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<document id="09FDF71EA16B15DA1D328CCA3B0A0A94" ID-DOI="10.11646/zootaxa.5047.4.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="5543116" ID-ZooBank="BF74A3DF-BFC4-43CB-9E3A-9AF5C0571AE8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1633071083356" checkinUser="plazi" docAuthor="Botero, Juan Pablo &amp; Vives, Eduard" docDate="2021" docId="03FC87BF2C08076357DCE2C0B3E8F816" docLanguage="en" docName="zootaxa.5047.4.6.pdf" docOrigin="Zootaxa 5047 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Philippistenia Botero &amp; Vives 2021, gen. nov." docType="treatment" docVersion="3" lastPageNumber="468" masterDocId="FFC5FFC72C090760574BE03BB757FB02" masterDocTitle="Philippistenia, a new Disteniini genus from the Philippines (Coleoptera: Disteniidae)" masterLastPageNumber="476" masterPageNumber="465" pageNumber="466" updateTime="1699416687718" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="D8348E26C30C2722ED260F73756BBC5D">
<mods:title id="B152B2031E9AEDFBD2B455E789DB06F4">Philippistenia, a new Disteniini genus from the Philippines (Coleoptera: Disteniidae)</mods:title>
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<mods:namePart id="01D4F2A6E8240856596D5D3FC3691D6C">Botero, Juan Pablo</mods:namePart>
<mods:affiliation id="7D81C33FBAD8A40C63B3996457326BCB">Museu de Zoologia, Universidade de São Paulo, São Paulo, Brazil</mods:affiliation>
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<mods:namePart id="D8EA89FFAD31283D8527C598D465A2E2">Vives, Eduard</mods:namePart>
<mods:affiliation id="6F3650D313A24A42121B6ED5B995E5F9">San Antoni, 73, 08221, Terrassa, Spain</mods:affiliation>
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<mods:title id="C3DDE868B9DE38C3E3A713AB35298508">Zootaxa</mods:title>
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<mods:part id="18E1D3BEEAFACDF489CB3284241D3865">
<mods:date id="1335EA5055949F505B7AA0D1D254AC75">2021</mods:date>
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<mods:number id="9AB846A0A0CFB411596A7CC329C7C563">2021-10-01</mods:number>
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<mods:number id="C8E8C9EFAF0B81C8A24F439BFEDC699B">5047</mods:number>
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<mods:identifier id="B96B1AF3F3A71BEAFF1ED61E9F6F207E" type="DOI">10.11646/zootaxa.5047.4.6</mods:identifier>
<mods:identifier id="56F2E379F5F88089FFD5CA75B4ECB015" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03FC87BF2C08076357DCE2C0B3E8F816" ID-DOI="http://doi.org/10.5281/zenodo.5543122" ID-Zenodo-Dep="5543122" LSID="urn:lsid:plazi:treatment:03FC87BF2C08076357DCE2C0B3E8F816" httpUri="http://treatment.plazi.org/id/03FC87BF2C08076357DCE2C0B3E8F816" lastPageId="3" lastPageNumber="468" pageId="1" pageNumber="466">
<subSubSection id="C34F65222C08076157DCE2C0B6E3F817" box="[151,436,763,790]" pageId="1" pageNumber="466" type="nomenclature">
<paragraph id="8BEA36A92C08076157DCE2C0B6E3F817" blockId="1.[151,436,763,790]" box="[151,436,763,790]" pageId="1" pageNumber="466">
<heading id="D0A281C52C08076157DCE2C0B6E3F817" bold="true" box="[151,436,763,790]" fontSize="11" level="1" pageId="1" pageNumber="466" reason="1">
<emphasis id="B921EABB2C08076157DCE2C0B6E3F817" bold="true" box="[151,436,763,790]" pageId="1" pageNumber="466">
<taxonomicName id="4C554D2A2C08076157DCE2C0B668F814" authority="Botero &amp; Vives, 2021" authorityName="Botero &amp; Vives" authorityYear="2021" box="[151,319,763,790]" class="Insecta" family="Disteniidae" genus="Philippistenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="466" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B921EABB2C08076157DCE2C0B668F814" bold="true" box="[151,319,763,790]" italics="true" pageId="1" pageNumber="466">Philippistenia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257C02C080761560DE2C0B6E3F817" box="[326,436,763,789]" pageId="1" pageNumber="466" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34F65222C08076157DCE37BB41FF85A" box="[151,840,831,856]" pageId="1" pageNumber="466" type="type_taxon">
<paragraph id="8BEA36A92C08076157DCE37BB41FF85A" blockId="1.[151,840,831,856]" box="[151,840,831,856]" pageId="1" pageNumber="466">
<emphasis id="B921EABB2C08076157DCE37BB674F855" bold="true" box="[151,291,831,856]" pageId="1" pageNumber="466">
<typeStatus id="54EE880B2C08076157DCE37BB79BF85A" box="[151,204,832,856]" pageId="1" pageNumber="466">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C554D2A2C0807615661E37BB5CFF85A" ID-CoL="36WZ4" authority="Heller, 1923" authorityName="Heller" authorityYear="1923" box="[298,664,832,856]" class="Insecta" family="Cerambycidae" genus="Distenia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="466" phylum="Arthropoda" rank="species" species="heterotarsalis">
<emphasis id="B921EABB2C0807615661E37BB545F85A" box="[298,530,832,856]" italics="true" pageId="1" pageNumber="466">Distenia heterotarsalis</emphasis>
<bibRefCitation id="EFC44B582C0807615553E37BB5CFF85A" author="Heller, K. M." box="[536,664,832,856]" pageId="1" pageNumber="466" pagination="33 - 48" refId="ref4961" refString="Heller, K. M. (1923) Neue malayische, meist philippinische Bockkafer und ein neuer Russelkafer. Tijdschrift voor Entomologie, 66, 33 - 48." type="journal article" year="1923">Heller, 1923</bibRefCitation>
</taxonomicName>
, here designated.
</paragraph>
</subSubSection>
<subSubSection id="C34F65222C08076157DCE3BEB37AFF0E" pageId="1" pageNumber="466" type="diagnosis">
<paragraph id="8BEA36A92C08076157DCE3BEB37AFF0E" blockId="1.[151,1437,901,1504]" pageId="1" pageNumber="466">
<emphasis id="B921EABB2C08076157DCE3BEB645F89D" bold="true" box="[151,274,901,927]" pageId="1" pageNumber="466">Diagnosis.</emphasis>
<taxonomicName id="4C554D2A2C0807615653E3BDB6E2F8A2" authority="Botero &amp; Vives, 2021" authorityName="Botero &amp; Vives" authorityYear="2021" box="[280,437,902,928]" class="Insecta" family="Disteniidae" genus="Philippistenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="466" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B921EABB2C0807615653E3BDB6E2F8A2" box="[280,437,902,928]" italics="true" pageId="1" pageNumber="466">Philippistenia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257C02C08076156F0E3BDB577F8A2" box="[443,544,902,928]" pageId="1" pageNumber="466" rank="genus">gen. nov.</taxonomicNameLabel>
can be differentiated from other genera of
<taxonomicName id="4C554D2A2C08076154B0E3BDB33EF8A2" box="[1019,1129,902,928]" class="Insecta" family="Cerambycidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="466" phylum="Arthropoda" rank="tribe" tribe="Disteniini">Disteniini</taxonomicName>
by the following characteristics: head distinctly narrowed behind eyes; lateral tubercles of prothorax directed upward; elytra pubescent, with longitudinal bands of dense black pubescence, always present on suture; metafemora linear, unarmed apically, and long, almost reaching elytral apex; and metatarsomere I at least as long as II + III.
</paragraph>
</subSubSection>
<subSubSection id="C34F65222C080762578CE42EB5B0FAAF" lastPageId="2" lastPageNumber="467" pageId="1" pageNumber="466" type="description">
<paragraph id="8BEA36A92C080761578CE42EB637FEE2" blockId="1.[151,1437,901,1504]" pageId="1" pageNumber="466">
<emphasis id="B921EABB2C080761578CE42EB60EFF2D" bold="true" box="[199,345,1045,1071]" pageId="1" pageNumber="466">Description.</emphasis>
Head prognathous, narrowed behind eyes (
<figureCitation id="136E2A2C2C0807615407E42DB4C3FF32" box="[844,916,1046,1072]" captionStart="FIGURES 1–5. 1-3" captionStartId="1.[151,264,1881,1905]" captionTargetBox="[151,1436,1529,1857]" captionTargetId="figure-403@1.[151,1436,1528,1857]" captionTargetPageId="1" captionText="FIGURES 1–5. 1-3. Detail of posterior area of head and pronotum. 1. Distenia (D.) columbina Lepeletier &amp; Audinet-Serville, 1828; 2. Philippistenia levitemporalis (Heller, 1924); 3. Typodryas callichromoides Thomson, 1864; 4–5. Detail of the elytra showing the longitudinal black bands of pubescence. 4. Philippistenia samarensis (Villiers, 1959); 5. Philippistenia heterotarsalis (Heller, 1923)." figureDoi="http://doi.org/10.5281/zenodo.5543118" httpUri="https://zenodo.org/record/5543118/files/figure.png" pageId="1" pageNumber="466">Fig. 2</figureCitation>
). Eyes somewhat coarsely faceted, upper eye lobes well-separated, distance between them at least 1.5 times width of upper lobe. Antennae longer than body in both sexes; antennomeres gradually thinner toward apex of antenna. Fourth maxillary palpomere sexually dimorphic, securiform in male, fusiform in female. Prothorax wider than long (including lateral tubercles), strongly constricted anteriorly and posteriorly, more evident anteriorly; lateral tubercles wide basally, acute apically, directed upward. Pronotum microsculptured, covered with dense, fine pubescence and sparse long setae, with five gibbosities: two subcircular, placed on each side of anterior half; two larger, subrounded, placed on each side of posterior half, subfused with anterolateral one; another elongate, slightly elevated, placed medially. Elytra long, at least 3.5 times as long as prothorax; slightly narrowed toward apex; surface of anterior half with coarse, dense punctation, on posterior half, punctation decreases and there is no punctation apically; with dense black pubescence on suture, forming longitudinal bands on some other regions of elytra (
<figureCitation id="136E2A2C2C0807615404E545B4EEFE9B" box="[847,953,1406,1433]" captionStart="FIGURES 1–5. 1-3" captionStartId="1.[151,264,1881,1905]" captionTargetBox="[151,1436,1529,1857]" captionTargetId="figure-403@1.[151,1436,1528,1857]" captionTargetPageId="1" captionText="FIGURES 1–5. 1-3. Detail of posterior area of head and pronotum. 1. Distenia (D.) columbina Lepeletier &amp; Audinet-Serville, 1828; 2. Philippistenia levitemporalis (Heller, 1924); 3. Typodryas callichromoides Thomson, 1864; 4–5. Detail of the elytra showing the longitudinal black bands of pubescence. 4. Philippistenia samarensis (Villiers, 1959); 5. Philippistenia heterotarsalis (Heller, 1923)." figureDoi="http://doi.org/10.5281/zenodo.5543118" httpUri="https://zenodo.org/record/5543118/files/figure.png" pageId="1" pageNumber="466">Figs 4–5</figureCitation>
); elytral apex with projection at inner and outer angle. Femora linear; apex unarmed; metafemora long, almost reaching elytral apex. Metatarsomere I at least as long as II + III.
</paragraph>
<caption id="DF2A66212C08076157DCE762B636FCDF" ID-DOI="http://doi.org/10.5281/zenodo.5543118" ID-Zenodo-Dep="5543118" httpUri="https://zenodo.org/record/5543118/files/figure.png" pageId="1" pageNumber="466" startId="1.[151,264,1881,1905]" targetBox="[151,1436,1529,1857]" targetPageId="1">
<paragraph id="8BEA36A92C08076157DCE762B636FCDF" blockId="1.[151,1437,1881,2013]" pageId="1" pageNumber="466">
<emphasis id="B921EABB2C08076157DCE762B63EFC73" bold="true" box="[151,361,1881,1905]" pageId="1" pageNumber="466">FIGURES 1–5. 1-3.</emphasis>
Detail of posterior area of head and pronotum.
<emphasis id="B921EABB2C0807615404E762B435FC73" bold="true" box="[847,866,1881,1905]" pageId="1" pageNumber="466">1.</emphasis>
<taxonomicName id="4C554D2A2C0807615422E761B4E8FC73" authorityName="Lepeletier &amp; Audinet-Serville" authorityYear="1828" box="[873,959,1882,1905]" class="Insecta" family="Cerambycidae" genus="Distenia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="466" phylum="Arthropoda" rank="genus">
<emphasis id="B921EABB2C0807615422E761B4E8FC73" box="[873,959,1882,1905]" italics="true" pageId="1" pageNumber="466">Distenia</emphasis>
</taxonomicName>
(
<emphasis id="B921EABB2C0807615486E761B488FC73" box="[973,991,1882,1905]" italics="true" pageId="1" pageNumber="466">D</emphasis>
.)
<emphasis id="B921EABB2C08076154BEE762B337FC73" box="[1013,1120,1881,1905]" italics="true" pageId="1" pageNumber="466">columbina</emphasis>
Lepeletier &amp; Audinet-Serville, 1828;
<emphasis id="B921EABB2C0807615792E746B7BBFC97" bold="true" box="[217,236,1917,1941]" pageId="1" pageNumber="466">2.</emphasis>
<taxonomicName id="4C554D2A2C08076157B8E746B5E2FC97" authority="(Heller, 1924)" authorityName="Botero &amp; Vives" authorityYear="2021" baseAuthorityName="Heller" baseAuthorityYear="1924" box="[243,693,1917,1941]" class="Insecta" family="Disteniidae" genus="Philippistenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="466" phylum="Arthropoda" rank="species" species="levitemporalis">
<emphasis id="B921EABB2C08076157B8E746B54DFC97" box="[243,538,1917,1941]" italics="true" pageId="1" pageNumber="466">Philippistenia levitemporalis</emphasis>
(
<bibRefCitation id="EFC44B582C0807615562E746B5F9FC97" author="Heller, K. M." box="[553,686,1917,1941]" pageId="1" pageNumber="466" pagination="163 - 180" refId="ref4991" refString="Heller, K. M. (1924) Neue philippinische Bock- und Russelkafer aus der Sammlung von Prof. C. F. Baker. Wiener Entomologische Zeitung, 41 (4 - 10), 163 - 180." type="journal article" year="1924">Heller, 1924</bibRefCitation>
)
</taxonomicName>
;
<emphasis id="B921EABB2C080761558AE746B583FC97" bold="true" box="[705,724,1917,1941]" pageId="1" pageNumber="466">3.</emphasis>
<taxonomicName id="4C554D2A2C0807615597E746B3CDFC97" authority="Thomson, 1864" authorityName="Thomson" authorityYear="1864" box="[732,1178,1917,1941]" class="Insecta" family="Cerambycidae" genus="Typodryas" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="466" phylum="Arthropoda" rank="species" species="callichromoides">
<emphasis id="B921EABB2C0807615597E746B4B9FC97" box="[732,1006,1917,1941]" italics="true" pageId="1" pageNumber="466">Typodryas callichromoides</emphasis>
Thomson, 1864
</taxonomicName>
;
<emphasis id="B921EABB2C08076153EEE746B385FC97" bold="true" box="[1189,1234,1917,1941]" pageId="1" pageNumber="466">4–5.</emphasis>
Detail of the elytra showing the longitudinal black bands of pubescence. 4.
<taxonomicName id="4C554D2A2C0807615599E79AB329FCBB" authority="(Villiers, 1959)" baseAuthorityName="Villiers" baseAuthorityYear="1959" box="[722,1150,1953,1977]" class="Insecta" family="Disteniidae" genus="Philippistenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="466" phylum="Arthropoda" rank="species" species="samarensis">
<emphasis id="B921EABB2C0807615599E79AB48FFCBB" box="[722,984,1953,1977]" italics="true" pageId="1" pageNumber="466">Philippistenia samarensis</emphasis>
(
<bibRefCitation id="EFC44B582C08076154ADE79AB322FCBB" author="Villiers, A." box="[998,1141,1953,1977]" pageId="1" pageNumber="466" pagination="65 - 68" refId="ref5678" refString="Villiers, A. (1959) Notes sur quelques Disteniinae des iles Philippines. (Col. Cerambycidae). Bulletin de la Societe entomologique de France, 64, 65 - 68." type="journal article" year="1959">Villiers, 1959</bibRefCitation>
)
</taxonomicName>
;
<emphasis id="B921EABB2C08076153C3E79AB3CCFCBB" bold="true" box="[1160,1179,1953,1977]" pageId="1" pageNumber="466">5.</emphasis>
<taxonomicName id="4C554D2A2C08076153E9E79AB609FCDF" authority="(Heller, 1923)" authorityName="Botero &amp; Vives" authorityYear="2021" baseAuthorityName="Heller" baseAuthorityYear="1923" class="Insecta" family="Disteniidae" genus="Philippistenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="466" phylum="Arthropoda" rank="species" species="heterotarsalis">
<emphasis id="B921EABB2C08076153E9E79AB792FCDF" italics="true" pageId="1" pageNumber="466">Philippistenia heterotarsalis</emphasis>
(
<bibRefCitation id="EFC44B582C0807615798E7FEB602FCDF" author="Heller, K. M." box="[211,341,1989,2013]" pageId="1" pageNumber="466" pagination="33 - 48" refId="ref4961" refString="Heller, K. M. (1923) Neue malayische, meist philippinische Bockkafer und ein neuer Russelkafer. Tijdschrift voor Entomologie, 66, 33 - 48." type="journal article" year="1923">Heller, 1923</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BEA36A92C0B0762578CE0ACB5B0FAAF" blockId="2.[151,1437,150,501]" pageId="2" pageNumber="467">
<emphasis id="B921EABB2C0B0762578CE0ACB6D0FBB2" bold="true" box="[199,391,150,177]" pageId="2" pageNumber="467">Male terminalia</emphasis>
(
<figureCitation id="136E2A2C2C0B076256DCE0ACB54EFBB3" box="[407,537,151,177]" captionStart="FIGURES 20–33" captionStartId="4.[151,264,1896,1920]" captionTargetBox="[213,1373,181,1871]" captionTargetId="figure-31@4.[211,1377,181,1872]" captionTargetPageId="4" captionText="FIGURES 20–33. Philippistenia levitemporalis (Heller, 1924). 20–21. Lectotype, female. 20. dorsal view; 21. Labels. 22–26. Paralectotype. 22. Labels; 23. Dorsal view; 24. Ventral view; 25. Lateral view; 26. Frontal view; 27–33. Male terminalia. 27. Tergite VIII; 28–30. Median lobe. 28. Dorsal view; 29. Lateral view; 30. Ventral view. 31–33. Tegmen. 31. Dorsal view; 32. Lateral view; 33. Ventral view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5543124" httpUri="https://zenodo.org/record/5543124/files/figure.png" pageId="2" pageNumber="467">Figs 27–33</figureCitation>
,
<figureCitation id="136E2A2C2C0B0762556FE0ACB53BFBB3" box="[548,620,151,177]" captionStart="FIGURES 34–46" captionStartId="6.[151,264,1309,1333]" captionTargetBox="[153,1436,182,1284]" captionTargetId="figure-31@6.[151,1436,181,1285]" captionTargetPageId="6" captionText="FIGURES 34–46. Philippistenia halconensis (Vives, 2012). 34. Holotype, male; 35–37. Female. 35. Dorsal view; 36. Lateral view; 37. Frontal view. 38–46. Male terminalia. 38. Tergite VIII; 39. Sternite VIII; 40. Ventral arc; 41–43. Median lobe. 41. Dorsal view; 42. Ventral view; 43. Lateral view. 44–46. Tegmen. 44. Dorsal view; 45. Ventral view; 46. Lateral view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5543126" httpUri="https://zenodo.org/record/5543126/files/figure.png" pageId="2" pageNumber="467">38–46</figureCitation>
,
<figureCitation id="136E2A2C2C0B07625532E0ACB596FBB0" box="[633,705,151,178]" captionStart="FIGURES 54–70" captionStartId="8.[151,264,1849,1873]" captionTargetBox="[244,1344,182,1822]" captionTargetId="figure-31@8.[242,1346,181,1825]" captionTargetPageId="8" captionText="FIGURES 54–70. Philippistenia heterotarsalis (Heller, 1923). 54–57. Holotype of Distenia samarensis Villiers, 1959, male, copyright MNHN Paris. 54. Dorsal view; 55. Ventral view; 56. Lateral view; 57. Labels. 58–70. Male specimen. 58. Frontal view; 59. Dorsal view; 60. Ventral view; 61. Lateral view; 62. Tergite VIII; 63. Sternite VIII; 64. Ventral arc; 65–67. Median lobe. 65. Dorsal view; 66. Ventral view; 67. Lateral view. 68–70. Tegmen. 68. Dorsal view; 69. Ventral view; 70. Lateral view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5543132" httpUri="https://zenodo.org/record/5543132/files/figure.png" pageId="2" pageNumber="467">62–70</figureCitation>
). Tergite VIII (
<emphasis id="B921EABB2C0B07625427E0ACB4C2FBB2" box="[876,917,151,176]" italics="true" pageId="2" pageNumber="467">e.g.</emphasis>
<figureCitation id="136E2A2C2C0B076254D7E0ACB4BAFBB3" box="[924,1005,151,177]" captionStart="FIGURES 34–46" captionStartId="6.[151,264,1309,1333]" captionTargetBox="[153,1436,182,1284]" captionTargetId="figure-31@6.[151,1436,181,1285]" captionTargetPageId="6" captionText="FIGURES 34–46. Philippistenia halconensis (Vives, 2012). 34. Holotype, male; 35–37. Female. 35. Dorsal view; 36. Lateral view; 37. Frontal view. 38–46. Male terminalia. 38. Tergite VIII; 39. Sternite VIII; 40. Ventral arc; 41–43. Median lobe. 41. Dorsal view; 42. Ventral view; 43. Lateral view. 44–46. Tegmen. 44. Dorsal view; 45. Ventral view; 46. Lateral view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5543126" httpUri="https://zenodo.org/record/5543126/files/figure.png" pageId="2" pageNumber="467">Fig. 38</figureCitation>
) with lateral margins strongly convergent at apical third, distal margin truncated and lined with long yellow setae. Sternite VIII (
<emphasis id="B921EABB2C0B076253EFE080B39BFBD6" box="[1188,1228,187,212]" italics="true" pageId="2" pageNumber="467">e.g.</emphasis>
<figureCitation id="136E2A2C2C0B0762539FE080B272FBD7" box="[1236,1317,187,213]" captionStart="FIGURES 34–46" captionStartId="6.[151,264,1309,1333]" captionTargetBox="[153,1436,182,1284]" captionTargetId="figure-31@6.[151,1436,181,1285]" captionTargetPageId="6" captionText="FIGURES 34–46. Philippistenia halconensis (Vives, 2012). 34. Holotype, male; 35–37. Female. 35. Dorsal view; 36. Lateral view; 37. Frontal view. 38–46. Male terminalia. 38. Tergite VIII; 39. Sternite VIII; 40. Ventral arc; 41–43. Median lobe. 41. Dorsal view; 42. Ventral view; 43. Lateral view. 44–46. Tegmen. 44. Dorsal view; 45. Ventral view; 46. Lateral view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5543126" httpUri="https://zenodo.org/record/5543126/files/figure.png" pageId="2" pageNumber="467">Fig. 39</figureCitation>
) subtransverse, distal margin with long sparse yellow setae; apophysis long, about 1,5 times as long as sternite. Ventral arc (
<emphasis id="B921EABB2C0B076257EAE138B79EFA1E" box="[161,201,259,284]" italics="true" pageId="2" pageNumber="467">e.g.</emphasis>
<figureCitation id="136E2A2C2C0B07625784E138B649FA1C" box="[207,286,259,286]" captionStart="FIGURES 54–70" captionStartId="8.[151,264,1849,1873]" captionTargetBox="[244,1344,182,1822]" captionTargetId="figure-31@8.[242,1346,181,1825]" captionTargetPageId="8" captionText="FIGURES 54–70. Philippistenia heterotarsalis (Heller, 1923). 54–57. Holotype of Distenia samarensis Villiers, 1959, male, copyright MNHN Paris. 54. Dorsal view; 55. Ventral view; 56. Lateral view; 57. Labels. 58–70. Male specimen. 58. Frontal view; 59. Dorsal view; 60. Ventral view; 61. Lateral view; 62. Tergite VIII; 63. Sternite VIII; 64. Ventral arc; 65–67. Median lobe. 65. Dorsal view; 66. Ventral view; 67. Lateral view. 68–70. Tegmen. 68. Dorsal view; 69. Ventral view; 70. Lateral view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5543132" httpUri="https://zenodo.org/record/5543132/files/figure.png" pageId="2" pageNumber="467">Fig. 64</figureCitation>
) fork-shaped, with apophysis long. Tegmen (
<emphasis id="B921EABB2C0B07625458E138B46BFA1E" box="[787,828,259,284]" italics="true" pageId="2" pageNumber="467">e.g.</emphasis>
<figureCitation id="136E2A2C2C0B07625409E138B4E8FA1F" box="[834,959,259,285]" captionStart="FIGURES 20–33" captionStartId="4.[151,264,1896,1920]" captionTargetBox="[213,1373,181,1871]" captionTargetId="figure-31@4.[211,1377,181,1872]" captionTargetPageId="4" captionText="FIGURES 20–33. Philippistenia levitemporalis (Heller, 1924). 20–21. Lectotype, female. 20. dorsal view; 21. Labels. 22–26. Paralectotype. 22. Labels; 23. Dorsal view; 24. Ventral view; 25. Lateral view; 26. Frontal view; 27–33. Male terminalia. 27. Tergite VIII; 28–30. Median lobe. 28. Dorsal view; 29. Lateral view; 30. Ventral view. 31–33. Tegmen. 31. Dorsal view; 32. Lateral view; 33. Ventral view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5543124" httpUri="https://zenodo.org/record/5543124/files/figure.png" pageId="2" pageNumber="467">Figs 31–33</figureCitation>
) with distal region completely divided into parameres; parameres (
<emphasis id="B921EABB2C0B076256D1E11CB695FA42" box="[410,450,295,320]" italics="true" pageId="2" pageNumber="467">e.g.</emphasis>
<figureCitation id="136E2A2C2C0B07625683E11CB512FA43" box="[456,581,295,321]" captionStart="FIGURES 54–70" captionStartId="8.[151,264,1849,1873]" captionTargetBox="[244,1344,182,1822]" captionTargetId="figure-31@8.[242,1346,181,1825]" captionTargetPageId="8" captionText="FIGURES 54–70. Philippistenia heterotarsalis (Heller, 1923). 54–57. Holotype of Distenia samarensis Villiers, 1959, male, copyright MNHN Paris. 54. Dorsal view; 55. Ventral view; 56. Lateral view; 57. Labels. 58–70. Male specimen. 58. Frontal view; 59. Dorsal view; 60. Ventral view; 61. Lateral view; 62. Tergite VIII; 63. Sternite VIII; 64. Ventral arc; 65–67. Median lobe. 65. Dorsal view; 66. Ventral view; 67. Lateral view. 68–70. Tegmen. 68. Dorsal view; 69. Ventral view; 70. Lateral view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5543132" httpUri="https://zenodo.org/record/5543132/files/figure.png" pageId="2" pageNumber="467">
Figs 68
<emphasis id="B921EABB2C0B07625551E11CB57FFA43" bold="true" box="[538,552,295,321]" pageId="2" pageNumber="467">–</emphasis>
70
</figureCitation>
) cylindrical-elongated, subparallel, bearing both short and long setae, mainly at apex; apex rounded; ring piece elongated, proximal part connected only by narrow membrane, without projection. Median lobe (
<emphasis id="B921EABB2C0B0762567DE154B609FA8A" box="[310,350,367,392]" italics="true" pageId="2" pageNumber="467">e.g.</emphasis>
<figureCitation id="136E2A2C2C0B0762562DE154B6B2FA8B" box="[358,485,367,393]" captionStart="FIGURES 34–46" captionStartId="6.[151,264,1309,1333]" captionTargetBox="[153,1436,182,1284]" captionTargetId="figure-31@6.[151,1436,181,1285]" captionTargetPageId="6" captionText="FIGURES 34–46. Philippistenia halconensis (Vives, 2012). 34. Holotype, male; 35–37. Female. 35. Dorsal view; 36. Lateral view; 37. Frontal view. 38–46. Male terminalia. 38. Tergite VIII; 39. Sternite VIII; 40. Ventral arc; 41–43. Median lobe. 41. Dorsal view; 42. Ventral view; 43. Lateral view. 44–46. Tegmen. 44. Dorsal view; 45. Ventral view; 46. Lateral view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5543126" httpUri="https://zenodo.org/record/5543126/files/figure.png" pageId="2" pageNumber="467">
Figs 41
<emphasis id="B921EABB2C0B076256F1E154B69FFA8B" bold="true" box="[442,456,367,393]" pageId="2" pageNumber="467">–</emphasis>
43
</figureCitation>
) about 1.3 times length of tegmen; curved at apical third in lateral view; dorsal lobe shorter than ventral lobe; basal apophysis very short.
</paragraph>
</subSubSection>
<subSubSection id="C34F65222C0B0762578CE18CB4BBFAF7" pageId="2" pageNumber="467" type="etymology">
<paragraph id="8BEA36A92C0B0762578CE18CB4BBFAF7" blockId="2.[151,1437,150,501]" pageId="2" pageNumber="467">
<emphasis id="B921EABB2C0B0762578CE18CB619FAD3" bold="true" box="[199,334,439,465]" pageId="2" pageNumber="467">Etymology.</emphasis>
The name
<taxonomicName id="4C554D2A2C0B07625684E18CB53BFAD3" authorityName="Botero &amp; Vives" authorityYear="2021" box="[463,620,439,465]" class="Insecta" family="Disteniidae" genus="Philippistenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="467" phylum="Arthropoda" rank="genus">
<emphasis id="B921EABB2C0B07625684E18CB53BFAD3" box="[463,620,439,465]" italics="true" pageId="2" pageNumber="467">Philippistenia</emphasis>
</taxonomicName>
is a compound of the words
<collectingCountry id="F34276392C0B076254F1E18CB36CFAD3" box="[954,1083,439,465]" name="Philippines" pageId="2" pageNumber="467">Philippines</collectingCountry>
, referring to the country where the species of the new genus are distributed, and
<taxonomicName id="4C554D2A2C0B076255F7E1E0B44CFAF6" authorityName="Lepeletier &amp; Audinet-Serville" authorityYear="1828" box="[700,795,475,500]" class="Insecta" family="Cerambycidae" genus="Distenia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="467" phylum="Arthropoda" rank="genus">
<emphasis id="B921EABB2C0B076255F7E1E0B44CFAF6" box="[700,795,475,500]" italics="true" pageId="2" pageNumber="467">Distenia</emphasis>
</taxonomicName>
. Feminine gender.
</paragraph>
</subSubSection>
<caption id="DF2A66212C0B076257DCE707B5B9FCC2" ID-DOI="http://doi.org/10.5281/zenodo.5543120" ID-Zenodo-Dep="5543120" httpUri="https://zenodo.org/record/5543120/files/figure.png" pageId="2" pageNumber="467" startId="2.[151,264,1852,1876]" targetBox="[151,1435,526,1826]" targetPageId="2">
<paragraph id="8BEA36A92C0B076257DCE707B5B9FCC2" blockId="2.[151,1436,1852,1984]" pageId="2" pageNumber="467">
<emphasis id="B921EABB2C0B076257DCE707B6D1FC56" bold="true" box="[151,390,1852,1876]" pageId="2" pageNumber="467">FIGURES 6–19. 6–18.</emphasis>
<taxonomicName id="4C554D2A2C0B076256C7E706B6B5FC56" authorityName="Lepeletier &amp; Audinet-Serville" authorityYear="1828" box="[396,482,1853,1876]" class="Insecta" family="Cerambycidae" genus="Distenia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="467" phylum="Arthropoda" rank="genus">
<emphasis id="B921EABB2C0B076256C7E706B6B5FC56" box="[396,482,1853,1876]" italics="true" pageId="2" pageNumber="467">Distenia</emphasis>
</taxonomicName>
(
<emphasis id="B921EABB2C0B076256BAE706B554FC56" box="[497,515,1853,1876]" italics="true" pageId="2" pageNumber="467">D</emphasis>
.)
<emphasis id="B921EABB2C0B07625553E707B5D4FC56" box="[536,643,1852,1876]" italics="true" pageId="2" pageNumber="467">columbina</emphasis>
Lepeletier &amp; Audinet-Serville, 1828, male.
<emphasis id="B921EABB2C0B0762530AE707B303FC56" bold="true" box="[1089,1108,1852,1876]" pageId="2" pageNumber="467">6.</emphasis>
Dorsal view;
<emphasis id="B921EABB2C0B076253A8E707B3A1FC56" bold="true" box="[1251,1270,1852,1876]" pageId="2" pageNumber="467">7.</emphasis>
Ventral view;
<emphasis id="B921EABB2C0B076252C2E707B2CBFC56" bold="true" box="[1417,1436,1852,1876]" pageId="2" pageNumber="467">8.</emphasis>
Lateral view;
<emphasis id="B921EABB2C0B0762566EE75BB66FFC7A" bold="true" box="[293,312,1888,1912]" pageId="2" pageNumber="467">9.</emphasis>
Frontal view;
<emphasis id="B921EABB2C0B07625684E75BB542FC7A" bold="true" box="[463,533,1888,1912]" pageId="2" pageNumber="467">10–18.</emphasis>
Male terminalia.
<emphasis id="B921EABB2C0B07625580E75BB5BCFC7A" bold="true" box="[715,747,1888,1912]" pageId="2" pageNumber="467">10.</emphasis>
Tergite VIII;
<emphasis id="B921EABB2C0B07625432E75BB4CFFC7A" bold="true" box="[889,920,1888,1912]" pageId="2" pageNumber="467">11.</emphasis>
Sternite VIII;
<emphasis id="B921EABB2C0B07625365E75BB319FC7A" bold="true" box="[1070,1102,1888,1912]" pageId="2" pageNumber="467">12.</emphasis>
Ventral arc;
<emphasis id="B921EABB2C0B07625399E75BB240FC7A" bold="true" box="[1234,1303,1888,1912]" pageId="2" pageNumber="467">13–15.</emphasis>
Tegmen.
<emphasis id="B921EABB2C0B07625236E75BB2CBFC7A" bold="true" box="[1405,1436,1888,1912]" pageId="2" pageNumber="467">13.</emphasis>
Dorsal view;
<emphasis id="B921EABB2C0B0762566AE7BFB616FC9E" bold="true" box="[289,321,1924,1948]" pageId="2" pageNumber="467">14.</emphasis>
Ventral view;
<emphasis id="B921EABB2C0B0762569DE7BFB6A1FC9E" bold="true" box="[470,502,1924,1948]" pageId="2" pageNumber="467">15.</emphasis>
Lateral view.
<emphasis id="B921EABB2C0B076255C3E7BFB599FC9E" bold="true" box="[648,718,1924,1948]" pageId="2" pageNumber="467">16–18.</emphasis>
Median lobe.
<emphasis id="B921EABB2C0B07625429E7BFB4D5FC9E" bold="true" box="[866,898,1924,1948]" pageId="2" pageNumber="467">16.</emphasis>
Dorsal view;
<emphasis id="B921EABB2C0B07625358E7BFB365FC9E" bold="true" box="[1043,1074,1924,1948]" pageId="2" pageNumber="467">17.</emphasis>
Ventral view;
<emphasis id="B921EABB2C0B07625383E7BFB3BFFC9E" bold="true" box="[1224,1256,1924,1948]" pageId="2" pageNumber="467">18.</emphasis>
Lateral view;
<emphasis id="B921EABB2C0B07625236E7BFB2CBFC9E" bold="true" box="[1405,1436,1924,1948]" pageId="2" pageNumber="467">19.</emphasis>
<taxonomicName id="4C554D2A2C0B076257DCE793B506FCC2" authority="Thomson, 1864" authorityName="Thomson" authorityYear="1864" box="[151,593,1960,1984]" class="Insecta" family="Cerambycidae" genus="Typodryas" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="467" phylum="Arthropoda" rank="species" species="callichromoides">
<emphasis id="B921EABB2C0B076257DCE793B6FEFCC2" box="[151,425,1960,1984]" italics="true" pageId="2" pageNumber="467">Typodryas callichromoides</emphasis>
Thomson, 1864
</taxonomicName>
. Scale = 1 mm.
</paragraph>
</caption>
<subSubSection id="C34F65222C0A0763578CE0ADB3E8F816" pageId="3" pageNumber="468" type="discussion">
<paragraph id="8BEA36A92C0A0763578CE0ADB325FAAF" blockId="3.[151,1437,150,789]" pageId="3" pageNumber="468">
<emphasis id="B921EABB2C0A0763578CE0ADB66CFBB2" bold="true" box="[199,315,150,176]" pageId="3" pageNumber="468">Remarks.</emphasis>
The genus
<taxonomicName id="4C554D2A2C0A076356F1E0ADB500FBB2" authorityName="Botero &amp; Vives" authorityYear="2021" box="[442,599,150,176]" class="Insecta" family="Disteniidae" genus="Philippistenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="468" phylum="Arthropoda" rank="genus">
<emphasis id="B921EABB2C0A076356F1E0ADB500FBB2" box="[442,599,150,176]" italics="true" pageId="3" pageNumber="468">Philippistenia</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C554D2A2C0A076355ACE0ACB411FBB2" authorityName="Lepeletier &amp; Audinet-Serville" authorityYear="1828" box="[743,838,151,176]" class="Insecta" family="Cerambycidae" genus="Distenia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="468" phylum="Arthropoda" rank="genus">
<emphasis id="B921EABB2C0A076355ACE0ACB411FBB2" box="[743,838,151,176]" italics="true" pageId="3" pageNumber="468">Distenia</emphasis>
</taxonomicName>
(
<figureCitation id="136E2A2C2C0A0763541FE0ADB4CDFBB2" box="[852,922,150,176]" captionStart="FIGURES 1–5. 1-3" captionStartId="1.[151,264,1881,1905]" captionTargetBox="[151,1436,1529,1857]" captionTargetId="figure-403@1.[151,1436,1528,1857]" captionTargetPageId="1" captionText="FIGURES 1–5. 1-3. Detail of posterior area of head and pronotum. 1. Distenia (D.) columbina Lepeletier &amp; Audinet-Serville, 1828; 2. Philippistenia levitemporalis (Heller, 1924); 3. Typodryas callichromoides Thomson, 1864; 4–5. Detail of the elytra showing the longitudinal black bands of pubescence. 4. Philippistenia samarensis (Villiers, 1959); 5. Philippistenia heterotarsalis (Heller, 1923)." figureDoi="http://doi.org/10.5281/zenodo.5543118" httpUri="https://zenodo.org/record/5543118/files/figure.png" pageId="3" pageNumber="468">Figs 1</figureCitation>
,
<figureCitation id="136E2A2C2C0A076354EDE0ADB4B7FBB3" box="[934,992,150,177]" captionStart="FIGURES 6–19. 6–18" captionStartId="2.[151,264,1852,1876]" captionTargetBox="[151,1435,526,1826]" captionTargetId="figure-203@2.[151,1436,525,1829]" captionTargetPageId="2" captionText="FIGURES 6–19. 6–18. Distenia (D.) columbina Lepeletier &amp; Audinet-Serville, 1828, male. 6. Dorsal view; 7. Ventral view; 8. Lateral view; 9. Frontal view; 10–18. Male terminalia. 10. Tergite VIII; 11. Sternite VIII; 12. Ventral arc; 13–15. Tegmen. 13. Dorsal view; 14. Ventral view; 15. Lateral view. 16–18. Median lobe. 16. Dorsal view; 17. Ventral view; 18. Lateral view; 19. Typodryas callichromoides Thomson, 1864. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5543120" httpUri="https://zenodo.org/record/5543120/files/figure.png" pageId="3" pageNumber="468">6–18</figureCitation>
) by the head distinctly narrowed behind the eyes (
<figureCitation id="136E2A2C2C0A07635649E081B610FBD6" box="[258,327,186,212]" captionStart="FIGURES 1–5. 1-3" captionStartId="1.[151,264,1881,1905]" captionTargetBox="[151,1436,1529,1857]" captionTargetId="figure-403@1.[151,1436,1528,1857]" captionTargetPageId="1" captionText="FIGURES 1–5. 1-3. Detail of posterior area of head and pronotum. 1. Distenia (D.) columbina Lepeletier &amp; Audinet-Serville, 1828; 2. Philippistenia levitemporalis (Heller, 1924); 3. Typodryas callichromoides Thomson, 1864; 4–5. Detail of the elytra showing the longitudinal black bands of pubescence. 4. Philippistenia samarensis (Villiers, 1959); 5. Philippistenia heterotarsalis (Heller, 1923)." figureDoi="http://doi.org/10.5281/zenodo.5543118" httpUri="https://zenodo.org/record/5543118/files/figure.png" pageId="3" pageNumber="468">Fig. 2</figureCitation>
); elytra with longitudinal bands of dense black pubescence, always present on the suture (
<emphasis id="B921EABB2C0A07635277E080B233FBD6" box="[1340,1380,187,212]" italics="true" pageId="3" pageNumber="468">e.g.</emphasis>
<figureCitation id="136E2A2C2C0A07635227E081B792FBFB" captionStart="FIGURES 1–5. 1-3" captionStartId="1.[151,264,1881,1905]" captionTargetBox="[151,1436,1529,1857]" captionTargetId="figure-403@1.[151,1436,1528,1857]" captionTargetPageId="1" captionText="FIGURES 1–5. 1-3. Detail of posterior area of head and pronotum. 1. Distenia (D.) columbina Lepeletier &amp; Audinet-Serville, 1828; 2. Philippistenia levitemporalis (Heller, 1924); 3. Typodryas callichromoides Thomson, 1864; 4–5. Detail of the elytra showing the longitudinal black bands of pubescence. 4. Philippistenia samarensis (Villiers, 1959); 5. Philippistenia heterotarsalis (Heller, 1923)." figureDoi="http://doi.org/10.5281/zenodo.5543118" httpUri="https://zenodo.org/record/5543118/files/figure.png" pageId="3" pageNumber="468">Figs 4–5</figureCitation>
); and the metafemora long, almost reaching the elytral apex. In
<taxonomicName id="4C554D2A2C0A076354E0E0E4B35DFBFA" authorityName="Lepeletier &amp; Audinet-Serville" authorityYear="1828" box="[939,1034,223,248]" class="Insecta" family="Cerambycidae" genus="Distenia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="468" phylum="Arthropoda" rank="genus">
<emphasis id="B921EABB2C0A076354E0E0E4B35DFBFA" box="[939,1034,223,248]" italics="true" pageId="3" pageNumber="468">Distenia</emphasis>
</taxonomicName>
, the head is not strongly narrowed behind the eyes (
<figureCitation id="136E2A2C2C0A07635616E139B6F5FA1E" box="[349,418,258,284]" captionStart="FIGURES 1–5. 1-3" captionStartId="1.[151,264,1881,1905]" captionTargetBox="[151,1436,1529,1857]" captionTargetId="figure-403@1.[151,1436,1528,1857]" captionTargetPageId="1" captionText="FIGURES 1–5. 1-3. Detail of posterior area of head and pronotum. 1. Distenia (D.) columbina Lepeletier &amp; Audinet-Serville, 1828; 2. Philippistenia levitemporalis (Heller, 1924); 3. Typodryas callichromoides Thomson, 1864; 4–5. Detail of the elytra showing the longitudinal black bands of pubescence. 4. Philippistenia samarensis (Villiers, 1959); 5. Philippistenia heterotarsalis (Heller, 1923)." figureDoi="http://doi.org/10.5281/zenodo.5543118" httpUri="https://zenodo.org/record/5543118/files/figure.png" pageId="3" pageNumber="468">Fig. 1</figureCitation>
); the elytra do not have longitudinal bands of dense black pubescence (
<figureCitation id="136E2A2C2C0A076353ACE139B278FA1F" box="[1255,1327,258,285]" captionStart="FIGURES 6–19. 6–18" captionStartId="2.[151,264,1852,1876]" captionTargetBox="[151,1435,526,1826]" captionTargetId="figure-203@2.[151,1436,525,1829]" captionTargetPageId="2" captionText="FIGURES 6–19. 6–18. Distenia (D.) columbina Lepeletier &amp; Audinet-Serville, 1828, male. 6. Dorsal view; 7. Ventral view; 8. Lateral view; 9. Frontal view; 10–18. Male terminalia. 10. Tergite VIII; 11. Sternite VIII; 12. Ventral arc; 13–15. Tegmen. 13. Dorsal view; 14. Ventral view; 15. Lateral view. 16–18. Median lobe. 16. Dorsal view; 17. Ventral view; 18. Lateral view; 19. Typodryas callichromoides Thomson, 1864. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5543120" httpUri="https://zenodo.org/record/5543120/files/figure.png" pageId="3" pageNumber="468">Fig. 6</figureCitation>
); and the metafemora are short, just reaching the apical third of the elytra. Comparing the male terminalia of
<taxonomicName id="4C554D2A2C0A076353B4E11DB7BFFA66" class="Insecta" family="Disteniidae" genus="Philippistenia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="468" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B921EABB2C0A076353B4E11DB2CBFA42" box="[1279,1436,294,320]" italics="true" pageId="3" pageNumber="468">Philippistenia</emphasis>
species
</taxonomicName>
with the male terminalia of
<taxonomicName id="4C554D2A2C0A0763556CE170B5D1FA66" authorityName="Lepeletier &amp; Audinet-Serville" authorityYear="1828" box="[551,646,331,356]" class="Insecta" family="Cerambycidae" genus="Distenia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="468" phylum="Arthropoda" rank="genus">
<emphasis id="B921EABB2C0A0763556CE170B5D1FA66" box="[551,646,331,356]" italics="true" pageId="3" pageNumber="468">Distenia</emphasis>
</taxonomicName>
(
<emphasis id="B921EABB2C0A076355DDE170B5FDFA66" box="[662,682,331,356]" italics="true" pageId="3" pageNumber="468">D</emphasis>
.)
<emphasis id="B921EABB2C0A07635589E171B46FFA66" box="[706,824,330,356]" italics="true" pageId="3" pageNumber="468">columbina</emphasis>
(
<figureCitation id="136E2A2C2C0A0763540CE171B4EEFA66" box="[839,953,330,356]" captionStart-0="FIGURES 1–5. 1-3" captionStart-1="FIGURES 6–19. 6–18" captionStartId-0="1.[151,264,1881,1905]" captionStartId-1="2.[151,264,1852,1876]" captionTargetBox-0="[151,1436,1529,1857]" captionTargetBox-1="[151,1435,526,1826]" captionTargetId-0="figure-403@1.[151,1436,1528,1857]" captionTargetId-1="figure-203@2.[151,1436,525,1829]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURES 1–5. 1-3. Detail of posterior area of head and pronotum. 1. Distenia (D.) columbina Lepeletier &amp; Audinet-Serville, 1828; 2. Philippistenia levitemporalis (Heller, 1924); 3. Typodryas callichromoides Thomson, 1864; 4–5. Detail of the elytra showing the longitudinal black bands of pubescence. 4. Philippistenia samarensis (Villiers, 1959); 5. Philippistenia heterotarsalis (Heller, 1923)." captionText-1="FIGURES 6–19. 6–18. Distenia (D.) columbina Lepeletier &amp; Audinet-Serville, 1828, male. 6. Dorsal view; 7. Ventral view; 8. Lateral view; 9. Frontal view; 10–18. Male terminalia. 10. Tergite VIII; 11. Sternite VIII; 12. Ventral arc; 13–15. Tegmen. 13. Dorsal view; 14. Ventral view; 15. Lateral view. 16–18. Median lobe. 16. Dorsal view; 17. Ventral view; 18. Lateral view; 19. Typodryas callichromoides Thomson, 1864. Scale = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5543118" figureDoi-1="http://doi.org/10.5281/zenodo.5543120" httpUri-0="https://zenodo.org/record/5543118/files/figure.png" httpUri-1="https://zenodo.org/record/5543120/files/figure.png" pageId="3" pageNumber="468">Figs 1–18</figureCitation>
) is possible to observe that in the latter the tergite VIII (
<figureCitation id="136E2A2C2C0A07635669E155B624FA8A" box="[290,371,366,392]" captionStart="FIGURES 6–19. 6–18" captionStartId="2.[151,264,1852,1876]" captionTargetBox="[151,1435,526,1826]" captionTargetId="figure-203@2.[151,1436,525,1829]" captionTargetPageId="2" captionText="FIGURES 6–19. 6–18. Distenia (D.) columbina Lepeletier &amp; Audinet-Serville, 1828, male. 6. Dorsal view; 7. Ventral view; 8. Lateral view; 9. Frontal view; 10–18. Male terminalia. 10. Tergite VIII; 11. Sternite VIII; 12. Ventral arc; 13–15. Tegmen. 13. Dorsal view; 14. Ventral view; 15. Lateral view. 16–18. Median lobe. 16. Dorsal view; 17. Ventral view; 18. Lateral view; 19. Typodryas callichromoides Thomson, 1864. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5543120" httpUri="https://zenodo.org/record/5543120/files/figure.png" pageId="3" pageNumber="468">Fig. 10</figureCitation>
) and the apophysis of sternite VIII (
<figureCitation id="136E2A2C2C0A07635449E155B406FA8A" box="[770,849,366,392]" captionStart="FIGURES 6–19. 6–18" captionStartId="2.[151,264,1852,1876]" captionTargetBox="[151,1435,526,1826]" captionTargetId="figure-203@2.[151,1436,525,1829]" captionTargetPageId="2" captionText="FIGURES 6–19. 6–18. Distenia (D.) columbina Lepeletier &amp; Audinet-Serville, 1828, male. 6. Dorsal view; 7. Ventral view; 8. Lateral view; 9. Frontal view; 10–18. Male terminalia. 10. Tergite VIII; 11. Sternite VIII; 12. Ventral arc; 13–15. Tegmen. 13. Dorsal view; 14. Ventral view; 15. Lateral view. 16–18. Median lobe. 16. Dorsal view; 17. Ventral view; 18. Lateral view; 19. Typodryas callichromoides Thomson, 1864. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5543120" httpUri="https://zenodo.org/record/5543120/files/figure.png" pageId="3" pageNumber="468">Fig. 11</figureCitation>
) are longer than in
<taxonomicName id="4C554D2A2C0A07635363E155B392FA8A" authorityName="Botero &amp; Vives" authorityYear="2021" box="[1064,1221,366,392]" class="Insecta" family="Disteniidae" genus="Philippistenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="468" phylum="Arthropoda" rank="genus">
<emphasis id="B921EABB2C0A07635363E155B392FA8A" box="[1064,1221,366,392]" italics="true" pageId="3" pageNumber="468">Philippistenia</emphasis>
</taxonomicName>
, and the parameres of tegmen are divergent apically (
<figureCitation id="136E2A2C2C0A0763555BE1A9B5DCFAAF" box="[528,651,402,429]" captionStart="FIGURES 6–19. 6–18" captionStartId="2.[151,264,1852,1876]" captionTargetBox="[151,1435,526,1826]" captionTargetId="figure-203@2.[151,1436,525,1829]" captionTargetPageId="2" captionText="FIGURES 6–19. 6–18. Distenia (D.) columbina Lepeletier &amp; Audinet-Serville, 1828, male. 6. Dorsal view; 7. Ventral view; 8. Lateral view; 9. Frontal view; 10–18. Male terminalia. 10. Tergite VIII; 11. Sternite VIII; 12. Ventral arc; 13–15. Tegmen. 13. Dorsal view; 14. Ventral view; 15. Lateral view. 16–18. Median lobe. 16. Dorsal view; 17. Ventral view; 18. Lateral view; 19. Typodryas callichromoides Thomson, 1864. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5543120" httpUri="https://zenodo.org/record/5543120/files/figure.png" pageId="3" pageNumber="468">Fig. 13–15</figureCitation>
) (in
<taxonomicName id="4C554D2A2C0A0763558AE1A9B4E1FAAE" box="[705,950,402,428]" class="Insecta" family="Disteniidae" genus="Philippistenia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="468" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B921EABB2C0A0763558AE1A9B409FAAE" box="[705,862,402,428]" italics="true" pageId="3" pageNumber="468">Philippistenia</emphasis>
species
</taxonomicName>
are subparallel).
</paragraph>
<paragraph id="8BEA36A92C0A0763578CE18DB6D5F93E" blockId="3.[151,1437,150,789]" pageId="3" pageNumber="468">
The new genus can be also differentiated from
<taxonomicName id="4C554D2A2C0A07635580E18DB4A5FAD2" authority="Thomson, 1864" authorityName="Thomson" authorityYear="1864" box="[715,1010,438,464]" class="Insecta" family="Cerambycidae" genus="Typodryas" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="468" phylum="Arthropoda" rank="genus">
<emphasis id="B921EABB2C0A07635580E18DB468FAD2" box="[715,831,438,464]" italics="true" pageId="3" pageNumber="468">Typodryas</emphasis>
Thomson, 1864
</taxonomicName>
(
<figureCitation id="136E2A2C2C0A0763534BE18DB307FAD2" box="[1024,1104,438,464]" captionStart="FIGURES 6–19. 6–18" captionStartId="2.[151,264,1852,1876]" captionTargetBox="[151,1435,526,1826]" captionTargetId="figure-203@2.[151,1436,525,1829]" captionTargetPageId="2" captionText="FIGURES 6–19. 6–18. Distenia (D.) columbina Lepeletier &amp; Audinet-Serville, 1828, male. 6. Dorsal view; 7. Ventral view; 8. Lateral view; 9. Frontal view; 10–18. Male terminalia. 10. Tergite VIII; 11. Sternite VIII; 12. Ventral arc; 13–15. Tegmen. 13. Dorsal view; 14. Ventral view; 15. Lateral view. 16–18. Median lobe. 16. Dorsal view; 17. Ventral view; 18. Lateral view; 19. Typodryas callichromoides Thomson, 1864. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5543120" httpUri="https://zenodo.org/record/5543120/files/figure.png" pageId="3" pageNumber="468">Fig. 19</figureCitation>
) by the absence of transversal wrinkles on the vertex and pronotum (
<figureCitation id="136E2A2C2C0A0763550FE1E1B5DFFAF6" box="[580,648,474,500]" captionStart="FIGURES 1–5. 1-3" captionStartId="1.[151,264,1881,1905]" captionTargetBox="[151,1436,1529,1857]" captionTargetId="figure-403@1.[151,1436,1528,1857]" captionTargetPageId="1" captionText="FIGURES 1–5. 1-3. Detail of posterior area of head and pronotum. 1. Distenia (D.) columbina Lepeletier &amp; Audinet-Serville, 1828; 2. Philippistenia levitemporalis (Heller, 1924); 3. Typodryas callichromoides Thomson, 1864; 4–5. Detail of the elytra showing the longitudinal black bands of pubescence. 4. Philippistenia samarensis (Villiers, 1959); 5. Philippistenia heterotarsalis (Heller, 1923)." figureDoi="http://doi.org/10.5281/zenodo.5543118" httpUri="https://zenodo.org/record/5543118/files/figure.png" pageId="3" pageNumber="468">Fig. 2</figureCitation>
), and the elytra with longitudinal bands of dense black pubescence. In
<taxonomicName id="4C554D2A2C0A076357DCE1C5B65CF91A" authorityName="Thomson" authorityYear="1864" box="[151,267,510,536]" class="Insecta" family="Cerambycidae" genus="Typodryas" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="468" phylum="Arthropoda" rank="genus">
<emphasis id="B921EABB2C0A076357DCE1C5B65CF91A" box="[151,267,510,536]" italics="true" pageId="3" pageNumber="468">Typodryas</emphasis>
</taxonomicName>
, the vertex and pronotum have transversal wrinkles (
<figureCitation id="136E2A2C2C0A07635413E1C5B4CDF91A" box="[856,922,510,536]" captionStart="FIGURES 1–5. 1-3" captionStartId="1.[151,264,1881,1905]" captionTargetBox="[151,1436,1529,1857]" captionTargetId="figure-403@1.[151,1436,1528,1857]" captionTargetPageId="1" captionText="FIGURES 1–5. 1-3. Detail of posterior area of head and pronotum. 1. Distenia (D.) columbina Lepeletier &amp; Audinet-Serville, 1828; 2. Philippistenia levitemporalis (Heller, 1924); 3. Typodryas callichromoides Thomson, 1864; 4–5. Detail of the elytra showing the longitudinal black bands of pubescence. 4. Philippistenia samarensis (Villiers, 1959); 5. Philippistenia heterotarsalis (Heller, 1923)." figureDoi="http://doi.org/10.5281/zenodo.5543118" httpUri="https://zenodo.org/record/5543118/files/figure.png" pageId="3" pageNumber="468">Fig. 3</figureCitation>
) and the elytra do not have longitudinal bands of black pubescence.
</paragraph>
<paragraph id="8BEA36A92C0A0763578CE27DB488F9F2" blockId="3.[151,1437,150,789]" pageId="3" pageNumber="468">
<taxonomicName id="4C554D2A2C0A0763578CE27DB633F962" authorityName="Botero &amp; Vives" authorityYear="2021" box="[199,356,582,608]" class="Insecta" family="Disteniidae" genus="Philippistenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="468" phylum="Arthropoda" rank="genus">
<emphasis id="B921EABB2C0A0763578CE27DB633F962" box="[199,356,582,608]" italics="true" pageId="3" pageNumber="468">Philippistenia</emphasis>
</taxonomicName>
can be differentiated from
<taxonomicName id="4C554D2A2C0A076355D7E27CB323F962" authority="Santos-Silva &amp; Hovore, 2007" authorityName="Santos-Silva &amp; Hovore" authorityYear="2007" box="[668,1140,582,609]" class="Insecta" family="Cerambycidae" genus="Novantinoe" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="468" phylum="Arthropoda" rank="genus">
<emphasis id="B921EABB2C0A076355D7E27CB44AF962" box="[668,797,583,608]" italics="true" pageId="3" pageNumber="468">Novantinoe</emphasis>
Santos-Silva &amp; Hovore, 2007
</taxonomicName>
by the pronotum with the medial tubercle slightly elevated, poorly visible; elytra with longitudinal bands of dense black pubescence; and the metafemora long, almost reaching the elytral apex. In
<taxonomicName id="4C554D2A2C0A076355B9E2B4B424F9AA" authorityName="Santos-Silva &amp; Hovore" authorityYear="2007" box="[754,883,655,680]" class="Insecta" family="Cerambycidae" genus="Novantinoe" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="468" phylum="Arthropoda" rank="genus">
<emphasis id="B921EABB2C0A076355B9E2B4B424F9AA" box="[754,883,655,680]" italics="true" pageId="3" pageNumber="468">Novantinoe</emphasis>
</taxonomicName>
, the medial tubercle of pronotum is well elevated and clearly visible; the elytra do not have longitudinal bands of black pubescence, and the metafemora do not reach the elytral apex.
<taxonomicName id="4C554D2A2C0A07635619E2ECB684F9F2" authorityName="Santos-Silva &amp; Hovore" authorityYear="2007" box="[338,467,727,752]" class="Insecta" family="Cerambycidae" genus="Novantinoe" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="468" phylum="Arthropoda" rank="genus">
<emphasis id="B921EABB2C0A07635619E2ECB684F9F2" box="[338,467,727,752]" italics="true" pageId="3" pageNumber="468">Novantinoe</emphasis>
</taxonomicName>
is geographically restricted to the New World.
</paragraph>
<paragraph id="8BEA36A92C0A0763578CE2C1B3E8F816" blockId="3.[151,1437,150,789]" box="[199,1215,762,789]" pageId="3" pageNumber="468">
Currently,
<taxonomicName id="4C554D2A2C0A07635674E2C1B68BF816" authorityName="Botero &amp; Vives" authorityYear="2021" box="[319,476,762,788]" class="Insecta" family="Disteniidae" genus="Philippistenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="468" phylum="Arthropoda" rank="genus">
<emphasis id="B921EABB2C0A07635674E2C1B68BF816" box="[319,476,762,788]" italics="true" pageId="3" pageNumber="468">Philippistenia</emphasis>
</taxonomicName>
is an endemic genus from the Islands of the
<collectingCountry id="F34276392C0A0763549AE2C1B318F816" box="[977,1103,762,788]" name="Philippines" pageId="3" pageNumber="468">Philippines</collectingCountry>
(
<figureCitation id="136E2A2C2C0A07635315E2C1B3E6F816" box="[1118,1201,762,788]" captionStart="FIGURE 71" captionStartId="9.[151,250,1868,1892]" captionTargetBox="[163,1423,194,1831]" captionTargetId="figure-19@9.[151,1436,181,1844]" captionTargetPageId="9" captionText="FIGURE 71. Geographical distribution of Philippistenia species." figureDoi="http://doi.org/10.5281/zenodo.5543136" httpUri="https://zenodo.org/record/5543136/files/figure.png" pageId="3" pageNumber="468">Fig. 71</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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