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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.734.21989" ID-PMC="PMC5904327" ID-Pensoft-Pub="1313-2970-734-43" ID-Pensoft-UUID="394D3A78FFD46838FF9BFFE8FB52C216" ID-PubMed="29674855" ID-Zenodo-Dep="1222331" ID-ZooBank="7C3C687B64BB42A5B9E4EC588BCD52D5" ModsDocID="1313-2970-734-43" checkinTime="1517868722990" checkinUser="pensoft" docAuthor="Assmann, Thorsten, Boutaud, Esteve, Buse, Joern, Gebert, Joerg, Drees, Claudia, Friedman, Ariel-Leib-Leonid, Khoury, Fares, Marcus, Tamar, Orbach, Eylon, Ittai Renan,, Schmidt, Constantin &amp; Zumstein, Pascale" docDate="2018" docId="25B9446AF51A08CAD44C33C2AA4ABCA7" docLanguage="en" docName="ZooKeys 734: 43-103" docOrigin="ZooKeys 734" docPubDate="2018-02-05" docSource="http://dx.doi.org/10.3897/zookeys.734.21989" docTitle="Calomera fischeri" docType="treatment" docVersion="6" id="394D3A78FFD46838FF9BFFE8FB52C216" lastPageNumber="59" masterDocId="394D3A78FFD46838FF9BFFE8FB52C216" masterDocTitle="The tiger beetles (Coleoptera, Cicindelidae) of the southern Levant and adjacent territories: from cybertaxonomy to conservation biology" masterLastPageNumber="103" masterPageNumber="43" pageNumber="59" updateTime="1668165402329" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>The tiger beetles (Coleoptera, Cicindelidae) of the southern Levant and adjacent territories: from cybertaxonomy to conservation biology</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Assmann, Thorsten</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-9203-769X</mods:nameIdentifier>
<mods:affiliation>Institute of Ecology, Leuphana University Lueneburg, Universitaetsallee 1, D- 21335 Lueneburg, Germany</mods:affiliation>
<mods:nameIdentifier type="email">assmann@uni.leuphana.de</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Boutaud, Esteve</mods:namePart>
<mods:affiliation>Institute of Ecology, Leuphana University Lueneburg, Universitaetsallee 1, D- 21335 Lueneburg, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Buse, Joern</mods:namePart>
<mods:affiliation>Ecosystem Monitoring, Research and Wildlife Conservation (SB 23 Invertebrates and Biodiversity), Black Forest National Park, Kniebisstrasse 67, D- 72250 Freudenstadt, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gebert, Joerg</mods:namePart>
<mods:affiliation>Karl-Liebknecht-Strasse 73, D- 01109 Dresden. Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Drees, Claudia</mods:namePart>
<mods:affiliation>Steinhardt Museum of Natural History, Tel Aviv University, Ramat-Aviv, Tel Aviv, IL- 69978, Israel &amp; Biocentre Grindel, Universitaet Hamburg, Martin-Luther-King-Platz 3, D- 20146 Hamburg, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Friedman, Ariel-Leib-Leonid</mods:namePart>
<mods:affiliation>Steinhardt Museum of Natural History, Tel Aviv University, Ramat-Aviv, Tel Aviv, IL- 69978, Israel</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Khoury, Fares</mods:namePart>
<mods:affiliation>Department of Biology and Biotechnology, American University of Madaba, P. O. Box 2882, Amman, JO- 11821, Jordan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Marcus, Tamar</mods:namePart>
<mods:affiliation>Institute of Ecology, Leuphana University Lueneburg, Universitaetsallee 1, D- 21335 Lueneburg, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Orbach, Eylon</mods:namePart>
<mods:affiliation>Remez St. 49, IL- 36044 Qiryat Tiv'on, Israel</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ittai Renan,</mods:namePart>
<mods:affiliation>Steinhardt Museum of Natural History, Tel Aviv University, Ramat-Aviv, Tel Aviv, IL- 69978, Israel</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Schmidt, Constantin</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3892-4255</mods:nameIdentifier>
<mods:affiliation>Deichstr. 13, D- 21354 Bleckede, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zumstein, Pascale</mods:namePart>
<mods:affiliation>Institute of Ecology, Leuphana University Lueneburg, Universitaetsallee 1, D- 21335 Lueneburg, Germany</mods:affiliation>
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<mods:date>2018</mods:date>
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<mods:number>2018-02-05</mods:number>
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<mods:detail type="volume">
<mods:number>734</mods:number>
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<mods:start>43</mods:start>
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<mods:url>http://dx.doi.org/10.3897/zookeys.734.21989</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.734.21989</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-734-43</mods:identifier>
<mods:identifier type="ZooBank">7C3C687B64BB42A5B9E4EC588BCD52D5</mods:identifier>
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<treatment ID-GBIF-Taxon="141125723" LSID="urn:lsid:plazi:treatment:25B9446AF51A08CAD44C33C2AA4ABCA7" httpUri="http://treatment.plazi.org/id/25B9446AF51A08CAD44C33C2AA4ABCA7" lastPageNumber="59" pageId="16" pageNumber="59">
<subSubSection pageId="16" pageNumber="59" type="nomenclature">
<paragraph pageId="16" pageNumber="59">
6.
<taxonomicName LSID="25B9446A-F51A-08CA-D44C-33C2AA4ABCA7" authority="(Adams, 1817)" baseAuthorityName="Adams" baseAuthorityYear="1817" class="Insecta" family="Cicindelidae" genus="Calomera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calomera fischeri" order="Coleoptera" pageId="16" pageNumber="59" phylum="Arthropoda" rank="species" species="fischeri">Calomera fischeri (Adams, 1817)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="59" type="habitat">
<paragraph pageId="16" pageNumber="59">Habitat.</paragraph>
<paragraph pageId="16" pageNumber="59">
On river banks and next to freshwater ponds with sparse vegetation on sandy, sometimes cohesive soil (
<bibRefCitation author="Arndt, E" editor="Arndt, E" journalOrPublisher="Pensoft, Sofia" pageId="38" pageNumber="81" pagination="73 - 76" refId="B8" refString="Arndt, E, 2011. Tribe Cicindelini Latreille, 1802. In: Arndt, E, Schnitter, P, Sfenthourakis, S, Wrase, DW, Eds., Ground beetles (Carabidae) of Greece. Pensoft, Sofia: 73 - 76" title="Tribe Cicindelini Latreille, 1802." volumeTitle="Ground beetles (Carabidae) of Greece." year="2011">Arndt 2011</bibRefCitation>
;
<bibRefCitation author="Avgin, SS" journalOrPublisher="BAUE Fen Bil Enst Degisi" pageId="38" pageNumber="81" pagination="23 - 28" refId="B14" refString="Avgin, SS, 2006. General information and tiger beetles Cicindela herbacea; Calomera fischeri fischeri and Homodela ismenia kilikiensis collected from southeast of Turkey (Coleoptera: Cicindelidae). BAUE Fen Bil Enst Degisi 8: 23 - 28" title="General information and tiger beetles Cicindela herbacea; Calomera fischeri fischeri and Homodela ismenia kilikiensis collected from southeast of Turkey (Coleoptera: Cicindelidae)." volume="8" year="2006">Avgin 2006</bibRefCitation>
;
<bibRefCitation author="Werner, K" journalOrPublisher="Science Nat, Venette" pageId="44" pageNumber="87" refId="B110" refString="Werner, K, 1992. Cicindelidae 2. Science Nat, Venette" title="Cicindelidae 2." year="1992">Werner 1992</bibRefCitation>
). In desert habitats the species can be widespread (cf.
<bibRefCitation author="Wiesner, J" journalOrPublisher="Entomologische Zeitschrift" pageId="45" pageNumber="88" pagination="382 - 384" refId="B115" refString="Wiesner, J, 1996. Weitere Mitteilungen ueber die Cicindelidae (Coleoptera) der Vereinigten Arabischen Emirate. Entomologische Zeitschrift 106: 382 - 384" title="Weitere Mitteilungen ueber die Cicindelidae (Coleoptera) der Vereinigten Arabischen Emirate." volume="106" year="1996">Wiesner 1996</bibRefCitation>
). Diurnal.
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="59" type="phenology">
<paragraph pageId="16" pageNumber="59">Phenology.</paragraph>
<paragraph pageId="16" pageNumber="59">
In Turkey, adults from the end of May to the beginning of September (
<bibRefCitation author="Avgin, SS" journalOrPublisher="BAUE Fen Bil Enst Degisi" pageId="38" pageNumber="81" pagination="23 - 28" refId="B14" refString="Avgin, SS, 2006. General information and tiger beetles Cicindela herbacea; Calomera fischeri fischeri and Homodela ismenia kilikiensis collected from southeast of Turkey (Coleoptera: Cicindelidae). BAUE Fen Bil Enst Degisi 8: 23 - 28" title="General information and tiger beetles Cicindela herbacea; Calomera fischeri fischeri and Homodela ismenia kilikiensis collected from southeast of Turkey (Coleoptera: Cicindelidae)." volume="8" year="2006">Avgin 2006</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="59" type="distribution">
<paragraph pageId="16" pageNumber="59">Distribution range.</paragraph>
<paragraph pageId="16" pageNumber="59">
from southeastern Europe to central Asia and India, southwards to Turkey and Syria (
<bibRefCitation author="Acciavatti, RE" journalOrPublisher="Annales of Carnegie Museum" pageId="38" pageNumber="81" pagination="77 - 353" refId="B4" refString="Acciavatti, RE, Pearson, DL, 1989. The tiger beetle genus Cicindela (Coleoptera, Insecta) from the Indian subcontinent. Annales of Carnegie Museum 58: 77 - 353" title="The tiger beetle genus Cicindela (Coleoptera, Insecta) from the Indian subcontinent." volume="58" year="1989">Acciavatti and Pearson 1989</bibRefCitation>
;
<bibRefCitation author="Werner, K" journalOrPublisher="Science Nat, Venette" pageId="44" pageNumber="87" refId="B109" refString="Werner, K, 1991. Cicindelidae 1. Science Nat, Venette" title="Cicindelidae 1." year="1991">Werner 1991</bibRefCitation>
;
<bibRefCitation author="Wiesner, J" journalOrPublisher="Erna Bauer, Keltern" pageId="45" pageNumber="88" refId="B114" refString="Wiesner, J, 1992. Checklist of the tiger beetles of the world. Erna Bauer, Keltern" title="Checklist of the tiger beetles of the world." year="1992">Wiesner 1992</bibRefCitation>
).
<bibRefCitation author="Austin, K" journalOrPublisher="Annales du Museum d'Histoire Naturelle de Nice" pageId="38" pageNumber="81" pagination="1 - 199" refId="B13" refString="Austin, K, Small, E, Lemaire, J-M, Jeanne, C, Makris, C, Georghiou, G, 2008. Revision du Catalogue des Carabidae (Coleoptera) de Chypre. A revised catalogue of the Carabidae (Coleoptera) of Cyprus. Annales du Museum d'Histoire Naturelle de Nice 23: 1 - 199" title="Revision du Catalogue des Carabidae (Coleoptera) de Chypre. A revised catalogue of the Carabidae (Coleoptera) of Cyprus." volume="23" year="2008">Austin et al. (2008</bibRefCitation>
: 22) questioned the occurrence on Cyprus. However, Horn and Roeschkle (1891) list the species for Cyprus, and old records exist in SDEI: &lt;Cyprus, Baudi&gt; (3 specimens in the collections of Kraatz and Rottenberg, Lutz Behne, pers. com.) Therefore, the species is listed correctly by
<bibRefCitation author="Putchkov, AV" editor="Loebl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="44" pageNumber="87" pagination="99 - 118" refId="B94" refString="Putchkov, AV, Matalin, AV, 2003. Subfamily Cicindelinae. In: Loebl, I, Smetana, A, Eds., Catalogue of Palaearctic Coleoptera Volume 1 Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup: 99 - 118" title="Subfamily Cicindelinae." volumeTitle="Catalogue of Palaearctic Coleoptera Volume 1 Archostemata - Myxophaga - Adephaga." year="2003">Putchkov and Matalin (2003</bibRefCitation>
;
<bibRefCitation author="Putchkov, AV" editor="Loebl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="44" pageNumber="87" pagination="217 - 249" refId="B95" refString="Putchkov, AV, Matalin, AV, 2017. Subfamily Cicindelinae. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera Volume 1 Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup: 217 - 249" title="Subfamily Cicindelinae." volumeTitle="Catalogue of Palaearctic Coleoptera Volume 1 Archostemata - Myxophaga - Adephaga." year="2017">2017</bibRefCitation>
: 219) for Cyprus.
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="59" type="distribution">
<paragraph pageId="16" pageNumber="59">Distribution in the southern Levant.</paragraph>
<paragraph pageId="16" pageNumber="59">
The distribution range of the species in the Middle East seems to be incorrectly reported. Despite the fact that numerous authors mention the species from Israel (
<bibRefCitation author="Avgin, SS" journalOrPublisher="BAUE Fen Bil Enst Degisi" pageId="38" pageNumber="81" pagination="23 - 28" refId="B14" refString="Avgin, SS, 2006. General information and tiger beetles Cicindela herbacea; Calomera fischeri fischeri and Homodela ismenia kilikiensis collected from southeast of Turkey (Coleoptera: Cicindelidae). BAUE Fen Bil Enst Degisi 8: 23 - 28" title="General information and tiger beetles Cicindela herbacea; Calomera fischeri fischeri and Homodela ismenia kilikiensis collected from southeast of Turkey (Coleoptera: Cicindelidae)." volume="8" year="2006">Avgin 2006</bibRefCitation>
;
<bibRefCitation author="Putchkov, AV" editor="Loebl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="44" pageNumber="87" pagination="99 - 118" refId="B94" refString="Putchkov, AV, Matalin, AV, 2003. Subfamily Cicindelinae. In: Loebl, I, Smetana, A, Eds., Catalogue of Palaearctic Coleoptera Volume 1 Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup: 99 - 118" title="Subfamily Cicindelinae." volumeTitle="Catalogue of Palaearctic Coleoptera Volume 1 Archostemata - Myxophaga - Adephaga." year="2003">Putchkov and Matalin 2003</bibRefCitation>
), we do not know of any verified record from the country. There are no specimens in SMNHTAU, and the species is mentioned neither by
<bibRefCitation author="Nussbaum, Y" journalOrPublisher="YES Quarterly" pageId="43" pageNumber="86" pagination="7 - 15" refId="B87" refString="Nussbaum, Y, 1987. Tiger beetles of Israel and Sinai (Coleoptera: Cicindelidae). YES Quarterly 4: 7 - 15" title="Tiger beetles of Israel and Sinai (Coleoptera: Cicindelidae)." volume="4" year="1987">Nussbaum (1987)</bibRefCitation>
nor by
<bibRefCitation author="Valdenberg, A" journalOrPublisher="Shappirit - Journal of the Israel Entomology Center" pageId="44" pageNumber="87" pagination="38 - 48" refId="B105" refString="Valdenberg, A, 1983. The tiger beetles family. General survey of the family and key to species in Israel and Sinai (in Hebrew). Shappirit - Journal of the Israel Entomology Center 1: 38 - 48" title="The tiger beetles family. General survey of the family and key to species in Israel and Sinai (in Hebrew)." volume="1" year="1983">Valdenberg (1983)</bibRefCitation>
. No verifiable records are known from Jordan (Putchkov pers. comm.). However, the species is still listed for Jordan in the latest version of the Palaearctic Catalogue of
<taxonomicName authorityName="Motschoulsky" authorityYear="1862" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="16" pageNumber="59" rank="order">Coleoptera</taxonomicName>
(
<bibRefCitation author="Putchkov, AV" editor="Loebl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="44" pageNumber="87" pagination="217 - 249" refId="B95" refString="Putchkov, AV, Matalin, AV, 2017. Subfamily Cicindelinae. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera Volume 1 Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup: 217 - 249" title="Subfamily Cicindelinae." volumeTitle="Catalogue of Palaearctic Coleoptera Volume 1 Archostemata - Myxophaga - Adephaga." year="2017">Putchkov and Matalin 2017</bibRefCitation>
).
</paragraph>
<paragraph pageId="16" pageNumber="59">
In Israel and Jordan, the
<normalizedToken originalValue="species">species'</normalizedToken>
typical habitats, such as river banks in dynamic floodplains or wet pioneer vegetation with patches of bare ground, have mostly been destroyed or are strongly influenced by human activity. Therefore, a recent occurrence of
<taxonomicName lsidName="C. fischeri" pageId="16" pageNumber="59" rank="species" species="fischeri">
<emphasis italics="true" pageId="16" pageNumber="59">C. fischeri</emphasis>
</taxonomicName>
in the Mediterranean part of the southern Levant is unlikely. There is a small chance that the species can be found in wadis or close to water reservoirs in the desert regions (cf.
<bibRefCitation author="Wiesner, J" journalOrPublisher="Entomologische Zeitschrift" pageId="45" pageNumber="88" pagination="382 - 384" refId="B115" refString="Wiesner, J, 1996. Weitere Mitteilungen ueber die Cicindelidae (Coleoptera) der Vereinigten Arabischen Emirate. Entomologische Zeitschrift 106: 382 - 384" title="Weitere Mitteilungen ueber die Cicindelidae (Coleoptera) der Vereinigten Arabischen Emirate." volume="106" year="1996">Wiesner 1996</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="59" type="taxonomic notes">
<paragraph pageId="16" pageNumber="59">Taxonomic notes.</paragraph>
<paragraph pageId="16" pageNumber="59">
The nominate subspecies occurs in the northern Levant, while the subspecies
<taxonomicName class="Insecta" family="Cicindelidae" genus="Calomera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calomera fischeri subsp. elongatosignata" order="Coleoptera" pageId="16" pageNumber="59" phylum="Arthropoda" rank="subSpecies" species="fischeri" subSpecies="elongatosignata">Calomera fischeri elongatosignata</taxonomicName>
(W. Horn, 1922) is found on the Arabian Peninsula (
<bibRefCitation author="Wiesner, J" journalOrPublisher="Erna Bauer, Keltern" pageId="45" pageNumber="88" refId="B114" refString="Wiesner, J, 1992. Checklist of the tiger beetles of the world. Erna Bauer, Keltern" title="Checklist of the tiger beetles of the world." year="1992">Wiesner 1992</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>