treatments-xml/data/03/85/87/038587C40B74FFFDF3E7FE15FD45FB64.xml
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<mods:title id="6D6634F08B6A7803E2156DC8250B826E">The Canacidae of the Arabian Peninsula (Diptera: Brachycera: Carnoidea)</mods:title>
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<paragraph id="8B9336D20B74FFFDF3E7FE15FCDEFE11" blockId="12.[151,698,440,466]" box="[151,698,440,466]" pageId="12" pageNumber="501">
<heading id="D0DB81BE0B74FFFDF3E7FE15FCDEFE11" bold="true" box="[151,698,440,466]" fontSize="11" level="1" pageId="12" pageNumber="501" reason="1">
<emphasis id="B958EAC00B74FFFDF3E7FE15FCDEFE11" bold="true" box="[151,698,440,466]" pageId="12" pageNumber="501">
The
<taxonomicName id="4C2C4D510B74FFFDF3BFFE15FF4CFE11" ID-CoL="646P4" ID-ENA="1944096" box="[207,296,440,466]" class="Insecta" family="Canacidae" genus="Tethina" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAC00B74FFFDF3BFFE15FF4CFE11" bold="true" box="[207,296,440,466]" italics="true" pageId="12" pageNumber="501">Tethina</emphasis>
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species of the
<taxonomicName id="4C2C4D510B74FFFDF2AEFE15FC02FE11" box="[478,614,440,466]" class="Insecta" family="Canacidae" genus="Tethina" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="501" phylum="Arthropoda" rank="species" species="alboguttata">
<emphasis id="B958EAC00B74FFFDF2AEFE15FC02FE11" bold="true" box="[478,614,440,466]" italics="true" pageId="12" pageNumber="501">alboguttata</emphasis>
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-group
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<paragraph id="8B9336D20B74FFFDF3E7FE53FC89FB9C" blockId="12.[151,1436,510,1191]" pageId="12" pageNumber="501">
This taxonomically very difficult group of species is characterised mainly by the depigmented crossveins of the wing, which are milky white, sometimes with a white halo surrounding them (not present in the species of the Arabian Peninsula); gena generally evenly microtomentose, sometimes with a faint longitudinal subshiny stripe; male terminalia with surstylus bearing an obvious mesal lobe (visible in posterior view); female cerci with many stout, upward curved, spine-like setae (pseudacanthophorites, according to Freidberg and Beschovski, 1996). Specimens from the Arabian Peninsula are also characterised by a pale whitish to grey microtomentum that generally coats the body cuticle. Munari (2010) pointed out a consistent, peculiar character occurring in most of the Arabian species of
<taxonomicName id="4C2C4D510B74FFFDF21DFD57FFA4FCD0" box="[365,448,762,787]" class="Insecta" family="Canacidae" genus="Tethina" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAC00B74FFFDF21DFD57FFA4FCD0" box="[365,448,762,787]" italics="true" pageId="12" pageNumber="501">Tethina</emphasis>
</taxonomicName>
of the
<taxonomicName id="4C2C4D510B74FFFDF17EFD57FCEBFCD0" box="[526,655,762,787]" class="Insecta" family="Canacidae" genus="Tethina" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="501" phylum="Arthropoda" rank="species" species="alboguttata">
<emphasis id="B958EAC00B74FFFDF17EFD57FCEBFCD0" box="[526,655,762,787]" italics="true" pageId="12" pageNumber="501">alboguttata</emphasis>
</taxonomicName>
-group, that is, a characteristic black macula clearly visible at the base of wing vein R1 (fig. 30). Except for
<taxonomicName id="4C2C4D510B74FFFDF15EFCB3FDF6FCF4" authority="Munari, 2008" authorityName="Munari" authorityYear="2008" box="[558,914,798,823]" class="Insecta" family="Canacidae" genus="Tethina" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="501" phylum="Arthropoda" rank="species" species="spinigera">
<emphasis id="B958EAC00B74FFFDF15EFCB3FC96FCF5" box="[558,754,798,823]" italics="true" pageId="12" pageNumber="501">Tethina spinigera</emphasis>
Munari, 2008
</taxonomicName>
a and
<taxonomicName id="4C2C4D510B74FFFDF0A4FCB2FB40FCF4" authority="Munari, 2008" authorityName="Munari" authorityYear="2008" box="[980,1316,798,823]" class="Insecta" family="Canacidae" genus="Tethina" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="501" phylum="Arthropoda" rank="species" species="callosirostris">
<emphasis id="B958EAC00B74FFFDF0A4FCB2FAE6FCF4" box="[980,1154,798,823]" italics="true" pageId="12" pageNumber="501">T. callosirostris</emphasis>
Munari, 2008
</taxonomicName>
a, in which this character does not occur or is poorly obvious, in all the other species belonging to this group (mostly Macaronesian-Mediterranean species) this peculiarity is lacking.
<collectingCountry id="F33B76420B74FFFDF0F2FCC7FDC5FC41" box="[898,929,874,898]" name="American Samoa" pageId="12" pageNumber="501">As</collectingCountry>
for
<taxonomicName id="4C2C4D510B74FFFDF0A5FCC7FA3AFC42" box="[981,1118,874,897]" class="Insecta" family="Canacidae" genus="Tethina" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="501" phylum="Arthropoda" rank="species" species="spinigera">
<emphasis id="B958EAC00B74FFFDF0A5FCC7FA3AFC42" box="[981,1118,874,897]" italics="true" pageId="12" pageNumber="501">T. spinigera</emphasis>
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(a species with no obvious black macula on R1), it exhibits an interesting and unique chaetotactic pattern of the epandrium, consisting of short, stout spinulae rather than long setae, that allows its taxonomic placement as a possible sister species of the remaining Arabian species of the
<taxonomicName id="4C2C4D510B74FFFDF17FFC75FC8EFC32" box="[527,746,984,1009]" class="Insecta" family="Canacidae" genus="Tethina" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="501" phylum="Arthropoda" rank="species" species="alboguttata">
<emphasis id="B958EAC00B74FFFDF17FFC75FC8EFC32" box="[527,746,984,1009]" italics="true" pageId="12" pageNumber="501">Tethina alboguttata</emphasis>
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-group. On the base of vein R1,
<taxonomicName id="4C2C4D510B74FFFDF73BFC74FA9DFC32" box="[1099,1273,984,1009]" class="Insecta" family="Canacidae" genus="Tethina" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="501" phylum="Arthropoda" rank="species" species="callosirostris">
<emphasis id="B958EAC00B74FFFDF73BFC74FA9DFC32" box="[1099,1273,984,1009]" italics="true" pageId="12" pageNumber="501">T. callosirostris</emphasis>
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shows instead a simple, transverse, fine, black line, like that occurring in some Mediterranean species. The unique occurrence of a black macula (an apomorphic character state) in the species from the Arabian Peninsula would indicate this region to be a possible, common centre of speciation.
</paragraph>
<paragraph id="8B9336D20B74FFFDF3B7FBC6FD45FB64" blockId="12.[151,1436,510,1191]" pageId="12" pageNumber="501">
The species listed below are distinguished from congeners of the
<taxonomicName id="4C2C4D510B74FFFDF0B5FBC6FA22FB47" box="[965,1094,1131,1156]" class="Insecta" family="Canacidae" genus="Tethina" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="501" phylum="Arthropoda" rank="species" species="alboguttata">
<emphasis id="B958EAC00B74FFFDF0B5FBC6FA22FB47" box="[965,1094,1131,1156]" italics="true" pageId="12" pageNumber="501">alboguttata</emphasis>
</taxonomicName>
-group by the combination of diagnostic characters specified in the following treatment.
</paragraph>
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