]" pageId="157" pageNumber="158">
<emphasis id="B96F064FFF38FF64FC6EF836AEE1EDDB" box="[951,966,1929,1954]" italics="true" pageId="157" pageNumber="158">–</emphasis>
<emphasis id="B96F064FFF38FF64FC14F837A92CEDDB" bold="true" box="[973,1035,1928,1954]" pageId="157" pageNumber="158">
<location id="8EC48C86FF38FF64FC14F837A92CEDDB" LSID="urn:lsid:plazi:treatment:03B26B4BFF38FF66FD8BF9EFAC9DEE0E:8EC48C86FF38FF64FC14F837A92CEDDB" box="[973,1035,1928,1954]" country="Peru" county="Museo de Historia Natural Javier Prado" municipality="Botanica" name="Type" pageId="157" pageNumber="158" stateProvince="Cajamarca">Type</location>
</emphasis>
:
<materialsCitation id="3B73D000FF38FF64FBC5F837AF04ED91" collectingDate="1904-04-26" collectionCode="G" collectorName="A. Weberbauer" country="Peru" elevation="2550" latitude="-7.116667" location="uber San Pablo" longLatPrecision="1303" longitude="-78.833336" pageId="157" pageNumber="158" specimenCode="G00034148" specimenCount="1" stateProvince="Cajamarca Region">
<collectingCountry id="F30C9ACDFF38FF64FBC5F837A940EDDB" box="[1052,1127,1928,1954]" name="Peru" pageId="157" pageNumber="158">PERU</collectingCountry>
–
<emphasis id="B96F064FFF38FF64FB50F837A85FEDDB" bold="true" box="[1161,1400,1928,1954]" pageId="157" pageNumber="158">
<collectingRegion id="49DF14BFFF38FF64FB50F837A85FEDDB" box="[1161,1400,1928,1954]" country="Peru" name="Cajamarca" pageId="157" pageNumber="158">Cajamarca Region</collectingRegion>
</emphasis>
•
<location id="8EC48C86FF38FF64FF26F814AC9CEDBC" LSID="urn:lsid:plazi:treatment:03B26B4BFF38FF66FD8BF9EFAC9DEE0E:8EC48C86FF38FF64FF26F814AC9CEDBC" box="[255,443,1963,1989]" country="Peru" latitude="-7.116667" longLatPrecision="1303" longitude="-78.833336" name="uber San Pablo" pageId="157" pageNumber="158" stateProvince="Cajamarca Region">uber San Pablo</location>
; [
<geoCoordinate id="EE2FBC9AFF38FF64FE08F813AF0AEDBC" box="[465,557,1962,1990]" degrees="7" direction="south" minutes="07" orientation="latitude" pageId="157" pageNumber="158" precision="925" value="-7.116667">7°07′ S</geoCoordinate>
,
<geoCoordinate id="EE2FBC9AFF38FF64FDE4F815AF92EDBF" box="[573,693,1962,1990]" degrees="78" direction="west" minutes="50" orientation="longitude" pageId="157" pageNumber="158" precision="925" value="-78.833336">78°50′ W</geoCoordinate>
];
<elevation id="00363D6EFF38FF64FD15F814AE93EDBC" box="[716,948,1963,1990]" metricMagnitude="3" metricUnit="m" metricValue="2.55" metricValueMax="2.7" metricValueMin="2.4" pageId="157" pageNumber="158" unit="m" value="2550.0" valueMax="2700.0" valueMin="2400.0">
<quantity id="4CE377B8FF38FF64FD15F814AE56EDBF" box="[716,881,1963,1990]" metricMagnitude="3" metricUnit="m" metricValue="2.55" metricValueMax="2.7" metricValueMin="2.4" pageId="157" pageNumber="158" unit="m" value="2550.0" valueMax="2700.0" valueMin="2400.0">2400–2700 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA5FC9DFF38FF64FC1DF814A941EDBC" box="[964,1126,1963,1989]" pageId="157" pageNumber="158" value="1904-04-26">
<collectingDate id="EFE10575FF38FF64FC1DF814A941EDBC" box="[964,1126,1963,1989]" pageId="157" pageNumber="158" value="1904-04-26">26 Apr. 1904</collectingDate>
</date>
;
<emphasis id="B96F064FFF38FF64FBACF814A849EDBC" box="[1141,1390,1963,1989]" italics="true" pageId="157" pageNumber="158">
<collectorName id="26EEBF8BFF38FF64FBACF814A80EEDBC" box="[1141,1321,1963,1989]" pageId="157" pageNumber="158">A. Weberbauer</collectorName>
3824
</emphasis>
; holotype:
<collectionCode id="ED0A4298FF38FF64FEB9F871AC52ED91" box="[352,373,1998,2024]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="157" pageNumber="158" type="Herbarium">G</collectionCode>
[
<specimenCode id="DBBD7226FF38FF64FE5DF871AF3FED91" box="[388,536,1998,2024]" collectionCode="G" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="157" pageNumber="158" type="Herbarium">G00034148</specimenCode>
];
</materialsCitation>
<materialsCitation id="3B73D000FF38FF64FDF2F871A929ED90" box="[555,1038,1998,2025]" collectingDate="1904-04-26" collectionCode="B" collectorName="A. Weberbauer" country="Peru" elevation="2550" latitude="-7.116667" location="uber San Pablo" longLatPrecision="1303" longitude="-78.833336" pageId="157" pageNumber="158" specimenCode="B100186581, B100186582" specimenCount="1" stateProvince="Cajamarca Region">
isotypes:
<collectionCode id="ED0A4298FF38FF64FD43F870AF89ED90" box="[666,686,1999,2025]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="157" pageNumber="158" type="Herbarium">B</collectionCode>
[
<specimenCode id="DBBD7226FF38FF64FD65F871AE7DED90" box="[700,858,1998,2025]" collectionCode="B" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="157" pageNumber="158" type="Herbarium">B100186581</specimenCode>
,
<specimenCode id="DBBD7226FF38FF64FCBCF871A923ED90" box="[869,1028,1998,2025]" collectionCode="B" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="157" pageNumber="158" type="Herbarium">B100186582</specimenCode>
].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF3BFF67FF64FEB6AF68EB5A" box="[189,591,265,291]" pageId="158" pageNumber="159" type="reference_group">
<paragraph id="8BA4DA5DFF3BFF67FF64FEB6AF68EB5A" blockId="158.[189,591,265,291]" box="[189,591,265,291]" pageId="158" pageNumber="159">
<bibRefCitation id="EF8AA7ACFF3BFF67FF64FEB6AF6BEB5A" author="Smith &amp; Wasshausen" box="[189,588,265,291]" firstAuthor="Smith" page="468" pageId="158" pageNumber="159" pagination="465 - 473" refId="ref231805" refString="Smith L. B. &amp; Wasshausen D. C. 1984. Notes on Begoniaceae, III. Phytologia 54 (7): 465 - 473. https: // doi. org / 10.5962 / bhl. part. 14431" type="journal article" year="1984">Smith &amp; Wasshausen (1984: 468)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF3BFF67FF64FEF3A80DEBEC" box="[189,1322,332,406]" pageId="158" pageNumber="159" type="etymology">
<paragraph id="8BA4DA5DFF3BFF67FF64FEF3AC62EB1F" blockId="158.[189,325,332,358]" box="[189,325,332,358]" pageId="158" pageNumber="159">
<heading id="D0EC6D31FF3BFF67FF64FEF3AC62EB1F" bold="true" box="[189,325,332,358]" fontSize="11" level="3" pageId="158" pageNumber="159" reason="3">
<emphasis id="B96F064FFF3BFF67FF64FEF3AC62EB1F" bold="true" box="[189,325,332,358]" pageId="158" pageNumber="159">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF3BFF67FF64FEC4A80DEBEC" blockId="158.[189,1322,379,406]" box="[189,1322,379,406]" pageId="158" pageNumber="159">
The
<typeStatus id="54A064FFFF3BFF67FF2BFEC3AC02EBEF" box="[242,293,380,406]" pageId="158" pageNumber="159">type</typeStatus>
collection was collected by August Weberbauer and the species is named in his honour.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF3BFF67FF64FE01AF68E99B" pageId="158" pageNumber="159" type="materials_examined">
<paragraph id="8BA4DA5DFF3BFF67FF64FE01AF09EBA1" blockId="158.[189,558,446,472]" box="[189,558,446,472]" pageId="158" pageNumber="159">
<heading id="D0EC6D31FF3BFF67FF64FE01AF09EBA1" bold="true" box="[189,558,446,472]" fontSize="11" level="3" pageId="158" pageNumber="159" reason="3">
<emphasis id="B96F064FFF3BFF67FF64FE01AF09EBA1" bold="true" box="[189,558,446,472]" pageId="158" pageNumber="159">Selected specimens examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF3BFF67FF64FE51AF68E99B" blockId="158.[188,1399,494,994]" pageId="158" pageNumber="159">
<materialsCitation id="3B73D000FF3BFF67FF64FE51A96FE855" collectingDate="1948-03-31" collectionCode="USM" collectorName="R. Ferreyra" country="Peru" county="Prov. Cajamarca" elevation="3200" latitude="-7.233333" location="Cumbre El Gavilan" longLatPrecision="1303" longitude="-78.46667" pageId="158" pageNumber="159" specimenCount="1" stateProvince="Cajamarca Region">
<collectingCountry id="F30C9ACDFF3BFF67FF64FE51AC2FE871" box="[189,264,494,520]" name="Peru" pageId="158" pageNumber="159">PERU</collectingCountry>
–
<emphasis id="B96F064FFF3BFF67FEF7FE51AE37E871" bold="true" box="[302,784,494,520]" pageId="158" pageNumber="159">
<collectingRegion id="49DF14BFFF3BFF67FEF7FE51AF04E871" box="[302,547,494,520]" country="Peru" name="Cajamarca" pageId="158" pageNumber="159">Cajamarca Region</collectingRegion>
:
<collectingCounty id="62C5A2D1FF3BFF67FDECFE51AE37E871" box="[565,784,494,520]" pageId="158" pageNumber="159">Prov. Cajamarca</collectingCounty>
</emphasis>
•
<location id="8EC48C86FF3BFF67FCE8FE51A906E871" LSID="urn:lsid:plazi:treatment:03B26B4BFF38FF66FD8BF9EFAC9DEE0E:8EC48C86FF3BFF67FCE8FE51A906E871" box="[817,1057,494,520]" country="Peru" county="Prov. Cajamarca" latitude="-7.233333" longLatPrecision="1303" longitude="-78.46667" name="Cumbre El Gavilan" pageId="158" pageNumber="159" stateProvince="Cajamarca Region">Cumbre El Gavilán</location>
,
<locationDeviation id="9A16C37FFF3BFF67FBE9FE51A855E871" box="[1072,1394,494,520]" pageId="158" pageNumber="159">entre Chilete y Cajamarca</locationDeviation>
; [
<geoCoordinate id="EE2FBC9AFF3BFF67FF1CFDAEAC06E855" box="[197,289,529,557]" degrees="7" direction="south" minutes="14" orientation="latitude" pageId="158" pageNumber="159" precision="925" value="-7.233333">7°14′ S</geoCoordinate>
,
<geoCoordinate id="EE2FBC9AFF3BFF67FEE8FDAEAC8FE854" box="[305,424,529,557]" degrees="78" direction="west" minutes="28" orientation="longitude" pageId="158" pageNumber="159" precision="925" value="-78.46667">78°28′ W</geoCoordinate>
];
<elevation id="00363D6EFF3BFF67FE66FDADAF7AE855" box="[447,605,530,557]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="158" pageNumber="159" unit="m" value="3200.0">
<quantity id="4CE377B8FF3BFF67FE66FDADAF3CE854" box="[447,539,530,557]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="158" pageNumber="159" unit="m" value="3200.0">3200 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA5FC9DFF3BFF67FDB4FDADAE34E855" box="[621,787,530,557]" pageId="158" pageNumber="159" value="1948-03-31">
<collectingDate id="EFE10575FF3BFF67FDB4FDADAE34E855" box="[621,787,530,557]" pageId="158" pageNumber="159" value="1948-03-31">31 Mar. 1948</collectingDate>
</date>
;
<emphasis id="B96F064FFF3BFF67FCFBFDACAED2E855" box="[802,1013,530,556]" italics="true" pageId="158" pageNumber="159">
<collectorName id="26EEBF8BFF3BFF67FCFBFDACAE97E855" box="[802,944,531,556]" pageId="158" pageNumber="159">R. Ferreyra</collectorName>
3282
</emphasis>
;
<collectionCode id="ED0A4298FF3BFF67FBD1FDADA96FE855" box="[1032,1096,530,556]" country="Malaysia" httpUri="http://grbio.org/cool/11pi-mg53" name="Universiti Sains Malaysia" pageId="158" pageNumber="159">USM</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B73D000FF3BFF67FBBFFDADAE9BE80C" collectingDate="2003-03-20" collectionCode="CPUN" collectorName="I. Sanchez V. &amp; M. Sanchez M. &amp; R. Cueva R. &amp; J. Montoya" elevation="3270" latitude="-7.23375" location="Dist. Cajamarca" longLatPrecision="15" longitude="-78.49" municipality="Dist. Cajamarca" pageId="158" pageNumber="159" specimenCount="1">
<collectingMunicipality id="6BC04027FF3BFF67FBBFFDADA80EE855" box="[1126,1321,530,556]" pageId="158" pageNumber="159">Dist. Cajamarca</collectingMunicipality>
,
<locationDeviation id="9A16C37FFF3BFF67FAEEFDADAF15E828" pageId="158" pageNumber="159">sobre la cima de la ladera occidental</locationDeviation>
;
<geoCoordinate id="EE2FBC9AFF3BFF67FD98FD8AAFFDE828" box="[577,730,565,593]" degrees="7" direction="south" minutes="14" orientation="latitude" pageId="158" pageNumber="159" precision="1" seconds="01.5" value="-7.23375">7°14′01.5″ S</geoCoordinate>
,
<geoCoordinate id="EE2FBC9AFF3BFF67FD33FD8AAEAFE828" box="[746,904,565,593]" degrees="78" direction="west" minutes="29" orientation="longitude" pageId="158" pageNumber="159" precision="15" seconds="24" value="-78.49">78°29′24″ W</geoCoordinate>
;
<elevation id="00363D6EFF3BFF67FC4EFD88A912E828" box="[919,1077,567,593]" metricMagnitude="3" metricUnit="m" metricValue="3.27" pageId="158" pageNumber="159" unit="m" value="3270.0">
<quantity id="4CE377B8FF3BFF67FC4EFD88AED5E828" box="[919,1010,567,593]" metricMagnitude="3" metricUnit="m" metricValue="3.27" pageId="158" pageNumber="159" unit="m" value="3270.0">3270 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA5FC9DFF3BFF67FB9CFD88A9CCE828" box="[1093,1259,567,593]" pageId="158" pageNumber="159" value="2003-03-20">
<collectingDate id="EFE10575FF3BFF67FB9CFD88A9CCE828" box="[1093,1259,567,593]" pageId="158" pageNumber="159" value="2003-03-20">20 Mar. 2003</collectingDate>
</date>
;
<emphasis id="B96F064FFF3BFF67FB23FD87AE7DE80C" italics="true" pageId="158" pageNumber="159">
<collectorName id="26EEBF8BFF3BFF67FB23FD87ADF1E80C" pageId="158" pageNumber="159">I. Sánchez V.</collectorName>
,
<collectorName id="26EEBF8BFF3BFF67FF3EFDE3AC84E80C" box="[231,419,603,629]" pageId="158" pageNumber="159">M. Sánchez M.</collectorName>
,
<collectorName id="26EEBF8BFF3BFF67FE6DFDE3AF6BE80C" box="[436,588,604,630]" pageId="158" pageNumber="159">R. Cueva R.</collectorName>
&amp;
<collectorName id="26EEBF8BFF3BFF67FDA2FDE3AE22E80C" box="[635,773,604,629]" pageId="158" pageNumber="159">J. Montoya</collectorName>
11766
</emphasis>
;
<collectionCode id="ED0A4298FF3BFF67FCB7FDE4AE9BE80C" box="[878,956,603,629]" country="Peru" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14355" lsid="urn:lsid:biocol.org:col:14355" name="Universidad Nacional de Cajamarca" pageId="158" pageNumber="159" type="Herbarium">CPUN</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B73D000FF3BFF67FC06FDE4AFB8E8C6" collectingDate="1975-05-22" collectionCode="HUT, MO, US" collectorName="A. Sagastegui A. &amp; C. Cabanillas S. &amp; O. Dias C." elevation="2320" latitude="-7.2833333" location="San Pablo" longLatPrecision="1303" longitude="-78.8" municipality="El Molino" pageId="158" pageNumber="159" specimenCode="MO-2228052, US00424972" specimenCount="1">
<collectingMunicipality id="6BC04027FF3BFF67FC06FDE4A97CE80C" box="[991,1115,603,629]" pageId="158" pageNumber="159">El Molino</collectingMunicipality>
(
<location id="8EC48C86FF3BFF67FBA9FDE4A9D5E80C" LSID="urn:lsid:plazi:treatment:03B26B4BFF38FF66FD8BF9EFAC9DEE0E:8EC48C86FF3BFF67FBA9FDE4A9D5E80C" box="[1136,1266,603,629]" country="United States of America" latitude="-7.2833333" longLatPrecision="1303" longitude="-78.8" municipality="El Molino" name="San Pablo" pageId="158" pageNumber="159">San Pablo</location>
); [
<geoCoordinate id="EE2FBC9AFF3BFF67FACBFDE5A856E80C" box="[1298,1393,602,630]" degrees="7" direction="south" minutes="17" orientation="latitude" pageId="158" pageNumber="159" precision="925" value="-7.2833333">7°17′ S</geoCoordinate>
,
<geoCoordinate id="EE2FBC9AFF3BFF67FF64FDC1AC13E8E3" box="[189,308,638,666]" degrees="78" direction="west" minutes="48" orientation="longitude" pageId="158" pageNumber="159" precision="925" value="-78.8">78°48′ W</geoCoordinate>
];
<elevation id="00363D6EFF3BFF67FE93FD3FACC0E8E3" box="[330,487,640,666]" metricMagnitude="3" metricUnit="m" metricValue="2.32" pageId="158" pageNumber="159" unit="m" value="2320.0">
<quantity id="4CE377B8FF3BFF67FE93FD3FAC82E8E3" box="[330,421,640,666]" metricMagnitude="3" metricUnit="m" metricValue="2.32" pageId="158" pageNumber="159" unit="m" value="2320.0">2320 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA5FC9DFF3BFF67FE2EFD3FAFBCE8E3" box="[503,667,639,666]" pageId="158" pageNumber="159" value="1975-05-22">
<collectingDate id="EFE10575FF3BFF67FE2EFD3FAFBCE8E3" box="[503,667,639,666]" pageId="158" pageNumber="159" value="1975-05-22">22 May 1975</collectingDate>
</date>
;
<emphasis id="B96F064FFF3BFF67FD70FD3FA804E8E3" box="[681,1315,640,666]" italics="true" pageId="158" pageNumber="159">
<collectorName id="26EEBF8BFF3BFF67FD70FD3FAE54E8E3" box="[681,883,640,666]" pageId="158" pageNumber="159">A. Sagástegui A.</collectorName>
,
<collectorName id="26EEBF8BFF3BFF67FC5AFD3FA96CE8E3" box="[899,1099,640,666]" pageId="158" pageNumber="159">C. Cabanillas S.</collectorName>
,
<collectorName id="26EEBF8BFF3BFF67FB82FD3FA9F9E8E3" box="[1115,1246,640,666]" pageId="158" pageNumber="159">O. Dias C.</collectorName>
8003
</emphasis>
;
<collectionCode id="ED0A4298FF3BFF67FAECFD3FA855E8E3" box="[1333,1394,640,666]" country="Japan" httpUri="http://grbio.org/cool/gtk4-xv5b" name="HUT Culture Collection" pageId="158" pageNumber="159">HUT</collectionCode>
,
<collectionCode id="ED0A4298FF3BFF67FF66FD1BADC9E8C7" box="[191,238,676,702]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="158" pageNumber="159" type="Herbarium">MO</collectionCode>
[
<specimenCode id="DBBD7226FF3BFF67FED9FD1BACB8E8C6" box="[256,415,676,703]" pageId="158" pageNumber="159">MO-2228052</specimenCode>
],
<collectionCode id="ED0A4298FF3BFF67FE63FD1BACF8E8C7" box="[442,479,676,702]" pageId="158" pageNumber="159">US</collectionCode>
[
<specimenCode id="DBBD7226FF3BFF67FE2AFD1BAFA9E8C7" box="[499,654,676,702]" pageId="158" pageNumber="159">US00424972</specimenCode>
].
</materialsCitation>
–
<materialsCitation id="3B73D000FF3BFF67FD1AFD1BAFECE97E" collectingDate="2007-04-24" collectionCode="E, K, USM" collectorName="T. E. Sarkinen &amp; C. E. Hughes &amp; A. Daza &amp; H. M. Baden" county="Prov. San Pablo" elevation="3280" latitude="-7.23" location="On the path to El Molino, caserillo El Cevil" longLatPrecision="21" longitude="-78.49027" pageId="158" pageNumber="159" specimenCode="E00607545" specimenCount="1">
<emphasis id="B96F064FFF3BFF67FD1AFD1BAEB6E8C7" bold="true" box="[707,913,676,702]" pageId="158" pageNumber="159">
<collectingCounty id="62C5A2D1FF3BFF67FD1AFD1BAEB6E8C7" box="[707,913,676,702]" pageId="158" pageNumber="159">Prov. San Pablo</collectingCounty>
</emphasis>
•
<location id="8EC48C86FF3BFF67FC74FD1BAC25E89B" LSID="urn:lsid:plazi:treatment:03B26B4BFF38FF66FD8BF9EFAC9DEE0E:8EC48C86FF3BFF67FC74FD1BAC25E89B" county="Prov. San Pablo" latitude="-7.23" longLatPrecision="21" longitude="-78.49027" name="On the path to El Molino, caserillo El Cevil" pageId="158" pageNumber="159">On the path to El Molino, caserillo El Cevil</location>
;
<geoCoordinate id="EE2FBC9AFF3BFF67FEC9FD78ACA8E89A" box="[272,399,711,739]" degrees="7" direction="south" minutes="13" orientation="latitude" pageId="158" pageNumber="159" precision="15" seconds="48" value="-7.23">7°13′48″ S</geoCoordinate>
,
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;
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ca
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east of and below San Pablo by Trail
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&amp;
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,
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[
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•
<materialsCitation id="3B73D000FF3BFF67FCA7FC89A90DE90C" collectingDate="1966-04-24" collectionCode="HUT" collectorName="A. Sagastegui A. &amp; M. Fukuskima" elevation="2700" latitude="-7.366667" location="Arriba de Cotumaza" longLatPrecision="1303" longitude="-78.8" municipality="Arriba de Cotumaza" pageId="158" pageNumber="159" specimenCount="1">
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<collectingDate id="EFE10575FF3BFF67FEBCFCE5AF26E90D" box="[357,513,858,885]" pageId="158" pageNumber="159" value="1966-04-24">24 Apr. 1966</collectingDate>
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&amp;
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6109
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;
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</materialsCitation>
•
<materialsCitation id="3B73D000FF3BFF67FB9DFCE5A850E9E0" collectingDate="2015-05-23" collectionCode="E, MOL" collectorName="M. C. Tebbitt &amp; A. Daza" elevation="2669" latitude="-7.4044447" longLatPrecision="21" longitude="-78.79139" pageId="158" pageNumber="159" specimenCount="1">
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,
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&amp;
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829
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–
<materialsCitation id="3B73D000FF3BFF67FF0DFC1CAF68E99B" collectingDate="1997-06-07" collectionCode="HUT" collectorName="M. Rodriguez E." county="Prov. Otuzco" elevation="3000" latitude="-7.9833336" location="San Pedro" longLatPrecision="1302" longitude="-78.5" municipality="Motil" pageId="158" pageNumber="159" specimenCount="1" stateProvince="La Libertad Region">
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:
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•
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(
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,
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a.s.l.
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155
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.
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</paragraph>
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<subSubSection id="C30189D6FF3BFF67FF64FBB5ACF0ED28" pageId="158" pageNumber="159" type="description">
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<paragraph id="8BA4DA5DFF3BFF67FF64FB85ACF0ED28" blockId="158.[187,1400,1082,1874]" pageId="158" pageNumber="159">
Acaulescent, rhizomatous herb, to
<quantity id="4CE377B8FF3BFF67FD87FB85AF8EEE2C" box="[606,681,1082,1109]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" pageId="158" pageNumber="159" unit="cm" value="25.0">25 cm</quantity>
high.
<emphasis id="B96F064FFF3BFF67FD21FB85AE79EE2D" box="[760,862,1082,1108]" italics="true" pageId="158" pageNumber="159">Rhizome</emphasis>
ellipsoid, 5–10 ×
<quantity id="4CE377B8FF3BFF67FBE4FB85A997EE2C" box="[1085,1200,1082,1109]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="158" pageNumber="159" unit="cm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 cm</quantity>
, with 1 growing point.
<emphasis id="B96F064FFF3BFF67FED0FBE1AC40EE01" box="[265,359,1118,1144]" italics="true" pageId="158" pageNumber="159">Stipules</emphasis>
persistent, triangular, 8–10 ×
<quantity id="4CE377B8FF3BFF67FD16FBE1AE30EE00" box="[719,791,1118,1145]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="158" pageNumber="159" unit="mm" value="6.0">6 mm</quantity>
, apex acute, opaque, dark brown, glabrous, margin entire, aciliate.
<emphasis id="B96F064FFF3BFF67FEAFFB3BACEFEEE4" box="[374,456,1156,1181]" italics="true" pageId="158" pageNumber="159">Leaves</emphasis>
1–3, alternate, basifixed; petiole
<quantity id="4CE377B8FF3BFF67FC8CFB3CAE9BEEE7" box="[853,956,1155,1182]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" metricValueMax="1.8" metricValueMin="0.7" pageId="158" pageNumber="159" unit="cm" value="12.5" valueMax="18.0" valueMin="7.0">7–18 cm</quantity>
long, pale green, sparsely to densely villous; blade subsymmetric, ovate, to 10 ×
<quantity id="4CE377B8FF3BFF67FD64FB17AE2FEEBB" box="[701,776,1192,1218]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" pageId="158" pageNumber="159" unit="cm" value="13.0">13 cm</quantity>
, succulent, apex acuminate, base cordate, basal lobes overlapping or not overlapping, sinus to
<quantity id="4CE377B8FF3BFF67FD4DFB73AFCAEE9F" box="[660,749,1228,1255]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="158" pageNumber="159" unit="mm" value="2.5">2.5 mm</quantity>
deep, margin serrate, with&gt; 10 to deeply-incised lobes, ciliate, upper surface pale green, densely pilose, lower surface pale green, glabrous, veins palmate-pinnate, 6–8 veined from the base.
<emphasis id="B96F064FFF3BFF67FD96FAAAAFD0EF56" box="[591,759,1301,1327]" italics="true" pageId="158" pageNumber="159">Inflorescences</emphasis>
1–3, bisexual, axillary, erect, cymose, with 3 branches, bearing up to 2 staminate flowers and 1 pistillate flowers, protandrous; peduncle to
<quantity id="4CE377B8FF3BFF67FB44FA86A9CEEF2D" box="[1181,1257,1337,1364]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="158" pageNumber="159" unit="cm" value="20.0">20 cm</quantity>
long, green to pink, glabrous to densely villous, bracts persistent, ovate, 3–10 ×
<quantity id="4CE377B8FF3BFF67FC28FAE1A970EF01" box="[1009,1111,1374,1400]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="9.0" metricValueMin="2.0" pageId="158" pageNumber="159" unit="mm" value="5.5" valueMax="9.0" valueMin="2.0">2–9 mm</quantity>
, opaque, glabrous, apex obtuse, margin entire, ciliate.
<emphasis id="B96F064FFF3BFF67FDF1FA3DAFD9EFE5" box="[552,766,1410,1436]" italics="true" pageId="158" pageNumber="159">Staminate flowers</emphasis>
: pedicels to
<quantity id="4CE377B8FF3BFF67FC47FA3DAED3EFE4" box="[926,1012,1410,1437]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="158" pageNumber="159" unit="mm" value="35.0">35 mm</quantity>
long, sparsely-villous; tepals 4, spreading, outer 2 ovate, 22–28 ×
<quantity id="4CE377B8FF3BFF67FD84FA19AFC6EFB8" box="[605,737,1446,1473]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" metricValueMax="1.9" metricValueMin="1.5" pageId="158" pageNumber="159" unit="mm" value="17.0" valueMax="19.0" valueMin="15.0">15–19 mm</quantity>
, apex rounded, white, glabrous, margin entire, aciliate, inner 2 ovate, 13–24 ×
<quantity id="4CE377B8FF3BFF67FE0EFA74AF7CEF9F" box="[471,603,1483,1510]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.7" metricValueMin="1.3" pageId="158" pageNumber="159" unit="mm" value="15.0" valueMax="17.0" valueMin="13.0">13–17 mm</quantity>
, apex truncate, white, glabrous, margin entire, aciliate; stamens ca 25, spreading, yellow, filaments
<quantity id="4CE377B8FF3BFF67FDE4FA4FAFBAEC73" box="[573,669,1520,1546]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="158" pageNumber="159" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity>
long, free, anthers ovoid, 0.8–1 ×
<quantity id="4CE377B8FF3BFF67FBECFA4FA9B7EC73" box="[1077,1168,1520,1546]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="158" pageNumber="159" unit="mm" value="0.8">0.8 mm</quantity>
long, dehiscing via lateral slits, connectives not extending, symmetrically basifixed.
<emphasis id="B96F064FFF3BFF67FC66F9ABA9AFEC57" box="[959,1160,1556,1582]" italics="true" pageId="158" pageNumber="159">Pistillate flowers</emphasis>
: pedicels to
<quantity id="4CE377B8FF3BFF67FAFAF9ABA850EC57" box="[1315,1399,1556,1582]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="158" pageNumber="159" unit="mm" value="10.0">10 mm</quantity>
long; bracteoles lacking; tepals 5, subequal, persistence in fruit, spreading, the largest ovate, ca 17 ×
<quantity id="4CE377B8FF3BFF67FF00F9E2AC03EC0E" box="[217,292,1629,1655]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="158" pageNumber="159" unit="mm" value="9.0">9 mm</quantity>
, apex rounded, white, glabrous, margin entire, aciliate, the smallest ovate, ca 10 ×
<quantity id="4CE377B8FF3BFF67FAFEF9E2A855EC0E" box="[1319,1394,1629,1655]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="158" pageNumber="159" unit="mm" value="9.0">9 mm</quantity>
, apex rounded, white, glabrous, margin entire, aciliate; ovary body ovoid, ca 10 ×
<quantity id="4CE377B8FF3BFF67FB45F93EA9C2ECE5" box="[1180,1253,1665,1692]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="158" pageNumber="159" unit="mm" value="8.0">8 mm</quantity>
, pale green, glabrous, unequally 3-winged, the largest wing triangular ca 10 ×
<quantity id="4CE377B8FF3BFF67FC09F919A93FECB9" box="[976,1048,1702,1728]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="158" pageNumber="159" unit="mm" value="7.0">7 mm</quantity>
, smallest marginal and
<quantity id="4CE377B8FF3BFF67FAE8F919A850ECB9" box="[1329,1399,1702,1728]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="158" pageNumber="159" unit="mm" value="1.0">1 mm</quantity>
wide; 3-locular, placentae branches divided, bearing ovules on both surfaces; styles 3, yellow, free, ca
<quantity id="4CE377B8FF3BFF67FF67F951AC24ED70" box="[190,259,1774,1801]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="158" pageNumber="159" unit="mm" value="5.0">5 mm</quantity>
long, 2-lobed, the lobes reniform, stigmatic papillae in a spirally-twisted band.
<emphasis id="B96F064FFF3BFF67FB6FF950A85FED71" box="[1206,1400,1774,1800]" italics="true" pageId="158" pageNumber="159">Fruiting pedicel</emphasis>
to
<quantity id="4CE377B8FF3BFF67FF07F8ACAC14ED57" box="[222,307,1811,1838]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="158" pageNumber="159" unit="mm" value="10.0">10 mm</quantity>
long.
<emphasis id="B96F064FFF3BFF67FE5DF8ABAF23ED54" box="[388,516,1811,1837]" italics="true" pageId="158" pageNumber="159">Fruit body</emphasis>
ovoid, to 10 ×
<quantity id="4CE377B8FF3BFF67FD1BF8ACAE2CED57" box="[706,779,1811,1838]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="158" pageNumber="159" unit="mm" value="8.0">8 mm</quantity>
, drying dark brown, wings same shape as in ovary, expanding to
<quantity id="4CE377B8FF3BFF67FE86F887AC84ED2B" box="[351,419,1848,1874]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="158" pageNumber="159" unit="mm" value="8.0">8 mm</quantity>
tall.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF3BFF66FF64F8C5A851EB3E" lastPageId="159" lastPageNumber="160" pageId="158" pageNumber="159" type="conservation">
<paragraph id="8BA4DA5DFF3BFF67FF64F8C5AF4AEDED" blockId="158.[189,621,1914,1940]" box="[189,621,1914,1940]" pageId="158" pageNumber="159">
<heading id="D0EC6D31FF3BFF67FF64F8C5AF4AEDED" bold="true" box="[189,621,1914,1940]" fontSize="11" level="3" pageId="158" pageNumber="159" reason="3">
<emphasis id="B96F064FFF3BFF67FF64F8C5AF4AEDED" bold="true" box="[189,621,1914,1940]" pageId="158" pageNumber="159">Proposed conservation assessment</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF3BFF67FF64F815A851ED90" blockId="158.[189,1399,1962,2025]" pageId="158" pageNumber="159">
Assessed by
<bibRefCitation id="EF8AA7ACFF3BFF67FE8AF815AF5CEDBD" author="Leon &amp; Monsalve" box="[339,635,1962,1988]" firstAuthor="Leon" pageId="158" pageNumber="159" pagination="165 - 170" refId="ref229226" refString="Leon B. &amp; Monsalve C. 2006. Begoniaceae endemicas del Peru. Revista Peruana de Biologia 13 (2): 165 - 170. https: // doi. org / 10.15381 / rpb. v 13 i 2.1810" type="journal article" year="2006">
<collectingRegion id="49DF14BFFF3BFF67FE8AF815ACA8EDBD" box="[339,399,1962,1988]" country="Nicaragua" name="Leon" pageId="158" pageNumber="159">León</collectingRegion>
&amp; Monsalve (2006)
</bibRefCitation>
as Data Deficient (DD). We have now identified specimens of the species from five localities in
<collectingRegion id="49DF14BFFF3BFF67FDCCF871AFB5ED91" box="[533,658,1998,2024]" country="Peru" name="Cajamarca" pageId="158" pageNumber="159">Cajamarca</collectingRegion>
and
<collectingRegion id="49DF14BFFF3BFF67FD11F870AE77ED91" box="[712,848,1998,2025]" country="Peru" name="La Libertad" pageId="158" pageNumber="159">La Libertad</collectingRegion>
Regions, with a combined EOO of&gt;
<quantity id="4CE377B8FF3BFF67FAD8F871A841ED91" box="[1281,1382,1998,2024]" metricMagnitude="6" metricUnit="m" metricValue="2.0" pageId="158" pageNumber="159" unit="km" value="2000.0">2000 km</quantity>
<superScript id="7C6E7715FF3BFF67FABEF871A857EDA5" attach="right" box="[1383,1392,1998,2012]" fontSize="6" pageId="158" pageNumber="159">2</superScript>
.
</paragraph>
<paragraph id="8BA4DA5DFF3AFF66FF64FEB6A851EB3E" blockId="159.[189,1398,265,327]" pageId="159" pageNumber="160">
The species prefers limestone outcrops in Dry Forest habitats, many of which are mined for concrete and are highly degraded throughout its range. We assess
<taxonomicName id="4C1BA1DEFF3AFF66FC8AFE92A92DEB3E" box="[851,1034,301,327]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="159" pageNumber="160" phylum="Tracheophyta" rank="species" species="weberbaueri">
<emphasis id="B96F064FFF3AFF66FC8AFE92A92DEB3E" box="[851,1034,301,327]" italics="true" pageId="159" pageNumber="160">B. weberbaueri</emphasis>
</taxonomicName>
as Endangered (EN B1ab(iii)).
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF3AFF66FF64FED1ACF9E8C1" pageId="159" pageNumber="160" type="discussion">
<paragraph id="8BA4DA5DFF3AFF66FF64FED1ACBAEBF0" blockId="159.[189,413,366,393]" box="[189,413,366,393]" pageId="159" pageNumber="160">
<heading id="D0EC6D31FF3AFF66FF64FED1ACBAEBF0" bold="true" box="[189,413,366,393]" fontSize="11" level="3" pageId="159" pageNumber="160" reason="3">
<emphasis id="B96F064FFF3AFF66FF64FED1ACBAEBF0" bold="true" box="[189,413,366,393]" pageId="159" pageNumber="160">Typification notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF3AFF66FF64FE21AC4BE85A" blockId="159.[187,1400,414,547]" pageId="159" pageNumber="160">
The protologue of
<taxonomicName id="4C1BA1DEFF3AFF66FE45FE20AF73EBC1" box="[412,596,414,440]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="159" pageNumber="160" phylum="Tracheophyta" rank="species" species="weberbaueri">
<emphasis id="B96F064FFF3AFF66FE45FE20AF73EBC1" box="[412,596,414,440]" italics="true" pageId="159" pageNumber="160">B. weberbaueri</emphasis>
</taxonomicName>
states that the type collection is held in
<collectingRegion id="49DF14BFFF3AFF66FBEBFE21A95CEBC1" box="[1074,1147,414,440]" country="Germany" name="Berlin" pageId="159" pageNumber="160">Berlin</collectingRegion>
(
<bibRefCitation id="EF8AA7ACFF3AFF66FB52FE21A84DEBC1" author="Irmscher" box="[1163,1386,414,441]" firstAuthor="Irmscher" page="78" pageId="159" pageNumber="160" pagination="1 - 102" refId="ref228521" refString="Irmscher E. 1953. Systematische Studien uber Begoniaceen des tropischen Sudamerikas, besonders Brasiliens. Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie 76 (1): 1 - 102." type="journal article" year="1953">Irmscher 1953: 78</bibRefCitation>
). This herbarium holds two duplicates of the type gathering thus one of these must be chosen as a
<typeStatus id="54A064FFFF3AFF66FAD5FE7EA85FEBA2" box="[1292,1400,449,475]" pageId="159" pageNumber="160">lectotype</typeStatus>
(
<bibRefCitation id="EF8AA7ACFF3AFF66FF1AFE5AAC4AEB86" author="McNeill" box="[195,365,485,511]" firstAuthor="McNeill" pageId="159" pageNumber="160" pagination="1112 - 1113" refId="ref229632" refString="McNeill J. 2014. Holotype specimens and type citations: general issues. Taxon 63 (5): 1112 - 1113. https: // doi. org / 10.12705 / 635.7" type="journal article" year="2014">McNeill 2014</bibRefCitation>
). We choose the specimen barcode B100186582 as it has a larger number of flowers than the alternative.
</paragraph>
<paragraph id="8BA4DA5DFF3AFF66FF64FDF4AC97E81C" blockId="159.[189,432,587,613]" box="[189,432,587,613]" pageId="159" pageNumber="160">
<heading id="D0EC6D31FF3AFF66FF64FDF4AC97E81C" bold="true" box="[189,432,587,613]" fontSize="11" level="3" pageId="159" pageNumber="160" reason="3">
<emphasis id="B96F064FFF3AFF66FF64FDF4AC97E81C" bold="true" box="[189,432,587,613]" pageId="159" pageNumber="160">Identification notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF3AFF66FF64FDC4ACF9E8C1" blockId="159.[188,1399,634,696]" pageId="159" pageNumber="160">
<taxonomicName id="4C1BA1DEFF3AFF66FF64FDC4AC9BE8ED" box="[189,444,634,660]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="159" pageNumber="160" phylum="Tracheophyta" rank="species" species="weberbaueri">
<emphasis id="B96F064FFF3AFF66FF64FDC4AC9BE8ED" box="[189,444,634,660]" italics="true" pageId="159" pageNumber="160">Begonia weberbaueri</emphasis>
</taxonomicName>
is the only species of Peruvian
<taxonomicName id="4C1BA1DEFF3AFF66FCF0FDC4AEABE8ED" box="[809,908,635,660]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="159" pageNumber="160" phylum="Tracheophyta" rank="genus">
<emphasis id="B96F064FFF3AFF66FCF0FDC4AEABE8ED" box="[809,908,635,660]" italics="true" pageId="159" pageNumber="160">Begonia</emphasis>
</taxonomicName>
with&gt; 10 deeply incised lobes around the edge of the leaf laminae.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF3AFF66FF64FD60AC9DEE0E" pageId="159" pageNumber="160" type="distribution">
<paragraph id="8BA4DA5DFF3AFF66FF64FD60ACD2E883" blockId="159.[189,501,735,762]" box="[189,501,735,762]" pageId="159" pageNumber="160">
<heading id="D0EC6D31FF3AFF66FF64FD60ACD2E883" bold="true" box="[189,501,735,762]" fontSize="11" level="3" pageId="159" pageNumber="160" reason="3">
<emphasis id="B96F064FFF3AFF66FF64FD60ACD2E883" bold="true" box="[189,501,735,762]" pageId="159" pageNumber="160">Distribution and ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF3AFF66FF64FCB0A932E909" blockId="159.[189,1399,783,881]" pageId="159" pageNumber="160">
Endemic to
<collectingCountry id="F30C9ACDFF3AFF66FE92FCB0ACA6E950" box="[331,385,783,809]" name="Peru" pageId="159" pageNumber="160">Peru</collectingCountry>
and known from
<collectingRegion id="49DF14BFFF3AFF66FD88FCB0AFE9E950" box="[593,718,783,809]" country="Peru" name="Cajamarca" pageId="159" pageNumber="160">Cajamarca</collectingRegion>
and
<collectingRegion id="49DF14BFFF3AFF66FCD1FCB0AEB6E950" box="[776,913,783,809]" country="Peru" name="La Libertad" pageId="159" pageNumber="160">La Libertad</collectingRegion>
Regions (
<figureCitation id="1320C6D8FF3AFF66FBD0FCB0A952E950" box="[1033,1141,783,809]" captionStart="Fig" captionStartId="147.[189,232,1892,1918]" captionTargetBox="[194,1392,1228,1847]" captionTargetId="figure-12@147.[189,1398,1218,1853]" captionTargetPageId="147" captionText="Fig. 48. Distribution of Begonia sect. Eupetalum (Lindl.) A.DC. in Peru and surrounding countries. A. B. geraniifolia Hook. (red) and B. joshii Moonlight (blue). B. B. neoharlingii L.B.Sm. &amp; Wassh. (red) and B. weberbaueri Irmsch. (blue). Black lines indicate country borders, grey lines indicate major administrative divisions, blue lines indicate rivers, and shading indicates elevation." figureDoi="http://doi.org/10.5281/zenodo.8178403" httpUri="https://zenodo.org/record/8178403/files/figure.png" pageId="159" pageNumber="160">Fig. 48B</figureCitation>
). Found in dry forests and scrublands at an elevation of
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a.s.l.
<taxonomicName id="4C1BA1DEFF3AFF66FC92FC8CA994E935" authority="Irmsch." authorityName="Irmsch." box="[843,1203,818,844]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="159" pageNumber="160" phylum="Tracheophyta" rank="species" species="weberbaueri">
<emphasis id="B96F064FFF3AFF66FC92FC8CA968E935" box="[843,1103,818,844]" italics="true" pageId="159" pageNumber="160">Begonia weberbaueri</emphasis>
Irmsch.
</taxonomicName>
dies back to its rhizome in the dry season and flowers in the wet season (March to June).
</paragraph>
<paragraph id="8BA4DA5DFF3AFF66FF64FC26AE75E9CF" blockId="159.[189,850,921,950]" box="[189,850,921,950]" pageId="159" pageNumber="160">
<emphasis id="B96F064FFF3AFF66FF64FC26AE75E9CF" bold="true" box="[189,850,921,950]" italics="true" pageId="159" pageNumber="160">
The
<taxonomicName id="4C1BA1DEFF3AFF66FF20FC26ACA5E9CF" box="[249,386,921,950]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="159" pageNumber="160" phylum="Tracheophyta" rank="species" species="octopetala">octopetala</taxonomicName>
group of
<taxonomicName id="4C1BA1DEFF3AFF66FDDDFC26AE75E9CF" box="[516,850,921,950]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="159" pageNumber="160" phylum="Tracheophyta" rank="section" section="Eupetalum">Begonia sect. Eupetalum</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BA4DA5DFF3AFF66FF64FC71AC9DEE0E" blockId="159.[189,1399,974,1144]" pageId="159" pageNumber="160">
<bibRefCitation id="EF8AA7ACFF3AFF66FF64FC71AC43E990" author="Tebbitt M. C." box="[189,356,974,1001]" journalOrPublisher="Novon" pageId="159" pageNumber="160" pagination="479 - 489" part="23" refId="ref232441" refString="Tebbitt M. C. 2015. Two new species of Andean tuberous Begonia in the B. octopetala group (Begoniaceae). Novon 23 (4): 479 - 489. https: // doi. org / 10.3417 / 2013027" title="Two new species of Andean tuberous Begonia in the B. octopetala group (Begoniaceae)" type="journal article" year="2015">Tebbitt (2015)</bibRefCitation>
defined the octopetala group of
<emphasis id="B96F064FFF3AFF66FD04FC70AFD0E991" box="[733,759,975,1000]" italics="true" pageId="159" pageNumber="160">B.</emphasis>
sect.
<taxonomicName id="4C1BA1DEFF3AFF66FCE3FC71AE9DE991" box="[826,954,974,1000]" class="Magnoliopsida" family="Begoniaceae" genus="Eupetalum" kingdom="Plantae" order="Cucurbitales" pageId="159" pageNumber="160" phylum="Tracheophyta" rank="genus">
<emphasis id="B96F064FFF3AFF66FCE3FC71AE9DE991" box="[826,954,974,1000]" italics="true" pageId="159" pageNumber="160">Eupetalum</emphasis>
</taxonomicName>
to include the members of the section with multifid styles, symmetric leaf blades, and no above ground stems. Most members of the group also have&gt; 4 tepals on the staminate flowers,&gt; 5 tepals on the pistillate flower, and asymmetric to subsymmetric wings on their ovaries and fruits. Of the ten species in the group, six are found in
<collectingCountry id="F30C9ACDFF3AFF66FA98FB85A850EE2D" box="[1345,1399,1082,1108]" name="Peru" pageId="159" pageNumber="160">Peru</collectingCountry>
and two are endemic.
</paragraph>
</subSubSection>
</treatment>
</document>
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