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<document id="3BB564B7583E1FDD53281546F63557DF" ID-DOI="10.11646/zootaxa.3584.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="10096816" ID-ZooBank="072032C4-63FC-499A-A61D-58B428051302" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629830673067" checkinUser="felipe" docAuthor="Michat, Mariano C., Alarie, Yves &amp; Watts, Chris H. S." docDate="2012" docId="256287ACFF842713FBE8FC10E35482D3" docLanguage="en" docName="zt03584p110.pdf" docOrigin="Zootaxa 3584 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.3584.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Limbodessus morgani" docType="treatment" docVersion="4" lastPageNumber="69" masterDocId="D95BFFD4FFC52757FB7FFF92E231831C" masterDocTitle="3584" masterLastPageNumber="110" masterPageNumber="1" pageNumber="66" updateTime="1699581498450" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:namePart id="542501538A526AA3C2E7B7EF0CBE1BFD">Michat, Mariano C.</mods:namePart>
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<mods:namePart id="266837F9FC056B8C92687AC83FB47B6A">Alarie, Yves</mods:namePart>
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<mods:namePart id="2744C52B91658CC74A274B44E38EBD1D">Watts, Chris H. S.</mods:namePart>
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<treatment id="256287ACFF842713FBE8FC10E35482D3" ID-DOI="http://doi.org/10.5281/zenodo.5462388" ID-Zenodo-Dep="5462388" LSID="urn:lsid:plazi:treatment:256287ACFF842713FBE8FC10E35482D3" httpUri="http://treatment.plazi.org/id/256287ACFF842713FBE8FC10E35482D3" lastPageId="68" lastPageNumber="69" pageId="65" pageNumber="66">
<subSubSection id="E5D16531FF842716FBE8FC10E1388080" box="[151,777,898,924]" pageId="65" pageNumber="66" type="nomenclature">
<paragraph id="AD7436BAFF842716FBE8FC10E1388080" blockId="65.[151,777,898,957]" box="[151,777,898,924]" pageId="65" pageNumber="66">
<heading id="F63C81D6FF842716FBE8FC10E1388080" bold="true" box="[151,777,898,924]" fontSize="11" level="1" pageId="65" pageNumber="66" reason="1">
<taxonomicName id="6ACB4D39FF842716FBE8FC10E1388080" ID-CoL="72BGY" authority="(Watts &amp; Humphreys, 2000)" baseAuthorityName="Watts &amp; Humphreys" baseAuthorityYear="2000" box="[151,777,898,924]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="morgani">
<emphasis id="9FBFEAA8FF842716FBE8FC10E1388080" bold="true" box="[151,777,898,924]" pageId="65" pageNumber="66">
<emphasis id="9FBFEAA8FF842716FBE8FC10E3908080" bold="true" box="[151,417,898,924]" italics="true" pageId="65" pageNumber="66">Limbodessus morgani</emphasis>
(Watts &amp; Humphreys, 2000)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E5D16531FF842716FBE8FC37E37380A1" box="[151,322,933,957]" pageId="65" pageNumber="66" type="description">
<paragraph id="AD7436BAFF842716FBE8FC37E37380A1" blockId="65.[151,777,898,957]" box="[151,322,933,957]" pageId="65" pageNumber="66">
(
<figureCitation id="35F02A3FFF842716FBE0FC37E30A80A1" box="[159,315,933,957]" captionStart-0="FIGURES 148–156" captionStart-1="FIGURES 157–161" captionStartId-0="66.[151,263,1893,1916]" captionStartId-1="67.[151,263,1921,1944]" captionTargetBox-0="[181,1410,197,1861]" captionTargetBox-1="[173,1424,196,1898]" captionTargetId-0="figure-80@66.[179,1410,195,1873]" captionTargetId-1="figure-57@67.[170,1426,193,1911]" captionTargetPageId-0="66" captionTargetPageId-1="67" captionText-0="FIGURES 148–156. Limbodessus morgani, first-instar larva. 148, cephalic capsule, dorsal aspect; 149, cephalic capsule, ventral aspect; 150, right antenna, dorsal aspect; 151, left antenna, ventral aspect; 152, right maxilla, dorsal aspect; 153, left" captionText-1="FIGURES 157–161. Limbodessus morgani, first-instar larva. 157, left metathoracic leg, anterior aspect; 158, right" figureDoi-0="http://doi.org/10.5281/zenodo.10096838" figureDoi-1="http://doi.org/10.5281/zenodo.10096840" httpUri-0="https://zenodo.org/record/10096838/files/figure.png" httpUri-1="https://zenodo.org/record/10096840/files/figure.png" pageId="65" pageNumber="66">Figs 148–161</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="E5D16531FF842716FBE8FC77E09C8779" pageId="65" pageNumber="66" type="materials_examined">
<paragraph id="AD7436BAFF842716FBE8FC77E09C8779" blockId="65.[151,1437,997,1924]" pageId="65" pageNumber="66">
<emphasis id="9FBFEAA8FF842716FBE8FC77E34C80E2" bold="true" box="[151,381,997,1022]" inLineHeading="true" pageId="65" pageNumber="66" reason="1">Source of material.</emphasis>
<materialsCitation id="1DA33CE7FF842716FAFBFC77E09C8779" collectingDate="2001-05-09" collectorName="NE Lake Way &amp; W. F. Humphreys &amp; Humphreys, C. H. S &amp; S. J. B. Cooper." country="Australia" county="No" latitude="-26.85444" location="One" longLatPrecision="1" longitude="120.33833" municipality="Lake Way" pageId="65" pageNumber="66" specimenCount="1">
<location id="A8146061FF842716FAFBFC77E38280E1" LSID="urn:lsid:plazi:treatment:256287ACFF842713FBE8FC10E35482D3:A8146061FF842716FAFBFC77E38280E1" box="[388,435,997,1021]" country="Australia" county="No" latitude="-26.85444" longLatPrecision="1" longitude="120.33833" municipality="Lake Way" name="One" pageId="65" pageNumber="66">One</location>
specimen of instar I and one of instar II were used for the description (
<tableCitation id="E0490301FF842716FF9FFC77E70480E1" box="[1248,1333,997,1021]" captionStart="TABLE 1" captionStartId="5.[151,238,608,632]" captionText="TABLE 1. Species and instars described in the present study. The numbers under each instar indicate the specimens studied." pageId="65" pageNumber="66">Table 1</tableCitation>
).
<location id="A8146061FF842716FE31FC74E7AD80E1" LSID="urn:lsid:plazi:treatment:256287ACFF842713FBE8FC10E35482D3:A8146061FF842716FE31FC74E7AD80E1" box="[1358,1436,998,1021]" country="Australia" county="No" latitude="-26.85444" longLatPrecision="1" longitude="120.33833" municipality="Lake Way" name="Larvae" pageId="65" pageNumber="66">Larvae</location>
were collected in association with adults at the following locality:
<emphasis id="9FBFEAA8FF842716F8DBFB95E73A873C" bold="true" box="[932,1291,1031,1056]" pageId="65" pageNumber="66">
<collectingCountry id="D5DC762AFF842716F8DBFB95E627873C" box="[932,1046,1031,1056]" name="Australia" pageId="65" pageNumber="66">Australia</collectingCountry>
,
<location id="A8146061FF842716FF5AFB95E65F873C" LSID="urn:lsid:plazi:treatment:256287ACFF842713FBE8FC10E35482D3:A8146061FF842716FF5AFB95E65F873C" box="[1061,1134,1031,1056]" country="Australia" county="No" latitude="-26.85444" longLatPrecision="1" longitude="120.33833" municipality="Lake Way" name="Carey" pageId="65" pageNumber="66">Carey</location>
palaeovalley
</emphasis>
,
<location id="A8146061FF842716FE62FB95E2FE875E" LSID="urn:lsid:plazi:treatment:256287ACFF842713FBE8FC10E35482D3:A8146061FF842716FE62FB95E2FE875E" country="Australia" county="No" latitude="-26.85444" longLatPrecision="1" longitude="120.33833" municipality="Lake Way" name="Uramurdah Lake" pageId="65" pageNumber="66">Uramurdah Lake</location>
calcrete,
<collectorName id="003E536CFF842716FA34FBB9E3CA875F" box="[331,507,1066,1091]" pageId="65" pageNumber="66">
NE
<collectingMunicipality id="4D10ACC0FF842716FAFFFBB8E3CA875F" box="[384,507,1066,1091]" pageId="65" pageNumber="66">Lake Way</collectingMunicipality>
</collectorName>
,
<collectingCounty id="44154E36FF842716F971FBB9E001875E" box="[526,560,1067,1090]" pageId="65" pageNumber="66">No</collectingCounty>
7, site 285, BES 6450,
<geoCoordinate id="C8FF507DFF842716F813FBB8E1D7875E" box="[876,998,1066,1090]" degrees="26.85444" direction="south" orientation="latitude" pageId="65" pageNumber="66" precision="1" value="-26.85444">26.85444S</geoCoordinate>
,
<geoCoordinate id="C8FF507DFF842716F885FBB8E6B4875E" box="[1018,1157,1066,1090]" degrees="120.33833" direction="east" orientation="longitude" pageId="65" pageNumber="66" precision="1" value="120.33833">120.33833E</geoCoordinate>
,
<date id="D975107AFF842716FFE6FBB8E737875E" box="[1177,1286,1066,1090]" pageId="65" pageNumber="66" value="2001-05-09">
<collectingDate id="C931E992FF842716FFE6FBB8E737875E" box="[1177,1286,1066,1090]" pageId="65" pageNumber="66" value="2001-05-09">9-V-2001</collectingDate>
</date>
, coll.
<collectorName id="003E536CFF842716FE26FBB9E32D8779" pageId="65" pageNumber="66">W. F. Humphreys</collectorName>
, C. H. S. Watts and
<collectorName id="003E536CFF842716FA82FBDFE09C8779" box="[509,685,1101,1125]" pageId="65" pageNumber="66">S. J. B. Cooper.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="E5D16531FF842713FBB8FBFDE3D283B3" lastPageId="68" lastPageNumber="69" pageId="65" pageNumber="66" type="diagnosis">
<paragraph id="AD7436BAFF842716FBB8FBFDE7388629" blockId="65.[151,1437,997,1924]" pageId="65" pageNumber="66">
<emphasis id="9FBFEAA8FF842716FBB8FBFDE38B8794" bold="true" box="[199,442,1135,1160]" inLineHeading="true" pageId="65" pageNumber="66" reason="1">Diagnosis (instar II).</emphasis>
Smaller species (HL less than
<quantity id="6A339B5FFF842716F869FBFDE14F879B" box="[790,894,1135,1159]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="65" pageNumber="66" unit="mm" value="0.4">0.40 mm</quantity>
); head subpentagonal; nasale subtriangular; halfcircle of dense spinulae on lateroventral margins of nasale absent; hole-like structure on ventrodistal surface of nasale absent; lateral margins of nasale not inflated in dorsal view; lateral branches of nasale minute; slender spinulae anterior to seta FR13 scarce (20 or less); occipital foramen well developed (HW/OCW less than 1.75); occipital suture present; lateral margins of parietal straight; secondary spiniform setae on lateral margins of parietal scarce; seta AN2 present; distal half of MN broad; setae LA3, LA4, LA5 and LA8 hair-like; secondary setae on U absent.
</paragraph>
<paragraph id="AD7436BAFF842716FBB8FAD3E732851A" blockId="65.[151,1437,997,1924]" pageId="65" pageNumber="66">
<emphasis id="9FBFEAA8FF842716FBB8FAD3E3108645" bold="true" box="[199,289,1345,1369]" pageId="65" pageNumber="66">Instar I</emphasis>
(
<figureCitation id="35F02A3FFF842716FA4FFAD2E3FD8644" box="[304,460,1344,1368]" captionStart-0="FIGURES 148–156" captionStart-1="FIGURES 157–161" captionStartId-0="66.[151,263,1893,1916]" captionStartId-1="67.[151,263,1921,1944]" captionTargetBox-0="[181,1410,197,1861]" captionTargetBox-1="[173,1424,196,1898]" captionTargetId-0="figure-80@66.[179,1410,195,1873]" captionTargetId-1="figure-57@67.[170,1426,193,1911]" captionTargetPageId-0="66" captionTargetPageId-1="67" captionText-0="FIGURES 148–156. Limbodessus morgani, first-instar larva. 148, cephalic capsule, dorsal aspect; 149, cephalic capsule, ventral aspect; 150, right antenna, dorsal aspect; 151, left antenna, ventral aspect; 152, right maxilla, dorsal aspect; 153, left" captionText-1="FIGURES 157–161. Limbodessus morgani, first-instar larva. 157, left metathoracic leg, anterior aspect; 158, right" figureDoi-0="http://doi.org/10.5281/zenodo.10096838" figureDoi-1="http://doi.org/10.5281/zenodo.10096840" httpUri-0="https://zenodo.org/record/10096838/files/figure.png" httpUri-1="https://zenodo.org/record/10096840/files/figure.png" pageId="65" pageNumber="66">Figs 148–161</figureCitation>
). Head (
<figureCitation id="35F02A3FFF842716F957FAD2E0F48644" box="[552,709,1344,1368]" captionStart="FIGURES 148–156" captionStartId="66.[151,263,1893,1916]" captionTargetBox="[181,1410,197,1861]" captionTargetId="figure-80@66.[179,1410,195,1873]" captionTargetPageId="66" captionText="FIGURES 148–156. Limbodessus morgani, first-instar larva. 148, cephalic capsule, dorsal aspect; 149, cephalic capsule, ventral aspect; 150, right antenna, dorsal aspect; 151, left antenna, ventral aspect; 152, right maxilla, dorsal aspect; 153, left" figureDoi="http://doi.org/10.5281/zenodo.10096838" httpUri="https://zenodo.org/record/10096838/files/figure.png" pageId="65" pageNumber="66">Figs 148–156</figureCitation>
). Cephalic capsule not strongly elongate (HL/HW less than 1.55); seta PA3 inserted contiguous to setae PA1 and PA2; A3 more than 2.80 times longer than A1; A3 less than 2.05 times longer than A2; MP2 more than 2.10 times longer than MP1; MP2 1.95–2.65 times longer than MP3; LP2 more than 2.40 times longer than LP1. Legs (
<figureCitation id="35F02A3FFF842716F970FA3BE09B86DD" box="[527,682,1449,1473]" captionStart="FIGURES 157–161" captionStartId="67.[151,263,1921,1944]" captionTargetBox="[173,1424,196,1898]" captionTargetId="figure-57@67.[170,1426,193,1911]" captionTargetPageId="67" captionText="FIGURES 157–161. Limbodessus morgani, first-instar larva. 157, left metathoracic leg, anterior aspect; 158, right" figureDoi="http://doi.org/10.5281/zenodo.10096840" httpUri="https://zenodo.org/record/10096840/files/figure.png" pageId="65" pageNumber="66">Figs 157–158</figureCitation>
). L3 less than 2.85 times longer than HW. Abdomen (
<figureCitation id="35F02A3FFF842716FF8AFA3BE7A186DD" box="[1269,1424,1449,1473]" captionStart="FIGURES 157–161" captionStartId="67.[151,263,1921,1944]" captionTargetBox="[173,1424,196,1898]" captionTargetId="figure-57@67.[170,1426,193,1911]" captionTargetPageId="67" captionText="FIGURES 157–161. Limbodessus morgani, first-instar larva. 157, left metathoracic leg, anterior aspect; 158, right" figureDoi="http://doi.org/10.5281/zenodo.10096840" httpUri="https://zenodo.org/record/10096840/files/figure.png" pageId="65" pageNumber="66">Figs 159–161</figureCitation>
). U more than 3.75 times longer than LAS; U less than 2.35 times longer than HW; U1 less than 1.05 times longer than U2. Chaetotaxy. Frontoclypeus with 12 lamellae clypeales; additional setae on U absent. Measurements
</paragraph>
<paragraph id="AD7436BAFF842716FBB9F983E04E84F0" blockId="65.[151,1437,997,1924]" lastBlockId="65.[151,1436,1934,2028]" pageId="65" pageNumber="66">
<emphasis id="9FBFEAA8FF842716FBB9F983E3028535" bold="true" box="[198,307,1553,1577]" pageId="65" pageNumber="66">Instar II.</emphasis>
Head. A3 less than 2.50 times longer than A1; A4 more than 0.75 times as long as A3; MN less than 4.70 times longer than broad; MP2 1.05–1.45 times longer than MP1; MP2 less than 2.05 times longer than MP3; LP2 more than 1.75 times longer than LP1. Legs. L3 less than 2.20 times longer than HW; CL(L3) more than 0.40 times as long as TA. Abdomen. U 3.00–4.00 times longer than LAS; U less than 2.15 times longer than HW; U1 less than 1.45 times longer than U2. Chaetotaxy. Anteroventral margin of nasale with 22 lamellae clypeales distributed in a single row; anterior secondary setae on proCO absent; meso- and metaCO with less than 5 posterodorsal secondary setae; ventral secondary setae on proCO absent; ventral secondary setae on mesoCO present; proFE with less than 3 posteroventral secondary setae; metaFE with less than 11 secondary setae; secondary setae on proTI absent; anterodorsal secondary setae on meso- and metaTI present; meso- and metaTI with less than 2 posterodorsal secondary setae; posteroventral secondary setae on meso- and metaTI absent; metaTI with less than 5 secondary setae; secondary setae on proTA absent; posterodorsal secondary setae on meso- and metaTA present; posteroventral secondary setae on meso- and metaTA absent; metaTA with less than 4 secondary setae; secondary setae on U absent. Measurements and ratios that characterize the body shape are shown in
<tableCitation id="E0490301FF842716FE3CF823E7A984D5" box="[1347,1432,1969,1993]" captionStart="TABLE 6" captionStartId="11.[151,239,408,432]" captionText="TABLE 6. Measurements and ratios for the second-instar larvae of species of Limbodessus." pageId="65" pageNumber="66">Table 6</tableCitation>
. Secondary leg setation detailed in
<tableCitation id="E0490301FF842716F967F846E04A84F0" box="[536,635,2004,2028]" captionStart="TABLE 14" captionStartId="53.[151,239,978,1002]" captionText="TABLE 14. Number and position of secondary setae on the legs of second-instar larvae of species of Limbodessus. Numbers between slash marks refer to pro-, meso- and metathoracic leg, respectively. A = anterior, D = dorsal, P = posterior, Pr = proximal, V = ventral, Total = total number of secondary setae on the segment (excluding primary setae)." pageId="65" pageNumber="66">Table 14</tableCitation>
.
</paragraph>
<caption id="F9B46632FF872715FBE8F8F7E7AD8486" ID-DOI="http://doi.org/10.5281/zenodo.10096838" ID-Zenodo-Dep="10096838" httpUri="https://zenodo.org/record/10096838/files/figure.png" pageId="66" pageNumber="67" startId="66.[151,263,1893,1916]" targetBox="[181,1410,197,1861]" targetPageId="66">
<paragraph id="AD7436BAFF872715FBE8F8F7E7AD8486" blockId="66.[151,1436,1893,1946]" pageId="66" pageNumber="67">
<emphasis id="9FBFEAA8FF872715FBE8F8F7E3408460" bold="true" box="[151,369,1893,1916]" pageId="66" pageNumber="67">FIGURES 148–156.</emphasis>
<taxonomicName id="6ACB4D39FF872715FA03F8F7E06F8467" baseAuthorityName="Watts &amp; Humphreys" baseAuthorityYear="2000" box="[380,606,1893,1915]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="morgani">
<emphasis id="9FBFEAA8FF872715FA03F8F7E06F8467" box="[380,606,1893,1915]" italics="true" pageId="66" pageNumber="67">Limbodessus morgani</emphasis>
</taxonomicName>
, first-instar larva. 148, cephalic capsule, dorsal aspect; 149, cephalic capsule, ventral aspect; 150, right antenna, dorsal aspect; 151, left antenna, ventral aspect; 152, right maxilla, dorsal aspect; 153, left
</paragraph>
</caption>
<caption id="F9B46632FF862714FBE8F813E7AD848B" ID-DOI="http://doi.org/10.5281/zenodo.10096840" ID-Zenodo-Dep="10096840" box="[151,1436,1921,1944]" httpUri="https://zenodo.org/record/10096840/files/figure.png" pageId="67" pageNumber="68" startId="67.[151,263,1921,1944]" targetBox="[173,1424,196,1898]" targetPageId="67">
<paragraph id="AD7436BAFF862714FBE8F813E7AD848B" blockId="67.[151,1436,1921,1944]" box="[151,1436,1921,1944]" pageId="67" pageNumber="68">
<emphasis id="9FBFEAA8FF862714FBE8F813E3478484" bold="true" box="[151,374,1921,1944]" pageId="67" pageNumber="68">FIGURES 157–161.</emphasis>
<taxonomicName id="6ACB4D39FF862714FAF9F813E05C848B" baseAuthorityName="Watts &amp; Humphreys" baseAuthorityYear="2000" box="[390,621,1921,1943]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="morgani">
<emphasis id="9FBFEAA8FF862714FAF9F813E05C848B" box="[390,621,1921,1943]" italics="true" pageId="67" pageNumber="68">Limbodessus morgani</emphasis>
</taxonomicName>
, first-instar larva. 157, left metathoracic leg, anterior aspect; 158, right
</paragraph>
</caption>
<paragraph id="AD7436BAFF812713FBB8FF0AE3D283B3" blockId="68.[151,1436,151,464]" box="[199,483,151,176]" pageId="68" pageNumber="69">
<emphasis id="9FBFEAA8FF812713FBB8FF0AE30F83AC" bold="true" box="[199,318,152,176]" pageId="68" pageNumber="69">Instar III.</emphasis>
Not available.
</paragraph>
</subSubSection>
<subSubSection id="E5D16531FF812713FBB0FF2EE35482D3" pageId="68" pageNumber="69" type="discussion">
<paragraph id="AD7436BAFF812713FBB0FF2EE35482D3" blockId="68.[151,1436,151,464]" pageId="68" pageNumber="69">
<emphasis id="9FBFEAA8FF812713FBB0FF2EE37283C8" bold="true" box="[207,323,188,212]" inLineHeading="true" pageId="68" pageNumber="69" reason="1">Remarks.</emphasis>
The absence of instar III specimens of
<taxonomicName id="6ACB4D39FF812713F87AFF2FE1B383C8" baseAuthorityName="Watts &amp; Humphreys" baseAuthorityYear="2000" box="[773,898,189,212]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="morgani">
<emphasis id="9FBFEAA8FF812713F87AFF2FE1B383C8" box="[773,898,189,212]" italics="true" pageId="68" pageNumber="69">L. morgani</emphasis>
</taxonomicName>
hampers the comparison with the other species described here, particularly regarding the chaetotaxic characters.
<taxonomicName id="6ACB4D39FF812713F8E3FF72E6A983EB" baseAuthorityName="Watts &amp; Humphreys" baseAuthorityYear="2000" box="[924,1176,224,248]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="morgani">
<emphasis id="9FBFEAA8FF812713F8E3FF72E6A983EB" box="[924,1176,224,248]" italics="true" pageId="68" pageNumber="69">Limbodessus morgani</emphasis>
</taxonomicName>
has a small size and belongs to the group of stygobitic species that lack secondary setae on the urogomphus (
<taxonomicName id="6ACB4D39FF812713FF0AFE97E7208200" baseAuthorityName="Watts &amp; Humphreys" baseAuthorityYear="2001" box="[1141,1297,260,284]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="challaensis">
<emphasis id="9FBFEAA8FF812713FF0AFE97E7208200" box="[1141,1297,260,284]" italics="true" pageId="68" pageNumber="69">L. challaensis</emphasis>
</taxonomicName>
,
<taxonomicName id="6ACB4D39FF812713FE60FE97E7468200" authorityName="Watts &amp; Humphreys" authorityYear="2006" box="[1311,1399,260,284]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="exilis">
<emphasis id="9FBFEAA8FF812713FE60FE97E7468200" box="[1311,1399,260,284]" italics="true" pageId="68" pageNumber="69">L. exilis</emphasis>
</taxonomicName>
,
<taxonomicName id="6ACB4D39FF812713FEFAFE97E370825C" baseAuthorityName="Watts &amp; Humphreys" baseAuthorityYear="2001" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="fridaywellensis">
<emphasis id="9FBFEAA8FF812713FEFAFE97E370825C" italics="true" pageId="68" pageNumber="69">L. fridaywellensis</emphasis>
</taxonomicName>
,
<taxonomicName id="6ACB4D39FF812713FA2EFEBBE033825C" baseAuthorityName="Watts &amp; Humphreys" baseAuthorityYear="2004" box="[337,514,296,320]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="hillviewensis">
<emphasis id="9FBFEAA8FF812713FA2EFEBBE033825C" box="[337,514,296,320]" italics="true" pageId="68" pageNumber="69">L. hillviewensis</emphasis>
</taxonomicName>
,
<taxonomicName id="6ACB4D39FF812713F96DFEBBE0B8825C" baseAuthorityName="Watts &amp; Humphreys" baseAuthorityYear="2000" box="[530,649,296,320]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="hinkleri">
<emphasis id="9FBFEAA8FF812713F96DFEBBE0B8825C" box="[530,649,296,320]" italics="true" pageId="68" pageNumber="69">L. hinkleri</emphasis>
</taxonomicName>
,
<taxonomicName id="6ACB4D39FF812713F9E6FEBBE0DD825C" authorityName="Watts &amp; Humphreys" authorityYear="2006" box="[665,748,296,320]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="leysi">
<emphasis id="9FBFEAA8FF812713F9E6FEBBE0DD825C" box="[665,748,296,320]" italics="true" pageId="68" pageNumber="69">L. leysi</emphasis>
</taxonomicName>
,
<taxonomicName id="6ACB4D39FF812713F982FEBBE1AF8223" baseAuthorityName="Watts &amp; Humphreys" baseAuthorityYear="2001" box="[765,926,296,320]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="masonensis">
<emphasis id="9FBFEAA8FF812713F982FEBBE1AF8223" box="[765,926,296,320]" italics="true" pageId="68" pageNumber="69">L. masonensis</emphasis>
</taxonomicName>
,
<taxonomicName id="6ACB4D39FF812713F8D1FEBBE66E825C" authorityName="Watts &amp; Humphreys" authorityYear="2006" box="[942,1119,296,320]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="millbilliensis">
<emphasis id="9FBFEAA8FF812713F8D1FEBBE66E825C" box="[942,1119,296,320]" italics="true" pageId="68" pageNumber="69">L. millbilliensis</emphasis>
</taxonomicName>
,
<taxonomicName id="6ACB4D39FF812713FF10FEBBE735825C" authorityName="Watts &amp; Humphreys" authorityYear="2009" box="[1135,1284,296,320]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="ordinarius">
<emphasis id="9FBFEAA8FF812713FF10FEBBE735825C" box="[1135,1284,296,320]" italics="true" pageId="68" pageNumber="69">L. ordinarius</emphasis>
</taxonomicName>
,
<taxonomicName id="6ACB4D39FF812713FE6AFEBBE744825C" baseAuthorityName="Watts &amp; Humphreys" baseAuthorityYear="1999" box="[1301,1397,296,320]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="pulpa">
<emphasis id="9FBFEAA8FF812713FE6AFEBBE744825C" box="[1301,1397,296,320]" italics="true" pageId="68" pageNumber="69">L. pulpa</emphasis>
</taxonomicName>
,
<taxonomicName id="6ACB4D39FF812713FEFAFEBBE2E68278" authorityName="Watts &amp; Humphreys" authorityYear="2006" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="raeae">
<emphasis id="9FBFEAA8FF812713FEFAFEBBE2E68278" italics="true" pageId="68" pageNumber="69">L. raeae</emphasis>
</taxonomicName>
,
<taxonomicName id="6ACB4D39FF812713FB9AFEDFE3928278" baseAuthorityName="Watts &amp; Humphreys" baseAuthorityYear="1999" box="[229,419,332,356]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="windarraensis">
<emphasis id="9FBFEAA8FF812713FB9AFEDFE3928278" box="[229,419,332,356]" italics="true" pageId="68" pageNumber="69">L. windarraensis</emphasis>
</taxonomicName>
,
<taxonomicName id="6ACB4D39FF812713FACEFEDFE05E8278" baseAuthorityName="Watts &amp; Humphreys" baseAuthorityYear="2003" box="[433,623,333,356]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="yuinmeryensis">
<emphasis id="9FBFEAA8FF812713FACEFEDFE05E8278" box="[433,623,333,356]" italics="true" pageId="68" pageNumber="69">L. yuinmeryensis</emphasis>
</taxonomicName>
). Within this group, it can be separated by the following combination of characters: lateral margins of the nasale not inflated in dorsal view, and claws relatively longer as compared to tarsus (ratio CL/TA&gt; 0.40). This last feature separates
<taxonomicName id="6ACB4D39FF812713F879FE07E1B382B0" baseAuthorityName="Watts &amp; Humphreys" baseAuthorityYear="2000" box="[774,898,405,428]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="morgani">
<emphasis id="9FBFEAA8FF812713F879FE07E1B382B0" box="[774,898,405,428]" italics="true" pageId="68" pageNumber="69">L. morgani</emphasis>
</taxonomicName>
from all other stygobitic species studied except
<taxonomicName id="6ACB4D39FF812713FBE8FE2BE36C82CC" baseAuthorityName="Watts &amp; Humphreys" baseAuthorityYear="2001" box="[151,349,440,464]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="fridaywellensis">
<emphasis id="9FBFEAA8FF812713FBE8FE2BE36C82CC" box="[151,349,440,464]" italics="true" pageId="68" pageNumber="69">L. fridaywellensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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