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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.912.47719" ID-GBIF-Dataset="26396469-500e-42b8-acba-f6a22f652923" ID-PMC="PMC7040049" ID-Pensoft-Pub="1313-2970-912-65" ID-Pensoft-UUID="D1FDF644344C5B5B83E6024904E40A05" ID-PubMed="32123500" ID-ZooBank="4F361E38B2684B70944DD6B6FCC66542" ModsDocID="1313-2970-912-65" checkinTime="1581958640118" checkinUser="pensoft" docAuthor="Lee, Chi-Feng &amp; Bezdek, Jan" docDate="2020" docId="97CB21600D935312BF008112928D35F2" docLanguage="en" docName="ZooKeys 912: 65-124" docOrigin="ZooKeys 912" docSource="http://dx.doi.org/10.3897/zookeys.912.47719" docTitle="Pseudotheopea boreri Lee &amp; Bezděk 2020, sp. nov." docType="treatment" docUuid="1646E14D-D30C-4144-813C-628D6B014FF8" docUuidSource="ZooBank" docVersion="4" id="D1FDF644344C5B5B83E6024904E40A05" lastPageNumber="65" masterDocId="D1FDF644344C5B5B83E6024904E40A05" masterDocTitle="Revision of the Theopea genus group (Coleoptera, Chrysomelidae, Galerucinae), part III: Descriptions of two new genera and nine new species" masterLastPageNumber="124" masterPageNumber="65" pageNumber="65" updateTime="1668168482312" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the Theopea genus group (Coleoptera, Chrysomelidae, Galerucinae), part III: Descriptions of two new genera and nine new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lee, Chi-Feng</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bezdek, Jan</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>912</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>65</mods:start>
<mods:end>124</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.912.47719</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.912.47719</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-912-65</mods:identifier>
<mods:identifier type="ZooBank">4F361E38B2684B70944DD6B6FCC66542</mods:identifier>
<mods:identifier type="Pensoft-UUID">D1FDF644344C5B5B83E6024904E40A05</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="162147250" LSID="urn:lsid:zoobank.org:act:1646E14D-D30C-4144-813C-628D6B014FF8" httpUri="http://treatment.plazi.org/id/97CB21600D935312BF008112928D35F2" lastPageNumber="65" pageId="0" pageNumber="65">
<subSubSection pageId="0" pageNumber="65" type="nomenclature">
<paragraph pageId="0" pageNumber="65">
<taxonomicName LSID="97CB2160-0D93-5312-BF00-8112928D35F2" authority="Lee &amp; Bezděk, 2020" authorityName="Lee &amp; Bezděk" authorityYear="2020" class="Insecta" family="Chrysomelidae" genus="Pseudotheopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudotheopea boreri" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="boreri" status="sp. nov.">Pseudotheopea boreri</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="65">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of Pseudotheopea boreri sp. nov. and P. clypealis. A P. boreri sp. nov., male, dorsal view B Same, ventral view C P. boreri, female, dorsal view D P. clypealis, male, dorsal view E Same, ventral view F P. clypealis, female, dorsal view." figureDoi="10.3897/zookeys.912.47719.figure8" httpUri="https://binary.pensoft.net/fig/380663" pageId="0" pageNumber="65">Figs 8A-C</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Heads of males of Pseudotheopea boreri sp. nov. and P. clypealis. A P. boreri sp. nov., dorsofrontal view B Same, front view C P. clypealis, dorsofrontal view D Same, front view." figureDoi="10.3897/zookeys.912.47719.figure9" httpUri="https://binary.pensoft.net/fig/380664" pageId="0" pageNumber="65">; 9A, B</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Diagnostic characters of Pseudotheopea boreri sp. nov. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure10" httpUri="https://binary.pensoft.net/fig/380665" pageId="0" pageNumber="65">; 10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="types">
<paragraph pageId="0" pageNumber="65">Types.</paragraph>
<paragraph pageId="0" pageNumber="65">
Holotype ♂ (NHMB), INDIA. Meghalaya, 9 km NW of Jowai,
<geoCoordinate degrees="25" direction="north" minutes="30" orientation="latitude" precision="925" value="25.5">25°30'N</geoCoordinate>
<geoCoordinate degrees="92" direction="east" minutes="10" orientation="longitude" precision="925" value="92.166664">92°10'E</geoCoordinate>
, 1400m, 12.V.1999, leg.
<normalizedToken originalValue="Dembický">Dembicky</normalizedToken>
and
<normalizedToken originalValue="Pacholátko">Pacholatko</normalizedToken>
. Paratypes. 1♂, 2♀♀ (NHMB), same as holotype;
<emphasis bold="true" pageId="0" pageNumber="65">INDIA</emphasis>
. Assam: 1♀ (NMPC), 5 km N of Umrongso, 700 m,
<geoCoordinate degrees="25" direction="north" minutes="27" orientation="latitude" precision="925" value="25.45">25°27'N</geoCoordinate>
<geoCoordinate degrees="92" direction="east" minutes="43" orientation="longitude" precision="925" value="92.71667">92°43'E</geoCoordinate>
, 17.-25.V.1999, leg. J.
<normalizedToken originalValue="Rolčík">Rolcik</normalizedToken>
; 1♀ (NHMB), same locality, 21.V.1999, leg.
<normalizedToken originalValue="Dembický">Dembicky</normalizedToken>
and
<normalizedToken originalValue="Pacholátko”">Pacholatko&quot;</normalizedToken>
; Meghalaya: 2♀♀ (JBCB), Nokrek N.P., 3km S Daribokgiri, 1400 m,
<geoCoordinate degrees="25" direction="north" minutes="27" orientation="latitude" precision="925" value="25.45">25°27'N</geoCoordinate>
<geoCoordinate degrees="90" direction="east" minutes="19" orientation="longitude" precision="925" value="90.316666">90°19'E</geoCoordinate>
, 26.IV.1999, leg.
<normalizedToken originalValue="Rolčík">Rolcik</normalizedToken>
; 1♀ (JBCB), 8 km N of Shillong, 1200 m,
<geoCoordinate degrees="25" direction="north" minutes="38" orientation="latitude" precision="925" value="25.633333">25°38'N</geoCoordinate>
<geoCoordinate degrees="91" direction="east" minutes="54" orientation="longitude" precision="925" value="91.9">91°54'E</geoCoordinate>
, 7.-9.V.2004, leg. R.
<normalizedToken originalValue="Businský">Businsky</normalizedToken>
; 1♂, 1♀ (NHMB), same but with &quot;leg. L.
<normalizedToken originalValue="Dembický”">Dembicky&quot;</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="description">
<paragraph pageId="0" pageNumber="65">Description.</paragraph>
<paragraph pageId="0" pageNumber="65">
Length 5.6-6.6 mm, width 2.0-2.6 mm. Body color (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of Pseudotheopea boreri sp. nov. and P. clypealis. A P. boreri sp. nov., male, dorsal view B Same, ventral view C P. boreri, female, dorsal view D P. clypealis, male, dorsal view E Same, ventral view F P. clypealis, female, dorsal view." figureDoi="10.3897/zookeys.912.47719.figure8" httpUri="https://binary.pensoft.net/fig/380663" pageId="0" pageNumber="65">8A-C</figureCitation>
) golden green, but legs yellowish brown but apices of tibiae and tarsi darker; mouth parts and antennae dark brown. Frontoclypeus (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Heads of males of Pseudotheopea boreri sp. nov. and P. clypealis. A P. boreri sp. nov., dorsofrontal view B Same, front view C P. clypealis, dorsofrontal view D Same, front view." figureDoi="10.3897/zookeys.912.47719.figure9" httpUri="https://binary.pensoft.net/fig/380664" pageId="0" pageNumber="65">9A, B</figureCitation>
) transverse and deeply excavated between eyes in males, concavity as wide as interspace between eyes, margined with long hair-like setae except along basal margin; with dense, long hair-like setae at center. Antennae filiform in males, (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Diagnostic characters of Pseudotheopea boreri sp. nov. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure10" httpUri="https://binary.pensoft.net/fig/380665" pageId="0" pageNumber="65">10A</figureCitation>
), relatively broader than females, antennomeres V and VI slightly curved, length ratios of antennomeres I-XI 1.0: 0.3: 0.9: 1.2: 1.1: 1.1: 1.1: 1.0: 1.0: 0.9: 1.1, length to width ratios of antennomeres I-XI 3.2: 1.3: 3.2: 4.1: 4.4: 4.5: 4.9: 5.1: 5.2: 5.0: 5.8; filiform in females (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Diagnostic characters of Pseudotheopea boreri sp. nov. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure10" httpUri="https://binary.pensoft.net/fig/380665" pageId="0" pageNumber="65">10B</figureCitation>
), length ratios of antennomeres I-XI 1.0: 0.3: 0.7: 0.9: 0.9: 0.8: 0.8: 0.8: 0.8: 0.8: 0.9, length to width ratios of antennomeres I-XI 3.8: 2.1: 3.7: 4.8: 4.9: 4.4: 4.7: 5.1: 5.3: 4.9: 6.6. Elytra elongate, parallel-sided, 1.8-2.0
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; disc with dense, coarse punctures, arranged into longitudinal rows, with one indistinct longitudinal ridge between two longitudinal rows of punctures; with distinct convex area behind scutellum in males. Tarsomeres I of front legs swollen in males; subparallel in females. Aedeagus (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Diagnostic characters of Pseudotheopea boreri sp. nov. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure10" httpUri="https://binary.pensoft.net/fig/380665" pageId="0" pageNumber="65">10C-E</figureCitation>
) extremely slender, 7.7
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; apex with shallow notch, both apices equal in length; tectum elongate from apical 1/10 to basal 2/5; moderately curved in lateral view, angular at apical 1/4, straight from apex to apical 1/4; triangular sclerites elongate; internal sac with elongate, endophallic sclerite complex, 0.4
<normalizedToken originalValue="×">x</normalizedToken>
as long as aedeagus, composed of two sclerites, apical piece (0.6
<normalizedToken originalValue="×">x</normalizedToken>
) much shorter than basal piece. Gonocoxae (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Diagnostic characters of Pseudotheopea boreri sp. nov. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure10" httpUri="https://binary.pensoft.net/fig/380665" pageId="0" pageNumber="65">10G</figureCitation>
) elongate, both gonocoxae fused from basal 1/4 to apical 1/5; apices convergent and narrowly rounded, each gonocoxa with eight setae along lateral margin from apex to apical 1/6, some setae extremely short; lateral processes reduced. Ventrite VIII (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Diagnostic characters of Pseudotheopea boreri sp. nov. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure10" httpUri="https://binary.pensoft.net/fig/380665" pageId="0" pageNumber="65">10F</figureCitation>
) elongate and well sclerotized; disc with several long setae at sides and near apical margin, and with dense, short setae along apical margin; spiculum extremely slender. Receptacle of spermatheca (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Diagnostic characters of Pseudotheopea boreri sp. nov. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure10" httpUri="https://binary.pensoft.net/fig/380665" pageId="0" pageNumber="65">10H</figureCitation>
) tightly joined with pump, pump slender and strongly curved; proximal spermathecal duct deeply inserted into receptacle, narrow and short.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="diagnosis">
<paragraph pageId="0" pageNumber="65">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="65">
<taxonomicName authorityName="Lee &amp; Bezděk" authorityYear="2020" class="Insecta" family="Chrysomelidae" genus="Pseudotheopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudotheopea boreri" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="boreri">
<emphasis italics="true" pageId="0" pageNumber="65">Pseudotheopea boreri</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of Pseudotheopea boreri sp. nov. and P. clypealis. A P. boreri sp. nov., male, dorsal view B Same, ventral view C P. boreri, female, dorsal view D P. clypealis, male, dorsal view E Same, ventral view F P. clypealis, female, dorsal view." figureDoi="10.3897/zookeys.912.47719.figure8" httpUri="https://binary.pensoft.net/fig/380663" pageId="0" pageNumber="65">8A-C</figureCitation>
),
<taxonomicName lsidName="P. clypealis" pageId="0" pageNumber="65" rank="species" species="clypealis">
<emphasis italics="true" pageId="0" pageNumber="65">P. clypealis</emphasis>
</taxonomicName>
(Medvedev) (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of Pseudotheopea boreri sp. nov. and P. clypealis. A P. boreri sp. nov., male, dorsal view B Same, ventral view C P. boreri, female, dorsal view D P. clypealis, male, dorsal view E Same, ventral view F P. clypealis, female, dorsal view." figureDoi="10.3897/zookeys.912.47719.figure8" httpUri="https://binary.pensoft.net/fig/380663" pageId="0" pageNumber="65">8D-F</figureCitation>
),
<taxonomicName lsidName="P. hsingtzungi" pageId="0" pageNumber="65" rank="species" species="hsingtzungi">
<emphasis italics="true" pageId="0" pageNumber="65">P. hsingtzungi</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Habitus of Pseudotheopea hsingtzungi sp. nov. and P. smaragdina. A P. hsingtzungi sp. nov., male, dorsal view B P. hsingtzungi sp. nov., female, dorsal view C Same, ventral view D P. smaragdina, male, dorsal view E P. smaragdina, female, dorsal view F Same, ventral view." figureDoi="10.3897/zookeys.912.47719.figure15" httpUri="https://binary.pensoft.net/fig/380670" pageId="0" pageNumber="65">15A-C</figureCitation>
), and
<taxonomicName lsidName="P. smaragdina" pageId="0" pageNumber="65" rank="species" species="smaragdina">
<emphasis italics="true" pageId="0" pageNumber="65">P. smaragdina</emphasis>
</taxonomicName>
(Gressitt and Kimoto) (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Habitus of Pseudotheopea hsingtzungi sp. nov. and P. smaragdina. A P. hsingtzungi sp. nov., male, dorsal view B P. hsingtzungi sp. nov., female, dorsal view C Same, ventral view D P. smaragdina, male, dorsal view E P. smaragdina, female, dorsal view F Same, ventral view." figureDoi="10.3897/zookeys.912.47719.figure15" httpUri="https://binary.pensoft.net/fig/380670" pageId="0" pageNumber="65">15D-F</figureCitation>
), are characterized by their golden green coloration. They can be identified based on their distribution:
<taxonomicName lsidName="P. boreri" pageId="0" pageNumber="65" rank="species" species="boreri">
<emphasis italics="true" pageId="0" pageNumber="65">P. boreri</emphasis>
</taxonomicName>
sp. nov. from India,
<taxonomicName lsidName="P. clypealis" pageId="0" pageNumber="65" rank="species" species="clypealis">
<emphasis italics="true" pageId="0" pageNumber="65">P. clypealis</emphasis>
</taxonomicName>
from Vietnam,
<taxonomicName lsidName="P. hsingtzungi" pageId="0" pageNumber="65" rank="species" species="hsingtzungi">
<emphasis italics="true" pageId="0" pageNumber="65">P. hsingtzungi</emphasis>
</taxonomicName>
sp. nov. from Laos, and
<taxonomicName lsidName="P. smaragdina" pageId="0" pageNumber="65" rank="species" species="smaragdina">
<emphasis italics="true" pageId="0" pageNumber="65">P. smaragdina</emphasis>
</taxonomicName>
from China.
<taxonomicName authorityName="Lee &amp; Bezděk" authorityYear="2020" class="Insecta" family="Chrysomelidae" genus="Pseudotheopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudotheopea boreri" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="boreri">
<emphasis italics="true" pageId="0" pageNumber="65">Pseudotheopea boreri</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of Pseudotheopea boreri sp. nov. and P. clypealis. A P. boreri sp. nov., male, dorsal view B Same, ventral view C P. boreri, female, dorsal view D P. clypealis, male, dorsal view E Same, ventral view F P. clypealis, female, dorsal view." figureDoi="10.3897/zookeys.912.47719.figure8" httpUri="https://binary.pensoft.net/fig/380663" pageId="0" pageNumber="65">8A</figureCitation>
) is similar to
<taxonomicName lsidName="P. hsingtzungi" pageId="0" pageNumber="65" rank="species" species="hsingtzungi">
<emphasis italics="true" pageId="0" pageNumber="65">P. hsingtzungi</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Habitus of Pseudotheopea hsingtzungi sp. nov. and P. smaragdina. A P. hsingtzungi sp. nov., male, dorsal view B P. hsingtzungi sp. nov., female, dorsal view C Same, ventral view D P. smaragdina, male, dorsal view E P. smaragdina, female, dorsal view F Same, ventral view." figureDoi="10.3897/zookeys.912.47719.figure15" httpUri="https://binary.pensoft.net/fig/380670" pageId="0" pageNumber="65">15A</figureCitation>
) and
<taxonomicName lsidName="P. smaragdina" pageId="0" pageNumber="65" rank="species" species="smaragdina">
<emphasis italics="true" pageId="0" pageNumber="65">P. smaragdina</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Habitus of Pseudotheopea hsingtzungi sp. nov. and P. smaragdina. A P. hsingtzungi sp. nov., male, dorsal view B P. hsingtzungi sp. nov., female, dorsal view C Same, ventral view D P. smaragdina, male, dorsal view E P. smaragdina, female, dorsal view F Same, ventral view." figureDoi="10.3897/zookeys.912.47719.figure15" httpUri="https://binary.pensoft.net/fig/380670" pageId="0" pageNumber="65">15D</figureCitation>
) by sharing the indistinct longitudinal ridges on the elytra (convex and distinct longitudinal ridges on the elytra in
<taxonomicName lsidName="P. clypealis" pageId="0" pageNumber="65" rank="species" species="clypealis">
<emphasis italics="true" pageId="0" pageNumber="65">P. clypealis</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of Pseudotheopea boreri sp. nov. and P. clypealis. A P. boreri sp. nov., male, dorsal view B Same, ventral view C P. boreri, female, dorsal view D P. clypealis, male, dorsal view E Same, ventral view F P. clypealis, female, dorsal view." figureDoi="10.3897/zookeys.912.47719.figure8" httpUri="https://binary.pensoft.net/fig/380663" pageId="0" pageNumber="65">8D</figureCitation>
)), but differs by the presence of convex area surrounding scutellum and with reduced longitudinal ridges on the elytra in males (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of Pseudotheopea boreri sp. nov. and P. clypealis. A P. boreri sp. nov., male, dorsal view B Same, ventral view C P. boreri, female, dorsal view D P. clypealis, male, dorsal view E Same, ventral view F P. clypealis, female, dorsal view." figureDoi="10.3897/zookeys.912.47719.figure8" httpUri="https://binary.pensoft.net/fig/380663" pageId="0" pageNumber="65">8A</figureCitation>
) (without convex area surrounding scutellum on the elytra in those of others (Figs
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of Pseudotheopea boreri sp. nov. and P. clypealis. A P. boreri sp. nov., male, dorsal view B Same, ventral view C P. boreri, female, dorsal view D P. clypealis, male, dorsal view E Same, ventral view F P. clypealis, female, dorsal view." figureDoi="10.3897/zookeys.912.47719.figure8" httpUri="https://binary.pensoft.net/fig/380663" pageId="0" pageNumber="65">8D</figureCitation>
,
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Habitus of Pseudotheopea hsingtzungi sp. nov. and P. smaragdina. A P. hsingtzungi sp. nov., male, dorsal view B P. hsingtzungi sp. nov., female, dorsal view C Same, ventral view D P. smaragdina, male, dorsal view E P. smaragdina, female, dorsal view F Same, ventral view." figureDoi="10.3897/zookeys.912.47719.figure15" httpUri="https://binary.pensoft.net/fig/380670" pageId="0" pageNumber="65">15A, D</figureCitation>
)) and concavity wide between eyes and without erect processes in males (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Heads of males of Pseudotheopea boreri sp. nov. and P. clypealis. A P. boreri sp. nov., dorsofrontal view B Same, front view C P. clypealis, dorsofrontal view D Same, front view." figureDoi="10.3897/zookeys.912.47719.figure9" httpUri="https://binary.pensoft.net/fig/380664" pageId="0" pageNumber="65">9A, B</figureCitation>
) (concavity wide between eyes with one erect process in those of
<taxonomicName lsidName="P. smaragdina" pageId="0" pageNumber="65" rank="species" species="smaragdina">
<emphasis italics="true" pageId="0" pageNumber="65">P. smaragdina</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Heads of males of Pseudotheopea hsingtzungi sp. nov. and P. smaragdina. A P. hsingtzungi sp. nov., dorsofrontal view B Same, front view C P. smaragdina, dorsofrontal view D Same, front view." figureDoi="10.3897/zookeys.912.47719.figure16" httpUri="https://binary.pensoft.net/fig/380671" pageId="0" pageNumber="65">16C, D</figureCitation>
); concavity narrowed between eyes and without erect processes in those of
<taxonomicName lsidName="P. hsingtzungi" pageId="0" pageNumber="65" rank="species" species="hsingtzungi">
<emphasis italics="true" pageId="0" pageNumber="65">P. hsingtzungi</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Heads of males of Pseudotheopea hsingtzungi sp. nov. and P. smaragdina. A P. hsingtzungi sp. nov., dorsofrontal view B Same, front view C P. smaragdina, dorsofrontal view D Same, front view." figureDoi="10.3897/zookeys.912.47719.figure16" httpUri="https://binary.pensoft.net/fig/380671" pageId="0" pageNumber="65">16A, B</figureCitation>
)). In males, the internal aedeagal sac of
<taxonomicName lsidName="P. boreri" pageId="0" pageNumber="65" rank="species" species="boreri">
<emphasis italics="true" pageId="0" pageNumber="65">P. boreri</emphasis>
</taxonomicName>
sp. nov. lacks additional sclerites except the median elongate sclerite (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Diagnostic characters of Pseudotheopea boreri sp. nov. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure10" httpUri="https://binary.pensoft.net/fig/380665" pageId="0" pageNumber="65">10C, D</figureCitation>
). This structure is shared with males of
<taxonomicName lsidName="P. gressitti" pageId="0" pageNumber="65" rank="species" species="gressitti">
<emphasis italics="true" pageId="0" pageNumber="65">P. gressitti</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Diagnostic characters of Pseudotheopea gressitti sp. nov. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure14" httpUri="https://binary.pensoft.net/fig/380669" pageId="0" pageNumber="65">14C, D</figureCitation>
), and
<taxonomicName lsidName="P. costata" pageId="0" pageNumber="65" rank="species" species="costata">
<emphasis italics="true" pageId="0" pageNumber="65">P. costata</emphasis>
</taxonomicName>
(Allard),
<taxonomicName authorityName="Lee &amp; Bezděk" authorityYear="2020" class="Insecta" family="Chrysomelidae" genus="Pseudotheopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudotheopea boreri" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="boreri">
<emphasis italics="true" pageId="0" pageNumber="65">Pseudotheopea boreri</emphasis>
</taxonomicName>
sp. nov. males differ from both species in possessing a dorso-ventrally flattened aedeagus with a sclerotized ventral surface (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Diagnostic characters of Pseudotheopea aeneipennis. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure7" httpUri="https://binary.pensoft.net/fig/380662" pageId="0" pageNumber="65">7C-E</figureCitation>
) (wide aedeagus with membranous ventral surface in
<taxonomicName lsidName="P. gressitti" pageId="0" pageNumber="65" rank="species" species="gressitti">
<emphasis italics="true" pageId="0" pageNumber="65">P. gressitti</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Diagnostic characters of Pseudotheopea gressitti sp. nov. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure14" httpUri="https://binary.pensoft.net/fig/380669" pageId="0" pageNumber="65">14C-E</figureCitation>
)).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="etymology">
<paragraph pageId="0" pageNumber="65">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="65">This new species is dedicated to Matthias Borer (Curator, NHMB), who encouraged the first author to focus his research on leaf beetles.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="distribution">
<paragraph pageId="0" pageNumber="65">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="65">India.</paragraph>
<caption doi="10.3897/zookeys.912.47719.figure8" httpUri="https://binary.pensoft.net/fig/380663" pageId="0" pageNumber="65" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="65">
<emphasis bold="true" pageId="0" pageNumber="65">Figure 8.</emphasis>
Habitus of
<taxonomicName authorityName="Lee &amp; Bezděk" authorityYear="2020" class="Insecta" family="Chrysomelidae" genus="Pseudotheopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudotheopea boreri" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="boreri">
<emphasis italics="true" pageId="0" pageNumber="65">Pseudotheopea boreri</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName lsidName="P. clypealis" pageId="0" pageNumber="65" rank="species" species="clypealis">
<emphasis italics="true" pageId="0" pageNumber="65">P. clypealis</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="0" pageNumber="65">A</emphasis>
<taxonomicName lsidName="P. boreri" pageId="0" pageNumber="65" rank="species" species="boreri">
<emphasis italics="true" pageId="0" pageNumber="65">P. boreri</emphasis>
</taxonomicName>
sp. nov., male, dorsal view
<emphasis bold="true" pageId="0" pageNumber="65">B</emphasis>
Same, ventral view
<emphasis bold="true" pageId="0" pageNumber="65">C</emphasis>
<taxonomicName lsidName="P. boreri" pageId="0" pageNumber="65" rank="species" species="boreri">
<emphasis italics="true" pageId="0" pageNumber="65">P. boreri</emphasis>
</taxonomicName>
, female, dorsal view
<emphasis bold="true" pageId="0" pageNumber="65">D</emphasis>
<taxonomicName lsidName="P. clypealis" pageId="0" pageNumber="65" rank="species" species="clypealis">
<emphasis italics="true" pageId="0" pageNumber="65">P. clypealis</emphasis>
</taxonomicName>
, male, dorsal view
<emphasis bold="true" pageId="0" pageNumber="65">E</emphasis>
Same, ventral view
<emphasis bold="true" pageId="0" pageNumber="65">F</emphasis>
<taxonomicName lsidName="P. clypealis" pageId="0" pageNumber="65" rank="species" species="clypealis">
<emphasis italics="true" pageId="0" pageNumber="65">P. clypealis</emphasis>
</taxonomicName>
, female, dorsal view.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.912.47719.figure9" httpUri="https://binary.pensoft.net/fig/380664" pageId="0" pageNumber="65" start="Figure 9" startId="F9">
<paragraph pageId="0" pageNumber="65">
<emphasis bold="true" pageId="0" pageNumber="65">Figure 9.</emphasis>
Heads of males of
<taxonomicName authorityName="Lee &amp; Bezděk" authorityYear="2020" class="Insecta" family="Chrysomelidae" genus="Pseudotheopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudotheopea boreri" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="boreri">
<emphasis italics="true" pageId="0" pageNumber="65">Pseudotheopea boreri</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName lsidName="P. clypealis" pageId="0" pageNumber="65" rank="species" species="clypealis">
<emphasis italics="true" pageId="0" pageNumber="65">P. clypealis</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="0" pageNumber="65">A</emphasis>
<taxonomicName lsidName="P. boreri" pageId="0" pageNumber="65" rank="species" species="boreri">
<emphasis italics="true" pageId="0" pageNumber="65">P. boreri</emphasis>
</taxonomicName>
sp. nov., dorsofrontal view
<emphasis bold="true" pageId="0" pageNumber="65">B</emphasis>
Same, front view
<emphasis bold="true" pageId="0" pageNumber="65">C</emphasis>
<taxonomicName lsidName="P. clypealis" pageId="0" pageNumber="65" rank="species" species="clypealis">
<emphasis italics="true" pageId="0" pageNumber="65">P. clypealis</emphasis>
</taxonomicName>
, dorsofrontal view
<emphasis bold="true" pageId="0" pageNumber="65">D</emphasis>
Same, front view.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.912.47719.figure10" httpUri="https://binary.pensoft.net/fig/380665" pageId="0" pageNumber="65" start="Figure 10" startId="F10">
<paragraph pageId="0" pageNumber="65">
<emphasis bold="true" pageId="0" pageNumber="65">Figure 10.</emphasis>
Diagnostic characters of
<taxonomicName authorityName="Lee &amp; Bezděk" authorityYear="2020" class="Insecta" family="Chrysomelidae" genus="Pseudotheopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudotheopea boreri" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="boreri">
<emphasis italics="true" pageId="0" pageNumber="65">Pseudotheopea boreri</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="65">A</emphasis>
Antenna, male
<emphasis bold="true" pageId="0" pageNumber="65">B</emphasis>
Antenna, female
<emphasis bold="true" pageId="0" pageNumber="65">C</emphasis>
Aedeagus, dorsal view
<emphasis bold="true" pageId="0" pageNumber="65">D</emphasis>
Aedeagus, lateral view
<emphasis bold="true" pageId="0" pageNumber="65">E</emphasis>
Aedeagus, ventral view
<emphasis bold="true" pageId="0" pageNumber="65">F</emphasis>
Abdominal ventrite VIII
<emphasis bold="true" pageId="0" pageNumber="65">G</emphasis>
Gonocoxae
<emphasis bold="true" pageId="0" pageNumber="65">H</emphasis>
Spermatheca.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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