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<document id="D0EF61E0C32051AAF91BF8290F54D971" ID-CLB-Dataset="5408" ID-DOI="10.11646/zootaxa.4987.1.1" ID-GBIF-Dataset="b7f231a2-b766-4f47-914c-c7d39518641f" ID-ISSN="1175-5326" ID-Zenodo-Dep="10086770" ID-ZooBank="33814755-4D9E-4587-BED2-E8B0ED19B371" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623996682541" checkinUser="plazi" docAuthor="Gardini, Giulio" docDate="2021" docId="03B21B453543FF9045C844BD375CFD65" docLanguage="en" docName="zootaxa.4987.1.1.pdf" docOrigin="Zootaxa 4987 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Chthonius inguscioi Gardini 2021, n. sp." docType="treatment" docVersion="10" lastPageNumber="75" masterDocId="FF8B633D3504FFDA455F474B3643FFE0" masterDocTitle="The Italian species of the Chthonius ischnocheles group (Arachnida, Pseudoscorpiones, Chthoniidae), with reference to neighbouring countries" masterLastPageNumber="131" masterPageNumber="1" pageNumber="72" updateTime="1699926180482" updateUser="ExternalLinkService">
<mods:mods id="8D222D7AC26C493C1F4A8CA803D99767" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="DC683716097E6F8835C8484914625144">The Italian species of the Chthonius ischnocheles group (Arachnida, Pseudoscorpiones, Chthoniidae), with reference to neighbouring countries</mods:title>
</mods:titleInfo>
<mods:name id="5C3EFEEAD9C8301E889E42961AB1505C" type="personal">
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<mods:namePart id="A9DFC37FBDC4C63305DFF2DA42AA8125">Gardini, Giulio</mods:namePart>
</mods:name>
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<mods:title id="B6031134B5F49FCF148BD8D3DE5836D8">Zootaxa</mods:title>
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<mods:part id="A313BADC2628AC09D2686171551582FC">
<mods:date id="7BF198D5CF89F606F064946E5DD49D28">2021</mods:date>
<mods:detail id="3F539D2CE94B61E8C481B409690941E1" type="pubDate">
<mods:number id="BD8681078CDE183C321D55F6307D8FBE">2021-06-17</mods:number>
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<mods:number id="03EE4145FFDDFD18BA9F1299473768B7">4987</mods:number>
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<mods:number id="A4293134064EB234462607E81BD817E3">1</mods:number>
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<mods:classification id="0778929581577CD013D4448629C285A4">journal article</mods:classification>
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<mods:identifier id="861FB0568265B72F3C8A570CDAA46516" type="DOI">10.11646/zootaxa.4987.1.1</mods:identifier>
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<treatment id="03B21B453543FF9045C844BD375CFD65" ID-DOI="http://doi.org/10.5281/zenodo.5110857" ID-GBIF-Taxon="183613349" ID-Zenodo-Dep="5110857" LSID="urn:lsid:plazi:treatment:03B21B453543FF9045C844BD375CFD65" httpUri="http://treatment.plazi.org/id/03B21B453543FF9045C844BD375CFD65" lastPageId="74" lastPageNumber="75" pageId="71" pageNumber="72">
<subSubSection id="C301F9D83543FF9D45C844BD3795FBF1" box="[151,470,1014,1041]" pageId="71" pageNumber="72" type="nomenclature">
<paragraph id="8BA4AA533543FF9D45C844BD3795FBF1" blockId="71.[151,470,1014,1077]" box="[151,470,1014,1041]" pageId="71" pageNumber="72">
<heading id="D0EC1D3F3543FF9D45C844BD3795FBF1" bold="true" box="[151,470,1014,1041]" fontSize="11" level="2" pageId="71" pageNumber="72" reason="4">
<emphasis id="B96F76413543FF9D45C844BD3795FBF1" bold="true" box="[151,470,1014,1041]" pageId="71" pageNumber="72">
<taxonomicName id="4C1BD1D03543FF9D45C844BD37CEFBF1" authority="Gardini, 2021" authorityName="Gardini" authorityYear="2021" box="[151,397,1014,1041]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="inguscioi" status="sp. nov.">
<emphasis id="B96F76413543FF9D45C844BD37CEFBF1" bold="true" box="[151,397,1014,1041]" italics="true" pageId="71" pageNumber="72">Chthonius inguscioi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25CCB3A3543FF9D44CB44BC3795FBF1" box="[404,470,1015,1041]" pageId="71" pageNumber="72" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C301F9D83543FF9D45C843503738FBD5" box="[151,379,1051,1077]" pageId="71" pageNumber="72" type="description">
<paragraph id="8BA4AA533543FF9D45C843503738FBD5" blockId="71.[151,470,1014,1077]" box="[151,379,1051,1077]" pageId="71" pageNumber="72">
(
<figureCitation id="1320B6D63543FF9D45C04350377EFBD5" box="[159,317,1051,1077]" captionStart-0="FIGURES 229–232" captionStart-1="FIGURES 233–236" captionStartId-0="72.[151,264,1935,1959]" captionStartId-1="73.[151,264,1943,1967]" captionTargetBox-0="[153,1434,1002,1905]" captionTargetBox-1="[201,1387,182,1920]" captionTargetId-0="figure-443@72.[151,1436,993,1912]" captionTargetId-1="figure-16@73.[200,1388,181,1920]" captionTargetPageId-0="72" captionTargetPageId-1="73" captionText-0="FIGURES 229–232. Chthonius inguscioi n. sp., male holotype. 229, anterior margin of carapace; 230, carapace; 231, right chelicera; 232, right chelal hand, dorsal view." captionText-1="FIGURES 233–236. Chthonius inguscioi n. sp., male holotype. 233, right chela, antiaxial view; 234, detail of dentition at the right chelal fingers tip, antiaxial view; 235, detail of fingers at level of trichobothria est-it and st-t, antiaxial view; 236, detail of fingers at level of trichobothrium sb, antiaxial view." figureDoi-0="http://doi.org/10.5281/zenodo.10121259" figureDoi-1="http://doi.org/10.5281/zenodo.10121261" httpUri-0="https://zenodo.org/record/10121259/files/figure.png" httpUri-1="https://zenodo.org/record/10121261/files/figure.png" pageId="71" pageNumber="72">Figs 229–236</figureCitation>
,
<figureCitation id="1320B6D63543FF9D441743503737FBD5" box="[328,372,1051,1077]" captionStart="FIGURES 410–413" captionStartId="127.[151,264,1800,1824]" captionTargetBox="[154,1432,184,1767]" captionTargetId="figure-16@127.[151,1435,181,1777]" captionTargetPageId="127" captionText="FIGURES 410–413. Distribution maps for the Italian species of the Chthonius ischnocheles group. 410, Chthonius caoduroi Callaini, 1987, Chthonius gallettii n. sp., Chthonius inguscioi n. sp., Chthonius marciai n. sp. and Chthonius multidentatus Beier, 1963; 411, Chthonius caprai Gardini, 1977, Chthonius jonicus Beier, 1931 and Chthonius leoi (Callaini, 1988); 412, Chthonius halberti Kew, 1916, Chthonius nicolosii n. sp., Chthonius ressli Beier, 1956 and Chthonius stammeri Beier, 1942; 413, Chthonius horridus Beier, 1934, Chthonius italicus Beier, 1930 and Chthonius lanai n. sp." figureDoi="http://doi.org/10.5281/zenodo.10121317" httpUri="https://zenodo.org/record/10121317/files/figure.png" pageId="71" pageNumber="72">410</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C301F9D83543FF9D45C84328376DFB42" pageId="71" pageNumber="72" type="materials_examined">
<paragraph id="8BA4AA533543FF9D45C84328376DFB42" blockId="71.[151,1437,1122,2013]" pageId="71" pageNumber="72">
<materialsCitation id="3B73A00E3543FF9D45C843283769FB42" ID-GBIF-Occurrence="3325145452" collectionCode="LE" country="Italy" county="Lecce Province" latitude="39.919445" location="Grotta Madonna della Rutta" longLatPrecision="19" longitude="18.273056" municipality="Presicce" pageId="71" pageNumber="72" specimenCount="1" stateProvince="Puglia" typeStatus="holotype">
<emphasis id="B96F76413543FF9D45C843283776FB9C" bold="true" box="[151,309,1122,1149]" pageId="71" pageNumber="72">
<typeStatus id="54A014F13543FF9D45C843283691FB9D" box="[151,210,1123,1149]" pageId="71" pageNumber="72">Type</typeStatus>
locality
</emphasis>
:
<collectingCountry id="F30CEAC33543FF9D44164328373EFB9D" box="[329,381,1123,1149]" name="Italy" pageId="71" pageNumber="72">Italy</collectingCountry>
,
<collectingRegion id="49DF64B13543FF9D44D24328379FFB9D" box="[397,476,1123,1149]" country="Italy" name="Puglia" pageId="71" pageNumber="72">Apulia</collectingRegion>
,
<collectingCounty id="62C5D2DF3543FF9D44B3432834E2FB9D" box="[492,673,1123,1149]" pageId="71" pageNumber="72">Lecce Province</collectingCounty>
,
<collectingMunicipality id="6BC030293543FF9D47ED43283551FB9D" box="[690,786,1123,1149]" pageId="71" pageNumber="72">Presicce</collectingMunicipality>
,
<location id="8EC4FC883543FF9D467C4328322FFB9D" LSID="urn:lsid:plazi:treatment:03B21B453543FF9045C844BD375CFD65:8EC4FC883543FF9D467C4328322FFB9D" box="[803,1132,1123,1149]" country="Italy" county="Lecce Province" latitude="39.919445" longLatPrecision="19" longitude="18.273056" municipality="Presicce" name="Grotta Madonna della Rutta" pageId="71" pageNumber="72" stateProvince="Puglia">Grotta Madonna della Rutta</location>
533
<location id="8EC4FC883543FF9D41F04328328FFB9D" LSID="urn:lsid:plazi:treatment:03B21B453543FF9045C844BD375CFD65:8EC4FC883543FF9D41F04328328FFB9D" box="[1199,1228,1123,1149]" country="Italy" county="Lecce Province" latitude="39.919445" longLatPrecision="19" longitude="18.273056" municipality="Presicce" name="Pu" pageId="71" pageNumber="72" stateProvince="Puglia">Pu</location>
/
<collectionCode id="ED0A32963543FF9D418A432832B4FB9D" box="[1237,1271,1123,1149]" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="71" pageNumber="72">LE</collectionCode>
(
<geoCoordinate id="EE2FCC943543FF9D4054432933DBFB9D" box="[1291,1432,1122,1149]" degrees="39" direction="north" minutes="55" orientation="latitude" pageId="71" pageNumber="72" precision="15" seconds="10" value="39.919445">39°55’10”N</geoCoordinate>
,
<geoCoordinate id="EE2FCC943543FF9D45C843CC3763FB42" box="[151,288,1159,1186]" degrees="18" direction="east" minutes="16" orientation="longitude" pageId="71" pageNumber="72" precision="15" seconds="23" value="18.273056">18°16’23”E</geoCoordinate>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C301F9D83543FF9D459843E1344BFB26" box="[199,520,1194,1222]" pageId="71" pageNumber="72" type="distribution">
<paragraph id="8BA4AA533543FF9D459843E1344BFB26" blockId="71.[151,1437,1122,2013]" box="[199,520,1194,1222]" pageId="71" pageNumber="72">
<emphasis id="B96F76413543FF9D459843E1371AFB24" bold="true" box="[199,345,1194,1220]" pageId="71" pageNumber="72">Distribution</emphasis>
.
<collectingCountry id="F30CEAC33543FF9D443843E037D9FB25" box="[359,410,1195,1221]" name="Italy" pageId="71" pageNumber="72">Italy</collectingCountry>
(
<collectingRegion id="49DF64B13543FF9D44F543E037BEFB26" box="[426,509,1195,1222]" country="Italy" name="Puglia" pageId="71" pageNumber="72">Apulia</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C301F9D83543FF9D45984385336BFA99" pageId="71" pageNumber="72" type="diagnosis">
<paragraph id="8BA4AA533543FF9D45984385336BFA99" blockId="71.[151,1437,1122,2013]" pageId="71" pageNumber="72">
<emphasis id="B96F76413543FF9D45984385372CFB0B" bold="true" box="[199,367,1230,1259]" pageId="71" pageNumber="72">
Diagnosis (
<specimenCount id="9D1D61DA3543FF9D4415439B371CFB0B" box="[330,351,1232,1259]" pageId="71" pageNumber="72" type="male">♂</specimenCount>
).
</emphasis>
A microphthalmic hypogean
<taxonomicName id="4C1BD1D03543FF9D47E94384356AFB09" authorityName="Rayi L. Koch" authorityYear="1873" box="[694,809,1231,1257]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="71" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="B96F76413543FF9D47E94384356AFB09" box="[694,809,1231,1257]" italics="true" pageId="71" pageNumber="72">Chthonius</emphasis>
</taxonomicName>
that differs from other species of the
<taxonomicName id="4C1BD1D03543FF9D419943843316FB09" authorityName=": Lazzeroni" authorityYear="1969" box="[1222,1365,1231,1257]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="ischnocheles">
<emphasis id="B96F76413543FF9D419943843316FB09" box="[1222,1365,1231,1257]" italics="true" pageId="71" pageNumber="72">ischnocheles</emphasis>
</taxonomicName>
group in the following combination of characters: anterior margin of carapace with 2 preocular microsetae on each side; posterior margin of carapace with 4 macrosetae (lateral setae shorter than the median ones); chelicerae with 3 lateral microsetae; chela length 0.88–1.0 mm; movable chelal finger length
<quantity id="4CE307B63543FF9D46CF42713200FAB5" box="[912,1091,1338,1365]" metricMagnitude="-4" metricUnit="m" metricValue="6.375" metricValueMax="6.8" metricValueMin="5.949999999999999" pageId="71" pageNumber="72" unit="mm" value="0.6375" valueMax="0.68" valueMin="0.595">0.595–0.68 mm</quantity>
; chelal fingers with contiguous teeth; both fixed and movable chelal fingers at level of
<emphasis id="B96F76413543FF9D465D42143579FA98" box="[770,826,1375,1400]" italics="true" pageId="71" pageNumber="72">est-it</emphasis>
and
<emphasis id="B96F76413543FF9D462F421435D7FA98" box="[880,916,1375,1400]" italics="true" pageId="71" pageNumber="72">st-t</emphasis>
with 8–11 teeth occupying
<quantity id="4CE307B63543FF9D419442143367FA99" box="[1227,1316,1375,1401]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="71" pageNumber="72" unit="mm" value="0.1">0.1 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C301F9D83543FF9D459842C834FDFA05" pageId="71" pageNumber="72" type="materials_examined">
<paragraph id="8BA4AA533543FF9D459842C834FDFA05" blockId="71.[151,1437,1122,2013]" pageId="71" pageNumber="72">
<emphasis id="B96F76413543FF9D459842C83735FA7C" bold="true" box="[199,374,1410,1437]" pageId="71" pageNumber="72">Type material.</emphasis>
<materialsCitation id="3B73A00E3543FF9D442042C834C5FA21" ID-GBIF-Occurrence="3325145555" collectingDate="1999-07-10" collectionCode="LE, R" collectorName="S. Inguscio &amp; Pepe" country="Italy" county="Lecce Prov." location="Grotta Madonna della Rutta" municipality="Presicce" pageId="71" pageNumber="72" specimenCount="1" specimenCount-male="1" stateProvince="Puglia" typeStatus="holotype">
<collectingCountry id="F30CEAC33543FF9D442042C8378CFA7D" box="[383,463,1411,1437]" name="Italy" pageId="71" pageNumber="72">ITALY</collectingCountry>
—
<emphasis id="B96F76413543FF9D44B542C83476FA7D" box="[490,565,1411,1437]" italics="true" pageId="71" pageNumber="72">
<collectingRegion id="49DF64B13543FF9D44B542C83476FA7D" box="[490,565,1411,1437]" country="Italy" name="Puglia" pageId="71" pageNumber="72">Apulia</collectingRegion>
</emphasis>
:
<specimenCount id="9D1D61DA3543FF9D471A42C83433FA7D" box="[581,624,1411,1437]" pageId="71" pageNumber="72" type="male">1 ♂</specimenCount>
(
<typeStatus id="54A014F13543FF9D47DF42C834ABFA7E" box="[640,744,1411,1438]" pageId="71" pageNumber="72" type="holotype">holotype</typeStatus>
),
<collectingCounty id="62C5D2DF3543FF9D47A242C835C6FA7D" box="[765,901,1411,1437]" pageId="71" pageNumber="72">Lecce Prov.</collectingCounty>
,
<collectingMunicipality id="6BC030293543FF9D46CE42C835B2FA7D" box="[913,1009,1411,1437]" pageId="71" pageNumber="72">Presicce</collectingMunicipality>
,
<location id="8EC4FC883543FF9D46A142C8337FFA7D" LSID="urn:lsid:plazi:treatment:03B21B453543FF9045C844BD375CFD65:8EC4FC883543FF9D46A142C8337FFA7D" box="[1022,1340,1411,1437]" country="Italy" county="Lecce Prov." municipality="Presicce" name="Grotta Madonna della Rutta" pageId="71" pageNumber="72" stateProvince="Puglia">Grotta Madonna della Rutta</location>
533
<location id="8EC4FC883543FF9D402842C833D7FA7D" LSID="urn:lsid:plazi:treatment:03B21B453543FF9045C844BD375CFD65:8EC4FC883543FF9D402842C833D7FA7D" box="[1399,1428,1411,1437]" country="Italy" county="Lecce Prov." municipality="Presicce" name="Pu" pageId="71" pageNumber="72" stateProvince="Puglia">Pu</location>
/
<collectionCode id="ED0A32963543FF9D45C842EC36FFFA21" box="[151,188,1447,1473]" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="71" pageNumber="72">LE</collectionCode>
,
<date id="FFA58C933543FF9D459842EC3711FA21" box="[199,338,1447,1473]" pageId="71" pageNumber="72" value="1999-07-10">
<collectingDate id="EFE1757B3543FF9D459842EC3711FA21" box="[199,338,1447,1473]" pageId="71" pageNumber="72" value="1999-07-10">10.VII.1999</collectingDate>
</date>
,
<collectorName id="26EECF853543FF9D440242EC3799FA21" box="[349,474,1447,1473]" pageId="71" pageNumber="72">S. Inguscio</collectorName>
&amp;
<collectionCode id="ED0A32963543FF9D44A242EC3450FA21" box="[509,531,1447,1473]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="71" pageNumber="72">R</collectionCode>
.
<collectorName id="26EECF853543FF9D474242EC3410FA21" box="[541,595,1447,1473]" pageId="71" pageNumber="72">Pepe</collectorName>
leg.
</materialsCitation>
;
<specimenCount id="9D1D61DA3543FF9D47CE42EC34F9FA21" box="[657,698,1447,1473]" pageId="71" pageNumber="72" type="male">2 ♂</specimenCount>
(
<typeStatus id="54A014F13543FF9D479742EC357AFA21" box="[712,825,1447,1473]" pageId="71" pageNumber="72" type="paratype">paratypes</typeStatus>
), id.,
<date id="FFA58C933543FF9D462942EC3242FA21" box="[886,1025,1447,1473]" pageId="71" pageNumber="72" value="2019-07-14">14.VII.2019</date>
, S. Inguscio leg.
<materialsCitation id="3B73A00E3543FF9D41E042EC34F9FA05" ID-GBIF-Occurrence="3325145755" collectionCode="MHNG" collectorName="G. Gardini &amp; Genoa" country="Italy" pageId="71" pageNumber="72" specimenCount="2" stateProvince="Apulia" typeStatus="holotype">
(
<typeStatus id="54A014F13543FF9D419842EC3368FA21" box="[1223,1323,1447,1473]" pageId="71" pageNumber="72" type="holotype">holotype</typeStatus>
deposited in
<collectionCode id="ED0A32963543FF9D45EB4280374FFA05" box="[180,268,1483,1509]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="71" pageNumber="72" type="Museum">MHNG</collectionCode>
,
<typeStatus id="54A014F13543FF9D4447428037C0FA05" box="[280,387,1483,1509]" pageId="71" pageNumber="72" type="paratype">paratypes</typeStatus>
in
<collectorName id="26EECF853543FF9D44F84280345CFA05" box="[423,543,1483,1509]" pageId="71" pageNumber="72">G. Gardini</collectorName>
coll.,
<collectorName id="26EECF853543FF9D473B428034F1FA05" box="[612,690,1483,1509]" pageId="71" pageNumber="72">Genoa</collectorName>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C301F9D83543FF9D459842A43425F9B1" pageId="71" pageNumber="72" type="etymology">
<paragraph id="8BA4AA533543FF9D459842A43425F9B1" blockId="71.[151,1437,1122,2013]" pageId="71" pageNumber="72">
<emphasis id="B96F76413543FF9D459842A437DEF9E8" box="[199,413,1519,1545]" italics="true" pageId="71" pageNumber="72">
<emphasis id="B96F76413543FF9D459842A437D5F9E9" bold="true" box="[199,406,1519,1545]" italics="true" pageId="71" pageNumber="72">Derivatio nominis</emphasis>
.
</emphasis>
Named after its collector, the naturalist and biospeleologist Salvatore Inguscio (Laboratorio Ipogeo Salentino di Biospeleologia “Sandro Ruffo”, Nardò, Lecce), in recognition of his contribution to the knowledge of the subterranean fauna of
<collectingRegion id="49DF64B13543FF9D474C417C3420F9B1" box="[531,611,1591,1617]" country="Italy" name="Puglia" pageId="71" pageNumber="72">Apulia</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C301F9D83543FF92459841113567FD11" lastPageId="72" lastPageNumber="73" pageId="71" pageNumber="72" type="description">
<paragraph id="8BA4AA533543FF9245984111335FFD65" blockId="71.[151,1437,1122,2013]" lastBlockId="72.[151,1437,150,969]" lastPageId="72" lastPageNumber="73" pageId="71" pageNumber="72">
<emphasis id="B96F76413543FF9D4598411134E6F994" bold="true" box="[199,677,1626,1655]" pageId="71" pageNumber="72">
Description of adults (
<specimenCount id="9D1D61DA3543FF9D4482411737B0F997" box="[477,499,1628,1655]" pageId="71" pageNumber="72" type="male">♂</specimenCount>
;
<specimenCount id="9D1D61DA3543FF9D475A41173459F997" box="[517,538,1628,1655]" pageId="71" pageNumber="72" type="female">♀</specimenCount>
unknown).
</emphasis>
Integument slightly pigmented, carapace, tergites, chelicerae and pedipalps pale brown; weak hispid granulation on lateral surfaces of carapace, on cheliceral palm and on base of fixed chelal finger. Carapace (
<figureCitation id="1320B6D63543FF9D44B441E8340FF95D" box="[491,588,1699,1725]" captionStart="FIGURES 229–232" captionStartId="72.[151,264,1935,1959]" captionTargetBox="[153,1434,1002,1905]" captionTargetId="figure-443@72.[151,1436,993,1912]" captionTargetPageId="72" captionText="FIGURES 229–232. Chthonius inguscioi n. sp., male holotype. 229, anterior margin of carapace; 230, carapace; 231, right chelicera; 232, right chelal hand, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10121259" httpUri="https://zenodo.org/record/10121259/files/figure.png" pageId="71" pageNumber="72">Fig. 230</figureCitation>
) 0.90 times longer than broad, trapezoidal, constricted posteriorly; anterior margin denticulate between median macrosetae and with a prominent, sharp epistome (
<figureCitation id="1320B6D63543FF9D4133418C328CF901" box="[1132,1231,1735,1761]" captionStart="FIGURES 229–232" captionStartId="72.[151,264,1935,1959]" captionTargetBox="[153,1434,1002,1905]" captionTargetId="figure-443@72.[151,1436,993,1912]" captionTargetPageId="72" captionText="FIGURES 229–232. Chthonius inguscioi n. sp., male holotype. 229, anterior margin of carapace; 230, carapace; 231, right chelicera; 232, right chelal hand, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10121259" httpUri="https://zenodo.org/record/10121259/files/figure.png" pageId="71" pageNumber="72">Fig. 229</figureCitation>
); ocular area as in fig. 230, anterior eyes reduced, with flat lens (diameter
<quantity id="4CE307B63543FF9D464941A135CEF8E5" box="[790,909,1770,1797]" metricMagnitude="-5" metricUnit="m" metricValue="3.5" pageId="71" pageNumber="72" unit="mm" value="0.035">0.035 mm</quantity>
), posterior eyes absent; distance from anterior eyes to anterior margin of carapace
<quantity id="4CE307B63543FF9D477E404534D6F8C9" box="[545,661,1806,1833]" metricMagnitude="-5" metricUnit="m" metricValue="4.5" pageId="71" pageNumber="72" unit="mm" value="0.045">0.045 mm</quantity>
; chaetotaxy mm
<quantity id="4CE307B63543FF9D460E404435D1F8CA" box="[849,914,1807,1834]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="71" pageNumber="72" unit="mm" value="4.0">4mm</quantity>
:6:4:2:4(24), length of anteromedian macrosetae
<quantity id="4CE307B63543FF9D45C840793748F8AD" box="[151,267,1842,1869]" metricMagnitude="-4" metricUnit="m" metricValue="1.35" pageId="71" pageNumber="72" unit="mm" value="0.135">0.135 mm</quantity>
, lateral setae of posterior row half as long as the median setae. Chaetotaxy of tergites 4:4:4:4:6:6:6:6:1T 2T1:4:1T2T1:0. Chaetotaxy of sternites 10:(3)10(3):(2)8(2):8:6:6:6:6:2T2T2:0:2; genital opening flanked by 7–7 setae. Chelicerae (
<figureCitation id="1320B6D63543FF9D443C40303782F875" box="[355,449,1915,1941]" captionStart="FIGURES 229–232" captionStartId="72.[151,264,1935,1959]" captionTargetBox="[153,1434,1002,1905]" captionTargetId="figure-443@72.[151,1436,993,1912]" captionTargetPageId="72" captionText="FIGURES 229–232. Chthonius inguscioi n. sp., male holotype. 229, anterior margin of carapace; 230, carapace; 231, right chelicera; 232, right chelal hand, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10121259" httpUri="https://zenodo.org/record/10121259/files/figure.png" pageId="71" pageNumber="72">Fig. 231</figureCitation>
) 2.15–2.3 times as long as broad, palm with 6 setae and 3 lateral microsetae; fixed finger with 2 distal large teeth followed by 7–8 small teeth; movable finger with an isolated subapical tooth (
<emphasis id="B96F76413543FF9D407D40D4337BF859" box="[1314,1336,1951,1977]" italics="true" pageId="71" pageNumber="72">di</emphasis>
) at level with the spinneret, followed by a large tooth and 8–9 teeth proximally reduced in size;
<emphasis id="B96F76413543FF9D410140883234F83C" box="[1118,1143,1987,2012]" italics="true" pageId="71" pageNumber="72">gs</emphasis>
ratio 0.56–0.62; spinneret weakly raised; rallum with 11 blades; serrulae interior and exterior both with 14–15 blades. Coxal setae: pedipalp 5 (including 2 on manducatory process), I 3 + 3 marginal microsetae, II 4, III 5, IV 6; coxa II with 7–9 coxal spines, coxa III with 4–6 coxal spines; intercoxal tubercle bisetose. Pedipalp: femur 5.4–5.65 times as long as broad, femoral chaetotaxy 3:6:2:5:1; chela (
<figureCitation id="1320B6D6354CFF9244A04648341CFEFD" box="[511,607,259,285]" captionStart="FIGURES 233–236" captionStartId="73.[151,264,1943,1967]" captionTargetBox="[201,1387,182,1920]" captionTargetId="figure-16@73.[200,1388,181,1920]" captionTargetPageId="73" captionText="FIGURES 233–236. Chthonius inguscioi n. sp., male holotype. 233, right chela, antiaxial view; 234, detail of dentition at the right chelal fingers tip, antiaxial view; 235, detail of fingers at level of trichobothria est-it and st-t, antiaxial view; 236, detail of fingers at level of trichobothrium sb, antiaxial view." figureDoi="http://doi.org/10.5281/zenodo.10121261" httpUri="https://zenodo.org/record/10121261/files/figure.png" pageId="72" pageNumber="73">Fig. 233</figureCitation>
) 6.25–6.3 times as long as deep; hand of chela 2.05–2.1 times as long as deep, with parallel sides in dorsal view (
<figureCitation id="1320B6D6354CFF92473D466C3486FEA1" box="[610,709,295,321]" captionStart="FIGURES 229–232" captionStartId="72.[151,264,1935,1959]" captionTargetBox="[153,1434,1002,1905]" captionTargetId="figure-443@72.[151,1436,993,1912]" captionTargetPageId="72" captionText="FIGURES 229–232. Chthonius inguscioi n. sp., male holotype. 229, anterior margin of carapace; 230, carapace; 231, right chelicera; 232, right chelal hand, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10121259" httpUri="https://zenodo.org/record/10121259/files/figure.png" pageId="72" pageNumber="73">Fig. 232</figureCitation>
); fixed chelal finger with 68–77 contiguous teeth, cuspidate and slightly inclined backwards in the distal two thirds of the finger, rounded and increasingly reduced in the proximal third (
<figureCitation id="1320B6D6354CFF92458446243738FE6A" box="[219,379,367,394]" captionStart="FIGURES 233–236" captionStartId="73.[151,264,1943,1967]" captionTargetBox="[201,1387,182,1920]" captionTargetId="figure-16@73.[200,1388,181,1920]" captionTargetPageId="73" captionText="FIGURES 233–236. Chthonius inguscioi n. sp., male holotype. 233, right chela, antiaxial view; 234, detail of dentition at the right chelal fingers tip, antiaxial view; 235, detail of fingers at level of trichobothria est-it and st-t, antiaxial view; 236, detail of fingers at level of trichobothrium sb, antiaxial view." figureDoi="http://doi.org/10.5281/zenodo.10121261" httpUri="https://zenodo.org/record/10121261/files/figure.png" pageId="72" pageNumber="73">Figs 234–236</figureCitation>
), all teeth with dental canal; fixed finger at level of
<emphasis id="B96F7641354CFF92469E462435BAFE68" box="[961,1017,367,392]" italics="true" pageId="72" pageNumber="73">est-it</emphasis>
with 11–12 teeth occupying
<quantity id="4CE307B6354CFF924060462433DBFE69" box="[1343,1432,367,393]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="72" pageNumber="73" unit="mm" value="0.1">0.1 mm</quantity>
, distance between successive apices
<quantity id="4CE307B6354CFF92477946D834D8FE4D" box="[550,667,403,429]" metricMagnitude="-6" metricUnit="m" metricValue="9.0" pageId="72" pageNumber="73" unit="mm" value="0.009">0.009 mm</quantity>
; tip of the fixed chelal finger with apical sensilla
<emphasis id="B96F7641354CFF9241E046D83296FE4D" box="[1215,1237,403,429]" italics="true" pageId="72" pageNumber="73">af</emphasis>
<subScript id="179FA816354CFF92418346EE32B2FE53" attach="right" box="[1244,1265,421,435]" fontSize="6" pageId="72" pageNumber="73">
<emphasis id="B96F7641354CFF92418346EE32AAFE53" box="[1244,1257,421,435]" italics="true" pageId="72" pageNumber="73">1-</emphasis>
2
</subScript>
, distal paraxial seta gradually curved and thin; movable chelal finger with 53–56 more widely spaced teeth, slightly scalene triangle-shaped, reclined and apically pointed from finger’s tip to halfway between trichobothria
<emphasis id="B96F7641354CFF9241934690334BFE14" box="[1228,1288,475,501]" italics="true" pageId="72" pageNumber="73">sb -st,</emphasis>
rounded and increasingly reduced towards finger base (
<figureCitation id="1320B6D6354CFF92473046B4354CFDFA" box="[623,783,511,538]" captionStart="FIGURES 233–236" captionStartId="73.[151,264,1943,1967]" captionTargetBox="[201,1387,182,1920]" captionTargetId="figure-16@73.[200,1388,181,1920]" captionTargetPageId="73" captionText="FIGURES 233–236. Chthonius inguscioi n. sp., male holotype. 233, right chela, antiaxial view; 234, detail of dentition at the right chelal fingers tip, antiaxial view; 235, detail of fingers at level of trichobothria est-it and st-t, antiaxial view; 236, detail of fingers at level of trichobothrium sb, antiaxial view." figureDoi="http://doi.org/10.5281/zenodo.10121261" httpUri="https://zenodo.org/record/10121261/files/figure.png" pageId="72" pageNumber="73">Figs 234–236</figureCitation>
); movable finger at level of
<emphasis id="B96F7641354CFF92411646B4322EFDF8" box="[1097,1133,511,536]" italics="true" pageId="72" pageNumber="73">st-t</emphasis>
with 9–10 teeth occupying
<quantity id="4CE307B6354CFF9245C8456836ADFDDD" box="[151,238,547,573]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="72" pageNumber="73" unit="mm" value="0.1">0.1 mm</quantity>
, distance between successive apices
<quantity id="4CE307B6354CFF9247DA45683506FDDD" box="[645,837,547,573]" metricMagnitude="-6" metricUnit="m" metricValue="9.999999999999998" metricValueMax="10.999999999999998" metricValueMin="9.0" pageId="72" pageNumber="73" unit="mm" value="0.009999999999999998" valueMax="0.011" valueMin="0.009">
<geoCoordinate id="EE2FCC94354CFF9247DA45683487FDDD" box="[645,708,547,573]" degrees="0.009" direction="north" orientation="latitude" pageId="72" pageNumber="73" precision="55" value="0.009">0.009</geoCoordinate>
<geoCoordinate id="EE2FCC94354CFF92479B45683553FDDD" box="[708,784,547,573]" degrees="0.011" direction="west" orientation="longitude" pageId="72" pageNumber="73" precision="55" value="-0.011">–0.011</geoCoordinate>
mm
</quantity>
; coupled sensilla
<emphasis id="B96F7641354CFF92415845683262FDDC" box="[1031,1057,547,572]" italics="true" pageId="72" pageNumber="73">pc</emphasis>
just distad
<emphasis id="B96F7641354CFF9241C0456832FBFDDD" box="[1183,1208,547,573]" italics="true" pageId="72" pageNumber="73">sb</emphasis>
; tip of movable chelal finger with apical sensilla
<emphasis id="B96F7641354CFF924483450C37BDFD80" box="[476,510,583,608]" italics="true" pageId="72" pageNumber="73">am</emphasis>
<subScript id="179FA816354CFF92475B4512345EFD87" attach="right" box="[516,541,601,615]" fontSize="6" pageId="72" pageNumber="73">
<emphasis id="B96F7641354CFF92475B45123456FD87" box="[516,533,601,615]" italics="true" pageId="72" pageNumber="73">1–</emphasis>
2
</subScript>
; trichobothria as in fig. 233; ratio
<emphasis id="B96F7641354CFF9246C8450C3247FD81" box="[919,1028,583,609]" italics="true" pageId="72" pageNumber="73">sb -st/sb -b</emphasis>
= 1.65; ratio of movable finger/hand of chela 2.0–2.05; ratio of pedipalpal femur/movable finger 1.0; ratio of pedipalpal femur/carapace 1.5.
</paragraph>
<paragraph id="8BA4AA53354CFF92459845C43567FD11" blockId="72.[151,1437,150,969]" pageId="72" pageNumber="73">
Measurements (in mm). Body length 1.5. Carapace 0.45 × 0.49 (0.47 anteriorly). Chelicerae 0.41–0.48 × 0.19– 0.21; movable finger length 0.22–0.26. Pedipalp: femur
<geoCoordinate id="EE2FCC94354CFF92464845F93515FD2D" box="[791,854,690,717]" degrees="0.595" direction="north" orientation="latitude" pageId="72" pageNumber="73" precision="55" value="0.595">0.595</geoCoordinate>
<geoCoordinate id="EE2FCC94354CFF92460A45F935E0FD2D" box="[853,931,690,717]" degrees="0.675" direction="west" orientation="longitude" pageId="72" pageNumber="73" precision="55" value="-0.675">–0.675</geoCoordinate>
×
<geoCoordinate id="EE2FCC94354CFF92469C45F93240FD2D" box="[963,1027,690,717]" degrees="0.105" direction="north" orientation="latitude" pageId="72" pageNumber="73" precision="55" value="0.105">0.105</geoCoordinate>
<geoCoordinate id="EE2FCC94354CFF92415C45F93211FD2D" box="[1027,1106,690,717]" degrees="0.125" direction="west" orientation="longitude" pageId="72" pageNumber="73" precision="55" value="-0.125">–0.125</geoCoordinate>
; chela 0.88–1.0 × 0.14–0.16; hand length 0.29–0.34; movable finger length 0.595–0.68.
</paragraph>
</subSubSection>
<subSubSection id="C301F9D8354CFF90459845B0375CFD65" lastPageId="74" lastPageNumber="75" pageId="72" pageNumber="73" type="discussion">
<paragraph id="8BA4AA53354CFF92459845B03315FCD9" blockId="72.[151,1437,150,969]" pageId="72" pageNumber="73">
<emphasis id="B96F7641354CFF92459845B03778FCF5" bold="true" box="[199,315,763,789]" pageId="72" pageNumber="73">Remarks.</emphasis>
Among the species of the
<taxonomicName id="4C1BD1D0354CFF92470045B0352BFCF5" authorityName=": Lazzeroni" authorityYear="1969" box="[607,872,763,789]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="ischnocheles">
<emphasis id="B96F7641354CFF92470045B0352BFCF5" box="[607,872,763,789]" italics="true" pageId="72" pageNumber="73">Chthonius ischnocheles</emphasis>
</taxonomicName>
group,
<taxonomicName id="4C1BD1D0354CFF9246E345B03200FCF4" authority="Gardini, 2021" authorityName="Gardini" authorityYear="2021" box="[956,1091,763,789]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="inguscioi" status="sp. nov.">
<emphasis id="B96F7641354CFF9246E345B03200FCF4" box="[956,1091,763,789]" italics="true" pageId="72" pageNumber="73">C. inguscioi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25CCB3A354CFF92411645B032C7FCF5" box="[1097,1156,763,789]" pageId="72" pageNumber="73" rank="species">n. sp.</taxonomicNameLabel>
is morphologically similar to the subterranean
<taxonomicName id="4C1BD1D0354CFF9244C744543413FCD9" authorityName="Beier" authorityYear="1963" box="[408,592,799,825]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="multidentatus">
<emphasis id="B96F7641354CFF9244C744543413FCD9" box="[408,592,799,825]" italics="true" pageId="72" pageNumber="73">C. multidentatus</emphasis>
</taxonomicName>
from
<collectingRegion id="49DF64B1354CFF9247CA44543495FCD9" box="[661,726,799,825]" country="Italy" name="Sicilia" pageId="72" pageNumber="73">Sicily</collectingRegion>
,
<taxonomicName id="4C1BD1D0354CFF9247BA4454352AFCD9" authorityName="Callaini" authorityYear="1987" box="[741,873,799,825]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="caoduroi">
<emphasis id="B96F7641354CFF9247BA4454352AFCD9" box="[741,873,799,825]" italics="true" pageId="72" pageNumber="73">C. caoduroi</emphasis>
</taxonomicName>
from
<collectingRegion id="49DF64B1354CFF9246F14454325FFCD9" box="[942,1052,799,825]" country="Italy" name="Basilicata" pageId="72" pageNumber="73">Basilicata</collectingRegion>
and
<taxonomicName id="4C1BD1D0354CFF92410C44543289FCD9" authorityName="Gardini" authorityYear="2021" box="[1107,1226,799,825]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="gallettii">
<emphasis id="B96F7641354CFF92410C44543289FCD9" box="[1107,1226,799,825]" italics="true" pageId="72" pageNumber="73">C. gallettii</emphasis>
</taxonomicName>
from
<collectingRegion id="49DF64B1354CFF92405044543311FCD9" box="[1295,1362,799,825]" country="Italy" name="Sicilia" pageId="72" pageNumber="73">Sicily</collectingRegion>
.
</paragraph>
<paragraph id="8BA4AA53354CFF924598440832C6FC29" blockId="72.[151,1437,150,969]" pageId="72" pageNumber="73">
<taxonomicName id="4C1BD1D0354CFF924598440837EBFCBC" authorityName="Gardini" authorityYear="2021" box="[199,424,835,861]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="inguscioi">
<emphasis id="B96F7641354CFF924598440837EBFCBC" box="[199,424,835,861]" italics="true" pageId="72" pageNumber="73">Chthonius inguscioi</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C1BD1D0354CFF924765440834B1FCBD" authorityName="Beier" authorityYear="1963" box="[570,754,835,861]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="multidentatus">
<emphasis id="B96F7641354CFF924765440834B1FCBD" box="[570,754,835,861]" italics="true" pageId="72" pageNumber="73">C. multidentatus</emphasis>
</taxonomicName>
in the following main characters: anterior eyes with flat lens, posterior eyes reduced to a pale cuticular area (no eyes in
<taxonomicName id="4C1BD1D0354CFF924666442C35B7FC61" authorityName="Beier" authorityYear="1963" box="[825,1012,871,897]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="multidentatus">
<emphasis id="B96F7641354CFF924666442C35B7FC61" box="[825,1012,871,897]" italics="true" pageId="72" pageNumber="73">C. multidentatus</emphasis>
</taxonomicName>
); anterior margin of carapace with 2 preocular microsetae on each side (1 preocular microseta in
<taxonomicName id="4C1BD1D0354CFF92466344C035B6FC45" authorityName="Beier" authorityYear="1963" box="[828,1013,907,933]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="multidentatus">
<emphasis id="B96F7641354CFF92466344C035B6FC45" box="[828,1013,907,933]" italics="true" pageId="72" pageNumber="73">C. multidentatus</emphasis>
</taxonomicName>
); fixed and movable chelal fingers of male with 68–77 and 53–56 teeth, respectively (male with 83 and
<quantity id="4CE307B6354CFF92462344E435F6FC29" box="[892,949,943,969]" metricMagnitude="0" metricUnit="m" metricValue="1.524" pageId="72" pageNumber="73" unit="in" value="60.0">60 in</quantity>
<taxonomicName id="4C1BD1D0354CFF9246E444E43237FC29" authorityName="Beier" authorityYear="1963" box="[955,1140,943,969]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="multidentatus">
<emphasis id="B96F7641354CFF9246E444E43237FC29" box="[955,1140,943,969]" italics="true" pageId="72" pageNumber="73">C. multidentatus</emphasis>
</taxonomicName>
).
</paragraph>
<caption id="DF64FADB354CFF9245C840C43426F82B" ID-DOI="http://doi.org/10.5281/zenodo.10121259" ID-Zenodo-Dep="10121259" httpUri="https://zenodo.org/record/10121259/files/figure.png" pageId="72" pageNumber="73" startId="72.[151,264,1935,1959]" targetBox="[153,1434,1002,1905]" targetPageId="72">
<paragraph id="8BA4AA53354CFF9245C840C43426F82B" blockId="72.[151,1436,1935,1995]" pageId="72" pageNumber="73">
<emphasis id="B96F7641354CFF9245C840C4372CF847" bold="true" box="[151,367,1935,1959]" pageId="72" pageNumber="73">FIGURES 229–232.</emphasis>
<taxonomicName id="4C1BD1D0354CFF92442740C43406F847" authority="Gardini, 2021" authorityName="Gardini" authorityYear="2021" box="[376,581,1935,1959]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="inguscioi" status="sp. nov.">
<emphasis id="B96F7641354CFF92442740C43406F847" box="[376,581,1935,1959]" italics="true" pageId="72" pageNumber="73">Chthonius inguscioi</emphasis>
</taxonomicName>
<emphasis id="B96F7641354CFF92471140C434CAF847" bold="true" box="[590,649,1935,1959]" pageId="72" pageNumber="73">
<taxonomicNameLabel id="A25CCB3A354CFF92471140C434CAF847" box="[590,649,1935,1959]" pageId="72" pageNumber="73" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male holotype. 229, anterior margin of carapace; 230, carapace; 231, right chelicera; 232, right chelal hand, dorsal view.
</paragraph>
</caption>
<caption id="DF64FADB354DFF9345C840DC34E6F818" ID-DOI="http://doi.org/10.5281/zenodo.10121261" ID-Zenodo-Dep="10121261" httpUri="https://zenodo.org/record/10121261/files/figure.png" pageId="73" pageNumber="74" startId="73.[151,264,1943,1967]" targetBox="[201,1387,182,1920]" targetPageId="73">
<paragraph id="8BA4AA53354DFF9345C840DC34E6F818" blockId="73.[151,1436,1943,2040]" pageId="73" pageNumber="74">
<emphasis id="B96F7641354DFF9345C840DC372EF84F" bold="true" box="[151,365,1943,1967]" pageId="73" pageNumber="74">FIGURES 233–236.</emphasis>
<taxonomicName id="4C1BD1D0354DFF93442B40D33403F84F" authority="Gardini, 2021" authorityName="Gardini" authorityYear="2021" box="[372,576,1944,1968]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="inguscioi" status="sp. nov.">
<emphasis id="B96F7641354DFF93442B40D33403F84F" box="[372,576,1944,1968]" italics="true" pageId="73" pageNumber="74">Chthonius inguscioi</emphasis>
</taxonomicName>
<emphasis id="B96F7641354DFF93471840D334C2F850" bold="true" box="[583,641,1944,1968]" pageId="73" pageNumber="74">
<taxonomicNameLabel id="A25CCB3A354DFF93471840D334C2F850" box="[583,641,1944,1968]" pageId="73" pageNumber="74" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male holotype. 233, right chela, antiaxial view; 234, detail of dentition at the right chelal fingers tip, antiaxial view; 235, detail of fingers at level of trichobothria
<emphasis id="B96F7641354DFF9346B440F7325DF833" box="[1003,1054,1980,2003]" italics="true" pageId="73" pageNumber="74">est-it</emphasis>
and
<emphasis id="B96F7641354DFF93411140F7322DF833" box="[1102,1134,1980,2003]" italics="true" pageId="73" pageNumber="74">st-t</emphasis>
, antiaxial view; 236, detail of fingers at level of trichobothrium
<emphasis id="B96F7641354DFF9344B140AB3446F818" box="[494,517,2016,2040]" italics="true" pageId="73" pageNumber="74">sb</emphasis>
, antiaxial view.
</paragraph>
</caption>
<paragraph id="8BA4AA53354EFF90459847DC376FFE69" blockId="74.[151,1436,151,645]" pageId="74" pageNumber="75">
<taxonomicName id="4C1BD1D0354EFF90459847DC37E4FF50" authorityName="Gardini" authorityYear="2021" box="[199,423,151,177]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="inguscioi">
<emphasis id="B96F7641354EFF90459847DC37E4FF50" box="[199,423,151,177]" italics="true" pageId="74" pageNumber="75">Chthonius inguscioi</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C1BD1D0354EFF90476847DC34F8FF51" authorityName="Callaini" authorityYear="1987" box="[567,699,151,177]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="caoduroi">
<emphasis id="B96F7641354EFF90476847DC34F8FF51" box="[567,699,151,177]" italics="true" pageId="74" pageNumber="75">C. caoduroi</emphasis>
</taxonomicName>
in the following main characters: anterior margin of carapace with 2 preocular microsetae on each side (1 preocular microseta in
<taxonomicName id="4C1BD1D0354EFF90461147F03591FF35" authorityName="Callaini" authorityYear="1987" box="[846,978,187,213]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="caoduroi">
<emphasis id="B96F7641354EFF90461147F03591FF35" box="[846,978,187,213]" italics="true" pageId="74" pageNumber="75">C. caoduroi</emphasis>
</taxonomicName>
); tergite I with 4 setae (2 or 3 setae in
<taxonomicName id="4C1BD1D0354EFF9040DC47F036B8FF19" authorityName="Callaini" authorityYear="1987" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="caoduroi">
<emphasis id="B96F7641354EFF9040DC47F036B8FF19" italics="true" pageId="74" pageNumber="75">C. caoduroi</emphasis>
</taxonomicName>
); male chela length 0.8–1.0 mm, 6.25–6.3 times as long as deep (
<quantity id="4CE307B6354EFF904683479432C0FF19" box="[988,1155,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="1.29" metricValueMax="1.3" metricValueMin="1.28" pageId="74" pageNumber="75" unit="mm" value="1.29" valueMax="1.3" valueMin="1.28">1.28–1.30 mm</quantity>
, 6.7–6.8 times as long as deep in
<taxonomicName id="4C1BD1D0354EFF9045B046483730FEFD" authorityName="Callaini" authorityYear="1987" box="[239,371,259,285]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="caoduroi">
<emphasis id="B96F7641354EFF9045B046483730FEFD" box="[239,371,259,285]" italics="true" pageId="74" pageNumber="75">C. caoduroi</emphasis>
</taxonomicName>
); fixed and movable chelal fingers at level of
<emphasis id="B96F7641354EFF90462B464835F4FEFC" box="[884,951,259,285]" italics="true" pageId="74" pageNumber="75">est -ist</emphasis>
and
<emphasis id="B96F7641354EFF9046B24648325BFEFC" box="[1005,1048,259,285]" italics="true" pageId="74" pageNumber="75">st -t,</emphasis>
respectively, with 11–12 and 9–10 teeth occupying
<quantity id="4CE307B6354EFF904410466C37E5FEA1" box="[335,422,295,321]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="74" pageNumber="75" unit="mm" value="0.1">0.1 mm</quantity>
, distance between successive apices 0.009 and
<quantity id="4CE307B6354EFF9046ED466C322CFEA1" box="[946,1135,295,321]" metricMagnitude="-6" metricUnit="m" metricValue="9.999999999999998" metricValueMax="10.999999999999998" metricValueMin="9.0" pageId="74" pageNumber="75" unit="mm" value="0.009999999999999998" valueMax="0.011" valueMin="0.009">
<geoCoordinate id="EE2FCC94354EFF9046ED466C35B2FEA1" box="[946,1009,295,321]" degrees="0.009" direction="north" orientation="latitude" pageId="74" pageNumber="75" precision="55" value="0.009">0.009</geoCoordinate>
<geoCoordinate id="EE2FCC94354EFF9046AE466C327EFEA1" box="[1009,1085,295,321]" degrees="0.011" direction="west" orientation="longitude" pageId="74" pageNumber="75" precision="55" value="-0.011">–0.011</geoCoordinate>
mm
</quantity>
respectively (with 6–7 and 5 teeth occupying
<quantity id="4CE307B6354EFF90443846003783FE85" box="[359,448,331,357]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="74" pageNumber="75" unit="mm" value="0.1">0.1 mm</quantity>
, distance between successive apices
<geoCoordinate id="EE2FCC94354EFF90463F460135DCFE85" box="[864,927,330,357]" degrees="0.014" direction="north" orientation="latitude" pageId="74" pageNumber="75" precision="55" value="0.014">0.014</geoCoordinate>
<geoCoordinate id="EE2FCC94354EFF9046FF460135AFFE85" box="[928,1004,330,357]" degrees="0.015" direction="west" orientation="longitude" pageId="74" pageNumber="75" precision="55" value="-0.015">–0.015</geoCoordinate>
and
<quantity id="4CE307B6354EFF90417B460032A5FE85" box="[1060,1254,331,357]" metricMagnitude="-5" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.7999999999999998" pageId="74" pageNumber="75" unit="mm" value="0.019" valueMax="0.02" valueMin="0.018">
<geoCoordinate id="EE2FCC94354EFF90417B46003221FE85" box="[1060,1122,331,357]" degrees="0.018" direction="north" orientation="latitude" pageId="74" pageNumber="75" precision="55" value="0.018">0.018</geoCoordinate>
<geoCoordinate id="EE2FCC94354EFF90413C460032F3FE85" box="[1123,1200,331,357]" degrees="0.020" direction="west" orientation="longitude" pageId="74" pageNumber="75" precision="55" value="-0.02">–0.020</geoCoordinate>
mm
</quantity>
, respectively, in
<taxonomicName id="4C1BD1D0354EFF9045C84624375FFE69" authorityName="Callaini" authorityYear="1987" box="[151,284,367,393]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="caoduroi">
<emphasis id="B96F7641354EFF9045C84624375FFE69" box="[151,284,367,393]" italics="true" pageId="74" pageNumber="75">C. caoduroi</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BA4AA53354EFF90459846D8375CFD65" blockId="74.[151,1436,151,645]" pageId="74" pageNumber="75">
<taxonomicName id="4C1BD1D0354EFF90459846D837E4FE4C" authorityName="Gardini" authorityYear="2021" box="[199,423,403,429]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="inguscioi">
<emphasis id="B96F7641354EFF90459846D837E4FE4C" box="[199,423,403,429]" italics="true" pageId="74" pageNumber="75">Chthonius inguscioi</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C1BD1D0354EFF90476846D834EDFE4D" authorityName="Gardini" authorityYear="2021" box="[567,686,403,429]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="gallettii">
<emphasis id="B96F7641354EFF90476846D834EDFE4D" box="[567,686,403,429]" italics="true" pageId="74" pageNumber="75">C. gallettii</emphasis>
</taxonomicName>
in the following main characters: posterior margin of carapace with 4 macrosetae (with 2 macrosetae and 1 lateral microseta on each side in
<taxonomicName id="4C1BD1D0354EFF90468946FC320CFE31" authorityName="Gardini" authorityYear="2021" box="[982,1103,439,465]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="gallettii">
<emphasis id="B96F7641354EFF90468946FC320CFE31" box="[982,1103,439,465]" italics="true" pageId="74" pageNumber="75">C. gallettii</emphasis>
</taxonomicName>
); male chela length 0.88–1.0 mm, 6.25–6.3 times as long as deep (
<quantity id="4CE307B6354EFF904764469134E0FE15" box="[571,675,474,501]" metricMagnitude="-3" metricUnit="m" metricValue="1.52" pageId="74" pageNumber="75" unit="mm" value="1.52">1.52 mm</quantity>
, 7.4 times as long as deep in
<taxonomicName id="4C1BD1D0354EFF9046B046903224FE15" authorityName="Gardini" authorityYear="2021" box="[1007,1127,475,501]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="gallettii">
<emphasis id="B96F7641354EFF9046B046903224FE15" box="[1007,1127,475,501]" italics="true" pageId="74" pageNumber="75">C. gallettii</emphasis>
</taxonomicName>
); fixed and movable chelal fingers at level of
<emphasis id="B96F7641354EFF90443246B437ECFDF8" box="[365,431,511,537]" italics="true" pageId="74" pageNumber="75">est -ist</emphasis>
and
<emphasis id="B96F7641354EFF9044B446B43455FDF8" box="[491,534,511,537]" italics="true" pageId="74" pageNumber="75">st -t,</emphasis>
respectively, with 11–12 and 9–10 teeth occupying
<quantity id="4CE307B6354EFF90412E46B4328FFDF9" box="[1137,1228,511,537]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="74" pageNumber="75" unit="mm" value="0.1">0.1 mm</quantity>
, distance between successive apices 0.009 and
<quantity id="4CE307B6354EFF90448A456834D0FDDD" box="[469,659,547,573]" metricMagnitude="-6" metricUnit="m" metricValue="9.999999999999998" metricValueMax="10.999999999999998" metricValueMin="9.0" pageId="74" pageNumber="75" unit="mm" value="0.009999999999999998" valueMax="0.011" valueMin="0.009">
<geoCoordinate id="EE2FCC94354EFF90448A45683457FDDD" box="[469,532,547,573]" degrees="0.009" direction="north" orientation="latitude" pageId="74" pageNumber="75" precision="55" value="0.009">0.009</geoCoordinate>
<geoCoordinate id="EE2FCC94354EFF90474A45683423FDDD" box="[533,608,547,573]" degrees="0.011" direction="west" orientation="longitude" pageId="74" pageNumber="75" precision="55" value="-0.011">–0.011</geoCoordinate>
mm
</quantity>
respectively (with 9 and 6 teeth occupying
<quantity id="4CE307B6354EFF90412545683292FDDD" box="[1146,1233,547,573]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="74" pageNumber="75" unit="mm" value="0.1">0.1 mm</quantity>
, distance between successive apices 0.011 and
<quantity id="4CE307B6354EFF90448A450C340AFD81" box="[469,585,583,609]" metricMagnitude="-5" metricUnit="m" metricValue="1.7" pageId="74" pageNumber="75" unit="mm" value="0.017">0.017 mm</quantity>
, respectively, in
<taxonomicName id="4C1BD1D0354EFF90465B450C353FFD81" authorityName="Gardini" authorityYear="2021" box="[772,892,583,609]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="gallettii">
<emphasis id="B96F7641354EFF90465B450C353FFD81" box="[772,892,583,609]" italics="true" pageId="74" pageNumber="75">C. gallettii</emphasis>
</taxonomicName>
); ratio of pedipalpal femur/carapace 1.5 (
<quantity id="4CE307B6354EFF904012450D33DFFD81" box="[1357,1436,582,609]" metricMagnitude="-2" metricUnit="m" metricValue="4.445" pageId="74" pageNumber="75" unit="in" value="1.75">1.75 in</quantity>
<taxonomicName id="4C1BD1D0354EFF9045C84520374CFD65" authorityName="Gardini" authorityYear="2021" box="[151,271,619,645]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="gallettii">
<emphasis id="B96F7641354EFF9045C84520374CFD65" box="[151,271,619,645]" italics="true" pageId="74" pageNumber="75">C. gallettii</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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