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<document id="8E9DD8559AF9C75AF34C60E6D14091E2" ID-CLB-Dataset="5408" ID-DOI="10.11646/zootaxa.4987.1.1" ID-GBIF-Dataset="b7f231a2-b766-4f47-914c-c7d39518641f" ID-ISSN="1175-5326" ID-Zenodo-Dep="10086770" ID-ZooBank="33814755-4D9E-4587-BED2-E8B0ED19B371" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623996682541" checkinUser="plazi" docAuthor="Gardini, Giulio" docDate="2021" docId="03B21B45355AFFBB45C845213313FCBD" docLanguage="en" docName="zootaxa.4987.1.1.pdf" docOrigin="Zootaxa 4987 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Chthonius lanai Gardini 2021, n. sp." docType="treatment" docVersion="10" lastPageNumber="98" masterDocId="FF8B633D3504FFDA455F474B3643FFE0" masterDocTitle="The Italian species of the Chthonius ischnocheles group (Arachnida, Pseudoscorpiones, Chthoniidae), with reference to neighbouring countries" masterLastPageNumber="131" masterPageNumber="1" pageNumber="95" updateTime="1699926180482" updateUser="ExternalLinkService">
<mods:mods id="073E60B876070AB450B4A8E76009AB49" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8FC7C7BCA4C3DBA5F7A105F131DDFF4F">
<mods:title id="97A5EEE0BC6CF1D98BCF8023CFCAC4FD">The Italian species of the Chthonius ischnocheles group (Arachnida, Pseudoscorpiones, Chthoniidae), with reference to neighbouring countries</mods:title>
</mods:titleInfo>
<mods:name id="2EF5D826D136CAFF390F4931C9A3655E" type="personal">
<mods:role id="254CC39C8B40D8093846C69F5FA4CEB6">
<mods:roleTerm id="23A567D1555502A942EF7DBDB2DCD5A9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4A9ECEF4322EF91698131439CBABB090">Gardini, Giulio</mods:namePart>
</mods:name>
<mods:typeOfResource id="0B8A2676D445AC5EF6C4604E89193B44">text</mods:typeOfResource>
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<mods:title id="443B24D792FA8E3C8DC3406E89E58480">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D0FC333570D4865A744956A36895E2EA">
<mods:date id="A98707F5319A1A22DC914E9F25D64484">2021</mods:date>
<mods:detail id="DA5239242A46356AF6CE838ACA33CDCE" type="pubDate">
<mods:number id="CD49359933FA882036303EAEB0F7D249">2021-06-17</mods:number>
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<mods:number id="CE6F108A1F917D1D7C774D078193A2F2">4987</mods:number>
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<mods:start id="34F9C0F9675C62DCFF2CAF75FB6F44EA">1</mods:start>
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<mods:classification id="E1A97A9C041B03A8AAD292B75258E613">journal article</mods:classification>
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<mods:identifier id="48E3A6BFE0524D982578C2D7051F4C82" type="DOI">10.11646/zootaxa.4987.1.1</mods:identifier>
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<treatment id="03B21B45355AFFBB45C845213313FCBD" ID-DOI="http://doi.org/10.5281/zenodo.5110863" ID-GBIF-Taxon="183613345" ID-Zenodo-Dep="5110863" LSID="urn:lsid:plazi:treatment:03B21B45355AFFBB45C845213313FCBD" httpUri="http://treatment.plazi.org/id/03B21B45355AFFBB45C845213313FCBD" lastPageId="97" lastPageNumber="98" pageId="94" pageNumber="95">
<subSubSection id="C301F9D8355AFF8445C8452137E6FD65" box="[151,421,618,645]" pageId="94" pageNumber="95" type="nomenclature">
<paragraph id="8BA4AA53355AFF8445C8452137E6FD65" blockId="94.[151,421,618,681]" box="[151,421,618,645]" pageId="94" pageNumber="95">
<heading id="D0EC1D3F355AFF8445C8452137E6FD65" bold="true" box="[151,421,618,645]" fontSize="11" level="2" pageId="94" pageNumber="95" reason="4">
<emphasis id="B96F7641355AFF8445C8452137E6FD65" bold="true" box="[151,421,618,645]" pageId="94" pageNumber="95">
<taxonomicName id="4C1BD1D0355AFF8445C84521371FFD65" authority="Gardini, 2021" authorityName="Gardini" authorityYear="2021" box="[151,348,618,645]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="lanai" status="sp. nov.">
<emphasis id="B96F7641355AFF8445C84521371FFD65" bold="true" box="[151,348,618,645]" italics="true" pageId="94" pageNumber="95">Chthonius lanai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25CCB3A355AFF84443C452037E6FD65" box="[355,421,619,645]" pageId="94" pageNumber="95" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C301F9D8355AFF8445C845C43738FD49" box="[151,379,655,681]" pageId="94" pageNumber="95" type="description">
<paragraph id="8BA4AA53355AFF8445C845C43738FD49" blockId="94.[151,421,618,681]" box="[151,379,655,681]" pageId="94" pageNumber="95">
(
<figureCitation id="1320B6D6355AFF8445C045C4377EFD49" box="[159,317,655,681]" captionStart-0="FIGURES 297–303" captionStart-1="FIGURES 304–308" captionStartId-0="95.[151,264,1879,1903]" captionStartId-1="96.[151,264,1889,1913]" captionTargetBox-0="[165,1424,210,1845]" captionTargetBox-1="[152,1433,181,1858]" captionTargetId-0="figure-32@95.[151,1436,201,1856]" captionTargetId-1="figure-32@96.[151,1436,181,1865]" captionTargetPageId-0="95" captionTargetPageId-1="96" captionText-0="FIGURES 297–303. Chthonius lanai n. sp. 297, female holotype, anterior margin of carapace; 298, male paratype, id.; 299, female holotype, right half of the carapace; 300, female holotype, right chelicera; 301, male paratype, movable finger of right chelicera; 302, male paratype, right chelal hand, dorsal view; 303, female holotype, id." captionText-1="FIGURES 304–308. Chthonius lanai n. sp. 304, female holotype, right chela, antiaxial view; 305, female holotype, detail of fingers at the chelal tip, antiaxial view; 306, female holotype, detail of fingers at level of trichobothria est-it and st-t, antiaxial view; 307, female holotype, detail of fingers at level of trichobothrium sb, antiaxial view; 308, male paratype, right chela, antiaxial view." figureDoi-0="http://doi.org/10.5281/zenodo.10121277" figureDoi-1="http://doi.org/10.5281/zenodo.10121279" httpUri-0="https://zenodo.org/record/10121277/files/figure.png" httpUri-1="https://zenodo.org/record/10121279/files/figure.png" pageId="94" pageNumber="95">Figs 297–308</figureCitation>
,
<figureCitation id="1320B6D6355AFF84441745C43730FD49" box="[328,371,655,681]" captionStart="FIGURES 410–413" captionStartId="127.[151,264,1800,1824]" captionTargetBox="[154,1432,184,1767]" captionTargetId="figure-16@127.[151,1435,181,1777]" captionTargetPageId="127" captionText="FIGURES 410–413. Distribution maps for the Italian species of the Chthonius ischnocheles group. 410, Chthonius caoduroi Callaini, 1987, Chthonius gallettii n. sp., Chthonius inguscioi n. sp., Chthonius marciai n. sp. and Chthonius multidentatus Beier, 1963; 411, Chthonius caprai Gardini, 1977, Chthonius jonicus Beier, 1931 and Chthonius leoi (Callaini, 1988); 412, Chthonius halberti Kew, 1916, Chthonius nicolosii n. sp., Chthonius ressli Beier, 1956 and Chthonius stammeri Beier, 1942; 413, Chthonius horridus Beier, 1934, Chthonius italicus Beier, 1930 and Chthonius lanai n. sp." figureDoi="http://doi.org/10.5281/zenodo.10121317" httpUri="https://zenodo.org/record/10121317/files/figure.png" pageId="94" pageNumber="95">413</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C301F9D8355AFF8445C8459C3763FCF5" pageId="94" pageNumber="95" type="materials_examined">
<paragraph id="8BA4AA53355AFF8445C8459C3763FCF5" blockId="94.[151,1437,726,2013]" pageId="94" pageNumber="95">
<materialsCitation id="3B73A00E355AFF8445C8459C375FFCF5" ID-GBIF-Occurrence="3325145639" collectionCode="CN" country="Italy" county="Cuneo Province" latitude="44.383892" location="Pertus" longLatPrecision="19" longitude="7.383611" municipality="Bernezzo" pageId="94" pageNumber="95" specimenCount="1" stateProvince="Piemonte" typeStatus="holotype">
<emphasis id="B96F7641355AFF8445C8459C377BFD10" bold="true" box="[151,312,726,753]" pageId="94" pageNumber="95">
<typeStatus id="54A014F1355AFF8445C8459C3691FD11" box="[151,210,727,753]" pageId="94" pageNumber="95">Type</typeStatus>
locality
</emphasis>
:
<collectingCountry id="F30CEAC3355AFF844410459C37C0FD11" box="[335,387,727,753]" name="Italy" pageId="94" pageNumber="95">Italy</collectingCountry>
,
<collectingRegion id="49DF64B1355AFF8444C9459C3447FD11" box="[406,516,727,753]" country="Italy" name="Piemonte" pageId="94" pageNumber="95">Piedmont</collectingRegion>
,
<collectingCounty id="62C5D2DF355AFF844747459C3495FD11" box="[536,726,727,753]" pageId="94" pageNumber="95">Cuneo Province</collectingCounty>
,
<collectingMunicipality id="6BC03029355AFF8447B6459C3515FD11" box="[745,854,727,753]" pageId="94" pageNumber="95">Bernezzo</collectingMunicipality>
,
<location id="8EC4FC88355AFF844636459C35ECFD11" LSID="urn:lsid:plazi:treatment:03B21B45355AFFBB45C845213313FCBD:8EC4FC88355AFF844636459C35ECFD11" box="[873,943,727,753]" country="Italy" county="Cuneo Province" latitude="44.383892" longLatPrecision="19" longitude="7.383611" municipality="Bernezzo" name="Pertus" pageId="94" pageNumber="95" stateProvince="Piemonte">Pertus</location>
d’la
<location id="8EC4FC88355AFF8446A6459C321BFD11" LSID="urn:lsid:plazi:treatment:03B21B45355AFFBB45C845213313FCBD:8EC4FC88355AFF8446A6459C321BFD11" box="[1017,1112,727,753]" country="Italy" county="Cuneo Province" latitude="44.383892" longLatPrecision="19" longitude="7.383611" municipality="Bernezzo" name="Kassetta" pageId="94" pageNumber="95" stateProvince="Piemonte">Kassetta</location>
1323
<location id="8EC4FC88355AFF8441F1459C3286FD11" LSID="urn:lsid:plazi:treatment:03B21B45355AFFBB45C845213313FCBD:8EC4FC88355AFF8441F1459C3286FD11" box="[1198,1221,727,753]" country="Italy" county="Cuneo Province" latitude="44.383892" longLatPrecision="19" longitude="7.383611" municipality="Bernezzo" name="Pi" pageId="94" pageNumber="95" stateProvince="Piemonte">Pi</location>
/
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(
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,
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)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C301F9D8355AFF84459844553464FCDA" box="[199,551,798,826]" pageId="94" pageNumber="95" type="distribution">
<paragraph id="8BA4AA53355AFF84459844553464FCDA" blockId="94.[151,1437,726,2013]" box="[199,551,798,826]" pageId="94" pageNumber="95">
<emphasis id="B96F7641355AFF8445984455371AFCD8" bold="true" box="[199,345,798,824]" pageId="94" pageNumber="95">Distribution</emphasis>
.
<collectingCountry id="F30CEAC3355AFF844438445437D9FCD9" box="[359,410,799,825]" name="Italy" pageId="94" pageNumber="95">Italy</collectingCountry>
(
<collectingRegion id="49DF64B1355AFF8444F644543458FCDA" box="[425,539,799,826]" country="Italy" name="Piemonte" pageId="94" pageNumber="95">Piedmont</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C301F9D8355AFF84459844093769FBB9" pageId="94" pageNumber="95" type="diagnosis">
<paragraph id="8BA4AA53355AFF84459844093769FBB9" blockId="94.[151,1437,726,2013]" pageId="94" pageNumber="95">
<emphasis id="B96F7641355AFF844598440937CBFCBF" bold="true" box="[199,392,834,863]" pageId="94" pageNumber="95">
Diagnosis (
<specimenCount id="9D1D61DA355AFF844410440F3727FCBF" box="[335,356,836,863]" pageId="94" pageNumber="95" type="male">♂</specimenCount>
<specimenCount id="9D1D61DA355AFF84443B440F373AFCBF" box="[356,377,836,863]" pageId="94" pageNumber="95" type="female">♀</specimenCount>
).
</emphasis>
An anophthalmic hypogean
<taxonomicName id="4C1BD1D0355AFF8447864408350FFCBD" authorityName="Rayi L. Koch" authorityYear="1873" box="[729,844,835,861]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="94" pageNumber="95" phylum="Arthropoda" rank="genus">
<emphasis id="B96F7641355AFF8447864408350FFCBD" box="[729,844,835,861]" italics="true" pageId="94" pageNumber="95">Chthonius</emphasis>
</taxonomicName>
that differs from other species of the
<taxonomicName id="4C1BD1D0355AFF844052440833DFFCBD" baseAuthorityName="Hermann" baseAuthorityYear="1804" box="[1293,1436,835,861]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="ischnocheles">
<emphasis id="B96F7641355AFF844052440833DFFCBD" box="[1293,1436,835,861]" italics="true" pageId="94" pageNumber="95">ischnocheles</emphasis>
</taxonomicName>
group in the following combination of characters: anterior margin of carapace without or with 1 or 2 preocular microsetae; posterior margin of carapace with 2 macrosetae and 1 lateral microseta on each side; chelicerae with 2 lateral microsetae; tergite I with 2 setae; chela length
<quantity id="4CE307B6355AFF8447AB44E435D8FC29" box="[756,923,943,969]" metricMagnitude="-3" metricUnit="m" metricValue="1.725" metricValueMax="1.74" metricValueMin="1.71" pageId="94" pageNumber="95" unit="mm" value="1.725" valueMax="1.74" valueMin="1.71">1.71–1.74 mm</quantity>
; movable chelal finger length
<quantity id="4CE307B6355AFF8441AE44E433DBFC29" box="[1265,1432,943,969]" metricMagnitude="-3" metricUnit="m" metricValue="1.19" metricValueMax="1.2" metricValueMin="1.18" pageId="94" pageNumber="95" unit="mm" value="1.19" valueMax="1.2" valueMin="1.18">1.18–1.20 mm</quantity>
; fixed chelal finger straight; chelal fingers with reclined, pointed and widely spaced teeth; fixed and movable chelal fingers with 60–64 and 49–52 teeth, respectively; both fixed and movable chelal fingers at level of
<emphasis id="B96F7641355AFF8441A244BC3303FBF0" box="[1277,1344,1015,1040]" italics="true" pageId="94" pageNumber="95">est-ist</emphasis>
and
<emphasis id="B96F7641355AFF84402744BC33DFFBF0" box="[1400,1436,1015,1040]" italics="true" pageId="94" pageNumber="95">st-t</emphasis>
with 5–6 teeth occupying
<quantity id="4CE307B6355AFF8444E543503451FBD5" box="[442,530,1051,1077]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="94" pageNumber="95" unit="mm" value="0.1">0.1 mm</quantity>
, distance between successive apices
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<geoCoordinate id="EE2FCC94355AFF8446EF435035ACFBD5" box="[944,1007,1051,1077]" degrees="0.016" direction="north" orientation="latitude" pageId="94" pageNumber="95" precision="55" value="0.016">0.016</geoCoordinate>
<geoCoordinate id="EE2FCC94355AFF8446B043503278FBD5" box="[1007,1083,1051,1077]" degrees="0.023" direction="west" orientation="longitude" pageId="94" pageNumber="95" precision="55" value="-0.023">–0.023</geoCoordinate>
mm
</quantity>
; ratio of pedipalpal femur/ carapace 1.9.
</paragraph>
</subSubSection>
<subSubSection id="C301F9D8355AFF84459843283229FB25" pageId="94" pageNumber="95" type="materials_examined">
<paragraph id="8BA4AA53355AFF84459843283229FB25" blockId="94.[151,1437,726,2013]" pageId="94" pageNumber="95">
<emphasis id="B96F7641355AFF84459843283736FB9C" bold="true" box="[199,373,1122,1149]" pageId="94" pageNumber="95">Type material.</emphasis>
<materialsCitation id="3B73A00E355AFF84442443283545FB9E" ID-GBIF-Occurrence="3325145760" box="[379,774,1123,1150]" country="Italy" location="Piedmont" pageId="94" pageNumber="95" specimenCount="1" specimenCount-female="1" stateProvince="Piemonte" typeStatus="holotype">
<collectingCountry id="F30CEAC3355AFF84442443283788FB9D" box="[379,459,1123,1149]" name="Italy" pageId="94" pageNumber="95">ITALY</collectingCountry>
—
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<collectingRegion id="49DF64B1355AFF8444B943283412FB9D" box="[486,593,1123,1149]" country="Italy" name="Piemonte" pageId="94" pageNumber="95">Piedmont</collectingRegion>
</emphasis>
:
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(
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)
</materialsCitation>
<materialsCitation id="3B73A00E355AFF84465343283594FB41" ID-GBIF-Occurrence="3325145901" collectingDate="2020-02-23" collectionCode="CN" collectorName="E. Lana" country="Italy" county="Cuneo Prov." elevation="1230" location="Vallone" municipality="Bernezzo" pageId="94" pageNumber="95" specimenCount="2" specimenCount-female="2" stateProvince="Piedmont" typeStatus="paratype">
<specimenCount id="9D1D61DA355AFF84465343283575FB9D" box="[780,822,1123,1149]" pageId="94" pageNumber="95" type="female">2 ♀</specimenCount>
(
<typeStatus id="54A014F1355AFF84461B432835F6FB9D" box="[836,949,1123,1149]" pageId="94" pageNumber="95" type="paratype">paratypes</typeStatus>
),
<collectingCounty id="62C5D2DF355AFF84469743283215FB9D" box="[968,1110,1123,1149]" pageId="94" pageNumber="95">Cuneo Prov.</collectingCounty>
,
<collectingMunicipality id="6BC03029355AFF84413F4328328EFB9D" box="[1120,1229,1123,1149]" pageId="94" pageNumber="95">Bernezzo</collectingMunicipality>
,
<location id="8EC4FC88355AFF8441884328336EFB9D" LSID="urn:lsid:plazi:treatment:03B21B45355AFFBB45C845213313FCBD:8EC4FC88355AFF8441884328336EFB9D" box="[1239,1325,1123,1149]" country="Italy" county="Cuneo Prov." municipality="Bernezzo" name="Vallone" pageId="94" pageNumber="95" stateProvince="Piedmont">Vallone</location>
del Cugino,
<location id="8EC4FC88355AFF84459F43CC3745FB41" LSID="urn:lsid:plazi:treatment:03B21B45355AFFBB45C845213313FCBD:8EC4FC88355AFF84459F43CC3745FB41" box="[192,262,1159,1185]" country="Italy" county="Cuneo Prov." municipality="Bernezzo" name="Pertus" pageId="94" pageNumber="95" stateProvince="Piedmont">Pertus</location>
d’la
<location id="8EC4FC88355AFF84446243CC37DFFB41" LSID="urn:lsid:plazi:treatment:03B21B45355AFFBB45C845213313FCBD:8EC4FC88355AFF84446243CC37DFFB41" box="[317,412,1159,1185]" country="Italy" county="Cuneo Prov." municipality="Bernezzo" name="Kassetta" pageId="94" pageNumber="95" stateProvince="Piedmont">Kassetta</location>
1323
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/
<collectionCode id="ED0A3296355AFF84475E43CC346AFB41" box="[513,553,1159,1185]" country="United Kingdom" name="Wellcome Collection of Bacteria, Burroughs Wellcome Research Laboratories" pageId="94" pageNumber="95">CN</collectionCode>
,
<elevation id="00364D60355AFF84476C43CC3481FB41" box="[563,706,1159,1185]" metricMagnitude="3" metricUnit="m" metricValue="1.23" pageId="94" pageNumber="95" unit="m" value="1230.0">
<quantity id="4CE307B6355AFF84476C43CC34C4FB41" box="[563,647,1159,1185]" metricMagnitude="3" metricUnit="m" metricValue="1.23" pageId="94" pageNumber="95" unit="m" value="1230.0">1230 m</quantity>
a.s.l.
</elevation>
,
<date id="FFA58C93355AFF84479243CC3506FB41" box="[717,837,1159,1185]" pageId="94" pageNumber="95" value="2020-02-23">
<collectingDate id="EFE1757B355AFF84479243CC3506FB41" box="[717,837,1159,1185]" pageId="94" pageNumber="95" value="2020-02-23">23.II.2020</collectingDate>
</date>
,
<collectorName id="26EECF85355AFF84461043CC35E6FB41" box="[847,933,1159,1185]" pageId="94" pageNumber="95">E. Lana</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B73A00E355AFF8446BD43CC3225FB25" ID-GBIF-Occurrence="3325145704" collectingDate="2013-06-15" collectionCode="MHNG" collectorName="G. Gardini" country="Italy" pageId="94" pageNumber="95" specimenCount="1" specimenCount-male="1" stateProvince="Piedmont" typeStatus="paratype">
<specimenCount id="9D1D61DA355AFF8446BD43CC3249FB41" box="[994,1034,1159,1185]" pageId="94" pageNumber="95" type="male">1 ♂</specimenCount>
(
<typeStatus id="54A014F1355AFF84414743CC323DFB41" box="[1048,1150,1159,1185]" pageId="94" pageNumber="95" type="paratype">paratype</typeStatus>
), id.,
<date id="FFA58C93355AFF8441E543CD337FFB41" box="[1210,1340,1158,1185]" pageId="94" pageNumber="95" value="2013-06-15">
<collectingDate id="EFE1757B355AFF8441E543CD337FFB41" box="[1210,1340,1158,1185]" pageId="94" pageNumber="95" value="2013-06-15">15.VI.2013</collectingDate>
</date>
,
<collectorName id="26EECF85355AFF84401943CC33DFFB41" box="[1350,1436,1159,1185]" pageId="94" pageNumber="95">E. Lana</collectorName>
&amp;
<collectorName id="26EECF85355AFF8445EB43E03764FB25" box="[180,295,1195,1221]" pageId="94" pageNumber="95">M. Chesta</collectorName>
leg. (
<typeStatus id="54A014F1355AFF84443943E03789FB25" box="[358,458,1195,1221]" pageId="94" pageNumber="95" type="holotype">holotype</typeStatus>
deposited in
<collectionCode id="ED0A3296355AFF84473F43E034FBFB25" box="[608,696,1195,1221]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="94" pageNumber="95" type="Museum">MHNG</collectionCode>
,
<typeStatus id="54A014F1355AFF84479C43E0356DFB25" box="[707,814,1195,1221]" pageId="94" pageNumber="95" type="paratype">paratypes</typeStatus>
in
<collectorName id="26EECF85355AFF84460D43E03588FB25" box="[850,971,1195,1221]" pageId="94" pageNumber="95">G. Gardini</collectorName>
coll.,
<collectorName id="26EECF85355AFF84414F43E0321DFB25" box="[1040,1118,1195,1221]" pageId="94" pageNumber="95">Genoa</collectorName>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C301F9D8355AFF8445A843843505FAD1" pageId="94" pageNumber="95" type="etymology">
<paragraph id="8BA4AA53355AFF8445A843843505FAD1" blockId="94.[151,1437,726,2013]" pageId="94" pageNumber="95">
<emphasis id="B96F7641355AFF8445A843843790FB08" box="[247,467,1231,1257]" italics="true" pageId="94" pageNumber="95">
<emphasis id="B96F7641355AFF8445A84384378FFB09" bold="true" box="[247,460,1231,1257]" italics="true" pageId="94" pageNumber="95">Derivatio nominis</emphasis>
.
</emphasis>
Named after the naturalist and biospeleologist Enrico Lana (Biologia Sotterranea
<collectingRegion id="49DF64B1355AFF8445C843B83747FAED" box="[151,260,1267,1293]" country="Italy" name="Piemonte" pageId="94" pageNumber="95">Piemonte</collectingRegion>
, Gruppo di Ricerca and Struttura Operativa Bossea, Club Alpino Italiano), in recognition of his contribution to the knowledge of the subterranean fauna of
<collectingRegion id="49DF64B1355AFF84478C425C3501FAD1" box="[723,834,1303,1329]" country="Italy" name="Piemonte" pageId="94" pageNumber="95">Piedmont</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C301F9D8355AFFBB4598427132F0FD65" lastPageId="97" lastPageNumber="98" pageId="94" pageNumber="95" type="description">
<paragraph id="8BA4AA53355AFFBB459842713477FE15" blockId="94.[151,1437,726,2013]" lastBlockId="97.[151,1437,150,861]" lastPageId="97" lastPageNumber="98" pageId="94" pageNumber="95">
<emphasis id="B96F7641355AFF84459842713449FAB7" bold="true" box="[199,522,1338,1367]" pageId="94" pageNumber="95">
Description of adults (
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<specimenCount id="9D1D61DA355AFF8444B9427737B8FAB7" box="[486,507,1340,1367]" pageId="94" pageNumber="95" type="female">♀</specimenCount>
).
</emphasis>
Integument slightly pigmented, chelicerae and chelal fingers reddish brown pale; hispid granulation on the lateral surfaces of carapace, on cheliceral palm and on the base of fixed chelal finger. Carapace (
<figureCitation id="1320B6D6355AFF84445142C8372DFA7D" box="[270,366,1411,1437]" captionStart="FIGURES 297–303" captionStartId="95.[151,264,1879,1903]" captionTargetBox="[165,1424,210,1845]" captionTargetId="figure-32@95.[151,1436,201,1856]" captionTargetPageId="95" captionText="FIGURES 297–303. Chthonius lanai n. sp. 297, female holotype, anterior margin of carapace; 298, male paratype, id.; 299, female holotype, right half of the carapace; 300, female holotype, right chelicera; 301, male paratype, movable finger of right chelicera; 302, male paratype, right chelal hand, dorsal view; 303, female holotype, id." figureDoi="http://doi.org/10.5281/zenodo.10121277" httpUri="https://zenodo.org/record/10121277/files/figure.png" pageId="94" pageNumber="95">Fig. 299</figureCitation>
) 0.90–0.95 times longer than broad, trapezoidal, constricted posteriorly; anterior margin strongly denticulate between the median macrosetae, without a prominent epistome (
<figureCitation id="1320B6D6355AFF84417B42EC3284FA21" box="[1060,1223,1447,1473]" captionStart="FIGURES 297–303" captionStartId="95.[151,264,1879,1903]" captionTargetBox="[165,1424,210,1845]" captionTargetId="figure-32@95.[151,1436,201,1856]" captionTargetPageId="95" captionText="FIGURES 297–303. Chthonius lanai n. sp. 297, female holotype, anterior margin of carapace; 298, male paratype, id.; 299, female holotype, right half of the carapace; 300, female holotype, right chelicera; 301, male paratype, movable finger of right chelicera; 302, male paratype, right chelal hand, dorsal view; 303, female holotype, id." figureDoi="http://doi.org/10.5281/zenodo.10121277" httpUri="https://zenodo.org/record/10121277/files/figure.png" pageId="94" pageNumber="95">Figs 297–298</figureCitation>
); ocular area as in fig. 299, anterior eyes reduced to a hardly noticeable cuticular pale area, posterior eyes absent; probable standard chaetotaxy m
<quantity id="4CE307B6355AFF84446D42A43719F9EA" box="[306,346,1519,1546]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="94" pageNumber="95" unit="m" value="4.0">4m</quantity>
:6:4:2:m
<quantity id="4CE307B6355AFF8444F142A43795F9EA" box="[430,470,1519,1546]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="94" pageNumber="95" unit="m" value="2.0">2m</quantity>
(22), anterior row also m4 or mm4, or without preocular microsetae (
<specimenCount id="9D1D61DA355AFF8441B642A432BFF9E9" box="[1257,1276,1519,1545]" pageId="94" pageNumber="95" type="female">♀</specimenCount>
<typeStatus id="54A014F1355AFF84405B42A43329F9E9" box="[1284,1386,1519,1545]" pageId="94" pageNumber="95" type="holotype">holotype</typeStatus>
: fig. 299), length of anteromedian macrosetae
<quantity id="4CE307B6355AFF84473441583552F9CD" box="[619,785,1555,1581]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.7999999999999998" pageId="94" pageNumber="95" unit="mm" value="0.19" valueMax="0.2" valueMin="0.18">0.18–0.20 mm</quantity>
. Chaetotaxy of tergites 2:3:4:4:6:6:6:6:1T2T1:4:1T2T1:0, tergites II 2 (
<specimenCount id="9D1D61DA355AFF844477417C370EF9B1" box="[296,333,1591,1617]" pageId="94" pageNumber="95" type="female">1 ♀</specimenCount>
), IV 3 (
<specimenCount id="9D1D61DA355AFF8444F7417C378DF9B1" box="[424,462,1591,1617]" pageId="94" pageNumber="95" type="female">1 ♀</specimenCount>
), V 5 (
<specimenCount id="9D1D61DA355AFF844742417C3405F9B1" box="[541,582,1591,1617]" pageId="94" pageNumber="95" type="female">2 ♀</specimenCount>
). Chaetotaxy of sternites II–X: 10–12:(3)8(3):(2)7–8(2):7–8:6:6:6:6:2T1T2; genital opening of male flanked by 9 setae on each side. Chelicerae (
<figureCitation id="1320B6D6355AFF8446EC41103255F995" box="[947,1046,1627,1653]" captionStart="FIGURES 297–303" captionStartId="95.[151,264,1879,1903]" captionTargetBox="[165,1424,210,1845]" captionTargetId="figure-32@95.[151,1436,201,1856]" captionTargetPageId="95" captionText="FIGURES 297–303. Chthonius lanai n. sp. 297, female holotype, anterior margin of carapace; 298, male paratype, id.; 299, female holotype, right half of the carapace; 300, female holotype, right chelicera; 301, male paratype, movable finger of right chelicera; 302, male paratype, right chelal hand, dorsal view; 303, female holotype, id." figureDoi="http://doi.org/10.5281/zenodo.10121277" httpUri="https://zenodo.org/record/10121277/files/figure.png" pageId="94" pageNumber="95">Fig. 300</figureCitation>
) 2.5 times as long as broad, palm with 6 setae and 2 lateral microsetae; fixed finger with 2 distal large teeth followed by 8–10 small teeth proximally increasingly reduced and 4–7 microtubercles; movable finger with an isolated subapical tooth (
<emphasis id="B96F7641355AFF84419D41E8329BF95D" box="[1218,1240,1699,1725]" italics="true" pageId="94" pageNumber="95">di</emphasis>
) at level with the spinneret (
<specimenCount id="9D1D61DA355AFF844451418C3767F901" box="[270,292,1735,1761]" pageId="94" pageNumber="95" type="male">♂</specimenCount>
) or more proximal with respect to the spinneret (
<specimenCount id="9D1D61DA355AFF844610418C3521F901" box="[847,866,1735,1761]" pageId="94" pageNumber="95" type="female">♀</specimenCount>
), followed by a large tooth and 9–10 teeth that are proximally reduced in size;
<emphasis id="B96F7641355AFF84448941A037ACF8E4" box="[470,495,1771,1796]" italics="true" pageId="94" pageNumber="95">gs</emphasis>
ratio 0.53–0.55; spinneret small, right angle-shaped in male, prominent and hemispherical in females (
<figureCitation id="1320B6D6355AFF8444D940443466F8C9" box="[390,549,1807,1833]" captionStart="FIGURES 297–303" captionStartId="95.[151,264,1879,1903]" captionTargetBox="[165,1424,210,1845]" captionTargetId="figure-32@95.[151,1436,201,1856]" captionTargetPageId="95" captionText="FIGURES 297–303. Chthonius lanai n. sp. 297, female holotype, anterior margin of carapace; 298, male paratype, id.; 299, female holotype, right half of the carapace; 300, female holotype, right chelicera; 301, male paratype, movable finger of right chelicera; 302, male paratype, right chelal hand, dorsal view; 303, female holotype, id." figureDoi="http://doi.org/10.5281/zenodo.10121277" httpUri="https://zenodo.org/record/10121277/files/figure.png" pageId="94" pageNumber="95">Figs 300–301</figureCitation>
); rallum with 11 blades; serrula interior with 14, exterior with 18 blades. Coxal setae: pedipalp 5 (including 2 on manducatory process), I (3)4 + 3(4) marginal microsetae, II (4)5, III 5(6), IV 6; coxa II with 16–17 coxal spines, coxa III with 6–8 coxal spines; intercoxal tubercle bisetose. Pedipalp: femur 8.9 (
<specimenCount id="9D1D61DA355AFF8445FF403036F5F875" box="[160,182,1915,1941]" pageId="94" pageNumber="95" type="male">♂</specimenCount>
) or 7.55 (
<specimenCount id="9D1D61DA355AFF8444794030377BF875" box="[294,312,1915,1941]" pageId="94" pageNumber="95" type="female">♀</specimenCount>
) times as long as broad, femoral chaetotaxy 3:5:3:(5)6:1; chela (
<figureCitation id="1320B6D6355AFF844154403032E4F875" box="[1035,1191,1915,1941]" captionStart="FIGURES 304–308" captionStartId="96.[151,264,1889,1913]" captionTargetBox="[152,1433,181,1858]" captionTargetId="figure-32@96.[151,1436,181,1865]" captionTargetPageId="96" captionText="FIGURES 304–308. Chthonius lanai n. sp. 304, female holotype, right chela, antiaxial view; 305, female holotype, detail of fingers at the chelal tip, antiaxial view; 306, female holotype, detail of fingers at level of trichobothria est-it and st-t, antiaxial view; 307, female holotype, detail of fingers at level of trichobothrium sb, antiaxial view; 308, male paratype, right chela, antiaxial view." figureDoi="http://doi.org/10.5281/zenodo.10121279" httpUri="https://zenodo.org/record/10121279/files/figure.png" pageId="94" pageNumber="95">Figs 304, 308</figureCitation>
) 7.25 (
<specimenCount id="9D1D61DA355AFF8441A84030334EF875" box="[1271,1293,1915,1941]" pageId="94" pageNumber="95" type="male">♂</specimenCount>
) or 7.1–7.25 (
<specimenCount id="9D1D61DA355AFF8445FD40D436F7F859" box="[162,180,1951,1977]" pageId="94" pageNumber="95" type="female">♀</specimenCount>
) times as long as deep; hand of chela 2.3 (
<specimenCount id="9D1D61DA355AFF8447F440D43482F859" box="[683,705,1951,1977]" pageId="94" pageNumber="95" type="male">♂</specimenCount>
) or 2.25–2.3 (
<specimenCount id="9D1D61DA355AFF84463640D43538F859" box="[873,891,1951,1977]" pageId="94" pageNumber="95" type="female">♀</specimenCount>
) times as long as deep, with parallel outline in dorsal view (
<figureCitation id="1320B6D6355AFF84447740883784F83D" box="[296,455,1987,2013]" captionStart="FIGURES 297–303" captionStartId="95.[151,264,1879,1903]" captionTargetBox="[165,1424,210,1845]" captionTargetId="figure-32@95.[151,1436,201,1856]" captionTargetPageId="95" captionText="FIGURES 297–303. Chthonius lanai n. sp. 297, female holotype, anterior margin of carapace; 298, male paratype, id.; 299, female holotype, right half of the carapace; 300, female holotype, right chelicera; 301, male paratype, movable finger of right chelicera; 302, male paratype, right chelal hand, dorsal view; 303, female holotype, id." figureDoi="http://doi.org/10.5281/zenodo.10121277" httpUri="https://zenodo.org/record/10121277/files/figure.png" pageId="94" pageNumber="95">Figs 302–303</figureCitation>
); fixed chelal finger straight, with 60–64 teeth inclined backwards: the 9–12 distal teeth pointed, thin and contiguous, followed by 44–46 reclined, pointed teeth that are increasingly widely spaced towards finger base and 8–10 proximal teeth that are low, rounded, and reduced in size (
<figureCitation id="1320B6D63565FFBB415247DC32EEFF51" box="[1037,1197,150,177]" captionStart="FIGURES 304–308" captionStartId="96.[151,264,1889,1913]" captionTargetBox="[152,1433,181,1858]" captionTargetId="figure-32@96.[151,1436,181,1865]" captionTargetPageId="96" captionText="FIGURES 304–308. Chthonius lanai n. sp. 304, female holotype, right chela, antiaxial view; 305, female holotype, detail of fingers at the chelal tip, antiaxial view; 306, female holotype, detail of fingers at level of trichobothria est-it and st-t, antiaxial view; 307, female holotype, detail of fingers at level of trichobothrium sb, antiaxial view; 308, male paratype, right chela, antiaxial view." figureDoi="http://doi.org/10.5281/zenodo.10121279" httpUri="https://zenodo.org/record/10121279/files/figure.png" pageId="97" pageNumber="98">Figs 305–307</figureCitation>
); all teeth with dental canal; base of fixed chelal finger with 4–8 microtubercles; fixed finger at level of
<emphasis id="B96F76413565FFBB416647F03232FF34" box="[1081,1137,187,212]" italics="true" pageId="97" pageNumber="98">est-it</emphasis>
with (5)6 teeth occupying
<quantity id="4CE307B63565FFBB45C8479436ADFF19" box="[151,238,223,249]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="97" pageNumber="98" unit="mm" value="0.1">0.1 mm</quantity>
, distance between successive apices 0.018 (
<specimenCount id="9D1D61DA3565FFBB478B479434A9FF19" box="[724,746,223,249]" pageId="97" pageNumber="98" type="male">♂</specimenCount>
) or
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(
<specimenCount id="9D1D61DA3565FFBB46EC47943586FF19" box="[947,965,223,249]" pageId="97" pageNumber="98" type="female">♀</specimenCount>
) mm; tip of fixed chelal finger with apical sensilla
<emphasis id="B96F76413565FFBB45AC4648374AFEFD" box="[243,265,259,285]" italics="true" pageId="97" pageNumber="98">af</emphasis>
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<emphasis id="B96F76413565FFBB444F465E375DFEC3" box="[272,286,277,291]" italics="true" pageId="97" pageNumber="98">1-</emphasis>
2
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, distal paraxial seta gradually curved and thin; movable chelal finger with 49–52 teeth that are large, squat, reclined apically, increasingly reduced towards finger base in the proximal half of the finger, and reaching back nearby
<emphasis id="B96F76413565FFBB44744600377AFE85" box="[299,313,331,357]" italics="true" pageId="97" pageNumber="98">b</emphasis>
(
<figureCitation id="1320B6D63565FFBB4415460037AFFE85" box="[330,492,330,357]" captionStart="FIGURES 304–308" captionStartId="96.[151,264,1889,1913]" captionTargetBox="[152,1433,181,1858]" captionTargetId="figure-32@96.[151,1436,181,1865]" captionTargetPageId="96" captionText="FIGURES 304–308. Chthonius lanai n. sp. 304, female holotype, right chela, antiaxial view; 305, female holotype, detail of fingers at the chelal tip, antiaxial view; 306, female holotype, detail of fingers at level of trichobothria est-it and st-t, antiaxial view; 307, female holotype, detail of fingers at level of trichobothrium sb, antiaxial view; 308, male paratype, right chela, antiaxial view." figureDoi="http://doi.org/10.5281/zenodo.10121279" httpUri="https://zenodo.org/record/10121279/files/figure.png" pageId="97" pageNumber="98">Figs 305–307</figureCitation>
); movable finger at level of
<emphasis id="B96F76413565FFBB466A4600351AFE84" box="[821,857,331,356]" italics="true" pageId="97" pageNumber="98">st-t</emphasis>
with 5 teeth occupying
<quantity id="4CE307B63565FFBB412D4600328FFE85" box="[1138,1228,331,357]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="97" pageNumber="98" unit="mm" value="0.1">0.1 mm</quantity>
, distance between successive apices
<geoCoordinate id="EE2FCC943565FFBB443C462437E1FE69" box="[355,418,367,393]" degrees="0.020" direction="north" orientation="latitude" pageId="97" pageNumber="98" precision="55" value="0.02">0.020</geoCoordinate>
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(
<specimenCount id="9D1D61DA3565FFBB475E46243459FE69" box="[513,538,367,393]" pageId="97" pageNumber="98" type="male">♂</specimenCount>
<specimenCount id="9D1D61DA3565FFBB474546243468FE69" box="[538,555,367,393]" pageId="97" pageNumber="98" type="female">♀</specimenCount>
); coupled sensilla
<emphasis id="B96F76413565FFBB47A04624355AFE68" box="[767,793,367,392]" italics="true" pageId="97" pageNumber="98">pc</emphasis>
at level of
<emphasis id="B96F76413565FFBB46C5462435F0FE69" box="[922,947,367,393]" italics="true" pageId="97" pageNumber="98">sb</emphasis>
or just distad
<emphasis id="B96F76413565FFBB410B4624322EFE69" box="[1108,1133,367,393]" italics="true" pageId="97" pageNumber="98">sb</emphasis>
; tip of movable chelal finger with apical sensilla
<emphasis id="B96F76413565FFBB44FD46D83787FE4C" box="[418,452,403,428]" italics="true" pageId="97" pageNumber="98">am</emphasis>
<subScript id="179FA8163565FFBB449346EE37A1FE53" attach="right" box="[460,482,421,435]" fontSize="6" pageId="97" pageNumber="98">
<emphasis id="B96F76413565FFBB449346EE379AFE53" box="[460,473,421,435]" italics="true" pageId="97" pageNumber="98">1-</emphasis>
2
</subScript>
; trichobothria as in figs 304, 308; ratio
<emphasis id="B96F76413565FFBB46FB46D83252FE4D" box="[932,1041,403,429]" italics="true" pageId="97" pageNumber="98">sb -st/sb -b</emphasis>
= 1.85 (
<specimenCount id="9D1D61DA3565FFBB412D46D832CBFE4D" box="[1138,1160,403,429]" pageId="97" pageNumber="98" type="male">♂</specimenCount>
) or 1.5–1.7 (
<specimenCount id="9D1D61DA3565FFBB404146D83372FE4D" box="[1310,1329,403,429]" pageId="97" pageNumber="98" type="female">♀</specimenCount>
); ratio of movable finger/hand of chela 2.15–2.2 (
<specimenCount id="9D1D61DA3565FFBB470246FC3435FE31" box="[605,630,439,465]" pageId="97" pageNumber="98" type="male">♂</specimenCount>
<specimenCount id="9D1D61DA3565FFBB472A46FC34C5FE31" box="[629,646,439,465]" pageId="97" pageNumber="98" type="female">♀</specimenCount>
); ratio of pedipalpal femur/movable finger 1.2 (
<specimenCount id="9D1D61DA3565FFBB41FB46FC32F9FE31" box="[1188,1210,439,465]" pageId="97" pageNumber="98" type="male">♂</specimenCount>
) or 1.0 (
<specimenCount id="9D1D61DA3565FFBB404046FC3371FE31" box="[1311,1330,439,465]" pageId="97" pageNumber="98" type="female">♀</specimenCount>
); ratio of pedipalpal femur/carapace 1.9 (
<specimenCount id="9D1D61DA3565FFBB44A446903457FE15" box="[507,532,475,501]" pageId="97" pageNumber="98" type="male">♂</specimenCount>
<specimenCount id="9D1D61DA3565FFBB474B46903466FE15" box="[532,549,475,501]" pageId="97" pageNumber="98" type="female">♀</specimenCount>
).
</paragraph>
<caption id="DF64FADB355BFF8545C8401C3245F857" ID-DOI="http://doi.org/10.5281/zenodo.10121277" ID-Zenodo-Dep="10121277" httpUri="https://zenodo.org/record/10121277/files/figure.png" pageId="95" pageNumber="96" startId="95.[151,264,1879,1903]" targetBox="[165,1424,210,1845]" targetPageId="95">
<paragraph id="8BA4AA53355BFF8545C8401C3245F857" blockId="95.[151,1436,1879,1976]" pageId="95" pageNumber="96">
<emphasis id="B96F7641355BFF8545C8401C372DF88F" bold="true" box="[151,366,1879,1903]" pageId="95" pageNumber="96">FIGURES 297–303.</emphasis>
<taxonomicName id="4C1BD1D0355BFF854429401C3459F88F" authority="Gardini, 2021" authorityName="Gardini" authorityYear="2021" box="[374,538,1879,1903]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="95" pageNumber="96" phylum="Arthropoda" rank="species" species="lanai" status="sp. nov.">
<emphasis id="B96F7641355BFF854429401C3459F88F" box="[374,538,1879,1903]" italics="true" pageId="95" pageNumber="96">Chthonius lanai</emphasis>
</taxonomicName>
<emphasis id="B96F7641355BFF85477D401C341FF88F" bold="true" box="[546,604,1879,1903]" pageId="95" pageNumber="96">
<taxonomicNameLabel id="A25CCB3A355BFF85477D401C341FF88F" box="[546,604,1879,1903]" pageId="95" pageNumber="96" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
297, female holotype, anterior margin of carapace; 298, male paratype, id.; 299, female holotype, right half of the carapace; 300, female holotype, right chelicera; 301, male paratype, movable finger of right chelicera; 302, male paratype, right chelal hand, dorsal view; 303, female holotype, id.
</paragraph>
</caption>
<caption id="DF64FADB3564FFBA45C8402A3757F805" ID-DOI="http://doi.org/10.5281/zenodo.10121279" ID-Zenodo-Dep="10121279" httpUri="https://zenodo.org/record/10121279/files/figure.png" pageId="96" pageNumber="97" startId="96.[151,264,1889,1913]" targetBox="[152,1433,181,1858]" targetPageId="96">
<paragraph id="8BA4AA533564FFBA45C8402A3757F805" blockId="96.[151,1436,1889,2021]" pageId="96" pageNumber="97">
<emphasis id="B96F76413564FFBA45C8402A372DF899" bold="true" box="[151,366,1889,1913]" pageId="96" pageNumber="97">FIGURES 304–308.</emphasis>
<taxonomicName id="4C1BD1D03564FFBA4429402A345AF899" authority="Gardini, 2021" authorityName="Gardini" authorityYear="2021" box="[374,537,1889,1913]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="lanai" status="sp. nov.">
<emphasis id="B96F76413564FFBA4429402A345AF899" box="[374,537,1889,1913]" italics="true" pageId="96" pageNumber="97">Chthonius lanai</emphasis>
</taxonomicName>
<emphasis id="B96F76413564FFBA477E402A3418F899" bold="true" box="[545,603,1889,1913]" pageId="96" pageNumber="97">
<taxonomicNameLabel id="A25CCB3A3564FFBA477E402A3418F899" box="[545,603,1889,1913]" pageId="96" pageNumber="97" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
304, female holotype, right chela, antiaxial view; 305, female holotype, detail of fingers at the chelal tip, antiaxial view; 306, female holotype, detail of fingers at level of trichobothria
<emphasis id="B96F76413564FFBA41ED40CD32A6F87D" box="[1202,1253,1926,1949]" italics="true" pageId="96" pageNumber="97">est-it</emphasis>
and
<emphasis id="B96F76413564FFBA404840CD3374F87D" box="[1303,1335,1926,1949]" italics="true" pageId="96" pageNumber="97">st-t</emphasis>
, antiaxial view; 307, female holotype, detail of fingers at level of trichobothrium
<emphasis id="B96F76413564FFBA462E40E235CBF821" box="[881,904,1961,1985]" italics="true" pageId="96" pageNumber="97">sb</emphasis>
, antiaxial view; 308, male paratype, right chela, antiaxial view.
</paragraph>
</caption>
<paragraph id="8BA4AA533565FFBB459846B432F0FD65" blockId="97.[151,1437,150,861]" pageId="97" pageNumber="98">
Measurements (in mm). Body length 2.15–2.0 (
<specimenCount id="9D1D61DA3565FFBB47AD46B43548FDF9" box="[754,779,511,537]" pageId="97" pageNumber="98" type="male">♂</specimenCount>
<specimenCount id="9D1D61DA3565FFBB465446B4355FFDF9" box="[779,796,511,537]" pageId="97" pageNumber="98" type="female">♀</specimenCount>
). Carapace 0.62 × 0.68 (0.66 anteriorly) (
<specimenCount id="9D1D61DA3565FFBB405846B4335EFDF9" box="[1287,1309,511,537]" pageId="97" pageNumber="98" type="male">♂</specimenCount>
) or 0.64 × 0.66–0.67 (0.63–0.64 anteriorly) (
<specimenCount id="9D1D61DA3565FFBB474B45683464FDDD" box="[532,551,547,573]" pageId="97" pageNumber="98" type="female">♀</specimenCount>
). Chelicerae 0.74 × 0.29 (
<specimenCount id="9D1D61DA3565FFBB46184568351EFDDD" box="[839,861,547,573]" pageId="97" pageNumber="98" type="male">♂</specimenCount>
) or 0.74 × 0.29 (
<specimenCount id="9D1D61DA3565FFBB414745683268FDDD" box="[1048,1067,547,573]" pageId="97" pageNumber="98" type="female">♀</specimenCount>
); movable finger length 0.37 (
<specimenCount id="9D1D61DA3565FFBB4021456833D7FDDD" box="[1406,1428,547,573]" pageId="97" pageNumber="98" type="male">♂</specimenCount>
) or 0.37–0.38 (
<specimenCount id="9D1D61DA3565FFBB4466450C370FFD81" box="[313,332,583,609]" pageId="97" pageNumber="98" type="female">♀</specimenCount>
). Pedipalp: femur 1.47 × 0.165 (
<specimenCount id="9D1D61DA3565FFBB47E0450C3496FD81" box="[703,725,583,609]" pageId="97" pageNumber="98" type="male">♂</specimenCount>
) or 1.21 × 0.16 (
<specimenCount id="9D1D61DA3565FFBB46C8450C35E9FD81" box="[919,938,583,609]" pageId="97" pageNumber="98" type="female">♀</specimenCount>
); chela 1.74 × 0.24 (
<specimenCount id="9D1D61DA3565FFBB41C8450C32EEFD81" box="[1175,1197,583,609]" pageId="97" pageNumber="98" type="male">♂</specimenCount>
) or 1.71–1.74 × 0.24 (
<specimenCount id="9D1D61DA3565FFBB45FD452036F6FD65" box="[162,181,619,645]" pageId="97" pageNumber="98" type="female">♀</specimenCount>
); hand length 0.56 (
<specimenCount id="9D1D61DA3565FFBB44C6452037ECFD65" box="[409,431,619,645]" pageId="97" pageNumber="98" type="male">♂</specimenCount>
) or 0.54–0.55 (
<specimenCount id="9D1D61DA3565FFBB470045203431FD65" box="[607,626,619,645]" pageId="97" pageNumber="98" type="female">♀</specimenCount>
); movable finger length 1.20 (
<specimenCount id="9D1D61DA3565FFBB4695452035A3FD65" box="[970,992,619,645]" pageId="97" pageNumber="98" type="male">♂</specimenCount>
) or 1.18–1.20 (
<specimenCount id="9D1D61DA3565FFBB41CF452032E0FD65" box="[1168,1187,619,645]" pageId="97" pageNumber="98" type="female">♀</specimenCount>
).
</paragraph>
</subSubSection>
<subSubSection id="C301F9D83565FFBB459845C43313FCBD" pageId="97" pageNumber="98" type="discussion">
<paragraph id="8BA4AA533565FFBB459845C43555FD2D" blockId="97.[151,1437,150,861]" pageId="97" pageNumber="98">
<emphasis id="B96F76413565FFBB459845C43778FD49" bold="true" box="[199,315,655,681]" pageId="97" pageNumber="98">Remarks.</emphasis>
Among the species of the
<taxonomicName id="4C1BD1D03565FFBB473545C43537FD49" baseAuthorityName="Hermann" baseAuthorityYear="1804" box="[618,884,655,681]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="ischnocheles">
<emphasis id="B96F76413565FFBB473545C43537FD49" box="[618,884,655,681]" italics="true" pageId="97" pageNumber="98">Chthonius ischnocheles</emphasis>
</taxonomicName>
group,
<taxonomicName id="4C1BD1D03565FFBB469345C4326BFD49" authorityName="Gardini" authorityYear="2021" box="[972,1064,655,681]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="lanai">
<emphasis id="B96F76413565FFBB469345C4326BFD49" box="[972,1064,655,681]" italics="true" pageId="97" pageNumber="98">C. lanai</emphasis>
</taxonomicName>
is morphologically very close to the subterranean
<taxonomicName id="4C1BD1D03565FFBB440945F8378AFD2D" authorityName="Beier" authorityYear="1930" box="[342,457,691,717]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="italicus">
<emphasis id="B96F76413565FFBB440945F8378AFD2D" box="[342,457,691,717]" italics="true" pageId="97" pageNumber="98">C. italicus</emphasis>
</taxonomicName>
from the south-western Alps.
</paragraph>
<paragraph id="8BA4AA533565FFBB4598459C3313FCBD" blockId="97.[151,1437,150,861]" pageId="97" pageNumber="98">
<taxonomicName id="4C1BD1D03565FFBB4598459C373EFD11" authority="Gardini, 2021" authorityName="Gardini" authorityYear="2021" box="[199,381,727,753]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="lanai" status="sp. nov.">
<emphasis id="B96F76413565FFBB4598459C373EFD11" box="[199,381,727,753]" italics="true" pageId="97" pageNumber="98">Chthonius lanai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25CCB3A3565FFBB44DA459C3780FD11" box="[389,451,727,753]" pageId="97" pageNumber="98" rank="species">n. sp.</taxonomicNameLabel>
differs from
<taxonomicName id="4C1BD1D03565FFBB4703459C3493FD11" authorityName="Beier" authorityYear="1930" box="[604,720,727,753]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="italicus">
<emphasis id="B96F76413565FFBB4703459C3493FD11" box="[604,720,727,753]" italics="true" pageId="97" pageNumber="98">C. italicus</emphasis>
</taxonomicName>
in the following characters: anterior eyes reduced to a smooth cuticular area (no eyes in
<taxonomicName id="4C1BD1D03565FFBB449F45B03477FCF5" authorityName="Beier" authorityYear="1930" box="[448,564,763,789]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="italicus">
<emphasis id="B96F76413565FFBB449F45B03477FCF5" box="[448,564,763,789]" italics="true" pageId="97" pageNumber="98">C. italicus</emphasis>
</taxonomicName>
); chaetotaxy of tergites I–IV 2:2–3:4:4 [(2–3)4:3–4:4:
<quantity id="4CE307B63565FFBB41C245B0328FFCF5" box="[1181,1228,763,789]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="97" pageNumber="98" unit="in" value="4.0">4 in</quantity>
<taxonomicName id="4C1BD1D03565FFBB418B45B0330AFCF5" authorityName="Beier" authorityYear="1930" box="[1236,1353,763,789]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="italicus">
<emphasis id="B96F76413565FFBB418B45B0330AFCF5" box="[1236,1353,763,789]" italics="true" pageId="97" pageNumber="98">C. italicus</emphasis>
</taxonomicName>
]; fixed chelal finger straight (slightly curved in
<taxonomicName id="4C1BD1D03565FFBB47034454348CFCD9" authorityName="Beier" authorityYear="1930" box="[604,719,799,825]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="italicus">
<emphasis id="B96F76413565FFBB47034454348CFCD9" box="[604,719,799,825]" italics="true" pageId="97" pageNumber="98">C. italicus</emphasis>
</taxonomicName>
); fixed and movable chelal fingers with larger and more widely spaced teeth (
<figureCitation id="1320B6D63565FFBB446F4408378FFCBD" box="[304,460,834,861]" captionStart="FIGURES 304–308" captionStartId="96.[151,264,1889,1913]" captionTargetBox="[152,1433,181,1858]" captionTargetId="figure-32@96.[151,1436,181,1865]" captionTargetPageId="96" captionText="FIGURES 304–308. Chthonius lanai n. sp. 304, female holotype, right chela, antiaxial view; 305, female holotype, detail of fingers at the chelal tip, antiaxial view; 306, female holotype, detail of fingers at level of trichobothria est-it and st-t, antiaxial view; 307, female holotype, detail of fingers at level of trichobothrium sb, antiaxial view; 308, male paratype, right chela, antiaxial view." figureDoi="http://doi.org/10.5281/zenodo.10121279" httpUri="https://zenodo.org/record/10121279/files/figure.png" pageId="97" pageNumber="98">Figs 305–307</figureCitation>
) [with smaller and more close-set teeth (
<figureCitation id="1320B6D63565FFBB46CB440832E2FCBD" box="[916,1185,834,861]" captionStart="FIGURES 274–281" captionStartId="89.[151,264,1675,1699]" captionTargetBox="[158,1422,188,1651]" captionTargetId="figure-16@89.[151,1436,181,1651]" captionTargetPageId="89" captionText="FIGURES 274–281. Chthonius italicus Beier, 1930, Grotta dei Surgentin, 29.III.2015 (Pigna, Imperia). 274, male, right chela, antiaxial view; 275, male, detail of fingers at the chelal tip, antiaxial view; 276, male, detail of fingers at level of trichobothria est-it and st-t, antiaxial view; 277, male, detail of fingers at level of trichobothrium sb, antiaxial view; 278, female, right chela, antiaxial view; 279, female, detail of fingers at the chelal tip, antiaxial view; 280, female, detail of fingers at level of trichobothria est-it and st-t, antiaxial view; 281, female, detail of fingers at level of trichobothrium sb, antiaxial view." figureDoi="http://doi.org/10.5281/zenodo.10121271" httpUri="https://zenodo.org/record/10121271/files/figure.png" pageId="97" pageNumber="98">Figs 275–277, 279–281</figureCitation>
) in
<taxonomicName id="4C1BD1D03565FFBB41934408337CFCBD" authorityName="Beier" authorityYear="1930" box="[1228,1343,835,861]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="italicus">
<emphasis id="B96F76413565FFBB41934408337CFCBD" box="[1228,1343,835,861]" italics="true" pageId="97" pageNumber="98">C. italicus</emphasis>
</taxonomicName>
].
</paragraph>
</subSubSection>
</treatment>
</document>
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