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<document id="D4C188525461A3656112355153BFB896" ID-ISSN="1175-5326" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629742400862" checkinUser="felipe" docAuthor="Fery, Hans" docDate="2011" docId="03FC87EDE30DFFFCFF300168D729EEDB" docLanguage="en" docName="zt03050p054.pdf" docOrigin="Zootaxa 3050" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Agabus puetzi Fery 2011, sp. n." docType="treatment" docVersion="4" lastPageNumber="44" masterDocId="FFC5FF95E30CFFFBFFA7077AD277EF5C" masterDocTitle="Agabus (Acatodes) puetzi sp. n., a new species of the confinis-group from the Qinghai-Tibet Plateau (Coleoptera, Dytiscidae, Agabini)" masterLastPageNumber="54" masterPageNumber="37" pageNumber="38" updateTime="1699105378929" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="A17875805DF98B8D13E1098E29E8477C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D9D95AFBE37E070E70844890BA18B1A2">
<mods:title id="F55649D3DADA434C944E39CB2D6E1B5C">Agabus (Acatodes) puetzi sp. n., a new species of the confinis-group from the Qinghai-Tibet Plateau (Coleoptera, Dytiscidae, Agabini)</mods:title>
</mods:titleInfo>
<mods:name id="44028406BB4A361E42556250F7CE2762" type="personal">
<mods:role id="99C644931B727C0BC5FCF10D328D55F6">
<mods:roleTerm id="BDFFDF5C3B2B9F31D6735FCAE140DBA6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="25704726D71A33DA17D8B92C0667DBEB">Fery, Hans</mods:namePart>
</mods:name>
<mods:typeOfResource id="73E1CE368D8C081601DDDE73AAEA8805">text</mods:typeOfResource>
<mods:relatedItem id="E1FC4443ED033BEE471C19CCA0A27ED9" type="host">
<mods:titleInfo id="EB186AD0FE7D77FF538F4457D56DEEAD">
<mods:title id="9620171ABE65A0901BC9C94C8187660B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="43BB3F3780AA537641DBD9D9591CF52D">
<mods:date id="2B5F9E0EAD7E3757881F379A2E4C3DA8">2011</mods:date>
<mods:detail id="9802903D9DA5A31BA53815DA1E28524C" type="pubDate">
<mods:number id="847E4A219E696CA5D00AB3EA7858D569">2011-10-06</mods:number>
</mods:detail>
<mods:detail id="6EF38D904E88BE1FC5857D440E5C97E3" type="volume">
<mods:number id="19AA93F996C5A9699D77AE507CB617D6">3050</mods:number>
</mods:detail>
<mods:extent id="7747F7F879E4D18A7EC2B6F24D69A1B2" unit="page">
<mods:start id="2F570793D934EA2226AFD63F888582FF">37</mods:start>
<mods:end id="FF7D0EB61408FD3F423AB50464CCB245">54</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="BCCFD79219A76025CB9316EF4F8C8BBF">journal article</mods:classification>
<mods:identifier id="E028D5071EA0FFC9F92414957049C6D5" type="ISSN">1175-5326</mods:identifier>
</mods:mods>
<treatment id="03FC87EDE30DFFFCFF300168D729EEDB" ID-DOI="http://doi.org/10.5281/zenodo.5282941" ID-Zenodo-Dep="5282941" LSID="urn:lsid:plazi:treatment:03FC87EDE30DFFFCFF300168D729EEDB" httpUri="http://treatment.plazi.org/id/03FC87EDE30DFFFCFF300168D729EEDB" lastPageId="7" lastPageNumber="44" pageId="1" pageNumber="38">
<subSubSection id="C34F6570E30DFFFAFF300168D3FDE970" box="[151,394,1554,1580]" pageId="1" pageNumber="38" type="nomenclature">
<paragraph id="8BEA36FBE30DFFFAFF300168D3FDE970" blockId="1.[151,394,1554,1580]" box="[151,394,1554,1580]" pageId="1" pageNumber="38">
<heading id="D0A28197E30DFFFAFF300168D3FDE970" bold="true" box="[151,394,1554,1580]" fontSize="11" level="1" pageId="1" pageNumber="38" reason="1">
<emphasis id="B921EAE9E30DFFFAFF300168D3FDE970" bold="true" box="[151,394,1554,1580]" pageId="1" pageNumber="38">
<taxonomicName id="4C554D78E30DFFFAFF300168D337E970" ID-CoL="8LZ74" authority="Fery, 2011" authorityName="Fery" authorityYear="2011" box="[151,320,1554,1580]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="38" phylum="Arthropoda" rank="species" species="puetzi" status="sp. nov.">
<emphasis id="B921EAE9E30DFFFAFF300168D337E970" bold="true" box="[151,320,1554,1580]" italics="true" pageId="1" pageNumber="38">Agabus puetzi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2125792E30DFFFAFEEF0168D3FDE970" box="[328,394,1554,1580]" pageId="1" pageNumber="38" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34F6570E30DFFFAFF300123D1C7E8CD" pageId="1" pageNumber="38" type="materials_examined">
<paragraph id="8BEA36FBE30DFFFAFF300123D3ACE9E5" blockId="1.[151,1436,1625,2009]" pageId="1" pageNumber="38">
<emphasis id="B921EAE9E30DFFFAFF300123D34FE92E" bold="true" box="[151,312,1625,1650]" inLineHeading="true" pageId="1" pageNumber="38" reason="1">Type locality.</emphasis>
<materialsCitation id="3B3D3CA6E30DFFFAFE9A0123D3A0E9E5" country="China" county="Batang County" elevation="4490" latitude="30.299" location="Shaluli Mountains" longLatPrecision="6" longitude="99.5527" municipality="Ganzi Tibetan Autonomic Prefecture" pageId="1" pageNumber="38" specimenCount="1" stateProvince="Sichuan">
<collectingCountry id="F342766BE30DFFFAFE9A0123D3F4E92D" box="[317,387,1625,1649]" name="China" pageId="1" pageNumber="38">China</collectingCountry>
,
<collectingRegion id="4991F819E30DFFFAFE280123D021E92D" box="[399,598,1625,1649]" country="China" name="Sichuan" pageId="1" pageNumber="38">Sichuan province</collectingRegion>
,
<collectingMunicipality id="6B8EAC81E30DFFFAFDC50123D673E92D" box="[610,1028,1625,1649]" pageId="1" pageNumber="38">Ganzi Tibetan Autonomic Prefecture</collectingMunicipality>
,
<collectingCounty id="628B4E77E30DFFFAFBB70120D6CDE92D" box="[1040,1210,1625,1650]" pageId="1" pageNumber="38">Batang County</collectingCounty>
,
<location id="8E8A6020E30DFFFAFB610123D7EFE92D" LSID="urn:lsid:plazi:treatment:03FC87EDE30DFFFCFF300168D729EEDB:8E8A6020E30DFFFAFB610123D7EFE92D" box="[1222,1432,1625,1649]" country="China" county="Batang County" latitude="30.299" longLatPrecision="6" longitude="99.5527" municipality="Ganzi Tibetan Autonomic Prefecture" name="Shaluli Mountains" pageId="1" pageNumber="38" stateProvince="Sichuan">Shaluli Mountains</location>
,
<location id="8E8A6020E30DFFFAFF300107D331E9C9" LSID="urn:lsid:plazi:treatment:03FC87EDE30DFFFCFF300168D729EEDB:8E8A6020E30DFFFAFF300107D331E9C9" box="[151,326,1661,1685]" country="China" county="Batang County" latitude="30.299" longLatPrecision="6" longitude="99.5527" municipality="Ganzi Tibetan Autonomic Prefecture" name="Haizi mountain" pageId="1" pageNumber="38" stateProvince="Sichuan">Haizi mountain</location>
,
<locationDeviation id="9A582FD9E30DFFFAFEF50107D069E9CA" box="[338,542,1661,1686]" location="Batang" pageId="1" pageNumber="38">
<quantity id="4CAD9B1EE30DFFFAFEF50107D3EEE9C9" box="[338,409,1661,1685]" metricMagnitude="4" metricUnit="m" metricValue="5.7" pageId="1" pageNumber="38" unit="km" value="57.0">57 km</quantity>
NE Batang
</locationDeviation>
, route 318, km 3219, border of &quot;Sister Lakes&quot;; ca.
<geoCoordinate id="EE61503CE30DFFFAFBF00107D6B0E9C9" box="[1111,1223,1661,1685]" degrees="30.2990" direction="north" orientation="latitude" pageId="1" pageNumber="38" precision="5" value="30.299">30.2990N</geoCoordinate>
<geoCoordinate id="EE61503CE30DFFFAFB690107D74BE9C9" box="[1230,1340,1661,1685]" degrees="99.5527" direction="east" orientation="longitude" pageId="1" pageNumber="38" precision="5" value="99.5527">99.5527E</geoCoordinate>
, altitude ca.
<quantity id="4CAD9B1EE30DFFFAFF1901DBD365E9E5" box="[190,274,1697,1721]" metricMagnitude="3" metricUnit="m" metricValue="4.49" pageId="1" pageNumber="38" unit="m" value="4490.0">
<elevation id="0078D1C8E30DFFFAFF1901DBD365E9E5" box="[190,274,1697,1721]" metricMagnitude="3" metricUnit="m" metricValue="4.49" pageId="1" pageNumber="38" unit="m" value="4490.0">4490 m</elevation>
</quantity>
(see
<figureCitation id="136E2A7EE30DFFFAFEEA01DBD3B9E9E5" box="[333,462,1697,1721]" captionStart="FIGURES 40–41" captionStartId="14.[151,264,1623,1646]" captionTargetBox="[264,1322,193,1602]" captionTargetId="figure-226@14.[264,1322,193,1602]" captionTargetPageId="14" captionText="FIGURES 40–41. Sister Lakes with Haizi mountain in the background; collecting site of Agabus puetzi sp. n. indicated by the arrow (for more details see text)." pageId="1" pageNumber="38">Figs 40, 41</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEA36FBE30DFFFAFF6001BFD1C7E8CD" blockId="1.[151,1436,1625,2009]" pageId="1" pageNumber="38">
<materialsCitation id="3B3D3CA6E30DFFFAFF6001BFD059E879" collectingDate="1999-07-03" collectionCode="CPE" collectorName="A. Putz" country="China" county="Batang Co." determinerName="Fery" elevation="4500" location="Water Pools" municipality="Shaluli Shan" pageId="1" pageNumber="38" specimenCount="1" specimenCount-male="1" stateProvince="Sichuan" typeStatus="holotype">
<emphasis id="B921EAE9E30DFFFAFF6001BFD34CE981" bold="true" box="[199,315,1733,1757]" inLineHeading="true" pageId="1" pageNumber="38" reason="1">
<typeStatus id="54EE8859E30DFFFAFF6001BFD341E981" box="[199,310,1733,1757]" pageId="1" pageNumber="38" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="9D53FD72E30DFFFAFEE601BCD321E981" box="[321,342,1734,1757]" pageId="1" pageNumber="38" type="male">♂</specimenCount>
, &quot;
<collectingCountry id="F342766BE30DFFFAFECA01BFD3C2E981" box="[365,437,1733,1757]" name="China" pageId="1" pageNumber="38">China</collectingCountry>
, Prov.
<collectingRegion id="4991F819E30DFFFAFDA601BFD029E981" box="[513,606,1733,1757]" country="China" name="Sichuan" pageId="1" pageNumber="38">Sichuan</collectingRegion>
, Ganzi Tibetian Auton. Pref.,
<collectingCounty id="628B4E77E30DFFFAFC0A01BCD65AE981" box="[941,1069,1733,1758]" pageId="1" pageNumber="38">Batang Co.</collectingCounty>
,
<collectingMunicipality id="6B8EAC81E30DFFFAFB9E01BFD6BCE981" box="[1081,1227,1733,1757]" pageId="1" pageNumber="38">Shaluli Shan</collectingMunicipality>
&quot;, &quot;
<location id="8E8A6020E30DFFFAFB4F01BFD702E981" LSID="urn:lsid:plazi:treatment:03FC87EDE30DFFFCFF300168D729EEDB:8E8A6020E30DFFFAFB4F01BFD702E981" box="[1256,1397,1733,1757]" country="China" county="Batang Co." municipality="Shaluli Shan" name="Water Pools" pageId="1" pageNumber="38" stateProvince="Sichuan">Water Pools</location>
,
<locationDeviation id="9A582FD9E30DFFFAFA2701BFD337E85E" location="Batang" pageId="1" pageNumber="38">
<quantity id="4CAD9B1EE30DFFFAFA2701BFD2CCE85D" metricMagnitude="4" metricUnit="m" metricValue="5.7" pageId="1" pageNumber="38" unit="km" value="57.0">57 km</quantity>
NE Batang
</locationDeviation>
, route 318, km 3219,
<quantity id="4CAD9B1EE30DFFFAFD900193D0F9E85D" box="[567,654,1769,1793]" metricMagnitude="3" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="38" unit="m" value="4500.0">
<elevation id="0078D1C8E30DFFFAFD900193D0F9E85D" box="[567,654,1769,1793]" metricMagnitude="3" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="38" unit="m" value="4500.0">4500 m</elevation>
</quantity>
,
<date id="FFEB103BE30DFFFAFD3D0193D160E85D" box="[666,791,1769,1793]" pageId="1" pageNumber="38" value="1999-07-03">
<collectingDate id="EFAFE9D3E30DFFFAFD3D0193D160E85D" box="[666,791,1769,1793]" pageId="1" pageNumber="38" value="1999-07-03">3.VII.1999</collectingDate>
</date>
, leg.
<collectorName id="26A0532DE30DFFFAFCF30193D1DDE85D" box="[852,938,1769,1793]" pageId="1" pageNumber="38">A. Pütz</collectorName>
&quot; [printed], &quot;
<typeStatus id="54EE8859E30DFFFAFB930193D6D6E85D" box="[1076,1185,1769,1793]" pageId="1" pageNumber="38" type="holotype">Holotype</typeStatus>
,
<taxonomicName id="4C554D78E30DFFFAFB0A0193D739E85D" authority="Fery, 2011" authorityName="Fery" authorityYear="2011" box="[1197,1358,1769,1793]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="38" phylum="Arthropoda" rank="species" species="puetzi" status="sp. nov.">Agabus puetzi</taxonomicName>
<emphasis id="B921EAE9E30DFFFAFAF20193D7E3E85D" bold="true" box="[1365,1428,1769,1793]" pageId="1" pageNumber="38">
<taxonomicNameLabel id="A2125792E30DFFFAFAF20193D7E3E85D" box="[1365,1428,1769,1793]" pageId="1" pageNumber="38" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<determinerName id="7E36DA3FE30DFFFAFF300074D2BDE87A" box="[151,202,1806,1830]" pageId="1" pageNumber="38">Fery</determinerName>
det. 2011&quot; [red, printed] (
<collectionCode id="ED44AE3EE30DFFFAFE4B0077D052E879" box="[492,549,1805,1829]" pageId="1" pageNumber="38">CPE</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B3D3CA6E30DFFFAFD9F0077D1DBE8CD" collectingDate="1999-07-03" collectionCode="CHF, CPE" collectorName="A. Putz" country="China" county="Batang Co." determinerName="Fery" elevation="4500" location="Water Pools" municipality="Shaluli Shan" pageId="1" pageNumber="38" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Sichuan" typeStatus="paratype">
<emphasis id="B921EAE9E30DFFFAFD9F0077D0C7E879" bold="true" box="[568,688,1805,1829]" pageId="1" pageNumber="38">
<typeStatus id="54EE8859E30DFFFAFD9F0077D0C7E879" box="[568,688,1805,1829]" pageId="1" pageNumber="38" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="9D53FD72E30DFFFAFD190077D09FE879" box="[702,744,1805,1829]" pageId="1" pageNumber="38" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D53FD72E30DFFFAFD510077D143E879" box="[758,820,1805,1829]" pageId="1" pageNumber="38" type="female">2 ♀♀</specimenCount>
, with same label data as holotype, but male with entire text on only a single label; all
<typeStatus id="54EE8859E30DFFFAFE540048D029E816" box="[499,606,1842,1866]" pageId="1" pageNumber="38" type="paratype">paratypes</typeStatus>
with the respective red
<typeStatus id="54EE8859E30DFFFAFCD50048D1A5E816" box="[882,978,1842,1866]" pageId="1" pageNumber="38" type="paratype">paratype</typeStatus>
label (
<collectionCode id="ED44AE3EE30DFFFAFB86004BD62DE815" box="[1057,1114,1841,1865]" pageId="1" pageNumber="38">CHF</collectionCode>
,
<collectionCode id="ED44AE3EE30DFFFAFBC0004BD6E8E815" box="[1127,1183,1841,1865]" pageId="1" pageNumber="38">CPE</collectionCode>
).
<emphasis id="B921EAE9E30DFFFAFB14004BD681E815" bold="true" box="[1203,1270,1841,1865]" pageId="1" pageNumber="38">Notes</emphasis>
: The
<typeStatus id="54EE8859E30DFFFAFA9D004BD7EBE815" box="[1338,1436,1841,1865]" pageId="1" pageNumber="38" type="holotype">holotype</typeStatus>
lacks the left mid leg; the male
<typeStatus id="54EE8859E30DFFFAFE52002CD022E832" box="[501,597,1878,1902]" pageId="1" pageNumber="38" type="paratype">paratype</typeStatus>
has the last three right protarsomeres and all left metatarsomeres disarticulated, and the sucker palettes of the protarsomeres are in part removed
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34F6570E30DFFFDFF6000E7D67DE95B" lastPageId="6" lastPageNumber="43" pageId="1" pageNumber="38" type="description">
<paragraph id="8BEA36FBE30DFFF9FF6000E7D665EF88" blockId="1.[151,1436,1625,2009]" lastBlockId="2.[151,1435,151,212]" lastPageId="2" lastPageNumber="39" pageId="1" pageNumber="38">
<emphasis id="B921EAE9E30DFFFAFF6000E7D3A2E8E9" box="[199,469,1949,1974]" inLineHeading="true" pageId="1" pageNumber="38" reason="1">
<emphasis id="B921EAE9E30DFFFAFF6000E7D326E8EA" bold="true" box="[199,337,1949,1974]" pageId="1" pageNumber="38">Description</emphasis>
(
<typeStatus id="54EE8859E30DFFFAFEC600E7D3BDE8E9" box="[353,458,1949,1973]" pageId="1" pageNumber="38" type="holotype">holotype</typeStatus>
).
</emphasis>
<emphasis id="B921EAE9E30DFFFAFE7B00E7D04CE8EA" bold="true" box="[476,571,1949,1974]" pageId="1" pageNumber="38">Habitus</emphasis>
rather flat, in dorsal view elongate oval, rather narrow (TL/MW: 2.08), with a distinct discontinuity in outline between pronotum and elytra (
<figureCitation id="136E2A7EE30DFFFAFCD900BBD1B5E885" box="[894,962,1985,2009]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1026,1049]" captionTargetBox="[293,1307,268,1013]" captionTargetId="figure-80@2.[283,1311,267,1013]" captionTargetPageId="2" captionText="FIGURES 1–2. Agabus puetzi sp. n.: habitus of: (1) male (paratype; disarticulated appendages re-arranged), (2) female." pageId="1" pageNumber="38">Fig. 1</figureCitation>
). Broadest near middle of elytral length. Head and pronotum black, elytra dark brown; entire surface shiny although provided with a primary reticulation of larger meshes and in large parts with an additional secondary microreticulation.
</paragraph>
<caption id="DF2A6673E30EFFF9FF300378D72DEB44" box="[151,1370,1026,1049]" pageId="2" pageNumber="39" startId="2.[151,264,1026,1049]" targetBox="[293,1307,268,1013]" targetPageId="2">
<paragraph id="8BEA36FBE30EFFF9FF300378D72DEB44" blockId="2.[151,1370,1026,1049]" box="[151,1370,1026,1049]" pageId="2" pageNumber="39">
<emphasis id="B921EAE9E30EFFF9FF300378D34DEB45" bold="true" box="[151,314,1026,1049]" pageId="2" pageNumber="39">FIGURES 1–2.</emphasis>
<taxonomicName id="4C554D78E30EFFF9FEE70379D3B8EB44" authority="Fery, 2011" authorityName="Fery" authorityYear="2011" box="[320,463,1027,1049]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="puetzi" status="sp. nov.">
<emphasis id="B921EAE9E30EFFF9FEE70379D3B8EB44" box="[320,463,1027,1049]" italics="true" pageId="2" pageNumber="39">Agabus puetzi</emphasis>
</taxonomicName>
<emphasis id="B921EAE9E30EFFF9FE720379D079EB44" bold="true" box="[469,526,1027,1048]" pageId="2" pageNumber="39">
<taxonomicNameLabel id="A2125792E30EFFF9FE720379D079EB44" box="[469,526,1027,1048]" pageId="2" pageNumber="39" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
: habitus of: (1) male (paratype; disarticulated appendages re-arranged), (2) female.
</paragraph>
</caption>
<caption id="DF2A6673E30EFFF9FF300017D663E8DF" box="[151,1044,1901,1924]" pageId="2" pageNumber="39" startId="2.[151,264,1901,1924]">
<paragraph id="8BEA36FBE30EFFF9FF300017D663E8DF" blockId="2.[151,1044,1901,1924]" box="[151,1044,1901,1924]" pageId="2" pageNumber="39">
<emphasis id="B921EAE9E30EFFF9FF300017D34DE8D8" bold="true" box="[151,314,1901,1924]" pageId="2" pageNumber="39">FIGURES 3–4.</emphasis>
<taxonomicName id="4C554D78E30EFFF9FEE70017D3B8E8DF" authority="Fery, 2011" authorityName="Fery" authorityYear="2011" box="[320,463,1901,1923]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="puetzi" status="sp. nov.">
<emphasis id="B921EAE9E30EFFF9FEE70017D3B8E8DF" box="[320,463,1901,1923]" italics="true" pageId="2" pageNumber="39">Agabus puetzi</emphasis>
</taxonomicName>
<emphasis id="B921EAE9E30EFFF9FE720014D079E8DF" bold="true" box="[469,526,1902,1923]" pageId="2" pageNumber="39">
<taxonomicNameLabel id="A2125792E30EFFF9FE720014D079E8DF" box="[469,526,1902,1923]" pageId="2" pageNumber="39" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
: left protibia and protarsus of (3) male, (4) female.
</paragraph>
</caption>
<caption id="DF2A6673E30FFFF8FF30015DD65DE900" pageId="3" pageNumber="40" startId="3.[151,263,1575,1598]" targetBox="[264,1323,193,1554]" targetPageId="3">
<paragraph id="8BEA36FBE30FFFF8FF30015DD65DE900" blockId="3.[151,1436,1575,1628]" pageId="3" pageNumber="40">
<emphasis id="B921EAE9E30FFFF8FF30015DD34CE962" bold="true" box="[151,315,1575,1598]" pageId="3" pageNumber="40">FIGURES 5–9.</emphasis>
<taxonomicName id="4C554D78E30FFFF8FEE4015DD3A4E961" authority="Fery, 2011" authorityName="Fery" authorityYear="2011" box="[323,467,1575,1597]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="puetzi" status="sp. nov.">
<emphasis id="B921EAE9E30FFFF8FEE4015DD3A4E961" box="[323,467,1575,1597]" italics="true" pageId="3" pageNumber="40">Agabus puetzi</emphasis>
</taxonomicName>
<emphasis id="B921EAE9E30FFFF8FE7D015DD063E960" bold="true" box="[474,532,1575,1596]" pageId="3" pageNumber="40">
<taxonomicNameLabel id="A2125792E30FFFF8FE7D015DD063E960" box="[474,532,1575,1596]" pageId="3" pageNumber="40" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
: median lobe of aedeagus: (5) right side, (6) left side, (7) detail of apex (weak swelling indicated by the arrow); (8) left paramere; female genitalia: (9) (a) gonocoxa, (b) tergite 9.
</paragraph>
</caption>
<paragraph id="8BEA36FBE30FFFFFFF6001F3D341EF88" blockId="3.[151,1436,1672,2019]" lastBlockId="4.[151,1437,151,2034]" lastPageId="4" lastPageNumber="41" pageId="3" pageNumber="40">
<emphasis id="B921EAE9E30FFFF8FF6001F3D370E9FD" bold="true" box="[199,263,1673,1697]" pageId="3" pageNumber="40">Head</emphasis>
black, except brown anterior margin and two indistinct round dark brown spots on vertex; labrum lighter brownish, contrasting distinctly with black clypeus. Bead on anterior margin of clypeus practically complete, consisting mostly of narrow, elongate, closely spaced and contiguous transverse punctures; bead slightly deviating backwards medially, interrupted over a very small distance, gap not much longer than diameter of meshes of primary reticulation on clypeus behind. Head behind clypeus with an oblique clypeal fovea on each side, consisting of about five impressed punctures and a distinct fine oblique groove stretching until lateral margin of clypeus; with several broad shallow pits and a few coarse punctures along inner side of eyes. Entire surface reticulated with not deeply impressed, more or less polygonal meshes, variable in shape and not isodiametric; strongly longitudinally stretched near eyes. Meshes behind clypeal foveae with microreticulation, but this becoming indistinct towards eyes and vertex. Punctation on head very sparse, some punctures inside meshes, others at intersections of reticulation lines and very few along these lines. Setae very sparse, confined to sides of anterior bead, clypeal foveae and pits near eyes.
</paragraph>
<paragraph id="8BEA36FBE308FFFFFF60079AD642EDB7" blockId="4.[151,1437,151,2034]" pageId="4" pageNumber="41">
<emphasis id="B921EAE9E308FFFFFF60079AD337EFA4" bold="true" box="[199,320,224,248]" pageId="4" pageNumber="41">Pronotum</emphasis>
completely black. In dorsal view sides slightly curved in posterior two thirds, anteriorly almost straight. Posterior angles slightly obtuse, very narrowly rounded. Lateral bead well marked, vanishing far before anterior angle. Anterior margin not completely beaded, only behind eyes with a short line, near anterior angles and rest of margin without bead. Sublaterally with a shallow oblique impression; here and also near lateral bead with a few punctures. Near each posterior angle with another, but smaller and even shallower impression. Somewhat behind anterior margin with rather broad row of very irregularly arranged coarse punctures, these slightly sparser medially, but row not interrupted; another row of punctures before posterior margin, but punctures less numerous and slightly smaller; row broadly interrupted medially, but sublaterally not interrupted again, at least punctures somewhat sparser.
<collectingRegion id="4991F819E308FFFFFECA0687D3C0ED49" box="[365,439,509,533]" country="France" name="Centre" pageId="4" pageNumber="41">Centre</collectingRegion>
of pronotal disc with a rather long distinct single scratch. Primary reticulation similar to that on centre of head; anterior margin narrowly without reticulation, between this margin and anterior row of punctures meshes rather small; secondary microreticulation near sides and in particular at anterior angles. Punctation at intersections of reticulation lines and along these lines similar to that on head, several meshes also with such rather distinct punctures inside, but most meshes with several additional very shallow and almost imperceptible punctures; these becoming slightly more impressed posteriorly. Setae long, but very sparse, present only in a few coarser punctures near anterior and posterior margin and in sublateral impressions.
</paragraph>
<paragraph id="8BEA36FBE308FFFFFF600582D64DEA5F" blockId="4.[151,1437,151,2034]" pageId="4" pageNumber="41">
<emphasis id="B921EAE9E308FFFFFF600582D364EC4C" bold="true" box="[199,275,760,784]" pageId="4" pageNumber="41">Elytra</emphasis>
dark brown; behind base, near lateral margins and in anterior half along suture diffusely delimited lighter brownish; scutellum, elytral base centrally and suture itself black. In dorsal view sides in anterior two thirds slightly rounded, not continuing curvature of pronotum, more strongly curved behind, conjointly rounded at apex. Margin of elytra in lateral view almost straight in anterior two thirds, not considerably ascending to shoulders. Lateral rim distinctly narrower than that of pronotum. Each elytron with a large subapical oblique impression, margin at apex thus appearing somewhat vaulted. Each elytron with four irregular puncture lines, starting short behind elytral base; in second third of elytral length with a few additional punctures between lines, in apical fourth lines becoming very irregular; sutural puncture lines only indicated by very few small punctures behind first third of elytral length; diameter of punctures much smaller than that of primary meshes. Directly beside lateral elytral rim with a fifth puncture line. Primary reticulation similar to that on head and pronotum; meshes smaller near suture and in subapical impressions. Secondary microreticulation present on entire surface, slightly more distinct in subapical impressions (reticulation similar to that in
<figureCitation id="136E2A7EE308FFFFFDDE03FAD171EBC4" box="[633,774,1152,1176]" captionStart="FIGURES 5–9" captionStartId="3.[151,263,1575,1598]" captionTargetBox="[264,1323,193,1554]" captionTargetId="figure-215@3.[264,1323,193,1554]" captionTargetPageId="3" captionText="FIGURES 5–9. Agabus puetzi sp. n.: median lobe of aedeagus: (5) right side, (6) left side, (7) detail of apex (weak swelling indicated by the arrow); (8) left paramere; female genitalia: (9) (a) gonocoxa, (b) tergite 9." pageId="4" pageNumber="41">
Figs 5 and
<quantity id="4CAD9B1EE308FFFFFD5F03FAD171EBC4" box="[760,774,1152,1176]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="4" pageNumber="41" unit="in" value="6.0">6</quantity>
</figureCitation>
in
<bibRefCitation id="EFC44B0AE308FFFFFC8B03FBD671EBC4" author="Larson" box="[812,1030,1152,1176]" firstAuthor="Larson" page="905" pageId="4" pageNumber="41" pagination="861 - 919" refId="ref10370" refString="Larson, D. J. (1989) Revision of North American Agabus Leach (Coleoptera: Dytiscidae): introduction, key to species groups, and classification of the ambiguus -, tristis -, and arcticus - groups. The Canadian Entomologist, 121, 861 - 919." type="journal article" year="1989">Larson (1989: 905)</bibRefCitation>
for
<taxonomicName id="4C554D78E308FFFFFB9003FAD353EBE0" authority="(Say, 1823))" authorityName="" baseAuthorityName="Say" baseAuthorityYear="1823" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="ambiguus" subGenus="Gaurodytes">
<emphasis id="B921EAE9E308FFFFFB9003FAD6FCEBC4" box="[1079,1163,1152,1176]" italics="true" pageId="4" pageNumber="41">Agabus</emphasis>
(
<emphasis id="B921EAE9E308FFFFFB3B03FAD769EBC4" box="[1180,1310,1152,1176]" italics="true" pageId="4" pageNumber="41">Gaurodytes</emphasis>
)
<emphasis id="B921EAE9E308FFFFFA8803FAD7EBEBC4" box="[1327,1436,1152,1176]" italics="true" pageId="4" pageNumber="41">ambiguus</emphasis>
(Say, 1823))
</taxonomicName>
. Setation very sparse and indistinct, more or less only perceptible with oblique illumination; restricted to punctures within lines. Setae often rather long (up to seven meshes in length), in particular in lateral puncture line and apical third of sublateral puncture line (also up to seven meshes in length).
</paragraph>
<paragraph id="8BEA36FBE308FFFFFF60026AD726E8AE" blockId="4.[151,1437,151,2034]" pageId="4" pageNumber="41">
<emphasis id="B921EAE9E308FFFFFF60026AD3F7EA74" bold="true" box="[199,384,1295,1320]" pageId="4" pageNumber="41">Ventral surface</emphasis>
with major parts black; apex of metacoxal processes and posterior margins of third to sixth abdominal ventrites transparently brownish; epipleura black in broad anterior half, near shoulders somewhat transparent dark brownish, in small posterior half dark brown. Head with most mouthparts brown, mentum, gula and genae black. Legs and trochanters reddish brown, femora largely darkened medially. Antennomeres reddish brown, progressively darkened distally, beginning with fifth; second antennomere short, third longer than second and fourth, fourth only indistinctly shorter than fifth. Maxillar and labial palpi reddish brown, apical palpomeres diffusely darkened medially. Before procoxae outline of prosternum and prosternal process in lateral view more or less straight, slightly descending backwards, almost imperceptibly impressed directly before procoxae, but process not stepped down from plane of prosternum; lanceolate blade bent upwards shortly behind middle of procoxae. Blade broad, rather flat, evenly vaulted in cross-section; sides not flattened; bead being only a thin but distinct line; apex bluntly pointed, not acuminate (
<figureCitation id="136E2A7EE308FFFFFD91010ED0FEE9D0" box="[566,649,1652,1676]" captionStart="FIGURE 10" captionStartId="5.[151,250,1577,1600]" captionTargetBox="[325,1261,483,1553]" captionTargetId="figure-325@5.[325,1261,483,1553]" captionTargetPageId="5" captionText="FIGURE 10. Ventral surface (in part) of Agabus puetzi sp. n.: a: blade of prosternal process; b: apex of metasternal process; c: furrow on anteromedial metasternal process; d: gula; e: bead of prosternal process; f: furrow of mesosternum (scale bar ca. 1.0 mm)." pageId="4" pageNumber="41">Fig. 10</figureCitation>
). Process between procoxae indistinctly tectiform in cross-section, but midline of convexity broadly rounded; before procoxae evenly and broadly rounded in cross-section. Distance between mesocoxae rather small, smaller than width of process between procoxae. Anteromedial metasternal process short, bluntly pointed anteriorly, not reaching forwards beyond middle of mesocoxae and not reaching mesosternum (
<figureCitation id="136E2A7EE308FFFFFF510079D33DE847" box="[246,330,1795,1819]" captionStart="FIGURE 10" captionStartId="5.[151,250,1577,1600]" captionTargetBox="[325,1261,483,1553]" captionTargetId="figure-325@5.[325,1261,483,1553]" captionTargetPageId="5" captionText="FIGURE 10. Ventral surface (in part) of Agabus puetzi sp. n.: a: blade of prosternal process; b: apex of metasternal process; c: furrow on anteromedial metasternal process; d: gula; e: bead of prosternal process; f: furrow of mesosternum (scale bar ca. 1.0 mm)." pageId="4" pageNumber="41">Fig. 10</figureCitation>
). Mesosternum before mesocoxae with a deep broad furrow for reception of prosternal blade (
<figureCitation id="136E2A7EE308FFFFFAC90079D2C1E863" captionStart="FIGURE 10" captionStartId="5.[151,250,1577,1600]" captionTargetBox="[325,1261,483,1553]" captionTargetId="figure-325@5.[325,1261,483,1553]" captionTargetPageId="5" captionText="FIGURE 10. Ventral surface (in part) of Agabus puetzi sp. n.: a: blade of prosternal process; b: apex of metasternal process; c: furrow on anteromedial metasternal process; d: gula; e: bead of prosternal process; f: furrow of mesosternum (scale bar ca. 1.0 mm)." pageId="4" pageNumber="41">Fig. 10</figureCitation>
). Apex of blade just contacting apex of anteromedial metasternal process, but not reaching further backwards than middle of mesocoxae, and, thus, not received in furrow on anteromedial metasternal process. This latter furrow being only a very shallow longitudinal impression, reaching more or less from apex of anteromedial metasternal process backwards until shortly before level of hind margin of mesocoxae (
<figureCitation id="136E2A7EE308FFFFFBC000E8D6B6E8F6" box="[1127,1217,1938,1962]" captionStart="FIGURE 10" captionStartId="5.[151,250,1577,1600]" captionTargetBox="[325,1261,483,1553]" captionTargetId="figure-325@5.[325,1261,483,1553]" captionTargetPageId="5" captionText="FIGURE 10. Ventral surface (in part) of Agabus puetzi sp. n.: a: blade of prosternal process; b: apex of metasternal process; c: furrow on anteromedial metasternal process; d: gula; e: bead of prosternal process; f: furrow of mesosternum (scale bar ca. 1.0 mm)." pageId="4" pageNumber="41">Fig. 10</figureCitation>
). Lateral lobes of metasternum (&quot;metasternal wings&quot;) rather broad (WC/WS: 2.26). Lines of metacoxal processes diverging anteriorly, reaching posterior margin of metasternum. Posterior margin of last abdominal ventrite evenly rounded.
</paragraph>
<paragraph id="8BEA36FBE309FFFEFF6007EDD37CEEF7" blockId="5.[151,1436,151,427]" pageId="5" pageNumber="42">Almost entire surface microreticulated, weakly shiny, but not matt; only midline of metasternum narrowly smooth, same with apical lobes of metacoxal processes. Meshes of primary reticulation only indicated on last abdominal ventrite, otherwise absent or incomplete on abdominal ventrites, in part replaced by some stronger impressed wrinkles. Lateral lobes of metasternum with some weak longitudinal rugae, sides of metacoxal plates with transverse rugae. Prosternum rugosely sculptured anteromedially; punctation of ventral surface very sparse and weak. Some fine punctures on blade of prosternal process, others at wrinkles of metacoxal plates; third to fifth abdominal ventrites with indistinct transverse row of setose punctures, last ventrite posteriorly with some stronger punctures.</paragraph>
<caption id="DF2A6673E309FFFEFF300153D2B9E921" pageId="5" pageNumber="42" startId="5.[151,250,1577,1600]" targetBox="[325,1261,483,1553]" targetPageId="5">
<paragraph id="8BEA36FBE309FFFEFF300153D2B9E921" blockId="5.[151,1436,1577,1661]" pageId="5" pageNumber="42">
<emphasis id="B921EAE9E309FFFEFF300153D368E91C" bold="true" box="[151,287,1577,1600]" pageId="5" pageNumber="42">FIGURE 10.</emphasis>
Ventral surface (in part) of
<taxonomicName id="4C554D78E309FFFEFD9F0150D0B0E91C" authority="Fery, 2011" authorityName="Fery" authorityYear="2011" box="[568,711,1578,1600]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="puetzi" status="sp. nov.">
<emphasis id="B921EAE9E309FFFEFD9F0150D0B0E91C" box="[568,711,1578,1600]" italics="true" pageId="5" pageNumber="42">Agabus puetzi</emphasis>
</taxonomicName>
<emphasis id="B921EAE9E309FFFEFD6A0150D171E963" bold="true" box="[717,774,1578,1599]" pageId="5" pageNumber="42">
<taxonomicNameLabel id="A2125792E309FFFEFD6A0150D171E963" box="[717,774,1578,1599]" pageId="5" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
: a: blade of prosternal process; b: apex of metasternal process; c: furrow on anteromedial metasternal process; d: gula; e: bead of prosternal process; f: furrow of mesosternum (scale bar ca. 1.0 mm).
</paragraph>
</caption>
<paragraph id="8BEA36FBE309FFFEFF6001D0D7E2E871" blockId="5.[151,1436,1706,2018]" pageId="5" pageNumber="42">Setae on ventral surface very sparse, short setae only present on mentum and on prosternum anteromedially; rather long setae emanating from a few punctures anteromedially on metasternum, apex of metacoxal processes, and third to sixth abdominal ventrites. Metafemora ventrally with a series of spaced rigid and transparent setae at posteroapical angle, typical for Agabini. Length of longer spur of metatibiae about 3/4 that of first metatarsomere.</paragraph>
<paragraph id="8BEA36FBE309FFFDFF600040D1A7ED60" blockId="5.[151,1436,1706,2018]" lastBlockId="6.[151,1437,151,2012]" lastPageId="6" lastPageNumber="43" pageId="5" pageNumber="42">
<emphasis id="B921EAE9E309FFFEFF600040D378E80E" bold="true" box="[199,271,1850,1874]" pageId="5" pageNumber="42">Males</emphasis>
: Protibiae proximally rather thin, distally broad; inner outline distinctly and very characteristically curved (
<figureCitation id="136E2A7EE309FFFEFF530024D34DE82A" box="[244,314,1886,1910]" captionStart="FIGURES 3–4" captionStartId="2.[151,264,1901,1924]" captionText="FIGURES 3–4. Agabus puetzi sp. n.: left protibia and protarsus of (3) male, (4) female." pageId="5" pageNumber="42">Fig. 3</figureCitation>
). First three pro- and mesotarsomeres strongly enlarged, provided with very dense setae on underside, each ending in a small elongate eccentrically stalked sucker palette; structure similar to that of
<taxonomicName id="4C554D78E309FFFEFAA600F8D330E8E2" authority="Thomson, 1856" authorityName="Thomson" authorityYear="1856" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="zetterstedti">
<emphasis id="B921EAE9E309FFFEFAA600F8D7EBE8C6" box="[1281,1436,1922,1946]" italics="true" pageId="5" pageNumber="42">A. zetterstedti</emphasis>
Thomson, 1856
</taxonomicName>
(cf. Fig.
<quantity id="4CAD9B1EE309FFFEFE0A00DCD391E8E2" box="[429,486,1958,1982]" metricMagnitude="0" metricUnit="m" metricValue="1.4731999999999998" pageId="5" pageNumber="42" unit="in" value="58.0">58 in</quantity>
<bibRefCitation id="EFC44B0AE309FFFEFE4B00DCD0C2E8E2" author="Larson" box="[492,693,1958,1982]" firstAuthor="Larson" page="914" pageId="5" pageNumber="42" pagination="861 - 919" refId="ref10370" refString="Larson, D. J. (1989) Revision of North American Agabus Leach (Coleoptera: Dytiscidae): introduction, key to species groups, and classification of the ambiguus -, tristis -, and arcticus - groups. The Canadian Entomologist, 121, 861 - 919." type="journal article" year="1989">Larson 1989: 914</bibRefCitation>
), but palettes more elongate and much more numerous in
<taxonomicName id="4C554D78E309FFFEFA9D00DCD7EBE8E2" authority="Fery, 2011" authorityName="Fery" authorityYear="2011" box="[1338,1436,1958,1982]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="puetzi" status="sp. nov.">
<emphasis id="B921EAE9E309FFFEFA9D00DCD7EBE8E2" box="[1338,1436,1958,1982]" italics="true" pageId="5" pageNumber="42">A. puetzi</emphasis>
</taxonomicName>
<emphasis id="B921EAE9E309FFFEFF3000B0D2AFE8BE" bold="true" box="[151,216,1994,2018]" pageId="5" pageNumber="42">
<taxonomicNameLabel id="A2125792E309FFFEFF3000B0D2AFE8BE" box="[151,216,1994,2018]" pageId="5" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; those of third protarsomeres reaching so far forwards that fourth ones also appear as to be provided with sucker setae; however, fourth tarsomeres less enlarged and without adhesive setae. Fifth pro- and mesotarsomeres long, almost as long as respective first four tarsomeres together; underside with two rows of small bristles, somewhat more irregular on fifth mesotarsomere; fourth protarsomeres on underside only with a few bristles distally, fourth mesotarsomeres with two distinct rows of bristles. Pro- and mesotarsal claws shorter than respective fifth tarsomere; claws very weakly curved proximally, more strongly curved near apex; posterior claws slightly shorter. Metatarsal claws evenly and slightly curved, equal in length, and only half as long as pro- and mesotarsal claws. Mesotibiae and first mesotarsomeres with natatorial setal fringe ventrally. Fifth to ninth antennomeres with anteroapical angle slightly produced and truncate, here with a sensory seta; structure less distinct on tenth antennomeres, third and fourth not produced. Median lobe asymmetric, bent to left before apex; ventral groove open over entire length. In lateral view right side of apical fourth broad, left side thin (
<figureCitation id="136E2A7EE30AFFFDFC7906A6D635EEA8" box="[990,1090,476,500]" captionStart="FIGURES 5–9" captionStartId="3.[151,263,1575,1598]" captionTargetBox="[264,1323,193,1554]" captionTargetId="figure-215@3.[264,1323,193,1554]" captionTargetPageId="3" captionText="FIGURES 5–9. Agabus puetzi sp. n.: median lobe of aedeagus: (5) right side, (6) left side, (7) detail of apex (weak swelling indicated by the arrow); (8) left paramere; female genitalia: (9) (a) gonocoxa, (b) tergite 9." pageId="6" pageNumber="43">Figs 5, 6</figureCitation>
,
<figureCitation id="136E2A7EE30AFFFDFBEA06A6D61BEEA8" box="[1101,1132,476,500]" captionStart="FIGURES 28–39" captionStartId="13.[151,264,1891,1914]" captionTargetBox="[193,1408,209,1856]" captionTargetId="figure-107@13.[159,1428,193,1870]" captionTargetPageId="13" captionText="FIGURES 28–39. Agabus, confinis-group, left side of median lobe of aedeagus (schematic) of: (28) A. matsumotoi, (29) A. moestus, (30) A. pseudoclypealis, (31) A. puetzi sp. n., (32) A. setulosus, (33) A. slovzovi, (34) A. suoduogangi, (35) A. thomsoni, (36) A. tibetanus, (37) A. turcmenus, (38) A. zetterstedti (a) with preapical spine strongly attached to the dorsal face of lobe, (b) with preapical spine less attached, (39) Agabus n. sp. from China to be described by J. Šťastný." pageId="6" pageNumber="43">31</figureCitation>
). Apex without a subapical dorsal spine, but right side before apex with a weak swelling (indicated by the arrow in
<figureCitation id="136E2A7EE30AFFFDFBD50685D6C1ED4B" box="[1138,1206,511,535]" captionStart="FIGURES 5–9" captionStartId="3.[151,263,1575,1598]" captionTargetBox="[264,1323,193,1554]" captionTargetId="figure-215@3.[264,1323,193,1554]" captionTargetPageId="3" captionText="FIGURES 5–9. Agabus puetzi sp. n.: median lobe of aedeagus: (5) right side, (6) left side, (7) detail of apex (weak swelling indicated by the arrow); (8) left paramere; female genitalia: (9) (a) gonocoxa, (b) tergite 9." pageId="6" pageNumber="43">Fig. 7</figureCitation>
). Parameres (
<figureCitation id="136E2A7EE30AFFFDFAF70685D7E5ED4B" box="[1360,1426,511,535]" captionStart="FIGURES 5–9" captionStartId="3.[151,263,1575,1598]" captionTargetBox="[264,1323,193,1554]" captionTargetId="figure-215@3.[264,1323,193,1554]" captionTargetPageId="3" captionText="FIGURES 5–9. Agabus puetzi sp. n.: median lobe of aedeagus: (5) right side, (6) left side, (7) detail of apex (weak swelling indicated by the arrow); (8) left paramere; female genitalia: (9) (a) gonocoxa, (b) tergite 9." pageId="6" pageNumber="43">Fig. 8</figureCitation>
) stylate (or style-like; in contrast to strap-like; cf.
<bibRefCitation id="EFC44B0AE30AFFFDFD1C055ED1B4ED60" author="Larson" box="[699,963,548,572]" etAl="et al." firstAuthor="Larson" page="481" pageId="6" pageNumber="43" refId="ref10501" refString="Larson, D. J., Alarie, Y. &amp; Roughley, R. E. (2000) Predaceous diving beetles (Coleoptera: Dytiscidae) of the Nearctic Region, with emphasis on the fauna of Canada and Alaska. Ottawa: NRC Research Press, 982 pp." type="book" year="2000">
Larson
<emphasis id="B921EAE9E30AFFFDFCB7055FD136ED60" box="[784,833,548,572]" italics="true" pageId="6" pageNumber="43">et al</emphasis>
. 2000: 481
</bibRefCitation>
).
</paragraph>
<paragraph id="8BEA36FBE30AFFFDFF600532D31AEBFC" blockId="6.[151,1437,151,2012]" pageId="6" pageNumber="43">
<emphasis id="B921EAE9E30AFFFDFF600532D35EED3C" bold="true" box="[199,297,584,608]" pageId="6" pageNumber="43">Females</emphasis>
: Both females studied with matt elytra; colour of elytra lighter than in males, head and pronotum as in males (
<figureCitation id="136E2A7EE30AFFFDFF4D0516D346EDD8" box="[234,305,620,644]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1026,1049]" captionTargetBox="[293,1307,268,1013]" captionTargetId="figure-80@2.[283,1311,267,1013]" captionTargetPageId="2" captionText="FIGURES 1–2. Agabus puetzi sp. n.: habitus of: (1) male (paratype; disarticulated appendages re-arranged), (2) female." pageId="6" pageNumber="43">Fig. 2</figureCitation>
). Reticulation double, principally as in males, but secondary one on elytra somewhat more deeply impressed and surface of micro-meshes not shiny. Head and pronotum shiny, secondary reticulation on pronotum extended on entire surface except a small area around discal scratch. Ventral surface same as in males, except central part of metasternum more matt. Inner outline of protibiae less curved than in males, nevertheless, characteristic shape well recognisable (
<figureCitation id="136E2A7EE30AFFFDFE140586D38EEC48" box="[435,505,764,788]" captionStart="FIGURES 3–4" captionStartId="2.[151,264,1901,1924]" captionText="FIGURES 3–4. Agabus puetzi sp. n.: left protibia and protarsus of (3) male, (4) female." pageId="6" pageNumber="43">Fig. 4</figureCitation>
). Pro- and mesotarsomeres not broadened, without sucker palettes; pro- and mesotarsal claws shorter and evenly curved; posterior protarsal claw slightly shorter than anterior one; metatarsal claws as in males, distinctly shorter than pro- and mesotarsal claws. Fifth pro- and mesotarsomeres rather long, but shorter than those of males. First pro- and mesotarsomeres anteriorly with one row of spines, posteriorly only distally with a few spines; on second to fourth pro- and mesotarsomeres both rows reduced to a few spines distally; fifth pro- and mesotarsomeres with two rows of spines. Mesotibiae and first mesotarsomeres ventrally without natatorial setal fringe (lack of this fringe in females being characteristic for part of Agabini, particularly for genus
<taxonomicName id="4C554D78E30AFFFDFF300482D29CEB4C" authorityName="Leach" authorityYear="1817" box="[151,235,1016,1040]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAE9E30AFFFDFF300482D29CEB4C" box="[151,235,1016,1040]" italics="true" pageId="6" pageNumber="43">Agabus</emphasis>
</taxonomicName>
; cf.
<bibRefCitation id="EFC44B0AE30AFFFDFEB9048DD397EB53" author="Nilsson" box="[286,480,1015,1039]" firstAuthor="Nilsson" page="18" pageId="6" pageNumber="43" pagination="17 - 36" refId="ref10812" refString="Nilsson, A. N. (2000) A new view of the generic classification of the Agabus - group of genera of the Agabini, aimed at solving the problem with a paraphyletic Agabus (Coleoptera: Dytiscidae). Koleopterologische Rundschau, 70, 17 - 36." type="journal article" year="2000">Nilsson 2000: 18</bibRefCitation>
). Fifth to ninth antennomeres with sensory seta at anteroapical angle, but characteristic shape of angle in males only very indistinctly indicated in females. Female genitalia (
<figureCitation id="136E2A7EE30AFFFDFB220366D6BBEB68" box="[1157,1228,1052,1076]" captionStart="FIGURES 5–9" captionStartId="3.[151,263,1575,1598]" captionTargetBox="[264,1323,193,1554]" captionTargetId="figure-215@3.[264,1323,193,1554]" captionTargetPageId="3" captionText="FIGURES 5–9. Agabus puetzi sp. n.: median lobe of aedeagus: (5) right side, (6) left side, (7) detail of apex (weak swelling indicated by the arrow); (8) left paramere; female genitalia: (9) (a) gonocoxa, (b) tergite 9." pageId="6" pageNumber="43">Fig. 9</figureCitation>
: (a) gonocoxa, (b) tergite 9) without conspicuous differences to those of e.g.,
<taxonomicName id="4C554D78E30AFFFDFC84033AD600EB04" authority="(Thunberg, 1794)" baseAuthorityName="Thunberg" baseAuthorityYear="1794" box="[803,1143,1088,1112]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="congener">
<emphasis id="B921EAE9E30AFFFDFC84033AD1DEEB04" box="[803,937,1088,1112]" italics="true" pageId="6" pageNumber="43">A. congener</emphasis>
(Thunberg, 1794)
</taxonomicName>
,
<taxonomicName id="4C554D78E30AFFFDFB25033AD2ADEB20" authority="(Thomson, 1867)" baseAuthorityName="Thomson" baseAuthorityYear="1867" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="lapponicus">
<emphasis id="B921EAE9E30AFFFDFB25033AD76CEB04" box="[1154,1307,1088,1112]" italics="true" pageId="6" pageNumber="43">A. lapponicus</emphasis>
(Thomson, 1867)
</taxonomicName>
,
<taxonomicName id="4C554D78E30AFFFDFF42031ED308EB20" authorityName="Thomson" authorityYear="1856" box="[229,383,1124,1148]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="zetterstedti">
<emphasis id="B921EAE9E30AFFFDFF42031ED308EB20" box="[229,383,1124,1148]" italics="true" pageId="6" pageNumber="43">A. zetterstedti</emphasis>
</taxonomicName>
or indeed
<taxonomicName id="4C554D78E30AFFFDFE53031ED155EB20" authority="(Paykull, 1798)" baseAuthorityName="Paykull" baseAuthorityYear="1798" box="[500,802,1124,1148]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="arcticus">
<emphasis id="B921EAE9E30AFFFDFE53031ED01BEB20" box="[500,620,1124,1148]" italics="true" pageId="6" pageNumber="43">A. arcticus</emphasis>
(Paykull, 1798)
</taxonomicName>
, latter being member of the
<taxonomicName id="4C554D78E30AFFFDFBFD031FD6C4EB20" baseAuthorityName="Paykull" baseAuthorityYear="1798" box="[1114,1203,1125,1148]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="arcticus">
<emphasis id="B921EAE9E30AFFFDFBFD031FD6C4EB20" box="[1114,1203,1125,1148]" italics="true" pageId="6" pageNumber="43">arcticus</emphasis>
</taxonomicName>
-group (cf. Fig.
<quantity id="4CAD9B1EE30AFFFDFAC4031ED7EBEB20" box="[1379,1436,1124,1148]" metricMagnitude="0" metricUnit="m" metricValue="1.3716" pageId="6" pageNumber="43" unit="in" value="54.0">54 in</quantity>
<bibRefCitation id="EFC44B0AE30AFFFDFF3003F2D317EBFC" author="Larson" box="[151,352,1160,1184]" firstAuthor="Larson" page="913" pageId="6" pageNumber="43" pagination="861 - 919" refId="ref10370" refString="Larson, D. J. (1989) Revision of North American Agabus Leach (Coleoptera: Dytiscidae): introduction, key to species groups, and classification of the ambiguus -, tristis -, and arcticus - groups. The Canadian Entomologist, 121, 861 - 919." type="journal article" year="1989">Larson 1989: 913</bibRefCitation>
).
</paragraph>
<paragraph id="8BEA36FBE30AFFFDFF6003D6D706EA50" blockId="6.[151,1437,151,2012]" pageId="6" pageNumber="43">
<emphasis id="B921EAE9E30AFFFDFF6003D6D30AEB98" bold="true" box="[199,381,1196,1220]" inLineHeading="true" pageId="6" pageNumber="43" reason="1">Measurements.</emphasis>
<typeStatus id="54EE8859E30AFFFDFE2303D6D39BEB98" box="[388,492,1196,1220]" pageId="6" pageNumber="43" type="holotype">Holotype</typeStatus>
(male): TL =
<quantity id="4CAD9B1EE30AFFFDFD2903D6D090EB9F" box="[654,743,1196,1220]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="6" pageNumber="43" unit="mm" value="7.5">7.5 mm</quantity>
, TL-h =
<quantity id="4CAD9B1EE30AFFFDFCEB03D6D1D2EB9F" box="[844,933,1196,1220]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="6" pageNumber="43" unit="mm" value="6.7">6.7 mm</quantity>
, MW =
<quantity id="4CAD9B1EE30AFFFDFBA303D6D62AEB9F" box="[1028,1117,1196,1220]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="6" pageNumber="43" unit="mm" value="3.6">3.6 mm</quantity>
, TL/MW = 2.08, WC/WS = 2.26; male
<typeStatus id="54EE8859E30AFFFDFEB303AAD30EEBB4" box="[276,377,1232,1256]" pageId="6" pageNumber="43" type="paratype">paratype</typeStatus>
: TL =
<quantity id="4CAD9B1EE30AFFFDFE6303AAD06BEBBB" box="[452,540,1232,1256]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="6" pageNumber="43" unit="mm" value="7.2">7.2 mm</quantity>
, TL-h =
<quantity id="4CAD9B1EE30AFFFDFDD803AAD0A0EBBB" box="[639,727,1232,1256]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="6" pageNumber="43" unit="mm" value="6.4">6.4 mm</quantity>
, MW =
<quantity id="4CAD9B1EE30AFFFDFC9303AAD1FBEBBB" box="[820,908,1232,1256]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="43" unit="mm" value="3.5">3.5 mm</quantity>
, TL/MW = 2.06, WC/WS = 2.13; female
<typeStatus id="54EE8859E30AFFFDFAC503AAD2A1EA50" pageId="6" pageNumber="43" type="paratype">paratypes</typeStatus>
: TL = 6.95/
<quantity id="4CAD9B1EE30AFFFDFEFC038ED3C6EA57" box="[347,433,1268,1292]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="6" pageNumber="43" unit="mm" value="7.1">7.1 mm</quantity>
, TL-h = 6.2/
<quantity id="4CAD9B1EE30AFFFDFD98038ED0E0EA57" box="[575,663,1268,1292]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="6" pageNumber="43" unit="mm" value="6.4">6.4 mm</quantity>
, MW = 3.45/
<quantity id="4CAD9B1EE30AFFFDFC89038ED1F2EA57" box="[814,901,1268,1292]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="43" unit="mm" value="3.5">3.5 mm</quantity>
, TL/MW = 2.01/2.03, WC/WS = 2.13/2.04.
</paragraph>
<paragraph id="8BEA36FBE30AFFFDFF60026DD67DE95B" blockId="6.[151,1437,151,2012]" pageId="6" pageNumber="43">
<emphasis id="B921EAE9E30AFFFDFF60026DD33AEA6C" bold="true" box="[199,333,1303,1328]" inLineHeading="true" pageId="6" pageNumber="43" reason="1">Variability.</emphasis>
I have not found considerably variable features in the
<specimenCount id="9D53FD72E30AFFFDFC650262D619EA6C" box="[962,1134,1304,1328]" pageId="6" pageNumber="43" type="generic">four specimens</specimenCount>
studied. The extent of the dark brown coloration on the elytra varies a little, the male
<typeStatus id="54EE8859E30AFFFDFC8B0246D1FBEA08" box="[812,908,1340,1364]" pageId="6" pageNumber="43" type="paratype">paratype</typeStatus>
has the elytra somewhat transparent anteriorly, and the sides of the pronotum can be slightly less curved. The male
<typeStatus id="54EE8859E30AFFFDFC0E021AD67EEA24" box="[937,1033,1376,1400]" pageId="6" pageNumber="43" type="paratype">paratype</typeStatus>
and
<specimenCount id="9D53FD72E30AFFFDFBE3021AD6B6EA24" box="[1092,1217,1376,1400]" pageId="6" pageNumber="43" type="female">one female</specimenCount>
have an additional shallow impression in the posterior half of the pronotum on each side of the mid line. The furrow on the anteromedial process of the metasternum is weaker in
<specimenCount id="9D53FD72E30AFFFDFD3B02D2D16CEA9C" box="[668,795,1448,1472]" pageId="6" pageNumber="43" type="female">one female</specimenCount>
, and more strongly impressed in the male
<typeStatus id="54EE8859E30AFFFDFAA602D2D712EA9C" box="[1281,1381,1448,1472]" pageId="6" pageNumber="43" type="paratype">paratype</typeStatus>
. The width of the metasternal wings also varies a little; according to the measurements, the WC/WS ratio is smaller in females, but the few specimens studied do not allow a well-founded statement.
</paragraph>
</subSubSection>
<subSubSection id="C34F6570E30AFFFDFF600169D0ADE8C8" pageId="6" pageNumber="43" type="distribution">
<paragraph id="8BEA36FBE30AFFFDFF600169D0ADE8C8" blockId="6.[151,1437,151,2012]" pageId="6" pageNumber="43">
<emphasis id="B921EAE9E30AFFFDFF600169D317E970" bold="true" box="[199,352,1555,1580]" inLineHeading="true" pageId="6" pageNumber="43" reason="1">Distribution.</emphasis>
Eastern Qinghai-Tibet Plateau; so far
<taxonomicName id="4C554D78E30AFFFDFC84016ED1FFE970" authority="Fery, 2011" authorityName="Fery" authorityYear="2011" box="[803,904,1556,1580]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="puetzi" status="sp. nov.">
<emphasis id="B921EAE9E30AFFFDFC84016ED1FFE970" box="[803,904,1556,1580]" italics="true" pageId="6" pageNumber="43">A. puetzi</emphasis>
</taxonomicName>
<emphasis id="B921EAE9E30AFFFDFC34016ED1A1E970" bold="true" box="[915,982,1556,1580]" pageId="6" pageNumber="43">
<taxonomicNameLabel id="A2125792E30AFFFDFC34016ED1A1E970" box="[915,982,1556,1580]" pageId="6" pageNumber="43" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is only known from the locus typicus. Lance He (
<collectingCountry id="F342766BE30AFFFDFEBB0142D315E90C" box="[284,354,1592,1616]" name="China" pageId="6" pageNumber="43">China</collectingCountry>
) communicated the following information about the area, which includes the locus typicus (slightly modified by me): The Shaluli Mountains cover a large area in western
<collectingRegion id="4991F819E30AFFFDFBBB0126D60EE928" box="[1052,1145,1628,1652]" country="China" name="Sichuan" pageId="6" pageNumber="43">Sichuan</collectingRegion>
, which is more than
<quantity id="4CAD9B1EE30AFFFDFAC30126D2CCE9C4" metricMagnitude="6" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="43" unit="km" value="1000.0">1000 km</quantity>
long. It's a part of the huge Hengduan Mountains, which lie across western
<collectingRegion id="4991F819E30AFFFDFBAA01FAD61DE9C4" box="[1037,1130,1664,1688]" country="China" name="Sichuan" pageId="6" pageNumber="43">Sichuan</collectingRegion>
, north-western
<collectingRegion id="4991F819E30AFFFDFAB001FAD71BE9CB" box="[1303,1388,1664,1687]" country="China" name="Yunnan" pageId="6" pageNumber="43">Yunnan</collectingRegion>
and eastern
<collectingRegion id="4991F819E30AFFFDFF5701DED359E9E0" box="[240,302,1700,1724]" country="China" name="Xizang" pageId="6" pageNumber="43">Tibet</collectingRegion>
. &quot;Haizi&quot; means in Tibetan &quot;small lake in the mountain&quot;. Our Haizi mountain, or better the &quot;Haizi peak&quot;, is just a small peak in southern Shaluli Mountains and it is named after the small twin lakes in front of it (it also has other names: Xiashe Peak, Peak Nitekhangri). There are several &quot;Haizi peaks&quot; known in western
<collectingRegion id="4991F819E30AFFFDFA9D0196D7E0E858" box="[1338,1431,1772,1796]" country="China" name="Sichuan" pageId="6" pageNumber="43">Sichuan</collectingRegion>
, our peak is located in Batang county, another one in Daocheng county etc. All of them have some small lakes closely situated, thus, &quot;Haizi peak&quot; is a common name for this kind of mountain. Our peak is so well known because it's so close to the
<collectingRegion id="4991F819E30AFFFDFE740022D05CE82C" box="[467,555,1880,1904]" country="China" name="Sichuan" pageId="6" pageNumber="43">Sichuan</collectingRegion>
–
<collectingRegion id="4991F819E30AFFFDFD9E0022D003E82C" box="[569,628,1880,1904]" country="China" name="Xizang" pageId="6" pageNumber="43">Tibet</collectingRegion>
highway. Actually there are plenty of gorgeous mountains in
<collectingRegion id="4991F819E30AFFFDFAE50022D28BE8C8" country="China" name="Sichuan" pageId="6" pageNumber="43">Sichuan province</collectingRegion>
, but many of them are not very accessible.
</paragraph>
</subSubSection>
<subSubSection id="C34F6570E30AFFFDFF6000E5D6E8E887" pageId="6" pageNumber="43" type="etymology">
<paragraph id="8BEA36FBE30AFFFDFF6000E5D6E8E887" blockId="6.[151,1437,151,2012]" pageId="6" pageNumber="43">
<emphasis id="B921EAE9E30AFFFDFF6000E5D3D1E8E4" bold="true" box="[199,422,1951,1976]" inLineHeading="true" pageId="6" pageNumber="43" reason="1">Derivatio nominis.</emphasis>
The new species is named after Andreas Pütz (Eisenhüttenstadt,
<collectingCountry id="F342766BE30AFFFDFB3400DAD688E8E4" box="[1171,1279,1952,1976]" name="Germany" pageId="6" pageNumber="43">Germany</collectingCountry>
), specialist in
<taxonomicName id="4C554D78E30AFFFDFF3000BED370E880" box="[151,263,1988,2012]" class="Insecta" family="Byrrhidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="family">Byrrhidae</taxonomicName>
and collector of the specimens. The specific epithet is a noun in the genitive case.
</paragraph>
</subSubSection>
<subSubSection id="C34F6570E30BFFFCFF6007EDD729EEDB" pageId="7" pageNumber="44" type="biology_ecology">
<paragraph id="8BEA36FBE30BFFFCFF6007EDD729EEDB" blockId="7.[151,1436,151,392]" pageId="7" pageNumber="44">
<emphasis id="B921EAE9E30BFFFCFF6007EDD351EFEC" bold="true" box="[199,294,151,176]" inLineHeading="true" pageId="7" pageNumber="44" reason="1">Biology.</emphasis>
The specimens were found near the border of the &quot;Sister Lakes&quot; (
<figureCitation id="136E2A7EE30BFFFCFBAB07EDD6F8EFF3" box="[1036,1167,151,175]" captionStart="FIGURES 40–41" captionStartId="14.[151,264,1623,1646]" captionTargetBox="[264,1322,193,1602]" captionTargetId="figure-226@14.[264,1322,193,1602]" captionTargetPageId="14" captionText="FIGURES 40–41. Sister Lakes with Haizi mountain in the background; collecting site of Agabus puetzi sp. n. indicated by the arrow (for more details see text)." pageId="7" pageNumber="44">Figs 40, 41</figureCitation>
), in small pools of variable size (diameter ca.
<quantity id="4CAD9B1EE30BFFFCFE3807C6D394EF88" box="[415,483,188,212]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="44" unit="m" value="0.5">0.5 m</quantity>
–
<quantity id="4CAD9B1EE30BFFFCFE5407C6D06AEF8F" box="[499,541,188,212]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="44" unit="m" value="1.0">1 m</quantity>
, depth not more than
<quantity id="4CAD9B1EE30BFFFCFCBF07C6D13FEF88" box="[792,840,188,212]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="44" unit="m" value="1.0">1 m</quantity>
), with coarse gravel on the ground, and without any vegetation, except some fine roots of small willows growing at the edge of the pools. The collector suspects that once during an earthquake big rocks fell down from the nearby mountain slopes and formed these little pools. The water was crystal clear and &quot;very very cold&quot; (temperature not measured). Other beetles found:
<taxonomicName id="4C554D78E30BFFFCFB1B0652D3ECEE38" authority="Regimbart, 1899" authorityName="Regimbart" authorityYear="1899" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="44" phylum="Arthropoda" rank="species" species="longissimus">
<emphasis id="B921EAE9E30BFFFCFB1B0652D7EBEE1C" box="[1212,1436,296,320]" italics="true" pageId="7" pageNumber="44">Agabus longissimus</emphasis>
Régimbart, 1899, 1908
</taxonomicName>
,
<taxonomicName id="4C554D78E30BFFFCFE010637D1E0EE38" authority="(Falkenstrom, 1936)" baseAuthorityName="Falkenstrom" baseAuthorityYear="1936" box="[422,919,332,356]" class="Insecta" family="Dytiscidae" genus="Boreonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="44" phylum="Arthropoda" rank="species" species="emmerichi">
<emphasis id="B921EAE9E30BFFFCFE010637D0DBEE38" box="[422,684,332,356]" italics="true" pageId="7" pageNumber="44">Boreonectes emmerichi</emphasis>
(Falkenström, 1936)
</taxonomicName>
(formerly in
<taxonomicName id="4C554D78E30BFFFCFB8B0636D7E0EE38" authority="Zimmermann, 1919" authorityName="Zimmermann" authorityYear="1919" box="[1068,1431,332,356]" class="Insecta" family="Dytiscidae" genus="Stictotarsus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAE9E30BFFFCFB8B0636D6D8EE38" box="[1068,1199,332,356]" italics="true" pageId="7" pageNumber="44">Stictotarsus</emphasis>
Zimmermann, 1919
</taxonomicName>
; see
<bibRefCitation id="EFC44B0AE30BFFFCFF650615D339EEDB" author="Angus" box="[194,334,367,391]" firstAuthor="Angus" pageId="7" pageNumber="44" pagination="123 - 131" refId="ref9931" refString="Angus, R. B. (2010) Boreonectes gen. n., a new genus for the Stictotarsus griseostriatus (De Geer) group of sibling species (Coleoptera: Dytiscidae), with additional karyosystematic data on the group. Comparative Cytogenetics, 4 (2), 123 - 131." type="journal article" year="2010">
<collectingRegion id="4991F819E30BFFFCFF650615D37CEEDB" box="[194,267,367,391]" country="United Kingdom" name="Angus" pageId="7" pageNumber="44">Angus</collectingRegion>
2010
</bibRefCitation>
),
<taxonomicName id="4C554D78E30BFFFCFEC6060AD081EEDB" authority="Zaitzev, 1953" authorityName="Zaitzev" authorityYear="1953" box="[353,758,367,392]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="44" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis id="B921EAE9E30BFFFCFEC6060AD022EED4" box="[353,597,368,392]" italics="true" pageId="7" pageNumber="44">Hydroporus tibetanus</emphasis>
Zaitzev, 1953
</taxonomicName>
, and
<taxonomicName id="4C554D78E30BFFFCFC96060AD181EEDB" box="[817,1014,368,392]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="44" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B921EAE9E30BFFFCFC96060AD1C2EED4" box="[817,949,368,392]" italics="true" pageId="7" pageNumber="44">Helophorus</emphasis>
spec.
</taxonomicName>
(
<taxonomicName id="4C554D78E30BFFFCFBA30615D6DDEEDB" authorityName="Latreille" authorityYear="1802" box="[1028,1194,367,391]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="44" phylum="Arthropoda" rank="family">Hydrophilidae</taxonomicName>
,
<taxonomicName id="4C554D78E30BFFFCFB120615D724EEDB" authorityName="Leach" authorityYear="1815" box="[1205,1363,367,391]" class="Insecta" family="Helophoridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="44" phylum="Arthropoda" rank="subFamily" subFamily="Helophorinae">Helophorinae</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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