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<document ID-DOI="http://dx.doi.org/10.3897/jhr.70.34206" ID-GBIF-Dataset="2be0b95e-7f6b-4817-b733-c6e2a4ebd224" ID-Pensoft-Pub="1314-2607-70-41" ID-Pensoft-UUID="043FFFE5D662FFB1FF8BFF88FF94FF84" ID-ZBK="B8F3FB3E723749E79AA71AAC9EB4C787" ID-Zenodo-Dep="3269963" ID-ZooBank="B8F3FB3E723749E79AA71AAC9EB4C787" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1314-2607-70-41" ModsDocOrigin="Journal of Hymenoptera Research 70" ModsDocTitle="Revision of the south Asian amisegine genus Cladobethylus Kieffer, 1922 (Hymenoptera, Chrysididae, Amiseginae)" checkinTime="1561791832741" checkinUser="pensoft" docAuthor="Kimsey, Lynn S." docDate="2019" docId="91778FD285221ACCF602A10D2E28EDB8" docLanguage="en" docName="JourHymenoptRes 70: 41-64" docOrigin="Journal of Hymenoptera Research 70" docPubDate="2019-06-28" docSource="http://dx.doi.org/10.3897/jhr.70.34206" docTitle="Cladobethylus Kieffer 1922" docType="treatment" docVersion="4" id="043FFFE5D662FFB1FF8BFF88FF94FF84" lastPageNumber="41" masterDocId="043FFFE5D662FFB1FF8BFF88FF94FF84" masterDocTitle="Revision of the south Asian amisegine genus Cladobethylus Kieffer, 1922 (Hymenoptera, Chrysididae, Amiseginae)" masterLastPageNumber="64" masterPageNumber="41" pageNumber="41" updateTime="1678751192564" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the south Asian amisegine genus Cladobethylus Kieffer, 1922 (Hymenoptera, Chrysididae, Amiseginae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kimsey, Lynn S.</mods:namePart>
<mods:affiliation>Bohart Museum of Entomology, Department of Entomology and Nematology, University of California, Davis, California, 95616, USA</mods:affiliation>
<mods:nameIdentifier type="email">lskimsey@ucdavis.edu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Journal of Hymenoptera Research</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="pubDate">
<mods:number>2019-06-28</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>70</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>41</mods:start>
<mods:end>64</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/jhr.70.34206</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/jhr.70.34206</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2607-70-41</mods:identifier>
<mods:identifier type="ZooBank">B8F3FB3E723749E79AA71AAC9EB4C787</mods:identifier>
<mods:identifier type="Pensoft-UUID">043FFFE5D662FFB1FF8BFF88FF94FF84</mods:identifier>
<mods:identifier type="Zenodo-Dep">3269963</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="158471887" LSID="urn:lsid:plazi:treatment:91778FD285221ACCF602A10D2E28EDB8" httpUri="http://treatment.plazi.org/id/91778FD285221ACCF602A10D2E28EDB8" lastPageNumber="41" pageId="0" pageNumber="41">
<subSubSection pageId="0" pageNumber="41" type="nomenclature">
<paragraph pageId="0" pageNumber="41">
Genus
<taxonomicName ID-CoL="8NPNX" LSID="5A3CB77E-9132-5CF0-81F2-FD3BCED1B949" authority="Kieffer, 1922" authorityName="Kieffer" authorityYear="1922" class="Insecta" family="Chrysididae" genus="Cladobethylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cladobethylus" order="Hymenoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">Cladobethylus Kieffer, 1922</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="reference_group">
<paragraph pageId="0" pageNumber="41">
<taxonomicName authorityName="Kieffer" authorityYear="1922" class="Insecta" family="Chrysididae" genus="Cladobethylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cladobethylus" order="Hymenoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">Cladobethylus</taxonomicName>
Kieffer, 1922: 67. Type species:
<taxonomicName authorityName="Kieffer" authorityYear="1922" class="Insecta" family="Chrysididae" genus="Cladobethylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cladobethylus cruciger" order="Hymenoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="cruciger">Cladobethylus cruciger</taxonomicName>
Kieffer, 1922:69. Original designation.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="diagnosis">
<paragraph pageId="0" pageNumber="41">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="41">
The most distinctive feature of the genus
<taxonomicName authorityName="Kieffer" authorityYear="1922" class="Insecta" family="Chrysididae" genus="Cladobethylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cladobethylus" order="Hymenoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="41">Cladobethylus</emphasis>
</taxonomicName>
is the impunctate welt that extends along the midline of the vertex from near the midocellus to the occipital carina. This feature is only shared with species of
<taxonomicName authorityName="Kimsey" authorityYear="1986" class="Insecta" family="Chrysididae" genus="Magdalium" kingdom="Animalia" lsidName="Magdalium" order="Hymenoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="41">Magdalium</emphasis>
</taxonomicName>
Kimsey and
<taxonomicName authorityName="Kimsey" authorityYear="2005" class="Insecta" family="Chrysididae" genus="Senesega" kingdom="Animalia" lsidName="Senesega" order="Hymenoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="41">Senesega</emphasis>
</taxonomicName>
Kimsey, although it is less well developed in these genera.
<taxonomicName authorityName="Kieffer" authorityYear="1922" class="Insecta" family="Chrysididae" genus="Cladobethylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cladobethylus" order="Hymenoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="41">Cladobethylus</emphasis>
</taxonomicName>
can be distinguished from those genera by the lack of an omaulus on the mesopleuron and the usually biangulate genal area, which is evenly curved in
<taxonomicName authorityName="Kimsey" authorityYear="1986" class="Insecta" family="Chrysididae" genus="Magdalium" kingdom="Animalia" lsidName="Magdalium" order="Hymenoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="41">Magdalium</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Kimsey" authorityYear="2005" class="Insecta" family="Chrysididae" genus="Senesega" kingdom="Animalia" lsidName="Senesega" order="Hymenoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="41">Senesega</emphasis>
</taxonomicName>
. In addition,
<taxonomicName authorityName="Kieffer" authorityYear="1922" class="Insecta" family="Chrysididae" genus="Cladobethylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cladobethylus" order="Hymenoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="41">Cladobethylus</emphasis>
</taxonomicName>
species have at most a weakly defined scrobal sulcus. The scrobal sulcus in
<taxonomicName authorityName="Kimsey" authorityYear="1986" class="Insecta" family="Chrysididae" genus="Magdalium" kingdom="Animalia" lsidName="Magdalium" order="Hymenoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="41">Magdalium</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Kimsey" authorityYear="2005" class="Insecta" family="Chrysididae" genus="Senesega" kingdom="Animalia" lsidName="Senesega" order="Hymenoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="41">Senesega</emphasis>
</taxonomicName>
is well defined.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="description">
<paragraph pageId="0" pageNumber="41">Description.</paragraph>
<paragraph pageId="0" pageNumber="41">Vertex with low impunctate welt along midline from midocellus to occiput; eye with short sparse setulae; scapal basin usually with at least some cross-ridging; malar space with vertical sulcus extending from ventral eye margin to mandibular articulation; occipital carina present; posterior margin of head behind eye usually biangulate in side view; male flagellum elongate and cylindrical; female flagellum short, fusiform and usually flattened on one surface; pronotum with narrow posteromedial groove; mesopleuron without scrobal sulcus or omaulus; scutum with notauli and parapsides; both sexes fully winged, forewing Rs extended by evenly curved dark streak, costal cell widest medially, stigma large and broad, R1 not indicated, medial vein arising before cu-a; propodeum with long dorsal surface and abrupt posterior declivity, lateral propodeal angle undeveloped; hindcoxa with dorsobasal carina; tarsal claws edentate in females, with one small subbasal tooth and subbasal angle in males.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="distribution">
<paragraph pageId="0" pageNumber="41">Distribution</paragraph>
<paragraph pageId="0" pageNumber="41">
(Fig.
<figureCitation captionStart="Figures 1–3" captionStartId="F1" captionText="Figures 1 - 3. 1 Distribution map of Cladobethylus species 2, 3 Cladobethylus acehensis male 2 Lateral view 3 dorsal view." figureDoi="10.3897/jhr.70.34206.figures1-3" httpUri="https://binary.pensoft.net/fig/313608" pageId="0" pageNumber="41">1</figureCitation>
). Tropical Asia: Sri Lanka, New Guinea, Malaysia, Vietnam, Sarawak, Philippines, Indonesia (Sumatra), Borneo, South Korea and Japan.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="host">
<paragraph pageId="0" pageNumber="41">Hosts.</paragraph>
<paragraph pageId="0" pageNumber="41">
<taxonomicName class="Insecta" family="Chrysididae" genus="Cladobethylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cladobethylus insularis" order="Hymenoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="insularis">
<emphasis italics="true" pageId="0" pageNumber="41">Cladobethylus insularis</emphasis>
</taxonomicName>
was reared from eggs of the oil palm stick insect,
<taxonomicName authorityName="Lucas" authorityYear="1869" class="Insecta" family="Lonchodidae" genus="Eurycantha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eurycantha insularis" order="Phasmatodea" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="insularis">
<emphasis italics="true" pageId="0" pageNumber="41">Eurycantha insularis</emphasis>
</taxonomicName>
Lucas (
<taxonomicName authorityName="Boisduval" authorityYear="1835" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Phasmatodea" pageId="0" pageNumber="41" phylum="Arthropoda" rank="order">Phasmatodea</taxonomicName>
:
<taxonomicName class="Insecta" family="Phasmatidae" kingdom="Animalia" lsidName="" order="Phasmida" pageId="0" pageNumber="41" phylum="Arthropoda" rank="family">Phasmatidae</taxonomicName>
), in Papua New Guinea (
<bibRefCitation DOI="https://doi.org/10.3897/jhr.30.4010" author="Kimsey, LS" journalOrPublisher="Journal of Hymenoptera Research" pageId="0" pageNumber="41" pagination="19 - 28" refId="B4" refString="Kimsey, LS, Dewhurst, CF, Nyaure, S, 2012. New species of egg parasites from the Oil Palm Stick Insect (Eurycantha insularis) in Papua New Guinea (Hymenoptera: Chrysididae; Phasmatodea: Phasmatidae). Journal of Hymenoptera Research 30: 19 - 28, DOI: https://doi.org/10.3897/jhr.30.4010" title="New species of egg parasites from the Oil Palm Stick Insect (Eurycantha insularis) in Papua New Guinea (Hymenoptera: Chrysididae; Phasmatodea: Phasmatidae)." url="https://doi.org/10.3897/jhr.30.4010" volume="30" year="2012">Kimsey et al. 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="key to the species of cladobethylus">
<paragraph pageId="0" pageNumber="41">
Key to the Species of
<taxonomicName authorityName="Kieffer" authorityYear="1922" class="Insecta" family="Chrysididae" genus="Cladobethylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cladobethylus" order="Hymenoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="41">Cladobethylus</emphasis>
</taxonomicName>
1
</paragraph>
<paragraph pageId="0" pageNumber="41">
<table inLine="true" pageId="0" pageNumber="41">
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Four visible metasomal segments; flagellum tapering, broadest submedially; females</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">2</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Five visible metasomal segments; flagellum slender, parallel-sided; males</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">11</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="3" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">Females</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Metasomal tergum II with punctures 0-2 puncture diameters apart</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">3</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Metasomal tergum II with tiny punctures 5-10 or more puncture tiny diameters apart</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">6</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">3</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Flagellomeres I-III whitish</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">4</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Flagellomeres I-III dark brown</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">5</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Malar space 3.5 MOD wide; femoral apices whitish, contrasting with rest of femur; body with bluish metallic highlights, particularly dorsally; Papua New Guinea</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">
<taxonomicName lsidName="Cl. insularis" pageId="0" pageNumber="41" rank="species" species="insularis">
<emphasis italics="true" pageId="0" pageNumber="41">Cl. insularis</emphasis>
</taxonomicName>
Kimsey
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Malar space 4.5 MOD wide; femoral apices not whitish or contrasting with rest of femur; body without bluish metallic highlights; Papua New Guinea</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">
<taxonomicName lsidName="Cl. aquilus" pageId="0" pageNumber="41" rank="species" species="aquilus">
<emphasis italics="true" pageId="0" pageNumber="41">Cl. aquilus</emphasis>
</taxonomicName>
Kimsey
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">5</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Underside of head with occipital fovea on either side of occipital suture; body with bluish metallic highlights, particularly dorsally; Philippines</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">
<taxonomicName lsidName="Cl. cruciger" pageId="0" pageNumber="41" rank="species" species="cruciger">
<emphasis italics="true" pageId="0" pageNumber="41">Cl. cruciger</emphasis>
</taxonomicName>
Kieffer
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Underside of head with occipital fovea on either side of occipital suture; body with bluish metallic highlights, particularly dorsally; Thailand, Viet Nam</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">
<taxonomicName lsidName="Cl. densepunctatus" pageId="0" pageNumber="41" rank="species" species="densepunctatus">
<emphasis italics="true" pageId="0" pageNumber="41">Cl. densepunctatus</emphasis>
</taxonomicName>
Kimsey, sp. nov.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">6</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Underside of head without occipital fovea on either side of occipital suture; metapleuron and propodeal side cross-ridged or wrinkled</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">7</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Underside of head with occipital fovea on either side of occipital suture; metapleuron and propodeal side smooth without cross-ridges or wrinkles</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">8</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">7</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Malar space less than 4 MOD wide; ocular setulae less than 1 MOD long; Thailand</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">
<taxonomicName lsidName="Cl. thailandicus" pageId="0" pageNumber="41" rank="species" species="thailandicus">
<emphasis italics="true" pageId="0" pageNumber="41">Cl. thailandicus</emphasis>
</taxonomicName>
Kimsey, sp. nov.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Malar space more than 4.5 MOD wide; ocular setulae 1 MOD long or longer; body with bluish metallic highlights, particularly dorsally; Korea</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">
<taxonomicName lsidName="Cl. koreensis" pageId="0" pageNumber="41" rank="species" species="koreensis">
<emphasis italics="true" pageId="0" pageNumber="41">Cl. koreensis</emphasis>
</taxonomicName>
Kimsey, sp. nov.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">8</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
Flagellomere I less than 1.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as broad; flagellomere II broader than long
</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">9</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Flagellomere I twice as long as broad or longer; flagellomere II as long as broad or longer</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">10</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">9</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Malar space more than 4 MOD wide; flagellomere X twice as long as broad; Malaysia</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">
<taxonomicName lsidName="Cl. gilbus" pageId="0" pageNumber="41" rank="species" species="gilbus">
<emphasis italics="true" pageId="0" pageNumber="41">Cl. gilbus</emphasis>
</taxonomicName>
Kimsey
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
Malar space less than 4 MOD wide; flagellomere X 1.5-1.6
<normalizedToken originalValue="×">x</normalizedToken>
as long as broad; Borneo
</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">
<taxonomicName lsidName="Cl. sarawakensis" pageId="0" pageNumber="41" rank="species" species="sarawakensis">
<emphasis italics="true" pageId="0" pageNumber="41">Cl. sarawakensis</emphasis>
</taxonomicName>
Kimsey, sp. nov.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">10</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Legs and antennae yellow; malar space less than 4.5 MOD wide; body without metallic highlights; Borneo</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">
<taxonomicName lsidName="Cl. darlingi" pageId="0" pageNumber="41" rank="species" species="darlingi">
<emphasis italics="true" pageId="0" pageNumber="41">Cl. darlingi</emphasis>
</taxonomicName>
Kimsey, sp. nov.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Legs and antennae brown; malar space more than 4.5 MOD wide; body with metallic highlights; Japan</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">
<taxonomicName lsidName="Cl. japonicus" pageId="0" pageNumber="41" rank="species" species="japonicus">
<emphasis italics="true" pageId="0" pageNumber="41">Cl. japonicus</emphasis>
</taxonomicName>
Kimsey
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="3" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">Males</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">11</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Metasomal tergum II with punctures 0-2 puncture diameters apart</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">12</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Metasomal tergum II with tiny punctures 10 or more puncture diameters apart</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">15</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">12</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Malar space less than 3 MOD wide; post ocular distance 2 MOD wide; body length 5-7 mm; Papua New Guinea</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">
<taxonomicName lsidName="Cl. insularis" pageId="0" pageNumber="41" rank="species" species="insularis">
<emphasis italics="true" pageId="0" pageNumber="41">Cl. insularis</emphasis>
</taxonomicName>
Kimsey &amp; Dewhurst
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Malar space 3 MOD wide or wider; post ocular distance less than 2 MOD wide; body length less than 4.5 mm</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">13</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">13</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Face and malar space coarsely punctate; head dorsolaterally strongly angulate behind eye in lateral view; Thailand, Viet Nam</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">
<taxonomicName lsidName="Cl. densepunctatus" pageId="0" pageNumber="41" rank="species" species="densepunctatus">
<emphasis italics="true" pageId="0" pageNumber="41">Cl. densepunctatus</emphasis>
</taxonomicName>
Kimsey, sp. nov.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Face and malar space largely impunctate, with few scattered punctures; head dorsolaterally rounded or obtusely angulate behind eye in lateral view</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">14</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">14</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Malar space more than 3.5 MOD wide; postocular distance 1.5 MOD wide or wider; Sri Lanka.</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">
<taxonomicName lsidName="Cl. ceylonicus" pageId="0" pageNumber="41" rank="species" species="ceylonicus">
<emphasis italics="true" pageId="0" pageNumber="41">Cl. ceylonicus</emphasis>
</taxonomicName>
Krombein
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Malar space 3 MOD wide; postocular distance less than 1.5 MOD wide; Papua New Guinea</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">
<taxonomicName lsidName="Cl. aquilus" pageId="0" pageNumber="41" rank="species" species="aquilus">
<emphasis italics="true" pageId="0" pageNumber="41">Cl. aquilus</emphasis>
</taxonomicName>
Kimsey
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">15</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
Postocular distance more than 2 MOD wide; scapal basin without cross ridging; flagellomere X less than 5.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as broad; Thailand
</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">
<taxonomicName lsidName="Cl. thailandicus" pageId="0" pageNumber="41" rank="species" species="thailandicus">
<emphasis italics="true" pageId="0" pageNumber="41">Cl. thailandicus</emphasis>
</taxonomicName>
Kimsey, sp. nov.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
Postocular distance less than 1.6 MOD wide; scapal basin with zones of cross-ridging; flagellomere X more than 5.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as broad
</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">16</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">16</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Malar space less than 3 MOD wide; postocular distance less than 1 MOD wide; Malaysia.</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">
<taxonomicName lsidName="Cl. gilbus" pageId="0" pageNumber="41" rank="species" species="gilbus">
<emphasis italics="true" pageId="0" pageNumber="41">Cl. gilbus</emphasis>
</taxonomicName>
Kimsey
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Malar space 3 or more MOD wide; postocular distance 1 MOD wide or wider</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">17</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">17</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
Flagellomere I 4
<normalizedToken originalValue="×">x</normalizedToken>
as long as broad or longer; Sumatra
</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">
<taxonomicName lsidName="Cl. acehensis" pageId="0" pageNumber="41" rank="species" species="acehensis">
<emphasis italics="true" pageId="0" pageNumber="41">Cl. acehensis</emphasis>
</taxonomicName>
Kimsey, sp. nov.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
Flagellomere I 3.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as broad or shorter
</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">18</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">18</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Least interocular distance twice greatest eye width or more in front view; Borneo</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">
<taxonomicName lsidName="Cl. darlingi" pageId="0" pageNumber="41" rank="species" species="darlingi">
<emphasis italics="true" pageId="0" pageNumber="41">Cl. darlingi</emphasis>
</taxonomicName>
Kimsey, sp. nov.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
Least interocular distance 1.5
<normalizedToken originalValue="×">x</normalizedToken>
greatest eye width or less, in front view; Philippines
</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">
<taxonomicName lsidName="Cl. cruciger" pageId="0" pageNumber="41" rank="species" species="cruciger">
<emphasis italics="true" pageId="0" pageNumber="41">Cl. cruciger</emphasis>
</taxonomicName>
Kieffer
</emphasis>
</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>
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