238 lines
31 KiB
XML
238 lines
31 KiB
XML
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<mods:title id="189DC99183E0F7683A306A44B385B635">Revision of the genus Orictites ANDREWES, 1931 (Coleoptera, Carabidae, Clivinini)</mods:title>
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<paragraph id="AD4F195BA850FFF7D619FF338DA9FF61" blockId="3.[121,616,168,453]" box="[121,236,168,192]" pageId="3" pageNumber="4">Catalogue</paragraph>
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<emphasis id="9F84C549A850FFF7D619FF568D9CFF44" box="[121,217,205,229]" italics="true" pageId="3" pageNumber="4">Orictites</emphasis>
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;
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<emphasis id="9F84C549A850FFF7D619FF698D9CFEAB" box="[121,217,242,266]" italics="true" pageId="3" pageNumber="4">Orictites</emphasis>
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ANDREWES
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;
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<treatmentCitation id="2C513F4AA850FFF7D706FF698F51FEAB" author="Csiki E." box="[358,532,242,266]" page="638" pageId="3" pageNumber="4" year="1933">
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<emphasis id="9F84C549A850FFF7D619FE838D9CFE91" box="[121,217,280,304]" italics="true" pageId="3" pageNumber="4">Orictites</emphasis>
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ANDREWES
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<treatmentCitation id="2C513F4AA850FFF7D706FE838F25FE91" author="Louwerens, C. J." box="[358,608,280,304]" page="172" pageId="3" pageNumber="4" year="1964">
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<bibRefCitation id="C96164AAA850FFF7D706FE838F25FE91" author="Louwerens, C. J." box="[358,608,280,304]" pageId="3" pageNumber="4" pagination="171 - 189" refId="ref27961" refString="Louwerens, C. J. (1964): An annotated list of the Carabidae, chiefly collected in East Borneo by Dr Eric MjOberg with descriptions of new species. - Entomologisk Tidskrift, Stockholm 85: 171 - 189." type="journal article" year="1964">Louwerens, 1964: 172</bibRefCitation>
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<emphasis id="9F84C549A850FFF7D619FEA68D9CFEF4" box="[121,217,317,341]" italics="true" pageId="3" pageNumber="4">Orictites</emphasis>
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ANDREWES
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;
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<treatmentCitation id="2C513F4AA850FFF7D706FEA68F04FEF4" author="Balkenohl, M." box="[358,577,317,341]" page="24" pageId="3" pageNumber="4" year="1996">
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<bibRefCitation id="C96164AAA850FFF7D706FEA68F04FEF4" author="Balkenohl, M." box="[358,577,317,341]" pageId="3" pageNumber="4" pagination="1 - 19" refId="ref27620" refString="Balkenohl, M. (1996): New Clivinini from the Oriental region (Coleoptera: Carabidae, Scaritinae). - Acta Zoologica Academiae Scientiarum Hungaricae, Budapest, 42 (1): 1 - 19." type="journal article" year="1996">Balkenohl 1996: 24</bibRefCitation>
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<paragraph id="AD4F195BA850FFF7D619FEF98F75FEDB" blockId="3.[121,616,168,453]" box="[121,560,354,378]" pageId="3" pageNumber="4">
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<emphasis id="9F84C549A850FFF7D619FEF98D9CFEDB" box="[121,217,354,378]" italics="true" pageId="3" pageNumber="4">Orictites</emphasis>
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ANDREWES
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;
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<treatmentCitation id="2C513F4AA850FFF7D706FEF98F6DFEDB" author="Lorenz, W." box="[358,552,354,378]" page="135" pageId="3" pageNumber="4" year="1998">
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<emphasis id="9F84C549A850FFF7D619FE138D9CFE01" box="[121,217,392,416]" italics="true" pageId="3" pageNumber="4">Orictites</emphasis>
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ANDREWES
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;
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<treatmentCitation id="2C513F4AA850FFF7D706FE138F5AFE01" author="Baehr, M." box="[358,543,392,416]" page="115" pageId="3" pageNumber="4" year="1999">
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<bibRefCitation id="C96164AAA850FFF7D706FE138F5AFE01" author="Baehr, M." box="[358,543,392,416]" pageId="3" pageNumber="4" pagination="115 - 126" refId="ref27552" refString="Baehr, M. (1999): Two new genera of clivinine Scaritinae from the Oriental Region (Coleoptera Carabidae). - In: A. Zamotajlov & R. Sciaky (eds.): Advances in Carabidology. Papers dedicated to the memory of Prof. Dr. Oleg L. Kryzhanovskij, MUISO publ., Krasnodar, 115 - 126 pp." type="book chapter" year="1999">Baehr 1999: 115</bibRefCitation>
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<emphasis id="9F84C549A850FFF7D619FE368D9CFE64" box="[121,217,429,453]" italics="true" pageId="3" pageNumber="4">Orictites</emphasis>
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ANDREWES
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;
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<treatmentCitation id="2C513F4AA850FFF7D706FE368F78FE64" author="Balkenohl, M." box="[358,573,429,453]" page="17" pageId="3" pageNumber="4" year="2001">
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<bibRefCitation id="C96164AAA850FFF7D706FE368F78FE64" author="Balkenohl, M." box="[358,573,429,453]" pageId="3" pageNumber="4" refId="ref27660" refString="Balkenohl, M. (2001): Key and Catalogue of the tribe Clivinini from the Oriental realm, with revisions of the genera Thliboclivina KULT, and Trilophidius JEANNEL (Insecta, Coleoptera, Carabidae, Scarititae, Clivinini). - 83 pp., Sofia, Moscow (Pensoft Series Faunistica 21)." type="book" year="2001">Balkenohl 2001: 17</bibRefCitation>
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.
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<typeStatus id="724BA7F9A850FFF7D619FD868DEBFD94" box="[121,174,541,565]" pageId="3" pageNumber="4">Type</typeStatus>
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species:
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<taxonomicName id="6AF062D8A850FFF7D777FD868FEFFD94" ID-CoL="8M924" authority="ANDREWES, 1931" authorityName="ANDREWES" authorityYear="1931" box="[279,682,541,565]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="minotaur">
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<emphasis id="9F84C549A850FFF7D777FD868CA1FD94" box="[279,484,541,565]" italics="true" pageId="3" pageNumber="4">Orictites minotaur</emphasis>
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<bibRefCitation id="C96164AAA850FFF7D78BFD868FEFFD94" author="Andrewes, H. E." box="[491,682,541,565]" pageId="3" pageNumber="4" pagination="431 - 485" refId="ref27519" refString="Andrewes, H. E. (1931): On the Carabidae of the Mount Kinabalu. - Journal of the FMS Museum, Kuala Lumpur 16: 431 - 485." type="journal article" year="1931">ANDREWES, 1931</bibRefCitation>
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, by original designation.
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Diagnosis: General appearance more or less like a
|
||
<taxonomicName id="6AF062D8A850FFF7D4AAFDF38E52FD21" authorityName="LATREILLE" authorityYear="1802" box="[714,791,616,640]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9F84C549A850FFF7D4AAFDF38E52FD21" box="[714,791,616,640]" italics="true" pageId="3" pageNumber="4">Clivina</emphasis>
|
||
</taxonomicName>
|
||
. Clypeus variable, straight anteriorly or excised or with teeth, with raised knob between the two clypeal setae or with blunt elevation, wings small or well-developed, more or less projecting, divided from supraantennal plates by distinct notches and from middle part of clypeus by distinct or indistinct notches. Frons separated from clypeus by band of big punctures and/or by broad flat furrow. Longitudinal frontal carinae distinct. Supraorbital carinae distinct. Eyes oval, flattened, enclosed posteriorly by distinctly vaulted genae of different length, temporae carinate; antennae relatively short, with some long setae among the usual pubescence. Labrum 7-setose. Mandibles strong, right or left one with tooth at middle or basally. Pronotum square, almost flattened with tooth at posterior angle, anterior angles formed by reflexed lateral margin, rounded and projecting anteriorly or angled. Reflexed lateral margin more or less crenulated. Lateral channel distinctly broad, separated from disc by longitudinal vault, median line conspicuously deep and broad, surface in most of the species covered with deep and broad punctures. Basal channel always broad and deep. Elytra subcylindrical, stria 1–4 free at base, intervals more or less convex, asetose or with setigerous punctures on interval 3, or 3, 5, and 7. Third interval of elytra broadened and raised at base with distinct tubercle, in most species the same is true for interval 4. Middle tibia with spur near apex furnished with seta. Terminal abdominal sternite with two widely separated apical setae at each side. Ventral surface almost covered with punctures. Distinguished from all other
|
||
<taxonomicName id="6AF062D8A850FFF7D740FA068C28FA14" authorityName="LATREILLE" authorityYear="1802" box="[288,365,1437,1461]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9F84C549A850FFF7D740FA068C28FA14" box="[288,365,1437,1461]" italics="true" pageId="3" pageNumber="4">Clivina</emphasis>
|
||
</taxonomicName>
|
||
-related genera by the following combination of characters: the flattened eyes with distinctly enlarged genae, the distinct carinae on the head, the transverse square and flattened pronotum with unusually broad lateral channel, the big and usually numerous punctures on the head, pronotum, and ventral surface, the broad and deep base of the pronotum, the distinctly convex intervals of the elytra, and the distinctly broadened, raised and tubercle possessing intervals 3 and 4 at base.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E5EA4AD0A857FFF1D6FFFE8C8E1EF9A1" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="description">
|
||
<paragraph id="AD4F195BA857FFF0D6FFFE8C8C2BFE91" blockId="4.[159,366,279,304]" box="[159,366,279,304]" pageId="4" pageNumber="5">
|
||
<emphasis id="9F84C549A857FFF0D6FFFE8C8C2BFE91" bold="true" box="[159,366,279,304]" pageId="4" pageNumber="5">Genus description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="AD4F195BA857FFF0D6FFFEF98F11FEDB" blockId="4.[159,1044,354,1573]" box="[159,596,354,378]" pageId="4" pageNumber="5">Size: Small to medium-sized, elongate.</paragraph>
|
||
<paragraph id="AD4F195BA857FFF0D6FFFE138FB9FE01" blockId="4.[159,1044,354,1573]" box="[159,764,392,416]" pageId="4" pageNumber="5">Colour: Shiny, without metallic lustre, black or brown.</paragraph>
|
||
<paragraph id="AD4F195BA857FFF0D6A4FE368C26FA14" blockId="4.[159,1044,354,1573]" pageId="4" pageNumber="5">
|
||
Head: Wide. Clypeus straight, excised, or with teeth anteriorly, limited laterally by small rounded or sharp tooth, with raised knob in between the two clypeal setae or with blunt elevation; wings always well-developed, rounded or angled, hollowed out, divided from supraantennal plates and from middle part of clypeus by notches. Supraantennal plates vaulted, with reflexed rounded margin, frons separated from clypeus by transverse furrow or depression, with or without big punctures, with nearly parallel running carinae at each side posteriorly, each carina with furrow mesially, carinae enclosing the convex frons; frons smooth or with punctures. Supraorbital setae located in broad furrow between eye and frontal carina, the posterior one arising from a small tubercle. Eyes oval, not big, orbit projecting laterally, tempora with distinct supraorbital carina, subtemporae and genae distinctly vaulted, rounded, nearly as high but slightly shorter than eyes, distinctly enclosing eyes posteriorly. Grooves for reception of antennae short. Neck constriction formed by punctures or indistinct. Labrum 7-setose, ciliate laterally, almost with isodiametric reticulation. Mandibles shorter than head, distinctly broadened at base, acute at apex, upper margin of scrobe of both mandibles sharply carinate, mandibular teeth differently developed in the two subgenera. Maxilla bent slightly sigmoidal. Apical maxillary palpomere slender, fusiform bottle-like. Apical labial palpomere slightly arcuate, fusiform bottle-like or securiform, second segment bisetose. Ligula somewhat truncated at apex, with one seta. Paraglossae slender, membranous.
|
||
<collectingRegion id="6F34D7B9A857FFF0D7B4FB268F77FB74" box="[468,562,1213,1237]" country="Myanmar" name="Chin" pageId="4" pageNumber="5">Mentum</collectingRegion>
|
||
and submentum divided by sharp furrow, with two pairs of setae each, median tooth broad, obtuse at apex, bisetose, lobes smooth or with wrinkles, longer than tooth, acute at apex, margins with carinae. Antennae short, not reaching over two thirds of pronotum, scapus with one seta dorsoapically, scapus and pedicellus in most of the cases convered with fine reticulation, segments 4–11 densely pubescent, with few additional long setae.
|
||
</paragraph>
|
||
<paragraph id="AD4F195BA857FFF1D6A4FA598C51FEF4" blockId="4.[159,1044,354,1573]" lastBlockId="5.[121,1006,93,1573]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">Pronotum: Square, as long as wide or broader than long, almost flattened. Anterior margin straight or slightly bisinuate. Reflexed lateral margin smooth or crenulated. Lateral channel distinctly broadened between the two lateral setigerous punctures, basal channel broad and deep. Anterior setigerous puncture located backwards, the posterior one located at level of posterior angle, removed from lateral channel. Anterior angles more or less projecting anteriorly, posterior angles well developed. Anterior transverse line sharp, punctured, or indistinct. Median line deep and conspicuously broad, often with blunt carina bilaterally. Surface nearly always covered with punctures of different size.</paragraph>
|
||
<paragraph id="AD4F195BA856FFF1D6FEFEF98E6EFCC1" blockId="5.[121,1006,93,1573]" pageId="5" pageNumber="6">
|
||
Elytra: Subcylindrical. Marginal channel broad, visible from above, with uninterrupted row of big setigerous punctures arising from broad tubercles, with second row of smaller punctures. Reflexed margin with more or less developed crenulation, more distinct at humerus. Reflexed lateral margin bending parallel to channel over rounded humerus up to stria 5, with or without humeral tooth. Basal tubercle big, with setigerous puncture, situated at declivity of 1
|
||
<superScript id="5A85B413A856FFF1D74EFDA48C7AFDF3" attach="left" box="[302,319,575,594]" fontSize="8" pageId="5" pageNumber="6">st</superScript>
|
||
stria. Striae 1–4 free at base, all striae deep, with more or less distinct punctures, striae 1–2 either ending free at apex or joining apically, 3–4 and 5–6 joined apically; striae 6 and 7 shortened at humerus. Striole distinct, interval between striole and sutura raised. Intervals conspicuously convex, with increasing distinctness laterally, 3
|
||
<superScript id="5A85B413A856FFF1D4FBFD4F8FEBFD46" attach="left" box="[667,686,724,743]" fontSize="8" pageId="5" pageNumber="6">rd</superScript>
|
||
interval and in some cases the4
|
||
<superScript id="5A85B413A856FFF1D6CDFD618D85FCAC" attach="left" box="[173,192,762,781]" fontSize="8" pageId="5" pageNumber="6">th</superScript>
|
||
as well broader and raised at base, with big tubercle at base, 7
|
||
<superScript id="5A85B413A856FFF1D5C4FD618EF2FCAC" attach="left" box="[932,951,762,781]" fontSize="8" pageId="5" pageNumber="6">th</superScript>
|
||
running to apex as more or less obtuse carina, 8
|
||
<superScript id="5A85B413A856FFF1D4E1FC848FD1FC93" attach="left" box="[641,660,799,818]" fontSize="8" pageId="5" pageNumber="6">th</superScript>
|
||
small, carinate. Intervals asetose or with setigerous punctures on interval 3, or 3, 5, and 7.
|
||
</paragraph>
|
||
<paragraph id="AD4F195BA856FFF1D6FEFCF68C9AFC24" blockId="5.[121,1006,93,1573]" box="[158,479,877,901]" pageId="5" pageNumber="6">Hind wings: Well developed.</paragraph>
|
||
<paragraph id="AD4F195BA856FFF1D6FEFC098C92FB11" blockId="5.[121,1006,93,1573]" pageId="5" pageNumber="6">
|
||
Ventral surface: Proepisternum with big scattered punctures and indistinct reticulation, in a few species with some small punctures, submarginal furrow engraved. Metasternum smooth at middle, punctured laterally. Epipleura somewhat broadened basally, with row of big punctures. Sternites 3
|
||
<superScript id="5A85B413A856FFF1D5F1FC648EE1FBB3" attach="left" box="[913,932,1023,1042]" fontSize="8" pageId="5" pageNumber="6">rd</superScript>
|
||
to 5
|
||
<superScript id="5A85B413A856FFF1D5BBFC648EABFBB3" attach="right" box="[987,1006,1023,1042]" fontSize="8" pageId="5" pageNumber="6">th</superScript>
|
||
with paralateral ambulatory setae at each side, covered densely and irregularly with big punctures or punctures limited to lateral margin, ventral strigae distinct, with transverse reticulation; terminal sternite with two widely separated apical setae at each side.
|
||
</paragraph>
|
||
<paragraph id="AD4F195BA856FFF1D6FEFB268E54FAE4" blockId="5.[121,1006,93,1573]" pageId="5" pageNumber="6">Legs: Anterior tibia with strong digitation, with two big and one small lateral preapical denticles, apical spine curved distinctly towards ventral and lateral. Basal tarsomere distinctly elongated. Mesotibia with long apical spine, furnished with strong seta. All tarsomeres relatively broad.</paragraph>
|
||
<paragraph id="AD4F195BA856FFF1D6FEFAC98EA7FACB" blockId="5.[121,1006,93,1573]" box="[158,994,1362,1386]" pageId="5" pageNumber="6">
|
||
Male genitalia: In general of
|
||
<taxonomicName id="6AF062D8A856FFF1D7BEFAC98F6EFACB" authorityName="LATREILLE" authorityYear="1802" box="[478,555,1362,1386]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9F84C549A856FFF1D7BEFAC98F6EFACB" box="[478,555,1362,1386]" italics="true" pageId="5" pageNumber="6">Clivina</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<typeStatus id="724BA7F9A856FFF1D453FAC98F2EFACB" box="[563,619,1362,1386]" pageId="5" pageNumber="6">type</typeStatus>
|
||
, with spatulate or stick-like apex.
|
||
</paragraph>
|
||
<paragraph id="AD4F195BA856FFF1D6FEFAE38C2EFA7B" blockId="5.[121,1006,93,1573]" pageId="5" pageNumber="6">Female genitalia: Stylomere more or less sigmoidal, with or without preapical setae, with one or two ensiform seta at middle and some nematiform setae laterally in basal half.</paragraph>
|
||
<paragraph id="AD4F195BA856FFF1D6FEFA738E1EF9A1" blockId="5.[121,1006,93,1573]" box="[158,859,1512,1536]" pageId="5" pageNumber="6">Sexual dimorphism: External sexual differences not observed.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E5EA4AD0A856FFF2D6FEF9968F71FF3B" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="distribution">
|
||
<paragraph id="AD4F195BA856FFF2D6FEF9968F71FF3B" blockId="5.[121,1006,93,1573]" lastBlockId="6.[159,1043,93,266]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
|
||
Distribution. South
|
||
<collectingCountry id="D5E759CBA856FFF1D7E7F9968CB0F984" box="[391,501,1549,1573]" name="China" pageId="5" pageNumber="6">East Asia</collectingCountry>
|
||
including
|
||
<collectingCountry id="D5E759CBA856FFF1D412F9968F99F984" box="[626,732,1549,1573]" name="Malaysia" pageId="5" pageNumber="6">Malaysia</collectingCountry>
|
||
, Sumatra, Borneo, and Sulawesi. Specimens were also found in North East
|
||
<collectingCountry id="D5E759CBA855FFF2D48CFFC68E61FFD4" box="[748,804,93,117]" name="India" pageId="6" pageNumber="7">India</collectingCountry>
|
||
(
|
||
<collectingRegion id="6F34D7B9A855FFF2D551FFC68EE7FFD4" box="[817,930,93,117]" country="India" name="Meghalaya" pageId="6" pageNumber="7">Megalaya</collectingRegion>
|
||
Hills) and in the West of Papua (
|
||
<figureCitation id="35CB05DEA855FFF2D7F4FF198CA1FF3B" box="[404,484,130,154]" captionStart="Fig" captionStartId="62.[159,188,1116,1134]" captionTargetBox="[159,1043,148,1081]" captionTargetId="figure-150@62.[159,1043,148,1042]" captionText="Fig. 41: Distribution of the species of the Genus Orictites in South East Asia (recorded localities of all species plotted)." figureDoi="http://doi.org/10.5281/zenodo.6284378" httpUri="https://zenodo.org/record/6284378/files/figure.png" pageId="6" pageNumber="7">Fig. 41</figureCitation>
|
||
; p. 63).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E5EA4AD0A855FFF2D6A4FF338CB0FEAB" pageId="6" pageNumber="7" type="biology_ecology">
|
||
<paragraph id="AD4F195BA855FFF2D6A4FF338CB0FEAB" blockId="6.[159,1043,93,266]" pageId="6" pageNumber="7">Habitat: Not well known. Specific labels of some specimens indicate finds in mountainous areas. Some species were found under bark of trees and rotten wood, e.g. in a pine forest.</paragraph>
|
||
</subSubSection>
|
||
</subSection>
|
||
</treatment>
|
||
</document> |