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<document id="AA5C0D7870F866EB518434C24FD82B0A" ID-CLB-Dataset="31852" ID-DOI="10.11646/zootaxa.4335.1.1" ID-GBIF-Dataset="f2ad3cb1-0bad-4be1-9e36-2170a6a08aaf" ID-ISSN="1175-5326" ID-Zenodo-Dep="1010014" ID-ZooBank="3429592D-23A8-4EBF-8412-BD57B3BC4314" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507877826645" checkinUser="plazi" docAuthor="Azevedo, Celso O." docDate="2017" docId="3B3687C07C6B9B3CFF67F6D29D8AF8BD" docLanguage="en" docName="zootaxa.4335.1.1.pdf" docOrigin="Zootaxa 4335 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dissomphalus strepsus Azevedo 2001" docType="treatment" docVersion="15" lastPageNumber="25" masterDocId="C70FFFB87C739B24FFF0F23F9F65FFDF" masterDocTitle="Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species" masterLastPageNumber="72" masterPageNumber="1" pageNumber="25" updateTime="1698455438235" updateUser="plazi">
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<mods:title id="DE605635AAAC62767E6AD5E04B6AACF6">Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species</mods:title>
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<treatment id="3B3687C07C6B9B3CFF67F6D29D8AF8BD" ID-DOI="http://doi.org/10.5281/zenodo.5020340" ID-GBIF-Taxon="135435233" ID-Zenodo-Dep="5020340" LSID="urn:lsid:plazi:treatment:3B3687C07C6B9B3CFF67F6D29D8AF8BD" httpUri="http://treatment.plazi.org/id/3B3687C07C6B9B3CFF67F6D29D8AF8BD" lastPageNumber="25" pageId="24" pageNumber="25">
<subSubSection id="FB85655D7C6B9B3CFF67F6D29E27FAF7" pageId="24" pageNumber="25" type="nomenclature">
<paragraph id="B32036D67C6B9B3CFF67F6D29D00FAD8" blockId="24.[151,613,1261,1320]" box="[151,613,1261,1287]" pageId="24" pageNumber="25">
<heading id="E86881BA7C6B9B3CFF67F6D29D00FAD8" bold="true" box="[151,613,1261,1287]" fontSize="11" level="1" pageId="24" pageNumber="25" reason="1">
<taxonomicName id="749F4D557C6B9B3CFF67F6D29D00FAD8" authority="Azevedo, 2001" authorityName="Azevedo" authorityYear="2001" box="[151,613,1261,1287]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="strepsus">
<emphasis id="81EBEAC47C6B9B3CFF67F6D29D00FAD8" bold="true" box="[151,613,1261,1287]" pageId="24" pageNumber="25">
<emphasis id="81EBEAC47C6B9B3CFF67F6D29ECEFAD8" bold="true" box="[151,427,1261,1287]" italics="true" pageId="24" pageNumber="25">Dissomphalus strepsus</emphasis>
<bibRefCitation id="D70E4B277C6B9B3CFE42F6D29D00FAD8" author="Azevedo" box="[434,613,1261,1287]" pageId="24" pageNumber="25" refString="Azevedo, C. O. (2001) Systematics of the Neotropical Dissomphalus Ashmead (Hymenoptera, Bethylidae) of the bicavatus group. Revista Brasileira de Entomologia, 45, 173 - 205." type="journal article" year="2001">Azevedo, 2001</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="B32036D67C6B9B3CFF67F7309E27FAF7" blockId="24.[151,613,1261,1320]" box="[151,322,1295,1320]" pageId="24" pageNumber="25">
(
<figureCitation id="2BA42A537C6B9B3CFF6FF7309E5CFAF7" box="[159,313,1295,1320]" captionStart="FIGURES 107 122" captionStartId="23.[151,264,1604,1627]" captionTargetBox="[157,1430,208,1562]" captionTargetId="figure@23.[151,1436,195,1583]" captionTargetPageId="23" captionText="FIGURES 107 122. Dissomphalus spp., ♂, genitalia, except otherwise indicated. 107 109. D. geniculatus 107. Dorsal view. 108. Ventral view. 109. Outer lobe and basal process of aedeagal dorsal body, lateral view. 110 112. D. intradentatus 110. Outer lobe of aedeagal dorsal body, lateral view. 111. Dorsal view. 112. Ventral view. 113 114. D. latus 113. Dorsal view. 114. Ventral view. 115 118. D. piscicercus 115. Dorsal view. 116. Ventral view. 117. apex of aedeagal ventral ramus, ventral view 118. Outer lobe of aedeagal dorsal body, lateral view. 119 121. D. strepsus 119. Dorsal view. 120. Outer lobe of aedeagal dorsal body, dorsal view. 121. Ventral view. 122. D. angulatus Dorsal view. Scale bars = 250 µm." httpUri="https://zenodo.org/record/1010036/files/figure.png" pageId="24" pageNumber="25">Figs 119121</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C6B9B3CFF67F7679BB4FAB1" box="[151,1233,1368,1390]" pageId="24" pageNumber="25" type="reference_group">
<paragraph id="B32036D67C6B9B3CFF67F7679BB4FAB1" blockId="24.[151,1233,1368,1390]" box="[151,1233,1368,1390]" pageId="24" pageNumber="25">
<materialsCitation id="03F73C8B7C6B9B3CFF67F7679BB4FAB1" ID-GBIF-Occurrence="1671715961" box="[151,1233,1368,1390]" collectionCode="EMUS" country="Costa Rica" location="Costa Rica" pageId="24" pageNumber="25" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<treatmentCitationGroup id="938F11F87C6B9B3CFF67F7679BB4FAB1" box="[151,1233,1368,1390]" pageId="24" pageNumber="25">
<taxonomicName id="749F4D557C6B9B3CFF67F7679D36FAB1" authority="Azevedo, 2001: 175" authorityName="Azevedo" authorityPageNumber="175" authorityYear="2001" box="[151,595,1368,1390]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="strepsus">
<emphasis id="81EBEAC47C6B9B3CFF67F7679E1AFAB1" box="[151,383,1368,1390]" italics="true" pageId="24" pageNumber="25">Dissomphalus strepsus</emphasis>
<treatmentCitation id="323E10C77C6B9B3CFE74F7679D36FAB1" author="Azevedo" box="[388,595,1368,1390]" page="175" pageId="24" pageNumber="25" year="2001">
<bibRefCitation id="D70E4B277C6B9B3CFE74F7679D44FAB1" author="Azevedo" box="[388,545,1368,1390]" pageId="24" pageNumber="25" refString="Azevedo, C. O. (2001) Systematics of the Neotropical Dissomphalus Ashmead (Hymenoptera, Bethylidae) of the bicavatus group. Revista Brasileira de Entomologia, 45, 173 - 205." type="journal article" year="2001">Azevedo, 2001</bibRefCitation>
: 175
</treatmentCitation>
</taxonomicName>
, 200201 (
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<typeStatus id="6C2488747C6B9B3CFD2EF7679C52FAB1" box="[734,823,1368,1390]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
from
<collectingCountry id="CB8876467C6B9B3CFC85F7679C86FAB1" box="[885,995,1368,1390]" name="Costa Rica" pageId="24" pageNumber="25">Costa Rica</collectingCountry>
<collectionCode id="D58EAE137C6B9B3CFC1AF7679B56FAB1" box="[1002,1075,1368,1390]" pageId="24" pageNumber="25">EMUS</collectionCode>
, figs 175179).
</treatmentCitationGroup>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C6B9B3CFF67F7A59FAEF970" pageId="24" pageNumber="25" type="diagnosis">
<paragraph id="B32036D67C6B9B3CFF67F7A59FAEF970" blockId="24.[151,1437,1434,1891]" pageId="24" pageNumber="25">
<emphasis id="81EBEAC47C6B9B3CFF67F7A59E74FA6C" bold="true" box="[151,273,1434,1459]" pageId="24" pageNumber="25">Diagnosis.</emphasis>
Mandible bidentate. Clypeus trapezoidal. Tergal process with pair of antero-lateral depressions, ovoid, deep, large with small pit covered by tuft of long setae. Hypopygium with posterior margin somewhat concave. Genitalia: paramere with apex oblique and concave; aedeagal dorsal ramus longer than aedeagal dorsal body, parallel, apical third directed downward and apex forward, sharpened and abruptly angled; inner margin with series of small teeth; aedeagal dorsal body with three pairs of apical lobes; outer pair with two rami, dorsal ramus parallel with apex narrow and angled downward, ventral ramus arched inward and dorsal ramus with apices nearly touching each other; inner pairs stout, membranose with rounded apex. Apodeme not extending beyond genital ring.
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C6B9B3CFF37F4859AF8F928" pageId="24" pageNumber="25" type="discussion">
<paragraph id="B32036D67C6B9B3CFF37F4859AF8F928" blockId="24.[151,1437,1434,1891]" pageId="24" pageNumber="25">
<emphasis id="81EBEAC47C6B9B3CFF37F4859E5EF90C" bold="true" box="[199,315,1722,1747]" pageId="24" pageNumber="25">Remarks.</emphasis>
This species is known to
<collectingCountry id="CB8876467C6B9B3CFD91F4849DB9F90C" box="[609,732,1722,1747]" name="Costa Rica" pageId="24" pageNumber="25">
<collectingRegion id="715BF8347C6B9B3CFD91F4849DC4F90C" box="[609,673,1723,1747]" country="New Zealand" name="West Coast" pageId="24" pageNumber="25">Costa</collectingRegion>
Rica
</collectingCountry>
and
<collectingRegion id="715BF8347C6B9B3CFCE4F4849C08F90D" box="[788,877,1723,1746]" country="Panama" name="Panama" pageId="24" pageNumber="25">
<collectingCountry id="CB8876467C6B9B3CFCE4F4849C08F90D" box="[788,877,1723,1746]" name="Panama" pageId="24" pageNumber="25">Panama</collectingCountry>
</collectingRegion>
(
<bibRefCitation id="D70E4B277C6B9B3CFC8CF4859B40F90C" author="Azevedo" box="[892,1061,1722,1747]" pageId="24" pageNumber="25" refString="Azevedo, C. O. (2001) Systematics of the Neotropical Dissomphalus Ashmead (Hymenoptera, Bethylidae) of the bicavatus group. Revista Brasileira de Entomologia, 45, 173 - 205." type="journal article" year="2001">Azevedo 2001</bibRefCitation>
). Now one specimen is added. In this series the specimens are quite similar to those previously studied, without any variation that worths to mention.
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C6B9B3CFF37F53D9AF0F8E0" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="B32036D67C6B9B3CFF37F53D9AF0F8E0" blockId="24.[151,1437,1434,1891]" pageId="24" pageNumber="25">
<materialsCitation id="03F73C8B7C6B9B3CFF37F53D9E2EF8E0" ID-GBIF-Occurrence="1671715960" collectingDate="1993-07-13" collectingDateMax="1993-07-31" collectingDateMin="1993-07-13" collectionCode="EMUS" collectorName="F. D. Parker" country="Costa Rica" location="Rio Naranjo" municipality="Material" pageId="24" pageNumber="25" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste" typeStatus="holotype">
<emphasis id="81EBEAC47C6B9B3CFF37F53D9ED4F8C4" bold="true" box="[199,433,1794,1819]" pageId="24" pageNumber="25">
<collectingMunicipality id="5344ACAC7C6B9B3CFF37F53D9E4AF8C4" box="[199,303,1794,1819]" pageId="24" pageNumber="25">Material</collectingMunicipality>
examined.
</emphasis>
<typeStatus id="6C2488747C6B9B3CFE4AF53D9D40F8C4" box="[442,549,1794,1819]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
,
<specimenCount id="A599FD5F7C6B9B3CFDC1F53C9D2DF8C5" box="[561,584,1795,1818]" pageId="24" pageNumber="25" type="male"></specimenCount>
,
<collectingCountry id="CB8876467C6B9B3CFDA6F53C9D98F8C4" box="[598,765,1795,1819]" name="Costa Rica" pageId="24" pageNumber="25">COSTA RICA</collectingCountry>
,
<collectingRegion id="715BF8347C6B9B3CFCF9F53C9CEAF8C5" box="[777,911,1795,1818]" country="Costa Rica" name="Guanacaste" pageId="24" pageNumber="25">
<emphasis id="81EBEAC47C6B9B3CFCF9F53C9CEAF8C5" box="[777,911,1795,1818]" italics="true" pageId="24" pageNumber="25">Guanacaste</emphasis>
</collectingRegion>
:
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, SE,
<location id="B640600D7C6B9B3CFBE0F53D9BFBF8C4" LSID="urn:lsid:plazi:treatment:3B3687C07C6B9B3CFF67F6D29D8AF8BD:B640600D7C6B9B3CFBE0F53D9BFBF8C4" box="[1040,1182,1794,1819]" country="Costa Rica" municipality="Material" name="Rio Naranjo" pageId="24" pageNumber="25" stateProvince="Guanacaste">Rio Naranjo</location>
,
<collectingDate id="D765E9FE7C6B9B3CFB5AF53C9A05F8C4" box="[1194,1376,1795,1819]" pageId="24" pageNumber="25" value="1993-07-13" valueMax="1993-07-31" valueMin="1993-07-13">
<date id="C72110167C6B9B3CFB5AF53C9A05F8C4" box="[1194,1376,1795,1819]" pageId="24" pageNumber="25" value="1993-07-13" valueMax="1993-07-31" valueMin="1993-07-13">1331.VII.1993</date>
</collectingDate>
,
<collectorName id="1E6A53007C6B9B3CFA9DF53C9F85F8E0" pageId="24" pageNumber="25">F.D. Parker</collectorName>
(
<collectionCode id="D58EAE137C6B9B3CFF1FF5189E26F8E0" box="[239,323,1831,1855]" pageId="24" pageNumber="25">EMUS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="03F73C8B7C6B9B3CFEA6F5189AF5F8E0" ID-GBIF-Occurrence="1671715964" box="[342,1424,1830,1855]" collectingDate="1996-06-04" collectingDateMax="1996-06-07" collectingDateMin="1996-06-04" collectionCode="UFES" collectorName="J. Ashe &amp; R. Brooks" country="Panama" elevation="1250" location="Cana Serrania de Pirre" pageId="24" pageNumber="25" specimenCount="1" specimenCount-male="1" stateProvince="Darien" typeStatus="holotype">
<collectingCountry id="CB8876467C6B9B3CFEA6F5189EB5F8E0" box="[342,464,1831,1855]" name="Panama" pageId="24" pageNumber="25">PANAMA</collectingCountry>
,
<collectingRegion id="715BF8347C6B9B3CFE2BF5189D4FF8E1" box="[475,554,1831,1854]" country="Panama" name="Darien" pageId="24" pageNumber="25">
<emphasis id="81EBEAC47C6B9B3CFE2BF5189D4FF8E1" box="[475,554,1831,1854]" italics="true" pageId="24" pageNumber="25">Darién</emphasis>
</collectingRegion>
:
<specimenCount id="A599FD5F7C6B9B3CFDC9F5189D39F8E0" box="[569,604,1831,1855]" pageId="24" pageNumber="25" type="male">1♂</specimenCount>
<location id="B640600D7C6B9B3CFD93F5189C07F8E0" LSID="urn:lsid:plazi:treatment:3B3687C07C6B9B3CFF67F6D29D8AF8BD:B640600D7C6B9B3CFD93F5189C07F8E0" box="[611,866,1830,1855]" country="Panama" name="Cana Serrania de Pirre" pageId="24" pageNumber="25" stateProvince="Darien">Cana Serrania de Pirre</location>
,
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<quantity id="74679B337C6B9B3CFC9DF5199CD8F8E0" box="[877,957,1830,1855]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="24" pageNumber="25" unit="m" value="1250.0">1250m</quantity>
</elevation>
,
<collectingDate id="D765E9FE7C6B9B3CFC39F5199B37F8E0" box="[969,1106,1830,1855]" pageId="24" pageNumber="25" value="1996-06-04" valueMax="1996-06-07" valueMin="1996-06-04">
<date id="C72110167C6B9B3CFC39F5199B37F8E0" box="[969,1106,1830,1855]" pageId="24" pageNumber="25" value="1996-06-04" valueMax="1996-06-07" valueMin="1996-06-04">47.vi.1996</date>
</collectingDate>
,
<collectorName id="1E6A53007C6B9B3CFBADF5189BD7F8E0" box="[1117,1202,1830,1855]" pageId="24" pageNumber="25">J. Ashe</collectorName>
,
<collectorName id="1E6A53007C6B9B3CFB4DF5189A4AF8E0" box="[1213,1327,1830,1855]" pageId="24" pageNumber="25">R. Brooks</collectorName>
(
<collectionCode id="D58EAE137C6B9B3CFACEF5189AEDF8E0" box="[1342,1416,1831,1855]" country="Brazil" httpUri="http://grbio.org/cool/n6nq-8nqh" name="Universidade Federal do Espirito Santo" pageId="24" pageNumber="25">UFES</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
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<paragraph id="B32036D67C6B9B3CFF37F5759D8AF8BD" blockId="24.[151,1437,1434,1891]" box="[199,751,1866,1891]" pageId="24" pageNumber="25">
<emphasis id="81EBEAC47C6B9B3CFF37F5759E3CF8BC" bold="true" box="[199,345,1866,1891]" pageId="24" pageNumber="25">Distribution</emphasis>
(
<figureCitation id="2BA42A537C6B9B3CFE98F5759EAEF8BC" box="[360,459,1866,1891]" captionStart="FIGURES 234 243" captionStartId="67.[151,264,1856,1879]" captionTargetBox="[174,1439,203,1822]" captionTargetId="figure@67.[162,1447,193,1827]" captionTargetPageId="67" captionText="FIGURES 234 243. Known distribution of Dissomphalus in Panama. 234. D. galeatus, D. geniculatus and D. geortus sp. nov.. 235. D. gilvipes, D. ibocius sp. nov. and D. ibrastus sp. nov.. 236. D. guttus and D. ilarius sp. nov.. 237. D. infissus, D. intradentatus and D. iozus sp. nov.. 238. D. isortus sp. nov., D. isteus sp. nov. and D. latus. 239. D. persontaus, D. pilus and D. pronus. 240. D. plaumanni and D. politus. 241. D. piscicercus, D. ramosus and D. raziogus sp. nov.. 242. D. punctatus, D. strepsus and D. subdeformis. 243. D. racarus sp. nov., D. strabus and D. tisinus sp. nov.." httpUri="https://zenodo.org/record/1010060/files/figure.png" pageId="24" pageNumber="25">Fig. 242</figureCitation>
).
<collectingCountry id="CB8876467C6B9B3CFE2EF5749D3DF8BC" box="[478,600,1866,1891]" name="Costa Rica" pageId="24" pageNumber="25">
<collectingRegion id="715BF8347C6B9B3CFE2EF5749D7BF8BC" box="[478,542,1867,1891]" country="New Zealand" name="West Coast" pageId="24" pageNumber="25">Costa</collectingRegion>
Rica
</collectingCountry>
and
<collectingRegion id="715BF8347C6B9B3CFD7EF5749D8FF8BD" box="[654,746,1867,1890]" country="Panama" name="Panama" pageId="24" pageNumber="25">
<collectingCountry id="CB8876467C6B9B3CFD7EF5749D8FF8BD" box="[654,746,1867,1890]" name="Panama" pageId="24" pageNumber="25">Panama</collectingCountry>
</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>