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<document id="FA3E82597C70E563C82ED2A93BC1A517" ID-DOI="10.5281/zenodo.211095" ID-GBIF-Dataset="9173a44b-d5cd-46a9-91fc-aef58e17f540" ID-ISSN="1175-5326" ID-Zenodo-Dep="211095" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460554602230" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2012" docId="82358795FF916446FF0DF9C4FA89FDB6" docLanguage="en" docName="zt03552p068.pdf" docOrigin="Zootaxa 3552" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Trichodromeus pusillus Coiffait 1983, comb. n." docType="treatment" docVersion="9" lastPageNumber="68" masterDocId="7E0CFFEDFF916444FF9AFF86FFD2FFBF" masterDocTitle="On the taxonomic placement of Geodromicus pusillus Coiffait, 1983 (Coleoptera: Staphylinidae: Omaliinae)" masterLastPageNumber="68" masterPageNumber="66" pageNumber="66" updateTime="1698318889573" updateUser="plazi">
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<mods:title id="668AF03C2BD1B31FCCC66EE4A67B04D7">On the taxonomic placement of Geodromicus pusillus Coiffait, 1983 (Coleoptera: Staphylinidae: Omaliinae)</mods:title>
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<treatment id="82358795FF916446FF0DF9C4FA89FDB6" ID-DOI="http://doi.org/10.5281/zenodo.6181902" ID-GBIF-Taxon="119555567" ID-Zenodo-Dep="6181902" LSID="urn:lsid:plazi:treatment:82358795FF916446FF0DF9C4FA89FDB6" httpUri="http://treatment.plazi.org/id/82358795FF916446FF0DF9C4FA89FDB6" lastPageId="2" lastPageNumber="68" pageId="0" pageNumber="66">
<subSubSection id="42866508FF916444FF0DF9C4FEDFF9C3" pageId="0" pageNumber="66" type="nomenclature">
<paragraph id="0A233683FF916444FF0DF9C4FD71F9E5" blockId="0.[151,675,1602,1660]" box="[151,675,1602,1626]" pageId="0" pageNumber="66">
<heading id="516B81EFFF916444FF0DF9C4FD71F9E5" bold="true" box="[151,675,1602,1626]" fontSize="10" level="2" pageId="0" pageNumber="66" reason="2">
<emphasis id="38E8EA91FF916444FF0DF9C4FD71F9E5" bold="true" box="[151,675,1602,1626]" pageId="0" pageNumber="66">
<taxonomicName id="CD9C4D00FF916444FF0DF9C4FDE9F9E5" ID-CoL="58CBQ" authority="Coiffait, 1983" authorityName="Coiffait" authorityYear="1983" box="[151,571,1602,1626]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="66" phylum="Arthropoda" rank="species" species="pusillus" status="comb. nov.">
<emphasis id="38E8EA91FF916444FF0DF9C4FE49F9E5" bold="true" box="[151,411,1602,1626]" italics="true" pageId="0" pageNumber="66">Trichodromeus pusillus</emphasis>
<bibRefCitation id="6E0D4B72FF916444FE38F9C4FDE9F9E5" author="Coiffait" box="[418,571,1602,1626]" pageId="0" pageNumber="66" refString="Coiffait, H. (1983) Nouveaux staphylinides du Cachemire (Coleoptera). Annales de la Societe entomologique de France (n. ser.), 19 (3), 273 - 289." type="journal article" year="1983">Coiffait, 1983</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="23DB57EAFF916444FDD9F9C4FD71F9E5" box="[579,675,1602,1626]" pageId="0" pageNumber="66">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="0A233683FF916444FF0DF9E3FEDFF9C3" blockId="0.[151,675,1602,1660]" box="[151,269,1637,1660]" pageId="0" pageNumber="66">
(
<figureCitation id="92A72A06FF916444FF05F9E3FED7F9C3" box="[159,261,1637,1660]" captionStart="FIGURES 1 3" captionStartId="1.[151,269,1669,1693]" captionTargetBox="[164,1428,844,1653]" captionTargetId="figure@1.[133,1450,833,1657]" captionTargetPageId="1" captionText="FIGURES 1 3. Trichodromeus pusillus: 1 — habitus, 2 — aedeagus, ventral view, 3 — aedeagus, lateral view. Scale bars: 1: 2.0 mm, 2, 3: 0.1 mm." httpUri="https://zenodo.org/record/211096/files/figure.png" pageId="0" pageNumber="66">Figs. 13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="42866508FF916444FF0DF92EFD98F900" box="[151,586,1704,1727]" pageId="0" pageNumber="66" type="reference_group">
<paragraph id="0A233683FF916444FF0DF92EFD98F900" blockId="0.[151,586,1704,1727]" box="[151,586,1704,1727]" pageId="0" pageNumber="66">
<treatmentCitationGroup id="2A8C11ADFF916444FF0DF92EFD98F900" box="[151,586,1704,1727]" pageId="0" pageNumber="66">
<taxonomicName id="CD9C4D00FF916444FF0DF92EFD98F900" ID-CoL="4NGRK" authority="Coiffait, 1983:279" authorityName="Coiffait" authorityPageNumber="279" authorityYear="1983" box="[151,586,1704,1727]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="66" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="38E8EA91FF916444FF0DF92EFEADF900" box="[151,383,1704,1727]" italics="true" pageId="0" pageNumber="66">Geodromicus pusillus</emphasis>
<treatmentCitation id="8B3D1092FF916444FE1CF92EFD98F900" author="Coiffait" box="[390,586,1704,1727]" page="279" pageId="0" pageNumber="66" year="1983">
<bibRefCitation id="6E0D4B72FF916444FE1CF92EFDC9F900" author="Coiffait" box="[390,539,1704,1727]" pageId="0" pageNumber="66" refString="Coiffait, H. (1983) Nouveaux staphylinides du Cachemire (Coleoptera). Annales de la Societe entomologique de France (n. ser.), 19 (3), 273 - 289." type="journal article" year="1983">Coiffait, 1983</bibRefCitation>
:279
</treatmentCitation>
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="42866508FF916444FF0DF96CFD3EF84B" pageId="0" pageNumber="66" type="materials_examined">
<paragraph id="0A233683FF916444FF0DF96CFA9CF833" blockId="0.[151,1437,1770,2036]" pageId="0" pageNumber="66">
<emphasis id="38E8EA91FF916444FF0DF96CFEEFF8BD" bold="true" box="[151,317,1770,1794]" pageId="0" pageNumber="66">
<typeStatus id="D5278821FF916444FF0DF96CFF1DF8BD" box="[151,207,1770,1794]" pageId="0" pageNumber="66">Type</typeStatus>
material.
</emphasis>
<typeStatus id="D5278821FF916444FEDEF96CFE75F8BE" box="[324,423,1770,1793]" pageId="0" pageNumber="66" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="CD9C4D00FF916444FE57F96CFD64F8BE" box="[461,694,1770,1793]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="66" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="38E8EA91FF916444FE57F96CFD64F8BE" box="[461,694,1770,1793]" italics="true" pageId="0" pageNumber="66">Geodromicus pusillus</emphasis>
</taxonomicName>
, male: [North
<collectingCountry id="728B7613FF916444FCCFF96CFC5DF8BE" box="[853,911,1770,1793]" name="India" pageId="0" pageNumber="66">India</collectingCountry>
] Cachemire [Kashmir] 8-80 Rohtang Pass
<quantity id="CD649B66FF916444FAFDF96CFF7EF89B" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="0" pageNumber="66" unit="m" value="4000.0">4000 m</quantity>
G.M[eurgues].—G. L[edoux]., TYPE &lt;printed red label&gt;,
<taxonomicName id="CD9C4D00FF916444FCC5F88BFAC1F89B" authority="H. Coiffait 1981" authorityName="H. Coiffait" authorityYear="1981" box="[863,1299,1805,1828]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="66" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="38E8EA91FF916444FCC5F88BFB9EF89B" box="[863,1100,1805,1828]" italics="true" pageId="0" pageNumber="66">Geodromicus pusillus</emphasis>
H. Coiffait 1981
</taxonomicName>
(
<collectionCode id="6C8DAE46FF916444FAB0F88BFA43F89B" box="[1322,1425,1805,1828]" pageId="0" pageNumber="66">MNHNP</collectionCode>
);
<typeStatus id="D5278821FF916444FF0DF8B6FF2DF8F8" box="[151,255,1840,1863]" pageId="0" pageNumber="66" type="paratype">Paratypes</typeStatus>
of
<taxonomicName id="CD9C4D00FF916444FEBEF8B6FDDFF8F8" box="[292,525,1840,1863]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="66" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="38E8EA91FF916444FEBEF8B6FDDFF8F8" box="[292,525,1840,1863]" italics="true" pageId="0" pageNumber="66">Geodromicus pusillus</emphasis>
</taxonomicName>
,
<specimenCount id="1C9AFD0AFF916444FD86F8B6FDA3F8F8" box="[540,625,1840,1863]" pageId="0" pageNumber="66" type="male">2 males</specimenCount>
,
<specimenCount id="1C9AFD0AFF916444FDE7F8B6FD0DF8F8" box="[637,735,1840,1863]" pageId="0" pageNumber="66" type="female">1 female</specimenCount>
: same labels as the previous except label Paratype &lt;printed red label&gt; (
<collectionCode id="6C8DAE46FF916444FF72F8D4FE9CF8D6" box="[232,334,1874,1897]" pageId="0" pageNumber="66">MNHNP</collectionCode>
);
<typeStatus id="D5278821FF916444FEFBF8D4FE1BF8D6" box="[353,457,1874,1897]" pageId="0" pageNumber="66" type="paratype">Paratypes</typeStatus>
of
<taxonomicName id="CD9C4D00FF916444FE74F8D4FD05F8D6" box="[494,727,1874,1897]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="66" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="38E8EA91FF916444FE74F8D4FD05F8D6" box="[494,727,1874,1897]" italics="true" pageId="0" pageNumber="66">Geodromicus pusillus</emphasis>
</taxonomicName>
,
<specimenCount id="1C9AFD0AFF916444FD7FF8D4FCE3F8D6" box="[741,817,1874,1897]" pageId="0" pageNumber="66" type="male">1 male</specimenCount>
,
<specimenCount id="1C9AFD0AFF916444FCA6F8D4FC4FF8D6" box="[828,925,1874,1897]" pageId="0" pageNumber="66" type="female">1 female</specimenCount>
: Cachemire,
<date id="7E221043FF916444FBA8F8D4FB50F8D6" box="[1074,1154,1874,1897]" pageId="0" pageNumber="66" value="1979-08">VIII.79</date>
Pahalgam 3800 Sheshnag G.M[eurgues].—G.L[edoux]., Paratype &lt;printed red label&gt;,
<taxonomicName id="CD9C4D00FF916444FCA0F8F3FB0FF833" authority="H. Coiffait 1981" authorityName="H. Coiffait" authorityYear="1981" box="[826,1245,1909,1932]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="66" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="38E8EA91FF916444FCA0F8F3FBF0F833" box="[826,1058,1909,1932]" italics="true" pageId="0" pageNumber="66">Geodromicus pusillus</emphasis>
H. Coiffait 1981
</taxonomicName>
(
<collectionCode id="6C8DAE46FF916444FB75F8F3FA90F833" box="[1263,1346,1909,1932]" httpUri="http://grbio.org/cool/a5xg-x4h4" name="Museum d'Histoire Naturelle Perpignan" pageId="0" pageNumber="66">MHNP</collectionCode>
).
</paragraph>
<paragraph id="0A233683FF916444FF5DF811FD3EF84B" blockId="0.[151,1437,1770,2036]" pageId="0" pageNumber="66">
<emphasis id="38E8EA91FF916444FF5DF811FE62F810" bold="true" box="[199,432,1943,1967]" pageId="0" pageNumber="66">Additional material.</emphasis>
<specimenCount id="1C9AFD0AFF916444FE22F81EFDDDF810" box="[440,527,1944,1967]" pageId="0" pageNumber="66" type="male">3 males</specimenCount>
,
<specimenCount id="1C9AFD0AFF916444FD86F81EFD5BF810" box="[540,649,1944,1967]" pageId="0" pageNumber="66" type="female">3 females</specimenCount>
: Cachemire, Godasar,
<date id="7E221043FF916444FCE4F81EFC02F810" box="[894,976,1944,1967]" pageId="0" pageNumber="66" value="81-07-21">21.7.81</date>
, G. Ledoux (CT, CS);
<specimenCount id="1C9AFD0AFF916444FB52F81EFAC4F810" box="[1224,1302,1944,1967]" pageId="0" pageNumber="66" type="male">1 male</specimenCount>
: Cachemire, Lac de Gangabal,
<date id="7E221043FF916444FEC4F83CFE62F86E" box="[350,432,1978,2001]" pageId="0" pageNumber="66" value="81-07-25">25.7.81</date>
, G. Ledoux leg. (CT);
<specimenCount id="1C9AFD0AFF916444FD30F83CFCDFF86E" box="[682,781,1978,2001]" pageId="0" pageNumber="66" type="female">1 female</specimenCount>
: Cachemire, Gangabal,
<date id="7E221043FF916444FB8AF83CFBB0F86E" box="[1040,1122,1978,2001]" pageId="0" pageNumber="66" value="81-07-27">27.7.81</date>
, G. Ledoux (CT);
<specimenCount id="1C9AFD0AFF916444FAB6F83CFA4BF86E" box="[1324,1433,1978,2001]" pageId="0" pageNumber="66" type="female">2 females</specimenCount>
: Cachemire, Glacier Harmukh,
<date id="7E221043FF916444FE45F85BFDE3F84B" box="[479,561,2013,2036]" pageId="0" pageNumber="66" value="81-07-25">25.7.81</date>
, G. Ledoux (CT).
</paragraph>
</subSubSection>
<subSubSection id="42866508FF906446FF5DFF11FA89FDB6" lastPageId="2" lastPageNumber="68" pageId="1" pageNumber="67" type="description">
<paragraph id="0A233683FF906445FF5DFF11FE36FEC0" blockId="1.[151,1437,151,799]" pageId="1" pageNumber="67">
<emphasis id="38E8EA91FF906445FF5DFF11FEBEFF10" bold="true" box="[199,364,151,175]" pageId="1" pageNumber="67">Redescription.</emphasis>
Measurements in mm (n=10). Maximum width of head including eyes: 0.60.68; length of head (from base of labrum to neck constriction along the head midline): 0.30.4; length of antenna: 1.51.6; longitudinal length of eye: 0.140.16; length of temple (from posterior margin of eye to neck constriction): 0.1; length of pronotum: 0.640.66; maximum width of pronotum: 0.80.86; minimum width of pronotum: 0.70.72; sutural length of elytra (length of elytra from apex of scutellum to posterior margin of sutural angle): 1.041.16; maximum width of elytra: 1.221.34; width of abdominal segment IV: 1.21.34; length of aedeagus: 0.6. Total length (from the base of labrum to the apex of abdomen): 3.74.4.
</paragraph>
<paragraph id="0A233683FF906445FF5DFE0CFA9FFE7B" blockId="1.[151,1437,151,799]" pageId="1" pageNumber="67">
Habitus as in
<figureCitation id="92A72A06FF906445FEC6FE0CFE4FFE1E" box="[348,413,394,417]" captionStart="FIGURES 1 3" captionStartId="1.[151,269,1669,1693]" captionTargetBox="[164,1428,844,1653]" captionTargetId="figure@1.[133,1450,833,1657]" captionTargetPageId="1" captionText="FIGURES 1 3. Trichodromeus pusillus: 1 — habitus, 2 — aedeagus, ventral view, 3 — aedeagus, lateral view. Scale bars: 1: 2.0 mm, 2, 3: 0.1 mm." httpUri="https://zenodo.org/record/211096/files/figure.png" pageId="1" pageNumber="67">Fig. 1</figureCitation>
. Dark brown to black; elytra, base and hind angles of pronotum brown; mouthparts, antennae, ocelli and legs brownish yellow. Pubescence very fine accumbent, white; pubescence of head longer, decumbent.
</paragraph>
<paragraph id="0A233683FF906445FF5DFE56FE93FCA0" blockId="1.[151,1437,151,799]" pageId="1" pageNumber="67">Head large, slightly transverse (twice as wide as long), with relatively large prominent eyes; eyes 1.61.7 times longer than temples; temples not extremely wide, convex; anteocellar depression deep; ocelli small, well visible, distance between ocelli 1.5 times the distance between ocellus and posterior margin of eye. Punctation very sparse, and irregular; interstices between punctures with well-defined microsculpture; clypeus glossy, without punctures or microsculpture. Maxillary palpi long, segment II curved in basal part and expanded to apex, segment III shorter than II and with long pubescence, segment IV slightly narrower than previous, and gradually tapered to apex from the anterior third of its length. Antennae relatively short, thick, reaching 1/3 length of elytra; antennal segments II XI with the same thickness, antennomere IV slightly shorter than antennomeres III and V, antennomere XI 2.1 times longer than previous; length/ width of antennomeres are: I, 0.18 × 0.09; II III, 0.14 × 0.08; IV, 0.12 × 0.08; V VIII, 0.13 × 0.08; IX X, 0.12 × 0.08; XI, 0.26 × 0.08.</paragraph>
<caption id="5EE3660BFF906445FF0DF903FEBFF902" httpUri="https://zenodo.org/record/211096/files/figure.png" pageId="1" pageNumber="67" targetBox="[164,1428,844,1653]" targetPageId="1">
<paragraph id="0A233683FF906445FF0DF903FEBFF902" blockId="1.[151,1436,1669,1725]" pageId="1" pageNumber="67">
<emphasis id="38E8EA91FF906445FF0DF903FE90F922" bold="true" box="[151,322,1669,1693]" pageId="1" pageNumber="67">FIGURES 13.</emphasis>
<taxonomicName id="CD9C4D00FF906445FED3F903FD91F923" box="[329,579,1669,1692]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="67" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="38E8EA91FF906445FED3F903FD91F923" box="[329,579,1669,1692]" italics="true" pageId="1" pageNumber="67">Trichodromeus pusillus</emphasis>
</taxonomicName>
: 1—habitus, 2—aedeagus, ventral view, 3—aedeagus, lateral view. Scale bars: 1: 2.0 mm, 2, 3: 0.1 mm.
</paragraph>
</caption>
<paragraph id="0A233683FF906445FF5DF96DFCD6F84B" blockId="1.[151,1436,1771,2036]" pageId="1" pageNumber="67">Pronotum transverse, convex, approximately as wide as head, 1.2 times wider than long, gradually narrowed posteriorly from widest anterior part and forming distinctly right angles, with transverse impression at base and the hind angles; punctation denser than that on head, evenly distributed and deep, distance between punctures as large as diameter of 12 punctures; interstices between punctures with isodiametric microsculpture. Scutellum triangular, without punctures, with distinct isodiametric microsculpture. Elytra trapezoidal, 1.61.7 times longer than pronotum, slightly convex and slightly widened to apex, posterior angles broadly rounded; punctation regular, punctures larger and deeper than those of pronotum, distance between punctures 12 times as large as diameter of a puncture; interstices between punctures without microsculpture. Wings fully developed.</paragraph>
<paragraph id="0A233683FF936446FF5DFF1EFCFAFF6E" blockId="2.[151,1437,152,521]" pageId="2" pageNumber="68">Abdomen convex, as wide as elytra; abdominal tergites with fine punctation and with distinct isodiametric microsculpture; tergites IVV with paired tomentose patches.</paragraph>
<paragraph id="0A233683FF936446FF5DFF5BFDBCFE7B" blockId="2.[151,1437,152,521]" pageId="2" pageNumber="68">
Male. Protarsi with dilated tarsomeres, profemur widened, inner edge of all tibiae with rows of small thick spines. Sternite VIII with small rounded emargination. Aedeagus (
<figureCitation id="92A72A06FF936446FC9CFE86FC9DFEA8" box="[774,847,256,279]" captionStart="FIGURES 1 3" captionStartId="1.[151,269,1669,1693]" captionTargetBox="[164,1428,844,1653]" captionTargetId="figure@1.[133,1450,833,1657]" captionTargetPageId="1" captionText="FIGURES 1 3. Trichodromeus pusillus: 1 — habitus, 2 — aedeagus, ventral view, 3 — aedeagus, lateral view. Scale bars: 1: 2.0 mm, 2, 3: 0.1 mm." httpUri="https://zenodo.org/record/211096/files/figure.png" pageId="2" pageNumber="68">Figs. 2</figureCitation>
) relatively short and wide, parallel in median part, base of bulbus large, median lobe strongly tapered from middle towards apex; endophallus with long flagellum, forming large spiral in basal part and with parallel structures in apical part, with pair of curved apically sclerotized appendages; parameres very thin, not reaching the apex of aedeagus, with four setae on each apex. Aedeagus laterally as in
<figureCitation id="92A72A06FF936446FAA8FEEEFAA0FEC0" box="[1330,1394,360,383]" captionStart="FIGURES 1 3" captionStartId="1.[151,269,1669,1693]" captionTargetBox="[164,1428,844,1653]" captionTargetId="figure@1.[133,1450,833,1657]" captionTargetPageId="1" captionText="FIGURES 1 3. Trichodromeus pusillus: 1 — habitus, 2 — aedeagus, ventral view, 3 — aedeagus, lateral view. Scale bars: 1: 2.0 mm, 2, 3: 0.1 mm." httpUri="https://zenodo.org/record/211096/files/figure.png" pageId="2" pageNumber="68">Fig. 3</figureCitation>
. Female. Protarsi with tarsomeres less dilated, profemur not widened, and inner edge of tibia without spines. Sternite VIII without emargination, and rounded apically.
</paragraph>
<paragraph id="0A233683FF936446FF5DFE49FA89FDB6" blockId="2.[151,1437,152,521]" pageId="2" pageNumber="68">
<emphasis id="38E8EA91FF936446FF5DFE49FE77FE58" bold="true" box="[199,421,463,487]" pageId="2" pageNumber="68">Comparative notes.</emphasis>
<taxonomicName id="CD9C4D00FF936446FE36FE56FDA3FE58" authority="Coiffait" authorityName="Coiffait" box="[428,625,464,487]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="68" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="38E8EA91FF936446FE36FE56FDCBFE58" box="[428,537,464,487]" italics="true" pageId="2" pageNumber="68">T. pusillus</emphasis>
Coiffait
</taxonomicName>
differs from all known species of the genus by the significant smaller body, thick antennae, less expanded temples and by the shape and internal structures of the median lobe of the aedeagus.
</paragraph>
</subSubSection>
</treatment>
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