313 lines
37 KiB
XML
313 lines
37 KiB
XML
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<mods:title id="CEEF3043D094F29A087B5482FB5201C8">Revision of the African cichlid fish genus Ctenochromis (Teleostei, Cichliformes), including a description of the new genus Shuja from Lake Tanganyika and the new species Ctenochromis scatebra from northern Tanzania</mods:title>
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<mods:namePart id="A6AC6AAF07CE0153A90B97996455824A">Genner, Martin J.</mods:namePart>
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<mods:affiliation id="8EF51421CF768C4480784A7A243A1CA4">Department of Biological and Marine Sciences, University of Hull, Hull, HU 5 7 RX, United Kingdom.</mods:affiliation>
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<mods:namePart id="CB7EF1907538375D2B99F87140C40A6E">Shechonge, Asilatu H.</mods:namePart>
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<mods:affiliation id="4E5B167639C50DA940B17016694AE323">Tanzania Fisheries Research Institute (TAFIRI) P. O. Box 9750, Dar es Salaam, Tanzania.</mods:affiliation>
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<mods:nameIdentifier id="B8528D19BBFB0594D2FF434C417C4242" type="email">ashechonge@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="B740B64C4ABE70E340B5E1201F187ED9">Ngatunga, Benjamin P.</mods:namePart>
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<mods:affiliation id="4213B5D0640E8A687E6FEB2B9B67587B">Tanzania Fisheries Research Institute (TAFIRI) P. O. Box 9750, Dar es Salaam, Tanzania.</mods:affiliation>
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<mods:nameIdentifier id="D67353D6BBCF2C6CBC77FCB45093F69B" type="email">bpngatunga@yahoo.co.uk</mods:nameIdentifier>
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<mods:namePart id="16C9B62C8422EB3C9B3AE9E99A1295A1">Turner, George F.</mods:namePart>
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<mods:affiliation id="7523043F689CBAAD3CA78B9DE0BDEDD0">School of Biological Sciences, Bangor University, Bangor, Gwynedd, LL 57 2 UW, United Kingdom.</mods:affiliation>
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<mods:nameIdentifier id="1C386E5A367E28D773E125388CB8F433" type="email">george.turner@bangor.ac.uk</mods:nameIdentifier>
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<mods:title id="77C5DA1B765116FED8B26732C466DD78">European Journal of Taxonomy</mods:title>
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<mods:part id="8D0EB318B017E2E25838BFED5DA2AB33">
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<mods:date id="D8D6072B213B959D439C092E63FF1617">2022</mods:date>
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<mods:identifier id="72B1A29B388CA8BE469CCC352A078678" type="DOI">10.5852/ejt.2022.819.1775</mods:identifier>
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<treatment id="0390B0596033EC2D5218FD1C6743322B" ID-DOI="http://doi.org/10.5281/zenodo.6554399" ID-GBIF-Taxon="195349788" ID-Zenodo-Dep="6554399" LSID="urn:lsid:plazi:treatment:0390B0596033EC2D5218FD1C6743322B" httpUri="http://treatment.plazi.org/id/0390B0596033EC2D5218FD1C6743322B" lastPageId="20" lastPageNumber="43" pageId="19" pageNumber="42">
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Genus
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<taxonomicName id="4C397ACC6033EC2A52AAFD1C6131305E" ID-CoL="3WYZ" ID-ENA="64552" authority="Pfeffer, 1893" authorityName="Pfeffer" authorityYear="1893" box="[707,1039,767,794]" class="Actinopterygii" family="Cichlidae" genus="Ctenochromis" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="42" phylum="Chordata" rank="genus">
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<emphasis id="B94DDD5D6033EC2A52AAFD1C6650305D" bold="true" box="[707,878,767,794]" italics="true" pageId="19" pageNumber="42">Ctenochromis</emphasis>
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<bibRefCitation id="EFA87CBE6033EC2A531CFD1C6131305E" author="Pfeffer G. J." box="[885,1039,767,793]" pageId="19" pageNumber="42" pagination="131 - 177" refId="ref15940" refString="Pfeffer G. J. 1893. Ostafrikanische Fische gesammelt von Herrn Dr. F. Stuhlmann im Jahre 1888 und 1889. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten 10 (2): 131 - 177." type="journal article" year="1893">Pfeffer, 1893</bibRefCitation>
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</taxonomicName>
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<subSubSection id="C32352C46033EC2A508AFCA266EB30CC" pageId="19" pageNumber="42" type="type_taxon">
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<paragraph id="8B86014F6033EC2A508AFCA26443301C" blockId="19.[227,381,833,859]" box="[227,381,833,859]" pageId="19" pageNumber="42">
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<heading id="D0CEB6236033EC2A508AFCA26443301C" bold="true" box="[227,381,833,859]" fontSize="11" level="3" pageId="19" pageNumber="42" reason="3">
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<emphasis id="B94DDD5D6033EC2A508AFCA26443301C" bold="true" box="[227,381,833,859]" pageId="19" pageNumber="42">
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<typeStatus id="5482BFED6033EC2A508AFCA2641E301C" box="[227,288,833,859]" pageId="19" pageNumber="42">Type</typeStatus>
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species
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B86014F6033EC2A508AFC9366EB30CC" blockId="19.[227,981,880,907]" box="[227,981,880,907]" pageId="19" pageNumber="42">
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<taxonomicName id="4C397ACC6033EC2A508AFC93679A30CD" ID-CoL="325L5" authority="Pfeffer, 1893" authorityName="Pfeffer" authorityYear="1893" box="[227,676,880,906]" class="Actinopterygii" family="Cichlidae" genus="Ctenochromis" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="42" phylum="Chordata" rank="species" species="pectoralis">
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<emphasis id="B94DDD5D6033EC2A508AFC93673D30CD" box="[227,515,880,906]" italics="true" pageId="19" pageNumber="42">Ctenochromis pectoralis</emphasis>
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<bibRefCitation id="EFA87CBE6033EC2A5263FC93679A30CD" author="Pfeffer G. J." box="[522,676,880,906]" pageId="19" pageNumber="42" pagination="131 - 177" refId="ref15940" refString="Pfeffer G. J. 1893. Ostafrikanische Fische gesammelt von Herrn Dr. F. Stuhlmann im Jahre 1888 und 1889. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten 10 (2): 131 - 177." type="journal article" year="1893">Pfeffer, 1893</bibRefCitation>
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</taxonomicName>
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(by original designation).
|
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</paragraph>
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</subSubSection>
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<subSubSection id="C32352C46033EC2A508AFC51606C37CE" pageId="19" pageNumber="42" type="diagnosis">
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<paragraph id="8B86014F6033EC2A508AFC516462308B" blockId="19.[227,348,946,972]" box="[227,348,946,972]" pageId="19" pageNumber="42">
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<heading id="D0CEB6236033EC2A508AFC516462308B" bold="true" box="[227,348,946,972]" fontSize="11" level="3" pageId="19" pageNumber="42" reason="3">
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<emphasis id="B94DDD5D6033EC2A508AFC516462308B" bold="true" box="[227,348,946,972]" pageId="19" pageNumber="42">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B86014F6033EC2A508AFC02606C37CE" blockId="19.[227,1439,993,1162]" pageId="19" pageNumber="42">
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<taxonomicName id="4C397ACC6033EC2A508AFC0264B830BC" authorityName="Pfeffer" authorityYear="1893" box="[227,390,993,1019]" class="Actinopterygii" family="Cichlidae" genus="Ctenochromis" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="42" phylum="Chordata" rank="genus">
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<emphasis id="B94DDD5D6033EC2A508AFC0264B830BC" box="[227,390,993,1019]" italics="true" pageId="19" pageNumber="42">Ctenochromis</emphasis>
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</taxonomicName>
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is a haplochromine cichlid genus restricted to species with the combination of four key characters, following
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<bibRefCitation id="EFA87CBE6033EC2A518EFBE767F23758" author="Greenwood P. H." box="[487,716,1028,1055]" pageId="19" pageNumber="42" pagination="265 - 322" refId="ref14894" refString="Greenwood P. H. 1979. Towards a phyletic classification of the ' genus' Haplochromis (Pisces, Cichlidae) and related taxa. Part I. Bulletin of the British Museum of Natural History (Zoology) 35: 265 - 322. https: // doi. org / 10.5962 / bhl. part. 20455" type="journal article" year="1979">Greenwood (1979)</bibRefCitation>
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: 1) “The abrupt size transition between the very small chest scales and the larger scales on the ventrolateral aspects of the anterior flanks”, 2) “a naked area on either side of the chest”, 3) “a failure of cheek squamation to reach the ventral margin of the cheek”, and 4) “anal fin markings of male fishes are in the form of one or two (rarely three) non-ocellate spots”.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32352C46033EC2D508AFB516743322B" lastPageId="20" lastPageNumber="43" pageId="19" pageNumber="42" type="discussion">
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<paragraph id="8B86014F6033EC2A508AFB516468378B" blockId="19.[227,342,1202,1228]" box="[227,342,1202,1228]" pageId="19" pageNumber="42">
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<heading id="D0CEB6236033EC2A508AFB516468378B" bold="true" box="[227,342,1202,1228]" fontSize="11" level="3" pageId="19" pageNumber="42" reason="3">
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<emphasis id="B94DDD5D6033EC2A508AFB516468378B" bold="true" box="[227,342,1202,1228]" pageId="19" pageNumber="42">Remarks</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B86014F6033EC2A508AFB02673336CE" blockId="19.[225,1436,1249,1418]" pageId="19" pageNumber="42">
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As noted by
|
||
<bibRefCitation id="EFA87CBE6033EC2A511BFB0267B537BC" author="Greenwood P. H." box="[370,651,1249,1275]" pageId="19" pageNumber="42" pagination="265 - 322" refId="ref14894" refString="Greenwood P. H. 1979. Towards a phyletic classification of the ' genus' Haplochromis (Pisces, Cichlidae) and related taxa. Part I. Bulletin of the British Museum of Natural History (Zoology) 35: 265 - 322. https: // doi. org / 10.5962 / bhl. part. 20455" type="journal article" year="1979">Greenwood (1979: 288)</bibRefCitation>
|
||
, non-ocellate spots are “without a dark margin or clear surround”. In
|
||
<bibRefCitation id="EFA87CBE6033EC2A5088FAE764803659" author="Greenwood P. H." box="[225,446,1284,1310]" pageId="19" pageNumber="42" pagination="265 - 322" refId="ref14894" refString="Greenwood P. H. 1979. Towards a phyletic classification of the ' genus' Haplochromis (Pisces, Cichlidae) and related taxa. Part I. Bulletin of the British Museum of Natural History (Zoology) 35: 265 - 322. https: // doi. org / 10.5962 / bhl. part. 20455" type="journal article" year="1979">Greenwood (1979)</bibRefCitation>
|
||
the significance of the non-ocellate male egg spots in the diagnosis of
|
||
<taxonomicName id="4C397ACC6033EC2A5490FAE760A23659" authorityName="Pfeffer" authorityYear="1893" box="[1273,1436,1284,1310]" class="Actinopterygii" family="Cichlidae" genus="Ctenochromis" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="42" phylum="Chordata" rank="genus">
|
||
<emphasis id="B94DDD5D6033EC2A5490FAE760A23659" box="[1273,1436,1284,1310]" italics="true" pageId="19" pageNumber="42">Ctenochromis</emphasis>
|
||
</taxonomicName>
|
||
is unclear, hence a rediagnosis has been provided here. Using the four diagnostic characters for
|
||
<taxonomicName id="4C397ACC6033EC2A508AFAA864B83622" authorityName="Pfeffer" authorityYear="1893" box="[227,390,1355,1381]" class="Actinopterygii" family="Cichlidae" genus="Ctenochromis" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="42" phylum="Chordata" rank="genus">
|
||
<emphasis id="B94DDD5D6033EC2A508AFAA864B83622" box="[227,390,1355,1381]" italics="true" pageId="19" pageNumber="42">Ctenochromis</emphasis>
|
||
</taxonomicName>
|
||
listed above, the genus currently includes only
|
||
<taxonomicName id="4C397ACC6033EC2A53D3FAAF616C3622" authorityName="Pfeffer" authorityYear="1893" box="[954,1106,1355,1382]" class="Actinopterygii" family="Cichlidae" genus="Ctenochromis" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="42" phylum="Chordata" rank="species" species="pectoralis">
|
||
<emphasis id="B94DDD5D6033EC2A53D3FAAF616C3622" box="[954,1106,1355,1382]" italics="true" pageId="19" pageNumber="42">C. pectoralis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C397ACC6033EC2A54E5FAA860A23622" authority="Genner & Hsu & Collins & Smith & Saxon & Shechonge & Ngatunga & Turner, 2022" authorityName="Genner & Hsu & Collins & Smith & Saxon & Shechonge & Ngatunga & Turner" authorityYear="2022" box="[1164,1436,1355,1381]" class="Actinopterygii" family="Cichlidae" genus="Ctenochromis" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="42" phylum="Chordata" rank="species" species="scatebra" status="sp. nov.">
|
||
<emphasis id="B94DDD5D6033EC2A54E5FAA860A23622" box="[1164,1436,1355,1381]" italics="true" pageId="19" pageNumber="42">Ctenochromis scatebra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27E60266033EC2A508AFA93640336CD" box="[227,317,1392,1418]" pageId="19" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
|
||
described herein.
|
||
</paragraph>
|
||
<paragraph id="8B86014F6033EC2A508AFA5160A23B49" blockId="19.[227,1438,1458,2063]" pageId="19" pageNumber="42">
|
||
Of
|
||
<bibRefCitation id="EFA87CBE6033EC2A5166FA516739368B" author="Greenwood P. H." box="[271,519,1458,1484]" pageId="19" pageNumber="42" pagination="265 - 322" refId="ref14894" refString="Greenwood P. H. 1979. Towards a phyletic classification of the ' genus' Haplochromis (Pisces, Cichlidae) and related taxa. Part I. Bulletin of the British Museum of Natural History (Zoology) 35: 265 - 322. https: // doi. org / 10.5962 / bhl. part. 20455" type="journal article" year="1979">Greenwood’s (1979)</bibRefCitation>
|
||
five species of
|
||
<taxonomicName id="4C397ACC6033EC2A52BCFA516644368B" authorityName="Pfeffer" authorityYear="1893" box="[725,890,1458,1484]" class="Actinopterygii" family="Cichlidae" genus="Ctenochromis" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="42" phylum="Chordata" rank="genus">
|
||
<emphasis id="B94DDD5D6033EC2A52BCFA516644368B" box="[725,890,1458,1484]" italics="true" pageId="19" pageNumber="42">Ctenochromis</emphasis>
|
||
</taxonomicName>
|
||
, namely
|
||
<taxonomicName id="4C397ACC6033EC2A539AFA5161AD368B" authorityName="Pfeffer" authorityYear="1893" box="[1011,1171,1458,1484]" class="Actinopterygii" family="Cichlidae" genus="Ctenochromis" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="42" phylum="Chordata" rank="species" species="pectoralis">
|
||
<emphasis id="B94DDD5D6033EC2A539AFA5161AD368B" box="[1011,1171,1458,1484]" italics="true" pageId="19" pageNumber="42">C. pectoralis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C397ACC6033EC2A54CEFA516033368B" authorityName="Genner & Hsu & Collins & Smith & Saxon & Shechonge & Ngatunga & Turner" authorityYear="2022" baseAuthorityName="Genner & Hsu & Collins & Smith & Saxon & Shechonge & Ngatunga & Turner" baseAuthorityYear="1894" box="[1191,1293,1458,1484]" class="Actinopterygii" family="Cichlidae" genus="Shuja" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="42" phylum="Chordata" rank="species" species="horei">
|
||
<emphasis id="B94DDD5D6033EC2A54CEFA516033368B" box="[1191,1293,1458,1484]" italics="true" pageId="19" pageNumber="42">C. horei</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C397ACC6033EC2A554BFA5160AB368B" box="[1314,1429,1458,1484]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="42" phylum="Chordata" rank="species" species="luluae">
|
||
<emphasis id="B94DDD5D6033EC2A554BFA5160AB368B" box="[1314,1429,1458,1484]" italics="true" pageId="19" pageNumber="42">C. luluae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C397ACC6033EC2A508AFA34649E36B7" box="[227,416,1494,1521]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="42" phylum="Chordata" rank="species" species="oligacanthus">
|
||
<emphasis id="B94DDD5D6033EC2A508AFA34649E36B7" box="[227,416,1494,1521]" italics="true" pageId="19" pageNumber="42">C. oligacanthus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C397ACC6033EC2A51B7FA3467EE36B7" authority=", Greenwood" authorityName="Greenwood" box="[478,720,1494,1521]" class="Actinopterygii" family="Cichlidae" genus="Ctenochromis" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="42" phylum="Chordata" rank="species" species="polli">
|
||
<emphasis id="B94DDD5D6033EC2A51B7FA34670936B7" box="[478,567,1494,1521]" italics="true" pageId="19" pageNumber="42">C. polli</emphasis>
|
||
, Greenwood
|
||
</taxonomicName>
|
||
notes that three species have non-ocellate egg spots but did not specifically mention which species, although it is likely that Greenwood considered his
|
||
<taxonomicName id="4C397ACC6033EC2A5529FA1860A23553" box="[1344,1436,1530,1557]" class="Actinopterygii" family="Cichlidae" genus="Ctenochromis" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="42" phylum="Chordata" rank="species" species="polli">
|
||
<emphasis id="B94DDD5D6033EC2A5529FA1860A23553" box="[1344,1436,1530,1557]" italics="true" pageId="19" pageNumber="42">C. polli</emphasis>
|
||
</taxonomicName>
|
||
to have this trait, given the mention of a colour photograph of the species in
|
||
<bibRefCitation id="EFA87CBE6033EC2A54EFF9FC6079357F" author="Voss J." box="[1158,1351,1566,1593]" pageId="19" pageNumber="42" pagination="34 - 80" refId="ref17301" refString="Voss J. 1977. Les livrees ou patrons de coloration chez les poissons Cichlides africains. Revue francaise d'Aquariologie 4: 34 - 80." type="journal article" year="1977">Voss (1977: 74)</bibRefCitation>
|
||
. Based on photographs of field collected specimens, or specimens kept in aquaria, we are aware of only one of Greenwood’s five species that unquestionably possesses non-ocellate egg spots, namely the
|
||
<typeStatus id="5482BFED6033EC2A5503F98460A335C6" box="[1386,1437,1639,1665]" pageId="19" pageNumber="42">type</typeStatus>
|
||
species
|
||
<taxonomicName id="4C397ACC6033EC2A512DF96864DA35E2" authorityName="Pfeffer" authorityYear="1893" box="[324,484,1675,1701]" class="Actinopterygii" family="Cichlidae" genus="Ctenochromis" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="42" phylum="Chordata" rank="species" species="pectoralis">
|
||
<emphasis id="B94DDD5D6033EC2A512DF96864DA35E2" box="[324,484,1675,1701]" italics="true" pageId="19" pageNumber="42">C. pectoralis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13021DCA6033EC2A5190F968677935E1" box="[505,583,1675,1702]" captionStart="Fig" captionStartId="20.[227,270,1859,1885]" captionTargetBox="[227,1436,895,1802]" captionTargetId="figure-184@20.[227,1436,895,1805]" captionTargetPageId="20" captionText="Fig. 6. a. Ctenochomis pectoralis Pfeffer, 1893, lectotype ZMH402 from Korogwe (imaged by Thilo Weddehage). b. C. pectoralis, paralectotype BMNH 1899.2.27.1 from Korogwe. c. C. pectoralis, ♂ from Ruvu River shortly after capture (part of BMNH 2021.7.15.1-3). d. C. pectoralis, ♂ from Ruvu River preserved state (part of BMNH 2021.7.15.1-3). e. C. scatebra Genner, Ngatunga & Turner sp. nov., ♂ from Chemka Springs shortly after capture (part of BMNH 2021.7.15.1-3). f. C. scatebra sp. nov. holotype BMNH 2021.7.15.4. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6554371" httpUri="https://zenodo.org/record/6554371/files/figure.png" pageId="19" pageNumber="42">Fig. 6</figureCitation>
|
||
). Aquarium specimens of Greenwood’s
|
||
<taxonomicName id="4C397ACC6033EC2A5452F968619E35E2" authorityName="Genner & Hsu & Collins & Smith & Saxon & Shechonge & Ngatunga & Turner" authorityYear="2022" baseAuthorityName="Genner & Hsu & Collins & Smith & Saxon & Shechonge & Ngatunga & Turner" baseAuthorityYear="1894" box="[1083,1184,1675,1701]" class="Actinopterygii" family="Cichlidae" genus="Shuja" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="42" phylum="Chordata" rank="species" species="horei">
|
||
<emphasis id="B94DDD5D6033EC2A5452F968619E35E2" box="[1083,1184,1675,1701]" italics="true" pageId="19" pageNumber="42">C. horei</emphasis>
|
||
</taxonomicName>
|
||
(herein
|
||
<taxonomicName id="4C397ACC6033EC2A5567F96860A235E2" authorityName="Genner & Hsu & Collins & Smith & Saxon & Shechonge & Ngatunga & Turner" authorityYear="2022" baseAuthorityName="Genner & Hsu & Collins & Smith & Saxon & Shechonge & Ngatunga & Turner" baseAuthorityYear="1894" box="[1294,1436,1675,1701]" class="Actinopterygii" family="Cichlidae" genus="Shuja" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="42" phylum="Chordata" rank="species" species="horei" status="gen. et comb. nov.">
|
||
<emphasis id="B94DDD5D6033EC2A5567F96860A235E2" box="[1294,1436,1675,1701]" italics="true" pageId="19" pageNumber="42">Shuja horei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27E60266033EC2A508AF94C64FC358E" box="[227,450,1711,1737]" pageId="19" pageNumber="42" rank="species">gen. et comb. nov.</taxonomicNameLabel>
|
||
) are clearly in possession of ocellate egg spots (see
|
||
<bibRefCitation id="EFA87CBE6033EC2A5452F94C61D6358D" author="Konings A." box="[1083,1256,1711,1738]" pageId="19" pageNumber="42" refId="ref15345" refString="Konings A. 2015. Tanganyika Cichlids in their Natural Habitat. 3 rd edition. Cichlid Press, El Paso, TX." type="book" year="2015">Konings 2015</bibRefCitation>
|
||
). In contrast to Greenwood, we consider
|
||
<bibRefCitation id="EFA87CBE6033EC2A5278F93767F435AA" author="Voss J." box="[529,714,1747,1774]" pageId="19" pageNumber="42" pagination="34 - 80" refId="ref17301" refString="Voss J. 1977. Les livrees ou patrons de coloration chez les poissons Cichlides africains. Revue francaise d'Aquariologie 4: 34 - 80." type="journal article" year="1977">Voss (1977: 74)</bibRefCitation>
|
||
to show a specimen of Greenwood’s
|
||
<taxonomicName id="4C397ACC6033EC2A54EBF93061E435AA" box="[1154,1242,1747,1773]" class="Actinopterygii" family="Cichlidae" genus="Ctenochromis" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="42" phylum="Chordata" rank="species" species="polli">
|
||
<emphasis id="B94DDD5D6033EC2A54EBF93061E435AA" box="[1154,1242,1747,1773]" italics="true" pageId="19" pageNumber="42">C. polli</emphasis>
|
||
</taxonomicName>
|
||
with an ocellate spot, as does a photograph in
|
||
<bibRefCitation id="EFA87CBE6033EC2A5222F91466013456" author="Lamboj A." box="[587,831,1783,1809]" pageId="19" pageNumber="42" refId="ref15502" refString="Lamboj A. 2004. The Cichlid Fishes of Western Africa. Birgit Schmettkamp Verlag, Bornheim, Germany." type="book" year="2004">Lamboj (2004: 212)</bibRefCitation>
|
||
. There is a photograph of a specimen of
|
||
<taxonomicName id="4C397ACC6033EC2A5546F91B60A03456" box="[1327,1438,1783,1810]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="42" phylum="Chordata" rank="species" species="luluae">
|
||
<emphasis id="B94DDD5D6033EC2A5546F91B60A03456" box="[1327,1438,1783,1810]" italics="true" pageId="19" pageNumber="42">C. luluae</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<bibRefCitation id="EFA87CBE6033EC2A516DF8F864CA3472" author="Lamboj A." box="[260,500,1819,1845]" pageId="19" pageNumber="42" refId="ref15502" refString="Lamboj A. 2004. The Cichlid Fishes of Western Africa. Birgit Schmettkamp Verlag, Bornheim, Germany." type="book" year="2004">Lamboj (2004: 211)</bibRefCitation>
|
||
with ocellate eggspots. We are unaware of any unambiguous evidence of the precise egg spot characteristics of Greenwood’s
|
||
<taxonomicName id="4C397ACC6033EC2A5372F8A366E9341E" box="[795,983,1855,1882]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="42" phylum="Chordata" rank="species" species="oligacanthus">
|
||
<emphasis id="B94DDD5D6033EC2A5372F8A366E9341E" box="[795,983,1855,1882]" italics="true" pageId="19" pageNumber="42">C. oligacanthus</emphasis>
|
||
</taxonomicName>
|
||
. Hence, we follow
|
||
<bibRefCitation id="EFA87CBE6033EC2A54D4F8A360A2341D" author="Daget J. & Gosse J. P. & Teugels G. G. & Thys Van Den Audenaerde D. F. E." box="[1213,1436,1855,1882]" pageId="19" pageNumber="42" refId="ref14343" refString="Daget J., Gosse J. P., Teugels G. G. & Thys Van Den Audenaerde D. F. E. 1991. Check List of the Freshwater Fishes of Africa = Catalogue des poissons d'eau douce d'Afrique. CLOFFA 4. Musee royal pour l'Afrique centrale, Tervuren." type="book" year="1991">
|
||
Daget
|
||
<emphasis id="B94DDD5D6033EC2A5562F8A36078341E" box="[1291,1350,1855,1881]" italics="true" pageId="19" pageNumber="42">et al.</emphasis>
|
||
(1991)
|
||
</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFA87CBE6033EC2A517CF88064C63439" author="Fricke R. & Eschmeyer W. N. & van der Laan R." box="[277,504,1891,1918]" pageId="19" pageNumber="42" refId="ref14735" refString="Fricke R., Eschmeyer W. N. & van der Laan R. 2021. Eschmeyer's Catalog of Fishes: Genera, Species, References. Available from http: // researcharchive. calacademy. org / research / ichthyology / catalog / fishcatmain. asp [accessed 4 Jul. 2021]." type="url" year="2021">
|
||
Fricke
|
||
<emphasis id="B94DDD5D6033EC2A510EF887649D343A" box="[359,419,1891,1917]" italics="true" pageId="19" pageNumber="42">et al.</emphasis>
|
||
(2021)
|
||
</bibRefCitation>
|
||
in assigning Greenwood’s
|
||
<taxonomicName id="4C397ACC6033EC2A5350F88766AC343A" box="[825,914,1891,1918]" class="Actinopterygii" family="Cichlidae" genus="Ctenochromis" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="42" phylum="Chordata" rank="species" species="polli">
|
||
<emphasis id="B94DDD5D6033EC2A5350F88766AC343A" box="[825,914,1891,1918]" italics="true" pageId="19" pageNumber="42">C. polli</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C397ACC6033EC2A53C8F8876130343A" box="[929,1038,1891,1918]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="42" phylum="Chordata" rank="species" species="luluae">
|
||
<emphasis id="B94DDD5D6033EC2A53C8F8876130343A" box="[929,1038,1891,1918]" italics="true" pageId="19" pageNumber="42">C. luluae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C397ACC6033EC2A542EF887603D343A" box="[1095,1283,1891,1918]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="42" phylum="Chordata" rank="species" species="oligacanthus">
|
||
<emphasis id="B94DDD5D6033EC2A542EF887603D343A" box="[1095,1283,1891,1918]" italics="true" pageId="19" pageNumber="42">C. oligacanthus</emphasis>
|
||
</taxonomicName>
|
||
to the catchall genus
|
||
<taxonomicName id="4C397ACC6033EC2A5138F86B64C734E5" authorityName="Hilgendorf" authorityYear="1888" box="[337,505,1928,1954]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="42" phylum="Chordata" rank="genus">
|
||
<emphasis id="B94DDD5D6033EC2A5138F86B64C734E5" box="[337,505,1928,1954]" italics="true" pageId="19" pageNumber="42">Haplochromis</emphasis>
|
||
</taxonomicName>
|
||
, as
|
||
<taxonomicName id="4C397ACC6033EC2A524EF86B663334E5" authorityName="Thys van den Audenaerde" authorityYear="1964" box="[551,781,1928,1954]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="42" phylum="Chordata" rank="species" species="polli">
|
||
<emphasis id="B94DDD5D6033EC2A524EF86B663334E5" box="[551,781,1928,1954]" italics="true" pageId="19" pageNumber="42">Haplochromis polli</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C397ACC6033EC2A5372F86B612B34E5" box="[795,1045,1928,1954]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="42" phylum="Chordata" rank="species" species="luluae">
|
||
<emphasis id="B94DDD5D6033EC2A5372F86B612B34E5" box="[795,1045,1928,1954]" italics="true" pageId="19" pageNumber="42">Haplochromis luluae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C397ACC6033EC2A5425F86B60AA34E5" box="[1100,1428,1928,1954]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="42" phylum="Chordata" rank="species" species="oligacanthus">
|
||
<emphasis id="B94DDD5D6033EC2A5425F86B60AA34E5" box="[1100,1428,1928,1954]" italics="true" pageId="19" pageNumber="42">Haplochromis oligacanthus</emphasis>
|
||
</taxonomicName>
|
||
, respectively. These three taxa will require a comprehensive taxonomic evaluation. In addition, the specimens we observed of
|
||
<taxonomicName id="4C397ACC6033EC2A5276F833679434AD" authorityName="Genner & Hsu & Collins & Smith & Saxon & Shechonge & Ngatunga & Turner" authorityYear="2022" baseAuthorityName="Genner & Hsu & Collins & Smith & Saxon & Shechonge & Ngatunga & Turner" baseAuthorityYear="1894" box="[543,682,2000,2026]" class="Actinopterygii" family="Cichlidae" genus="Shura" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="42" phylum="Chordata" rank="species" species="horei" status="gen. et comb. nov.">
|
||
<emphasis id="B94DDD5D6033EC2A5276F833679434AD" box="[543,682,2000,2026]" italics="true" pageId="19" pageNumber="42">Shura horei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27E60266033EC2A52DBF83266B734AC" box="[690,905,2000,2027]" pageId="19" pageNumber="42" rank="species">gen. et comb. nov.</taxonomicNameLabel>
|
||
do not possess the “naked area on either side of the chest” characteristic of
|
||
<taxonomicName id="4C397ACC6033EC2A5223F81767D03B49" authorityName="Pfeffer" authorityYear="1893" box="[586,750,2036,2062]" class="Actinopterygii" family="Cichlidae" genus="Ctenochromis" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="42" phylum="Chordata" rank="genus">
|
||
<emphasis id="B94DDD5D6033EC2A5223F81767D03B49" box="[586,750,2036,2062]" italics="true" pageId="19" pageNumber="42">Ctenochromis</emphasis>
|
||
</taxonomicName>
|
||
, but instead have a single scaleless area at the anterior of
|
||
</paragraph>
|
||
<paragraph id="8B86014F6034EC2D508AFECC6743322B" blockId="20.[227,1437,303,364]" pageId="20" pageNumber="43">
|
||
the chest. This single scaleless area at the anterior of the chest is also shared with the phylogenetically proximate
|
||
<taxonomicName id="4C397ACC6034EC2D5108FEB26748322B" baseAuthorityName="Boulenger" baseAuthorityYear="1898" box="[353,630,337,364]" class="Actinopterygii" family="Cichlidae" genus="Gnathochromis" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="43" phylum="Chordata" rank="species" species="pfefferi">
|
||
<emphasis id="B94DDD5D6034EC2D5108FEB26748322B" box="[353,630,337,364]" italics="true" pageId="20" pageNumber="43">Gnathochromis pfefferi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |